treatments-xml/data/03/A2/87/03A2878B004BF70FDE6A1B30FD5FF79F.xml

<document id="247D2318EF81F283BC5A21BF4D1B6334" ID-DOI="10.37520/aemnp.2020.040" ID-ISSN="1804-6487" ID-Zenodo-Dep="5177782" ID-ZooBank="BBEBE695-58A2-4CF2-821D-E2C60DF9EA57" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628619056934" checkinUser="carolina" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2020" docId="03A2878B004BF70FDE6A1B30FD5FF79F" docLanguage="en" docName="ActaEntMusNatPrag.60.2.591-598.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 60 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2020.040" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Caloboreaphilus concisus Shavrin &amp; Smetana 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="597" masterDocId="FF9BFFF30048F709DF661C37FFB0FF8A" masterDocTitle="New species and records of Coryphiini from China (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="598" masterPageNumber="591" pageNumber="594" updateTime="1699048029042" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="91788B3BE123A30B394C52C17EC98E62">New species and records of Coryphiini from China (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="3270B7F31F3180C9885A15FF894C51A0">Shavrin, Alexey V.</mods:namePart>
<mods:affiliation id="ABA08DDEE76979F6B56128D25689DE4A">Institute of Life Sciences and Technologies, Daugavpils University, Vienibas 13, Daugavpils, LV- 5401, Latvia; e-mail: ashavrin @ hotmail. com</mods:affiliation>
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<mods:namePart id="D5FCB4A7D092745972C9E383F4E2B330">Smetana, Aleš</mods:namePart>
<mods:affiliation id="58AB3AB1F7FB5FA52BDB6C6EF9834B4F">Agriculture and Agri-Food Canada, Biodiversity, Central Experimental Farm, K. W. Neatby Bldg., Ottawa, ON K</mods:affiliation>
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<taxonomicName id="4C0B4D1E004BF70ADE6A1B30FD97F895" ID-CoL="8XWDK" authority="Shavrin &amp; Smetana, 2020" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[268,551,1799,1823]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="concisus" status="sp. nov.">
<emphasis id="B97FEA8F004BF70ADE6A1B30FD97F895" bold="true" box="[268,551,1799,1823]" italics="true" pageId="3" pageNumber="594">Caloboreaphilus concisus</emphasis>
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<paragraph id="8BB4369D004BF70ADEF71B11FE4EF8B3" blockId="3.[401,510,1830,1849]" box="[401,510,1830,1849]" pageId="3" pageNumber="594">
(
<figureCitation id="13302A18004BF70ADEF11B11FE48F8B3" box="[407,504,1830,1849]" captionStart-0="Figs 11–14" captionStart-1="Figs 15–27" captionStartId-0="3.[151,188,1479,1498]" captionStartId-1="4.[151,188,1520,1539]" captionTargetBox-0="[171,1416,196,1459]" captionTargetBox-1="[204,1415,208,1497]" captionTargetId-0="figure-13@3.[171,1417,189,1459]" captionTargetId-1="figure-220@4.[159,1429,189,1519]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs 11–14. Boreaphilus japonicus Sharp, 1874 (Figs 11–13) and map of distribution of Coryphiini in China (Fig. 14). 11 – habitus; 12 – labrum, dorsal view.13 – aedeagus, parameral view (internal sac is not shown); 14 – map: Archaeoboreaphilus paradoxus sp. nov. (black circle), Boreaphilus japonicus (white circles), Caloboreaphilus hammondi Zerche, 1990 (black star), C. concisus sp. nov. (white star), Coryphium taibaiense Li, Li &amp; Zhao, 2007 (black square), C. tangi Li, Li &amp; Zhao, 2007 (white square), Haida argonautarum Smetana, 2003 (black diamond), H. satoi Smetana, 2003 (white diamond). Scale bars: 1.0 mm (Fig. 11); 0.1 mm (Figs 12–13)." captionText-1="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi-0="http://doi.org/10.5281/zenodo.5177786" figureDoi-1="http://doi.org/10.5281/zenodo.5177788" httpUri-0="https://zenodo.org/record/5177786/files/figure.png" httpUri-1="https://zenodo.org/record/5177788/files/figure.png" pageId="3" pageNumber="594">Figs 14–27</figureCitation>
)
</paragraph>
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<typeStatus id="54B0883F004BF70ADFF11B67FF75F8E9" box="[151,197,1872,1891]" pageId="3" pageNumber="594">Type</typeStatus>
locality.
