231 lines
26 KiB
XML
231 lines
26 KiB
XML
<document id="8CF6A9558BD10474CF900D25F6284653" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. & Sikes, Derek S." docDate="2018" docId="03971078FFDFBC74FD77FA18FCDECDEF" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus smetanai Mullen and Campbell 2018, new species" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="48" updateTime="1699007718751" updateUser="plazi">
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<mods:titleInfo id="15977B18F82A392A559A7D81EE76A33A">
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<mods:title id="8D59DE54E498336ADBEA595D0CA1D785">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:name id="BA6E717187E7B35FE1B85B0F245A4000" type="personal">
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<mods:roleTerm id="96CF66FB100DE28BB9B6BA851D87B0F7">Author</mods:roleTerm>
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<mods:namePart id="7ACCF86B10FF4D913D8D1497BFAC9C7F">Mullen, Logan J.</mods:namePart>
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<mods:roleTerm id="B6C65E97F4D34848DC4167C46B98FB89">Author</mods:roleTerm>
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<mods:namePart id="EE698F5F473691DE6606B400BCFB0F0D">Campbell, J. M.</mods:namePart>
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<mods:name id="FFA666DEA388360B9C247CEB3504609A" type="personal">
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<mods:roleTerm id="27D831EB77A7F7DD0A6E3040875723B2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="651EC3EE1E285F5DFE9BEEC90FB3F2BA">Sikes, Derek S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DAEEB2D182C5599348459F09811126A9">text</mods:typeOfResource>
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<mods:titleInfo id="D53580819A376731DCB8F51661AF7B1F">
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<mods:title id="A58C0264CFDC32042FDD164A733E6BBE">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="CAC09F9CBFEB68A9DA8587A2BCB54837">
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<mods:date id="455F378FAFCFC51D0E6FB1561952BCAF">2018</mods:date>
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<mods:detail id="732FA4EAA105A0EF41D1AF1ECBCEE3DE" type="pubDate">
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<mods:number id="4F8E1C617D0FC74C5FD2C9EBF68D9AB6">2018-12-28</mods:number>
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<mods:detail id="4F29A174475B294682D5E0D7CD060899" type="volume">
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<mods:number id="FED3FE358AE83E27D2F8BC67F7D2B12B">72</mods:number>
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<mods:start id="F8C46B9545CDF3BA4D224400C44BF951">1</mods:start>
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<mods:url id="9CA0E4B71CB7070D1AA67597CC146709">http://dx.doi.org/10.1649/0010-065x-72.mo4.1</mods:url>
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</mods:location>
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<mods:classification id="561E7D89B70EA977073FBFAC901DF809">journal article</mods:classification>
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<mods:identifier id="846D950D5E31E9691D90770CA36EE039" type="DOI">10.1649/0010-065X-72.mo4.1</mods:identifier>
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<mods:identifier id="1A6A62EFF16B2E17B502CE55DD8EC3D8" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03971078FFDFBC74FD77FA18FCDECDEF" LSID="urn:lsid:plazi:treatment:03971078FFDFBC74FD77FA18FCDECDEF" httpUri="http://treatment.plazi.org/id/03971078FFDFBC74FD77FA18FCDECDEF" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
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<subSubSection id="C324F2E5FFDFBC6BFD77FA18FC66CBD8" pageId="47" pageNumber="48" type="nomenclature">
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<paragraph id="8B81A16EFFDFBC6BFD77FA18FC66CBD8" blockId="47.[685,1172,1470,1598]" pageId="47" pageNumber="48">
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<heading id="D0C91602FFDFBC6BFD77FA18FC66CBD8" centered="true" fontSize="10" level="2" pageId="47" pageNumber="48" reason="2">
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<emphasis id="B94A7D7CFFDFBC6BFD77FA18FC66CBD8" bold="true" pageId="47" pageNumber="48">
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18.
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<taxonomicName id="4C3EDAEDFFDFBC6BFD0AFA18FB32CBE5" ID-CoL="8Y8D8" authority="Mullen and Campbell" authorityName="Mullen and Campbell" authorityYear="2018" box="[720,1167,1470,1491]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
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<emphasis id="B94A7D7CFFDFBC6BFD0AFA18FC12CBE5" bold="true" box="[720,943,1470,1491]" italics="true" pageId="47" pageNumber="48">Phlaeopterus smetanai</emphasis>
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Mullen and Campbell
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</taxonomicName>
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,
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<taxonomicNameLabel id="A279C007FFDFBC6BFCBCFA7FFC66CBD8" box="[870,987,1497,1518]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFDFBC6BFD2DFA55FB38C808" pageId="47" pageNumber="48" type="description">
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||
<paragraph id="8B81A16EFFDFBC6BFD2DFA55FB38C808" blockId="47.[685,1172,1470,1598]" pageId="47" pageNumber="48">
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(
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<figureCitation id="1305BDEBFFDFBC6BFD24FA55FCEFC83E" box="[766,850,1523,1544]" captionStart="Fig" captionStartId="14.[197,228,957,976]" captionTargetBox="[177,1188,196,940]" captionTargetId="figure-363@14.[175,1190,194,942]" captionTargetPageId="14" captionText="Fig. 5. Phlaeopterus species, habitus. A) P. olympicus, female (UAMObs:Ento:232960), B) P. smetanai, male" figureDoi="http://doi.org/10.5281/zenodo.5102853" httpUri="https://zenodo.org/record/5102853/files/figure.png" pageId="47" pageNumber="48">Figs. 5B</figureCitation>
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,
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<figureCitation id="1305BDEBFFDFBC6BFC84FA55FC34C83E" box="[862,905,1523,1544]" captionStart="Fig" captionStartId="19.[709,740,1028,1047]" captionTargetBox="[155,1169,195,1011]" captionTargetPageId="19" captionText="Fig. 14. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. olympicus, B) P. smetanai. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." figureDoi="http://doi.org/10.5281/zenodo.5102863" httpUri="https://zenodo.org/record/5102863/files/figure.png" pageId="47" pageNumber="48">14B</figureCitation>
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, 16E, 23B–C, 33H) Zoobank.org/
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<uri id="FFAFAD6CFFDFBC6BFCAAF9A8FB38C808" pageId="47" pageNumber="48">
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urn:lsid:zoobank.org:act:
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<uuid id="FF989BBBFFDFBC6BFD66F98FFB38C808" box="[700,1157,1577,1598]" pageId="47" pageNumber="48">B4AB0AFD-A8C3-4922-8957-C875B57264E8</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFDFBC6BFD1FF9FBFCABC8BB" pageId="47" pageNumber="48" type="materials_examined">
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<paragraph id="8B81A16EFFDFBC6BFD1FF9FBFCABC8BB" blockId="47.[685,1173,1629,1730]" pageId="47" pageNumber="48">
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<materialsCitation id="3B56AB33FFDFBC6BFD1FF9FBFCAFC8BB" country="United States of America" location="Mt. Hood National Forest" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
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<emphasis id="B94A7D7CFFDFBC6BFD1FF9FBFCEAC844" bold="true" box="[709,855,1629,1650]" pageId="47" pageNumber="48">
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<typeStatus id="54851FCCFFDFBC6BFD1FF9FBFD4AC844" box="[709,759,1629,1650]" pageId="47" pageNumber="48">Type</typeStatus>
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Locality.