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<collectingCountry id="F31C760D004BF70ADE461B67FEE7F8E9" box="[288,343,1872,1891]" name="China" pageId="3" pageNumber="594">China</collectingCountry>
, 
<collectingRegion id="49CFF87F004BF70ADE051B67FE1DF8E9" box="[355,429,1872,1891]" country="China" name="Sichuan" pageId="3" pageNumber="594">Sichuan</collectingRegion>
, 
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E of Songpan, 32°37′47″N, 103°49′25″E, 
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a.s.l.
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Hඈඅඈඍඒඉൾ: J, ‘ 
<collectingCountry id="F31C760D004BF70ADECE1BB2FE43F812" box="[424,499,1925,1944]" name="China" pageId="3" pageNumber="594">CHINA</collectingCountry>
: 
<collectingRegion id="49CFF87F004BF70ADE9A1BB2FDF3F812" box="[508,579,1925,1944]" country="China" name="Sichuan" pageId="3" pageNumber="594">Sichuan</collectingRegion>
, 
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| E Songpan, 
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| 
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, 
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| 
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, sifting 11, | V. Grebennikov || 
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| 
<taxonomicName id="4C0B4D1E004BF70ADE381B8DFE5AF847" authorityName="Zerche" authorityYear="1990" box="[350,490,1978,1997]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">Caloboreaphilus</taxonomicName>
| concisus 
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| Shavrin A. &amp; Smetana A.des. 2020 [red rectangular label, printed]’ (
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). Pൺ*©ൺඍඒඉൾඌ: 2 JJ (
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dissected), same data as the 
<typeStatus id="54B0883F004BF70ADD5C1BC7FD35F789" box="[570,645,2032,2051]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
(1 J: 
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, 1 J: 
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); 
<specimenCount id="9D0DFD14004BF70ADE62143DFE95F797" box="[260,293,2052,2077]" pageId="3" pageNumber="594" type="female">1 ♀</specimenCount>
, ‘ 
<collectingCountry id="F31C760D004BF70ADE55143DFECCF797" box="[307,380,2058,2077]" name="China" pageId="3" pageNumber="594">CHINA</collectingCountry>
, 
<collectingRegion id="49CFF87F004BF70ADEE1143DFE7EF797" box="[391,462,2058,2077]" country="China" name="Sichuan" pageId="3" pageNumber="594">Sichuan</collectingRegion>
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| E Songpan, 
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| 
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, 
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, | 
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, sifting 10, | V. Grebennikov’ (
<collectionCode id="ED1AAE58004BF70ADA041AB1FA22F913" box="[1378,1426,1670,1689]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="594">CNC</collectionCode>
); 1 J, ‘ 
<collectingCountry id="F31C760D004BF70ADC001A97FC00F939" box="[870,944,1696,1715]" name="China" pageId="3" pageNumber="594">CHINA</collectingCountry>
: 
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| E Songpan, 
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| 
<geoCoordinate id="EE3F505A004BF70ADA461A97FA28F939" box="[1312,1432,1696,1715]" degrees="103" direction="east" minutes="49" orientation="longitude" pageId="3" pageNumber="594" precision="15" seconds="25" value="103.82361">E 103°49’25’’</geoCoordinate>
, 