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</emphasis>
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<location id="8EE1F7B5FFDFBC6BFC85F9FBFBE1C844" LSID="urn:lsid:plazi:treatment:03971078FFDFBC74FD77FA18FCDECDEF:8EE1F7B5FFDFBC6BFC85F9FBFBE1C844" box="[863,1116,1629,1650]" country="United States of America" name="Mt. Hood National Forest" pageId="47" pageNumber="48" stateProvince="Oregon">Mt. Hood National Forest</location>
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,
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<collectingRegion id="49FA6F8CFFDFBC6BFBB3F9FBFD6EC8BB" country="United States of America" name="Oregon" pageId="47" pageNumber="48">Oregon</collectingRegion>
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,
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<collectingCountry id="F329E1FEFFDFBC6BFD05F9DEFCAFC8BB" box="[735,786,1656,1677]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFDFBC74FD1FF934FEA0CBE4" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
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<paragraph id="8B81A16EFFDFBC6BFD1FF934FB2EC8F4" blockId="47.[685,1173,1629,1730]" pageId="47" pageNumber="48">
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<emphasis id="B94A7D7CFFDFBC6BFD1FF934FC14C891" bold="true" box="[709,937,1682,1703]" pageId="47" pageNumber="48">Description. Habitus:</emphasis>
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Length 6.0–9.0 mm. Black, usually darker than any other
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<taxonomicName id="4C3EDAEDFFDFBC6BFBCFF90BFB2EC8F4" box="[1045,1171,1709,1730]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B94A7D7CFFDFBC6BFBCFF90BFB2EC8F4" box="[1045,1171,1709,1730]" italics="true" pageId="47" pageNumber="48">Phlaeopterus</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="8B81A16EFFC0BC74FF77FF64FEA0CBE4" blockId="48.[173,660,194,1729]" pageId="48" pageNumber="49">
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species; legs, palpi, and antennae sometimes lighter, dark brown (
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<figureCitation id="1305BDEBFFC0BC74FEF2FF7AFEC9CEC7" box="[296,372,220,241]" captionStart="Fig" captionStartId="14.[197,228,957,976]" captionTargetBox="[177,1188,196,940]" captionTargetId="figure-363@14.[175,1190,194,942]" captionTargetPageId="14" captionText="Fig. 5. Phlaeopterus species, habitus. A) P. olympicus, female (UAMObs:Ento:232960), B) P. smetanai, male" figureDoi="http://doi.org/10.5281/zenodo.5102853" httpUri="https://zenodo.org/record/5102853/files/figure.png" pageId="48" pageNumber="49">Fig. 5B</figureCitation>
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).
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<emphasis id="B94A7D7CFFC0BC74FE5FFF7AFE7ECEC7" bold="true" box="[389,451,220,241]" pageId="48" pageNumber="49">Head:</emphasis>
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Broad, ratio of width across eyes to head length about 5:4. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5–10 at least 2 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Epipharynx as in Fig. 33H. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3
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<superScript id="7C4B0C26FFC0BC74FE19FE5DFE6FCC3F" attach="left" box="[451,466,507,521]" fontSize="6" pageId="48" pageNumber="49">rd</superScript>
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palpomere 1.6–1.8 times longer than palpomere 2.
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<emphasis id="B94A7D7CFFC0BC74FE2BFDBDFDFACC06" bold="true" box="[497,583,539,560]" pageId="48" pageNumber="49">Thorax:</emphasis>
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Pronotum somewhat broad (Fig. 16E), length to width ratio = 0.63–0.77; ratio of pronotal width to head width = 1.49–1.67; maximum width subequal to elytral width at humeral angles; punctures on dorsal surface separated by average distance slightly greater than diameter of a puncture; lateral margins explanate posterad lateral foveae, progressively more narrowly explanate from lateral foveae to just before apex; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.3–2.6 times longer than pronotum; apical margins broadly convex. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth, tooth more strongly notched on posterior margin than on anterior margin; longitudinal carina along midline of mesosternum complete.
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<emphasis id="B94A7D7CFFC0BC74FE64FC5FFE45CA38" bold="true" box="[446,504,1017,1038]" pageId="48" pageNumber="49">Legs:</emphasis>
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All tibiae with dense pubescence to apex or with small glabrous region at apex, if with glabrous region, length of glabrous apex of mesotibia shorter than length of basal mesotarsomere. Metatrochanter without tooth on apical margin.
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<emphasis id="B94A7D7CFFC0BC74FE43FBD8FDBBCAA5" bold="true" box="[409,518,1150,1171]" pageId="48" pageNumber="49">Abdomen:</emphasis>
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Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wingfolding patches.
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<emphasis id="B94A7D7CFFC0BC74FE90FB68FE0FCAD5" bold="true" box="[330,434,1230,1251]" pageId="48" pageNumber="49">Aedeagus:</emphasis>
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Length
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<quantity id="4CC60C8BFFC0BC74FDD9FB68FD2CCAD5" box="[515,657,1229,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.67" metricValueMin="1.28" pageId="48" pageNumber="49" unit="mm" value="1.475" valueMax="1.67" valueMin="1.28">1.28–1.67 mm</quantity>
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. Median lobe roughly triangular, evenly to convexly narrowed from base to apex. Parameres slightly diverging from base to near apex then converging; ending just beyond to distinctly beyond apex of median lobe (
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<figureCitation id="1305BDEBFFC0BC74FE67FAF4FD89CB51" box="[445,564,1362,1384]" captionStart="Fig" captionStartId="28.[197,228,1491,1510]" captionTargetBox="[177,1188,196,1463]" captionTargetId="figure-70@28.[175,1190,194,1475]" captionTargetPageId="28" captionText="Fig. 23. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. bakerensis, B) P. smetanai, C) P. smetanai, D) P. occidentalis." figureDoi="http://doi.org/10.5281/zenodo.5102877" httpUri="https://zenodo.org/record/5102877/files/figure.png" pageId="48" pageNumber="49">Fig. 23B–C</figureCitation>
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). Internal sac elongate, roughly rectangular, narrowed from base to apex; apex subtruncate; lightly sclerotized; more densely covered in microspinules on apical half.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C324F2E5FFC0BC74FF1FFA71FC3ACEC7" pageId="48" pageNumber="49" type="materials_examined">
|
||
<paragraph id="8B81A16EFFC0BC74FF1FFA71FC3ACEC7" blockId="48.[173,660,194,1729]" lastBlockId="48.[707,1194,194,985]" pageId="48" pageNumber="49">
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||
<emphasis id="B94A7D7CFFC0BC74FF1FFA71FEC8CBDA" bold="true" box="[197,373,1495,1516]" pageId="48" pageNumber="49">Type Specimens.</emphasis>
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||
<materialsCitation id="3B56AB33FFC0BC74FE5EFA71FEB5C861" collectingDate="1979-07-31" collectorName="Mt. Hood N. F. & Tilly Jane Cr. & J. M. & B. A. Campbell" location="Mt. Hood" pageId="48" pageNumber="49" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="54851FCCFFC0BC74FE5EFA71FE61CBDA" box="[388,476,1495,1516]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
|
||
male (UAMObs: Ento:235828) and
|
||
<typeStatus id="54851FCCFFC0BC74FE80FA54FE16C831" box="[346,427,1522,1543]" pageId="48" pageNumber="49" type="paratype">paratype</typeStatus>
|
||
female (UAMObs:Ento: 235829) labeled as follows: ORE.,
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||
<collectorName id="26CBC4B8FFC0BC74FE21F9AAFD32C817" box="[507,655,1548,1569]" pageId="48" pageNumber="49">
|
||
<location id="8EE1F7B5FFC0BC74FE21F9AAFDE5C817" LSID="urn:lsid:plazi:treatment:03971078FFDFBC74FD77FA18FCDECDEF:8EE1F7B5FFC0BC74FE21F9AAFDE5C817" box="[507,600,1548,1569]" name="Mt. Hood" pageId="48" pageNumber="49">Mt. Hood</location>
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N. F.