<date id="FFB5105D004BF70ADC5A1A8CFC23F944" box="[828,915,1723,1742]" pageId="3" pageNumber="594" value="2012-05-28">28.V.2012</date>
| 
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, sifting 14. | V.Grebennikov’ (
<collectionCode id="ED1AAE58004BF70ADBBC1A8CFABAF944" box="[1242,1290,1723,1742]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="594">CNC</collectionCode>
); 1 J (dissected), 
<specimenCount id="9D0DFD14004BF70ADC5A1AE1FCD9F962" box="[828,873,1743,1769]" pageId="3" pageNumber="594" type="female">5 ♀♀</specimenCount>
, ‘ 
<collectingCountry id="F31C760D004BF70ADC131AE1FC0EF963" box="[885,958,1750,1769]" name="China" pageId="3" pageNumber="594">CHINA</collectingCountry>
, 
<collectingRegion id="49CFF87F004BF70ADCA31AE1FBBBF963" box="[965,1035,1750,1769]" country="China" name="Sichuan" pageId="3" pageNumber="594">Sichuan</collectingRegion>
, 
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| E Songpan, 
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| 
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, 
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, | 
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, sifting 12, | V. Grebennikov’ (
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: 
<collectionCode id="ED1AAE58004BF70ADA451AC7FAEEF889" box="[1315,1374,1776,1795]" pageId="3" pageNumber="594">ASCD</collectionCode>
, 1 J 
<specimenCount id="9D0DFD14004BF70ADAF71AC7FCEBF897" pageId="3" pageNumber="594" type="female">4 ♀♀</specimenCount>
: 
<collectionCode id="ED1AAE58004BF70ADC011B3CFC27F894" box="[871,919,1803,1822]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="594">CNC</collectionCode>
). All 
<typeStatus id="54B0883F004BF70ADCA31B3CFBA7F894" box="[965,1047,1803,1822]" pageId="3" pageNumber="594" type="paratype">paratypes</typeStatus>
with additional red rectangular printed label: ‘ 
<typeStatus id="54B0883F004BF70ADC591B11FC1DF8B3" box="[831,941,1830,1849]" pageId="3" pageNumber="594" type="paratype">PARATYPE</typeStatus>
| 
<taxonomicName id="4C0B4D1E004BF70ADCDA1B11FBF8F8B3" authorityName="Zerche" authorityYear="1990" box="[956,1096,1830,1849]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="genus">Caloboreaphilus</taxonomicName>
| concisus 
<taxonomicNameLabel id="A24C57F4004BF70ADBC31B11FB7EF8B3" box="[1189,1230,1830,1849]" pageId="3" pageNumber="594" rank="species">sp.n.</taxonomicNameLabel>
| Shavrin A. &amp; Smetana A. des. 2020’.
</paragraph>
</subSubSection>
<subSubSection id="C3116516004BF70FDC5A1B5BFD91F8BF" lastPageId="6" lastPageNumber="597" pageId="3" pageNumber="594" type="description">
<paragraph id="8BB4369D004BF70ADC5A1B5BFA26F7A9" blockId="3.[828,1437,1900,2083]" pageId="3" pageNumber="594">
<emphasis id="B97FEA8F004BF70ADC5A1B5BFC77F809" bold="true" box="[828,967,1900,1923]" pageId="3" pageNumber="594">Description.</emphasis>
Measurements (n = 11, in mm): HW: 0.75– 0.97; HL: 0.57–0.75; AL (
<typeStatus id="54B0883F004BF70ADB2D1BBBFB1FF829" box="[1099,1199,1932,1955]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
): 1.25; OL: 0.17–0.20; LT: 0.25–0.30; PL: 0.52–0.62; PW: 0.68–0.92; ESL: 0.93– 1.15; EW: 1.06–1.40; AW: 1.17–1.60; MTbL (
<typeStatus id="54B0883F004BF70ADA4B1BFBFA21F869" box="[1325,1425,1996,2019]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
): 0.82; MTrL (
<typeStatus id="54B0883F004BF70ADCAC1BDBFB9EF789" box="[970,1070,2028,2051]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
): 0.49 (MTrL 1–4: 0.32; MTrL 5: 0.17); AedL: 0.80–0.82; TL: 3.50–4.40 (
<typeStatus id="54B0883F004BF70ADB8C143BFAFDF7A9" box="[1258,1357,2060,2083]" pageId="3" pageNumber="594" type="holotype">holotype</typeStatus>
: 4.10).
</paragraph>
<caption id="DF746615004CF70DDFF119C7FB56F9D9" ID-DOI="http://doi.org/10.5281/zenodo.5177788" ID-Zenodo-Dep="5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="4" pageNumber="595" startId="4.[151,188,1520,1539]" targetBox="[204,1415,208,1497]" targetPageId="4">
<paragraph id="8BB4369D004CF70DDFF119C7FB56F9D9" blockId="4.[151,1437,1520,1619]" pageId="4" pageNumber="595">
Figs 15–27. 
<taxonomicName id="4C0B4D1E004CF70DDE6619C7FE6DF989" authority="Shavrin &amp; Smetana, 2020" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[256,477,1520,1539]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="concisus" status="sp. nov.">
<emphasis id="B97FEA8F004CF70DDE6619C7FE6DF989" box="[256,477,1520,1539]" italics="true" pageId="4" pageNumber="595">Caloboreaphilus concisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57F4004CF70DDE8419C7FD94F989" box="[482,548,1520,1539]" pageId="4" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27).
</paragraph>
</caption>
<paragraph id="8BB4369D004CF70DDFD31ABFFDBCF795" blockId="4.[151,760,1672,2079]" pageId="4" pageNumber="595">
Forebody reddish-brown, sometimes with darker indistinct spot in middle of head, paler basal margin of pronotum, lateral and basal margins of elytra; middle portion of head (ventrally), thoracic sclerites (ventrally) and abdomen brown, with yellow-brown paratergites and intersegmental membranes between tergites III–VI; mouthparts, legs and antennomeres yellow to yellow-brown. Body shiny, without microsculpture, except of scutellum with fine transverse meshes in middle and abdomen with dense, indistinct or distinct, transverse meshes between punctures. Body covered by dense semierect setae, sparser and longer on anterior portion of head and somewhat finer on abdomen. Habitus as in 
<figureCitation id="13302A18004CF70DDEDF143FFDB7F795" box="[441,519,2056,2079]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="4" pageNumber="595">Fig. 15</figureCitation>
.
</paragraph>
<paragraph id="8BB4369D004CF70CDC3C1ABFFB8AF7AB" blockId="4.[828,1437,1672,2079]" lastBlockId="5.[828,1437,1770,2081]" lastPageId="5" lastPageNumber="596" pageId="4" pageNumber="595">
Head large and convex, 1.2–1.3 times as broad as long, slightly broader than pronotum, with evenly elevated middle portion and distinctly elevated supra-antennal prominences; basal portions of clypeus with very deep narrow impressions; mediobasal margin of head deeply, subtriangularly or semicircularly concave. Punctation very dense and moderately deep, sparser in medioanterior portion, denser in middle and around eyes. Temples distinctly longer than length of eyes, convex. Ocelli large, convex, situated significantly behind level of posterior margins of eyes, distance between ocelli about twice as long as distance between ocellus and posterior margin of eye. Eyes small, strongly convex. Neck with dense and fine microreticulati- on, without visible punctures. Labrum transverse, with three latero-apical and two apical long setae (
<figureCitation id="13302A18004DF70CDD551B3CFD30F8A8" box="[563,640,1803,1826]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="5" pageNumber="596">Fig. 16</figureCitation>
). Mandible as in 
<figureCitation id="13302A18004DF70CDFAD1B1CFEA6F8C8" box="[203,278,1835,1858]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="5" pageNumber="596">Fig. 18</figureCitation>