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||
</collectorName>
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,
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||
<collectorName id="26CBC4B8FFC0BC74FF77F981FE88C80A" box="[173,309,1575,1596]" pageId="48" pageNumber="49">Tilly Jane Cr.</collectorName>
|
||
, 7000’
|
||
<date id="FF8087AEFFC0BC74FE58F981FE45C80A" box="[386,504,1575,1596]" pageId="48" pageNumber="49" value="1979-07-31">
|
||
<collectingDate id="EFC47E46FFC0BC74FE58F981FE45C80A" box="[386,504,1575,1596]" pageId="48" pageNumber="49" value="1979-07-31">31.VII.1979</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26CBC4B8FFC0BC74FDDFF981FEB5C861" pageId="48" pageNumber="49">J. M. & B. A. Campbell</collectorName>
|
||
</materialsCitation>
|
||
/
|
||
<typeStatus id="54851FCCFFC0BC74FEC5F9E4FE21C861" box="[287,412,1602,1623]" pageId="48" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
|
||
³ (or
|
||
<typeStatus id="54851FCCFFC0BC74FE2EF9E4FDD3C861" box="[500,622,1602,1623]" pageId="48" pageNumber="49" type="allotype">ALLOTYPE</typeStatus>
|
||
<specimenCount id="9D386AE7FFC0BC74FDA4F9E2FD36C860" box="[638,651,1604,1622]" pageId="48" pageNumber="49" type="female">♀</specimenCount>
|
||
)
|
||
<taxonomicName id="4C3EDAEDFFC0BC74FF77F9FAFE2DC847" authorityName="Mullen and Campbell" authorityYear="2018" box="[173,400,1628,1649]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">
|
||
<emphasis id="B94A7D7CFFC0BC74FF77F9FAFE2DC847" box="[173,400,1628,1649]" italics="true" pageId="48" pageNumber="49">Phlaeopterus smetanai</emphasis>
|
||
</taxonomicName>
|
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, desig. L.J. Mullen and J. M. Campbell CNC No. 18465 (red label). Both specimens are in the CNC.
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<typeStatus id="54851FCCFFC0BC74FE1FF937FD9CC890" box="[453,545,1681,1702]" pageId="48" pageNumber="49" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<emphasis id="B94A7D7CFFC0BC74FDE9F937FD82C890" box="[563,575,1681,1702]" italics="true" pageId="48" pageNumber="49">n</emphasis>
|
||
= 504) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, NHMUK, OSAC, RBC, ROM, UCRC, and USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFC0BC74FD01FF51FC5ECC23" pageId="48" pageNumber="49" type="distribution">
|
||
<paragraph id="8B81A16EFFC0BC74FD01FF51FBA4CF6D" blockId="48.[707,1194,194,985]" pageId="48" pageNumber="49">
|
||
<emphasis id="B94A7D7CFFC0BC74FD01FF51FCE1CF3A" bold="true" box="[731,860,247,268]" pageId="48" pageNumber="49">Distribution.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFC0BC74FCBEFF51FBFCCF3A" authorityName="Mullen and Campbell" authorityYear="2018" box="[868,1089,247,268]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">
|
||
<emphasis id="B94A7D7CFFC0BC74FCBEFF51FBFCCF3A" box="[868,1089,247,268]" italics="true" pageId="48" pageNumber="49">Phlaeopterus smetanai</emphasis>
|
||
</taxonomicName>
|
||
is broadly distributed in the northwestern
|
||
<collectingCountry id="F329E1FEFFC0BC74FC21FEB7FB96CF10" box="[1019,1067,273,294]" name="United States of America" pageId="48" pageNumber="49">USA</collectingCountry>
|
||
from
|
||
<collectingRegion id="49FA6F8CFFC0BC74FBB7FEB7FCBDCF77" country="United States of America" name="Washington" pageId="48" pageNumber="49">Washington</collectingRegion>
|
||
east to
|
||
<collectingRegion id="49FA6F8CFFC0BC74FC9EFE8AFCC7CF77" box="[836,890,300,321]" country="United States of America" name="Idaho" pageId="48" pageNumber="49">Idaho</collectingRegion>
|
||
and south to
|
||
<collectingRegion id="49FA6F8CFFC0BC74FC2EFE8AFBF2CF77" box="[1012,1103,300,321]" country="United States of America" name="Colorado" pageId="48" pageNumber="49">Colorado</collectingRegion>
|
||
,
|
||
<collectingRegion id="49FA6F8CFFC0BC74FB8DFE8AFB1BCF77" box="[1111,1190,300,321]" country="United States of America" name="Arizona" pageId="48" pageNumber="49">Arizona</collectingRegion>
|
||
, and southern
|
||
<collectingRegion id="49FA6F8CFFC0BC74FC9CFEE0FC18CF6D" box="[838,933,326,347]" country="United States of America" name="California" pageId="48" pageNumber="49">California</collectingRegion>
|
||
(
|
||
<figureCitation id="1305BDEBFFC0BC74FC6EFEE0FBB0CF6D" box="[948,1037,326,347]" captionStart="Fig" captionStartId="19.[709,740,1028,1047]" captionTargetBox="[155,1169,195,1011]" captionTargetPageId="19" captionText="Fig. 14. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. olympicus, B) P. smetanai. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." figureDoi="http://doi.org/10.5281/zenodo.5102863" httpUri="https://zenodo.org/record/5102863/files/figure.png" pageId="48" pageNumber="49">Fig. 14B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFC0BC74FD01FEC7FC5ECC23" blockId="48.[707,1194,194,985]" pageId="48" pageNumber="49">
|
||
<emphasis id="B94A7D7CFFC0BC74FD01FEC7FCF7CF40" bold="true" box="[731,842,353,374]" pageId="48" pageNumber="49">Bionomics.</emphasis>
|
||
Adults have been collected during April–October at elevations of
|
||
<quantity id="4CC60C8BFFC0BC74FC39FEDDFBDFCFA6" box="[995,1122,379,401]" metricMagnitude="3" metricUnit="m" metricValue="1.68" metricValueMax="3.0" metricValueMin="0.36" pageId="48" pageNumber="49" unit="m" value="1680.0" valueMax="3000.0" valueMin="360.0">360–3,000 m</quantity>
|
||
. Adults have been collected under rocks and in moss from the edges of cold streams or in the splash zones of waterfalls, as well as in gravel at the edges of springs and in one case from under rocks near a spring that had recently stopped flowing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFC0BC74FD01FDBDFCADCDBF" pageId="48" pageNumber="49" type="discussion">
|
||
<paragraph id="8B81A16EFFC0BC74FD01FDBDFCADCDBF" blockId="48.[707,1194,194,985]" pageId="48" pageNumber="49">
|
||
<emphasis id="B94A7D7CFFC0BC74FD01FDBDFC80CC06" bold="true" box="[731,829,539,560]" pageId="48" pageNumber="49">Remarks.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFC0BC74FC9DFDBDFB98CC06" authorityName="Mullen and Campbell" authorityYear="2018" box="[839,1061,539,560]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="smetanai">
|
||
<emphasis id="B94A7D7CFFC0BC74FC9DFDBDFB98CC06" box="[839,1061,539,560]" italics="true" pageId="48" pageNumber="49">Phlaeopterus smetanai</emphasis>
|
||
</taxonomicName>
|
||
is more uniformly black than any other
|
||
<taxonomicName id="4C3EDAEDFFC0BC74FC13FD93FB2ACC7C" box="[969,1175,565,586]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFC0BC74FC13FD93FBFACC7C" box="[969,1095,565,586]" italics="true" pageId="48" pageNumber="49">Phlaeopterus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
It is most similar to
|
||