. Mentum large, trapezoidal, with rounded laterobasal margins, strongly narrowed apicad; labium narrow, with elongate labial palpomeres, apical palpomere narrow, about as long as preapical palpomere, glossa narrow, strongly protruded apicad (
<figureCitation id="13302A18004DF70CDE1D1B9CFE77F848" box="[379,455,1963,1986]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="5" pageNumber="596">Fig. 17</figureCitation>
). Preapical palpomere of maxillary palp wide, club-like, slightly longer than preceding palpomere; galea moderately narrow; lacinia moderately narrow, slightly longer than galea, with elongate spine- like apex, five long preapical spines and long setae along medial margin (
<figureCitation id="13302A18004DF70CDC841B3DFB80F8AB" box="[994,1072,1802,1825]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="5" pageNumber="596">Fig. 19</figureCitation>
). Antenna exceeding basal third of elytra when reclined, with shortened antennomeres IV–IX; antennomeres IV–X covered by dense pubescence; basal antennomere wide, 1.6 times as long as II, antennomere II ovoid, III about as long as and indistinctly narrower than II, IV short, about as long as broad, V–VI slightly broader than IV, VII–VIII slightly transverse, IX–X indistinctly longer than VIII, apical antennomere about twice as long as X, with conical apex.
</paragraph>
<caption id="DF746615004DF70CDFF11A0EFEBBF916" ID-DOI="http://doi.org/10.5281/zenodo.5177790" ID-Zenodo-Dep="5177790" httpUri="https://zenodo.org/record/5177790/files/figure.png" pageId="5" pageNumber="596" startId="5.[151,188,1593,1612]" targetBox="[153,1431,199,1560]" targetPageId="5">
<paragraph id="8BB4369D004DF70CDFF11A0EFEBBF916" blockId="5.[151,1436,1593,1692]" pageId="5" pageNumber="596">
Figs 28–37. 
<taxonomicName id="4C0B4D1E004DF70CDE641A0EFDC5F9C6" authority="Li, Li &amp; Zhao, 2007" authorityName="Li, Li &amp; Zhao" authorityYear="2007" box="[258,629,1593,1612]" class="Insecta" family="Staphylinidae" genus="Coryphium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="596" phylum="Arthropoda" rank="species" species="taibaiense">
<emphasis id="B97FEA8F004DF70CDE641A0EFE0FF9C6" box="[258,447,1593,1612]" italics="true" pageId="5" pageNumber="596">Coryphium taibaiense</emphasis>
<bibRefCitation id="EF9A4B6C004DF70CDEA21A0EFDC5F9C6" author="LI J. - W. &amp; LI L. - Z. &amp; ZHAO M. - J." box="[452,629,1593,1612]" pageId="5" pageNumber="596" pagination="89 - 93" refId="ref5452" refString="LI J. - W., LI L. - Z. &amp; ZHAO M. - J. 2007: A review on the genus Coryphium Stephens (Coleoptera, Staphylinidae) of China. Mitteilungen aus dem Museum fur Naturkunde Berlin. Deutsche Entomologische Zeitschrift, Neue Folge 54: 89 - 93." type="journal article" year="2007">Li, Li &amp; Zhao, 2007</bibRefCitation>
</taxonomicName>
. 28 – habitus (paratype); 29 – labrum, dorsal view; 30 – labium and mentum, ventral view; 31 – left mandible, dorsal view; 32 – left maxilla, ventral view; 33 – aedeagus, parameral view; 34 – aedeagus, lateral view; 35 – male abdominal tergite VIII, dorsal view; 36 – male abdominal sternite IX, ventral view; 37 – male abdominal sternite VIII, ventral view. Scale bars: 1.0 mm (Fig. 28); 0.1 mm (Figs 29–37).
</paragraph>
</caption>
<paragraph id="8BB4369D004EF70FDFD31C89FD49FD1F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">Pronotum 1.3–1.4 times as broad as long, significantly convex in middle, with deep and wide medial impression, stretching from about apical margin toward middle, and with moderately deep transverse or semicircular impression in mediobasal third, mediolateral portions with very wide and deep depressions; pronotum widest at anterior third, significantly narrowed posteriad toward widely rounded hind angles; latero-apical portions widely rounded; apical margin rounded, distinctly narower than widely rounded posterior margin; lateral edges with regular rounded crenulation, distinctly larger on latero-apical and finer on latero-basal margins. Punctation denser and coarser than that on middle of head, sometimes more irregular and sparser on bottom of medioapical and mediobasal impressions, distance between punctures in middle usually about length of nearest puncture.</paragraph>
<paragraph id="8BB4369D004EF70FDFD31EA9FD49FD5F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">Scutellum large, with rounded apex and indistinct semicircular impression in middle, with several small punctures.</paragraph>
<paragraph id="8BB4369D004EF70FDFD31EE9FEBCFC7F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">Elytra convex, short, 1.7–1.8 times as long as pronotum, about as long as wide or slightly wider, gradually widened apicad, reaching apical margin of abdominal tergite IV or basal margin of abdominal tergite V; apical margin straight; lateral portions narrowly flattened, with bordered margins; laterobasal margins with fine acute crenulation. Punctation slightly sparser and larger than that on pronotum, denser and coarser in basal portions near scutellum. Wings fully developed.</paragraph>
<paragraph id="8BB4369D004EF70FDFD31FC9FD47FBBF" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">Metatarsus about 1.6 times as long as metatibia; apical metatarsomere about as long three preceding tarsomeres.</paragraph>
<paragraph id="8BB4369D004EF70FDFD31809FDFDFB5F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">Abdomen distinctly wider than elytra, with fine and dense punctation, with wide transverse wing-folding spots in middle of tergite IV and small, rounded, sometimes indistinct spots in middle of tergite V; apical margin of tergite VII with very fine palisade fringe.</paragraph>
<paragraph id="8BB4369D004EF70FDFD318EAFD69FA7F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFD318EAFF44FB7F" bold="true" box="[181,244,1245,1269]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFD318EAFF5CFB7F" bold="true" box="[181,236,1245,1269]" italics="true" pageId="6" pageNumber="597">Male</emphasis>
.
</emphasis>
First four protarsomeres slightly widened. Apical margins of abdominal tergite VIII (
<figureCitation id="13302A18004EF70FDE9C18C9FDF4FA9F" box="[506,580,1278,1301]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 22</figureCitation>
) and sternite VIII (
<figureCitation id="13302A18004EF70FDFF91929FF58FABF" box="[159,232,1310,1333]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 24</figureCitation>
) sinuate.Abdominal sternite IX as in 
<figureCitation id="13302A18004EF70FDD001929FD01FABF" box="[614,689,1310,1333]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 23</figureCitation>
. Aedeagus (
<figureCitation id="13302A18004EF70FDFB31909FE91FADF" box="[213,289,1342,1365]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 20</figureCitation>
) long, with median lobe gradually narrowing toward acute apex; parameres narrow, cylindrical, reaching apex of median lobe, with two minute apical and three fine lateral setae along inner edge; internal sac long, moderately wide, with apical field of moderately large and two basal fields of finer spiculae. Aedeagus laterally as in 
<figureCitation id="13302A18004EF70FDDEE19E9FD65FA7F" box="[648,725,1502,1525]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 21</figureCitation>
.
</paragraph>
<paragraph id="8BB4369D004EF70FDFD319CAFD91F8BF" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFD319CAFEBDF99F" bold="true" box="[181,269,1533,1557]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFD319CAFEB6F99F" bold="true" box="[181,262,1533,1557]" italics="true" pageId="6" pageNumber="597">Female</emphasis>
.
</emphasis>
First four protarsomeres narrow.Apical margins of abdominal tergite VIII (
<figureCitation id="13302A18004EF70FDED41A29FE4FF9BF" box="[434,511,1566,1589]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 25</figureCitation>
) and sternite VIII (
<figureCitation id="13302A18004EF70FDDAB1A29FF03F9DF" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 26</figureCitation>
) obtusely triangular. Genital segment (
<figureCitation id="13302A18004EF70FDD261A09FD39F9DF" box="[576,649,1598,1621]" captionStart="Figs 15–27" captionStartId="4.[151,188,1520,1539]" captionTargetBox="[204,1415,208,1497]" captionTargetId="figure-220@4.[159,1429,189,1519]" captionTargetPageId="4" captionText="Figs 15–27. Caloboreaphilus concisus sp. nov. 15 – habitus (paratype); 16 – labrum, dorsal view; 17 – labium and mentum, ventral view; 18 – left mandible, dorsal view; 19 – left maxilla, ventral view; 20 – aedeagus, parameral view; 21 – aedeagus, lateral view; 22 – male abdominal tergite VIII, dorsal view; 23 – male abdominal sternite IX, ventral view; 24 – male abdominal sternite VIII, ventral view; 25 – female abdominal tergite VIII, dorsal view; 26 – female abdominal sternite IX; 27 – female genital segment, dorsal view. Scale bars: 1.0 mm (Fig. 15); 0.1 mm (Figs 16–27)." figureDoi="http://doi.org/10.5281/zenodo.5177788" httpUri="https://zenodo.org/record/5177788/files/figure.png" pageId="6" pageNumber="597">Fig. 27</figureCitation>
) with long gonocoxites and elongate styli, each with long apical seta. 
<emphasis id="B97FEA8F004EF70FDFF11A49FE3DF91F" bold="true" box="[151,397,1662,1685]" pageId="6" pageNumber="597">Differential diagnosis.</emphasis>
<taxonomicName id="4C0B4D1E004EF70FDEF71A49FD12F91F" authority="Shavrin &amp; Smetana, 2020" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[401,674,1662,1685]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="concisus" status="sp. nov.">
<emphasis id="B97FEA8F004EF70FDEF71A49FD12F91F" box="[401,674,1662,1685]" italics="true" pageId="6" pageNumber="597">Caloboreaphilus concisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C57F4004EF70FDDCE1A49FD4AF91F" box="[680,762,1662,1685]" pageId="6" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from 
<taxonomicName id="4C0B4D1E004EF70FDE811AA9FDC3F93F" authorityName="Zerche" authorityYear="1990" box="[487,627,1694,1717]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="B97FEA8F004EF70FDE811AA9FDC3F93F" box="[487,627,1694,1717]" italics="true" pageId="6" pageNumber="597">C. hammondi</emphasis>
</taxonomicName>
by the larger body, wider head, larger crenulation of lateral sides of the pronotum, shorter elytra, wider abdomen, the shape of the narrower median lobe (asymmetrical in 
<taxonomicName id="4C0B4D1E004EF70FDD511AC9FD74F89F" authorityName="Zerche" authorityYear="1990" box="[567,708,1790,1813]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="hammondi">
<emphasis id="B97FEA8F004EF70FDD511AC9FD74F89F" box="[567,708,1790,1813]" italics="true" pageId="6" pageNumber="597">C. hammondi</emphasis>
</taxonomicName>
) and different structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="C3116516004EF70FDFF11B09FD8EF85F" pageId="6" pageNumber="597" type="etymology">
<paragraph id="8BB4369D004EF70FDFF11B09FEBBF81F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFF11B09FEAAF8DF" bold="true" box="[151,282,1854,1877]" pageId="6" pageNumber="597">Etymology.</emphasis>
The specific epithet is the Latin adjective 
<taxonomicName id="4C0B4D1E004EF70FDFF11B69FF43F8FF" authorityName="Shavrin &amp; Smetana" authorityYear="2020" box="[151,243,1886,1909]" class="Insecta" family="Staphylinidae" genus="Caloboreaphilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="597" phylum="Arthropoda" rank="species" species="concisus">
<emphasis id="B97FEA8F004EF70FDFF11B69FF43F8FF" box="[151,243,1886,1909]" italics="true" pageId="6" pageNumber="597">concisus</emphasis>
</taxonomicName>
, - 
<emphasis id="B97FEA8F004EF70FDE6A1B69FEA9F8FF" box="[268,281,1886,1909]" italics="true" pageId="6" pageNumber="597">a</emphasis>
, - 
<emphasis id="B97FEA8F004EF70FDE541B69FEE3F8FF" box="[306,339,1886,1909]" italics="true" pageId="6" pageNumber="597">um</emphasis>
(short). It refers to the short elytra of the species
</paragraph>
<paragraph id="8BB4369D004EF70FDFF11BA9FD8EF85F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFF11BA9FF46F83F" bold="true" box="[151,246,1950,1973]" pageId="6" pageNumber="597">Habitat.</emphasis>
Specimens were collected by sifting forest litter at elevations from 
<quantity id="4CF39B78004EF70FDE381B89FDB7F85F" box="[350,519,1982,2005]" metricMagnitude="3" metricUnit="m" metricValue="3.8105" metricValueMax="3.86" metricValueMin="3.761" pageId="6" pageNumber="597" unit="m" value="3810.5" valueMax="3860.0" valueMin="3761.0">3761 to 3860 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C3116516004EF70FDFF11BE9FD5FF79F" pageId="6" pageNumber="597" type="distribution">
<paragraph id="8BB4369D004EF70FDFF11BE9FD5FF79F" blockId="6.[151,762,190,2069]" pageId="6" pageNumber="597">
<emphasis id="B97FEA8F004EF70FDFF11BE9FE99F87F" bold="true" box="[151,297,2014,2037]" pageId="6" pageNumber="597">Distribution.</emphasis>
The species is known from several locations close to each other in northern 
<collectingRegion id="49CFF87F004EF70FDE871BC9FD8AF79F" box="[481,570,2046,2069]" country="China" name="Sichuan" pageId="6" pageNumber="597">Sichuan</collectingRegion>
, 
<collectingCountry id="F31C760D004EF70FDD231BC9FD35F79F" box="[581,645,2046,2069]" name="China" pageId="6" pageNumber="597">China</collectingCountry>
(
<figureCitation id="13302A18004EF70FDDF51BC9FD53F79F" box="[659,739,2046,2069]" captionStart="Figs 11–14" captionStartId="3.[151,188,1479,1498]" captionTargetBox="[171,1416,196,1459]" captionTargetId="figure-13@3.[171,1417,189,1459]" captionTargetPageId="3" captionText="Figs 11–14. Boreaphilus japonicus Sharp, 1874 (Figs 11–13) and map of distribution of Coryphiini in China (Fig. 14). 11 – habitus; 12 – labrum, dorsal view.13 – aedeagus, parameral view (internal sac is not shown); 14 – map: Archaeoboreaphilus paradoxus sp. nov. (black circle), Boreaphilus japonicus (white circles), Caloboreaphilus hammondi Zerche, 1990 (black star), C. concisus sp. nov. (white star), Coryphium taibaiense Li, Li &amp; Zhao, 2007 (black square), C. tangi Li, Li &amp; Zhao, 2007 (white square), Haida argonautarum Smetana, 2003 (black diamond), H. satoi Smetana, 2003 (white diamond). Scale bars: 1.0 mm (Fig. 11); 0.1 mm (Figs 12–13)." figureDoi="http://doi.org/10.5281/zenodo.5177786" httpUri="https://zenodo.org/record/5177786/files/figure.png" pageId="6" pageNumber="597">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>