<taxonomicName id="4C3EDAEDFFC0BC74FCB6FDF6FC48CC53" authorityName="Mullen and Campbell" authorityYear="2018" box="[876,1013,592,613]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B94A7D7CFFC0BC74FCB6FDF6FC48CC53" box="[876,1013,592,613]" italics="true" pageId="48" pageNumber="49">P. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
, from which it can be distinguished by its darker color, the subglabrous region at the apex of the mesotibia shorter, and by aedeagal characters, including lacking the subapical transverse fold of the latter species. Members of a population near Payson,
|
||
<collectingRegion id="49FA6F8CFFC0BC74FBC7FD73FBD6CCDC" box="[1053,1131,725,746]" country="United States of America" name="Arizona" pageId="48" pageNumber="49">Arizona</collectingRegion>
|
||
differ from other specimens by their shape and larger size of the aedeagus (
|
||
<quantity id="4CC60C8BFFC0BC74FCBEFCACFC4CCD29" box="[868,1009,778,800]" metricMagnitude="-3" metricUnit="m" metricValue="1.635" metricValueMax="1.67" metricValueMin="1.6" pageId="48" pageNumber="49" unit="mm" value="1.635" valueMax="1.67" valueMin="1.6">1.60–1.67 mm</quantity>
|
||
<emphasis id="B94A7D7CFFC0BC74FC2CFCACFBB7CD29" box="[1014,1034,778,799]" italics="true" pageId="48" pageNumber="49">vs</emphasis>
|
||
.
|
||
<quantity id="4CC60C8BFFC0BC74FBCCFCACFB18CD29" box="[1046,1189,778,800]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.52" metricValueMin="1.28" pageId="48" pageNumber="49" unit="mm" value="1.4" valueMax="1.52" valueMin="1.28">1.28–1.52 mm</quantity>
|
||
; Payson population aedeagus in
|
||
<figureCitation id="1305BDEBFFC0BC74FBD6FC83FBD4CD0C" box="[1036,1129,805,826]" captionStart="Fig" captionStartId="28.[197,228,1491,1510]" captionTargetBox="[177,1188,196,1463]" captionTargetId="figure-70@28.[175,1190,194,1475]" captionTargetPageId="28" captionText="Fig. 23. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. bakerensis, B) P. smetanai, C) P. smetanai, D) P. occidentalis." figureDoi="http://doi.org/10.5281/zenodo.5102877" httpUri="https://zenodo.org/record/5102877/files/figure.png" pageId="48" pageNumber="49">Fig. 23C</figureCitation>
|
||
, main population aedeagus in
|
||
<figureCitation id="1305BDEBFFC0BC74FC7EFC99FC46CD62" box="[932,1019,831,852]" captionStart="Fig" captionStartId="28.[197,228,1491,1510]" captionTargetBox="[177,1188,196,1463]" captionTargetId="figure-70@28.[175,1190,194,1475]" captionTargetPageId="28" captionText="Fig. 23. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. bakerensis, B) P. smetanai, C) P. smetanai, D) P. occidentalis." figureDoi="http://doi.org/10.5281/zenodo.5102877" httpUri="https://zenodo.org/record/5102877/files/figure.png" pageId="48" pageNumber="49">Fig. 23B</figureCitation>
|
||
). Genetic analysis could help determine if this population is a distinct species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFC0BC74FD01FC29FCDECDEF" pageId="48" pageNumber="49" type="etymology">
|
||
<paragraph id="8B81A16EFFC0BC74FD01FC29FCDECDEF" blockId="48.[707,1194,194,985]" pageId="48" pageNumber="49">
|
||
<emphasis id="B94A7D7CFFC0BC74FD01FC29FCF0CD92" bold="true" box="[731,845,911,932]" pageId="48" pageNumber="49">Etymology.</emphasis>
|
||
This species is named in honor of Dr. Aleš Smetana, Biosystematics Research Institute, Ottawa,
|
||
<collectingCountry id="F329E1FEFFC0BC74FCCFFC62FCE2CDEF" box="[789,863,964,985]" name="Canada" pageId="48" pageNumber="49">Canada</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |