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<document id="048D4ECDB2234E1B73EA4C0213848D7E" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. &amp; Sikes, Derek S." docDate="2018" docId="03971078FFDCBC6AFD3FFAEAFBD1C8F4" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus occidentalis Mullen and Campbell 2018, new species" docType="treatment" docUuid="7D45457E-C5D0-4CF2-83FB-BAC51055CDCF" docUuidSource="ZooBank" docVersion="3" lastPageNumber="47" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="45" updateTime="1699007718751" updateUser="plazi">
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<mods:title id="5B95367DED03180FA4FC7CF4F28D9BB0">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="E4779B53E6A7ED176C8192F0B6880308">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="CE8DED3949D277857E3416FCA1632FE1">Campbell, J. M.</mods:namePart>
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<mods:namePart id="01BD573353849F9F91CD066960F583F6">Sikes, Derek S.</mods:namePart>
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<mods:title id="0FA523297D1B847D493179FFCF3CE653">The Coleopterists Bulletin</mods:title>
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<heading id="D0C91602FFDCBC68FD3FFAEAFB95CB4D" centered="true" fontSize="10" level="2" pageId="44" pageNumber="45" reason="2">
<emphasis id="B94A7D7CFFDCBC68FD3FFAEAFB95CB4D" bold="true" pageId="44" pageNumber="45">
16.
<taxonomicName id="4C3EDAEDFFDCBC68FCD0FAEDFC1ACB4D" ID-CoL="8Y8D3" authority="Mullen and Campbell" authorityName="Mullen and Campbell" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="B94A7D7CFFDCBC68FCD0FAEDFBB4CB56" bold="true" box="[778,1033,1355,1376]" italics="true" pageId="44" pageNumber="45">Phlaeopterus occidentalis</emphasis>
Mullen and Campbell
</taxonomicName>
,
<taxonomicNameLabel id="A279C007FFDCBC68FC69FAC0FB95CB4D" box="[947,1064,1382,1403]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B81A16EFFDCBC68FCDAFA27FB1DCBFD" blockId="44.[716,1184,1355,1483]" pageId="44" pageNumber="45">
(
<figureCitation id="1305BDEBFFDCBC68FCDDFA27FCE3CBA0" box="[775,862,1409,1430]" captionStart="Fig" captionStartId="13.[175,206,1742,1761]" captionTargetBox="[157,1166,196,1725]" captionTargetId="figure-1@13.[155,1168,194,1727]" captionTargetPageId="13" captionText="Fig. 4. Phlaeopterus species, habitus. A) P. loganensis, female (UAMObs:Ento:233072), B) P. longipennis, male" figureDoi="http://doi.org/10.5281/zenodo.5102849" httpUri="https://zenodo.org/record/5102849/files/figure.png" pageId="44" pageNumber="45">Figs. 4D</figureCitation>
,
<figureCitation id="1305BDEBFFDCBC68FCB3FA27FC2ECBA0" box="[873,915,1409,1430]" captionStart="Fig" captionStartId="19.[175,206,1027,1046]" captionText="Fig. 13. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. obsoletus, B) P. occidentalis. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="44" pageNumber="45">13B</figureCitation>
, 16F, 20E, 23D, 24A) Zoobank.org/
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urn:lsid:zoobank.org:act:
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<paragraph id="8B81A16EFFDCBC68FD1AFA46FB14CBC3" blockId="44.[704,1193,1504,1525]" box="[704,1193,1504,1525]" pageId="44" pageNumber="45">
<bibRefCitation id="EFAFDC9FFFDCBC68FD1AFA46FCE2CBC3" author="Casey" box="[704,863,1504,1525]" pageId="44" pageNumber="45" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey 1893: 402</bibRefCitation>
[misidentification as
<taxonomicName id="4C3EDAEDFFDCBC68FBF0FA46FB26CBC3" baseAuthorityName="Fauvel" baseAuthorityYear="1878" box="[1066,1179,1504,1525]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="cavicollis">
<emphasis id="B94A7D7CFFDCBC68FBF0FA46FB26CBC3" box="[1066,1179,1504,1525]" italics="true" pageId="44" pageNumber="45">T. cavicollis</emphasis>
</taxonomicName>
].
</paragraph>
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<paragraph id="8B81A16EFFDCBC68FD01F9A8FC3FC80B" blockId="44.[707,1194,1550,1730]" pageId="44" pageNumber="45">
<materialsCitation id="3B56AB33FFDCBC68FD01F9A8FCC2C80B" country="United States of America" location="Mt. Rainier National Park" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="B94A7D7CFFDCBC68FD01F9A8FCC7C815" bold="true" box="[731,890,1550,1571]" pageId="44" pageNumber="45">
<typeStatus id="54851FCCFFDCBC68FD01F9A8FCB2C815" box="[731,783,1550,1571]" pageId="44" pageNumber="45">Type</typeStatus>
Locality.
</emphasis>
<location id="8EE1F7B5FFDCBC68FC52F9A8FB1BC815" LSID="urn:lsid:plazi:treatment:03971078FFDCBC6AFD3FFAEAFBD1C8F4:8EE1F7B5FFDCBC68FC52F9A8FB1BC815" box="[904,1190,1550,1571]" country="United States of America" name="Mt. Rainier National Park" pageId="44" pageNumber="45" stateProvince="Washington">Mt. Rainier National Park</location>
,
<collectingRegion id="49FA6F8CFFDCBC68FD19F98EFC83C80B" box="[707,830,1576,1597]" country="United States of America" name="Washington" pageId="44" pageNumber="45">Washington</collectingRegion>
,
<collectingCountry id="F329E1FEFFDCBC68FC90F98EFCC2C80B" box="[842,895,1576,1597]" name="United States of America" pageId="44" pageNumber="45">USA</collectingCountry>
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.
</paragraph>
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<subSubSection id="C324F2E5FFDCBC6AFD01F9E5FE2DC8BB" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="description">
<paragraph id="8B81A16EFFDCBC68FD01F9E5FB17C8F4" blockId="44.[707,1194,1550,1730]" pageId="44" pageNumber="45">
<emphasis id="B94A7D7CFFDCBC68FD01F9E5FC03C86E" bold="true" box="[731,958,1603,1624]" pageId="44" pageNumber="45">Description. Habitus:</emphasis>
Length
<quantity id="4CC60C8BFFDCBC68FBFFF9E5FB1BC86E" box="[1061,1190,1603,1624]" metricMagnitude="-3" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="8.2" metricValueMin="6.1" pageId="44" pageNumber="45" unit="mm" value="7.1499999999999995" valueMax="8.2" valueMin="6.1">6.18.2 mm</quantity>
. Reddish brown to black; elytra sometimes lighter than pronotum; palpi, legs, and antennae reddishbrown to black (
<figureCitation id="1305BDEBFFDCBC68FC84F934FC17C891" box="[862,938,1682,1703]" captionStart="Fig" captionStartId="13.[175,206,1742,1761]" captionTargetBox="[157,1166,196,1725]" captionTargetId="figure-1@13.[155,1168,194,1727]" captionTargetPageId="13" captionText="Fig. 4. Phlaeopterus species, habitus. A) P. loganensis, female (UAMObs:Ento:233072), B) P. longipennis, male" figureDoi="http://doi.org/10.5281/zenodo.5102849" httpUri="https://zenodo.org/record/5102849/files/figure.png" pageId="44" pageNumber="45">Fig. 4D</figureCitation>
).
<emphasis id="B94A7D7CFFDCBC68FC60F934FC4AC891" bold="true" box="[954,1015,1682,1703]" pageId="44" pageNumber="45">Head:</emphasis>
Width across eyes to head length subequal to slightly wider than long.
</paragraph>
<caption id="DF41F1E6FFDDBC69FF75FB3FFE67CAC2" pageId="45" pageNumber="46" startId="45.[175,206,1177,1196]">
<paragraph id="8B81A16EFFDDBC69FF75FB3FFE67CAC2" blockId="45.[151,639,1177,1268]" pageId="45" pageNumber="46">
<emphasis id="B94A7D7CFFDDBC69FF75FB3FFF4ECA9A" bold="true" box="[175,243,1177,1196]" pageId="45" pageNumber="46">Fig. 40.</emphasis>
Hypopharynges, dorsal view, of A)
<taxonomicName id="4C3EDAEDFFDDBC69FDE4FB3FFF64CAF2" authorityName="LeConte" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B94A7D7CFFDDBC69FDE4FB3FFF64CAF2" italics="true" pageId="45" pageNumber="46">Lesteva pallipes</emphasis>
</taxonomicName>
, B)
<taxonomicName id="4C3EDAEDFFDDBC69FECCFB17FE6ACAF2" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[278,471,1201,1220]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="longoelytrata">
<emphasis id="B94A7D7CFFDDBC69FECCFB17FE6ACAF2" box="[278,471,1201,1220]" italics="true" pageId="45" pageNumber="46">Lesteva longoelytrata</emphasis>
</taxonomicName>
, C)
<taxonomicName id="4C3EDAEDFFDDBC69FDC9FB17FDC2CAF2" box="[531,639,1201,1220]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFDDBC69FDC9FB17FDE8CAF2" box="[531,597,1201,1220]" italics="true" pageId="45" pageNumber="46">Unamis</emphasis>
sp.
</taxonomicName>
undescribed, D)
<taxonomicName id="4C3EDAEDFFDDBC69FE97FB6FFE7ECAEA" box="[333,451,1225,1244]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFDDBC69FE97FB6FFE32CAEA" box="[333,399,1225,1244]" italics="true" pageId="45" pageNumber="46">Unamis</emphasis>
sp.
</taxonomicName>
, E)
<taxonomicName id="4C3EDAEDFFDDBC69FDD4FB6FFF40CAC2" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFDDBC69FDD4FB6FFF40CAC2" italics="true" pageId="45" pageNumber="46">Phlaeopterus lagrandeuri</emphasis>
</taxonomicName>
, F)
<taxonomicName id="4C3EDAEDFFDDBC69FEF8FB47FE6BCAC2" authorityName="Hatch" authorityYear="1957" box="[290,470,1249,1268]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="houkae">
<emphasis id="B94A7D7CFFDDBC69FEF8FB47FE6BCAC2" box="[290,470,1249,1268]" italics="true" pageId="45" pageNumber="46">Phlaeopterus houkae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF41F1E6FFDDBC69FD1FFB07FC05CB22" ID-DOI="http://doi.org/10.5281/zenodo.5102909" ID-Zenodo-Dep="5102909" httpUri="https://zenodo.org/record/5102909/files/figure.png" pageId="45" pageNumber="46" startId="45.[709,740,1185,1204]" targetBox="[154,1169,195,1168]" targetPageId="45">
<paragraph id="8B81A16EFFDDBC69FD1FFB07FC05CB22" blockId="45.[685,1173,1185,1300]" pageId="45" pageNumber="46">
<emphasis id="B94A7D7CFFDDBC69FD1FFB07FCA4CA82" bold="true" box="[709,793,1185,1204]" pageId="45" pageNumber="46">Fig. 41.</emphasis>
<taxonomicName id="4C3EDAEDFFDDBC69FCF4FB07FC49CA82" box="[814,1012,1185,1204]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFDDBC69FCF4FB07FC23CA82" box="[814,926,1185,1204]" italics="true" pageId="45" pageNumber="46">Phlaeopterus</emphasis>
species
</taxonomicName>
, hypopharynges, dorsal view, of A)
<taxonomicName id="4C3EDAEDFFDDBC69FC89FB1FFC05CAFA" authorityName="Mullen and Campbell" authorityYear="2018" box="[851,952,1209,1228]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="B94A7D7CFFDDBC69FC89FB1FFC05CAFA" box="[851,952,1209,1228]" italics="true" pageId="45" pageNumber="46">P. obsoletus</emphasis>
</taxonomicName>
, B)
<taxonomicName id="4C3EDAEDFFDDBC69FC38FB1FFBE6CAFA" authorityName="Mullen and Campbell" authorityYear="2018" box="[994,1115,1209,1228]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="B94A7D7CFFDDBC69FC38FB1FFBE6CAFA" box="[994,1115,1209,1228]" italics="true" pageId="45" pageNumber="46">P. kavanaughi</emphasis>
</taxonomicName>
, C)
<taxonomicName id="4C3EDAEDFFDDBC69FB5EFB1FFD42CAD2" authorityName="Mullen and Campbell" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B94A7D7CFFDDBC69FB5EFB1FFD42CAD2" italics="true" pageId="45" pageNumber="46">P. elongatus</emphasis>
</taxonomicName>
. D)
<taxonomicName id="4C3EDAEDFFDDBC69FCF1FB77FBB4CAD2" authorityName="Hatch" authorityYear="1957" box="[811,1033,1233,1252]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFDDBC69FCF1FB77FBB4CAD2" box="[811,1033,1233,1252]" italics="true" pageId="45" pageNumber="46">Phlaeopterus lagrandeuri</emphasis>
</taxonomicName>
, metatrochanter with apical tooth. Heads, ventral view, of E)
<taxonomicName id="4C3EDAEDFFDDBC69FB8EFB4FFD52CB22" authorityName="LeConte" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B94A7D7CFFDDBC69FB8EFB4FFD52CB22" italics="true" pageId="45" pageNumber="46">Lesteva pallipes</emphasis>
</taxonomicName>
, F)
<taxonomicName id="4C3EDAEDFFDDBC69FCCFFAA7FC0ECB22" authorityName="Mannerheim" authorityYear="1830" box="[789,947,1281,1300]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="B94A7D7CFFDDBC69FCCFFAA7FC0ECB22" box="[789,947,1281,1300]" italics="true" pageId="45" pageNumber="46">Lesteva pubescens</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B81A16EFFDDBC6AFF4DFA9FFE2DC8BB" blockId="45.[151,638,1337,1730]" lastBlockId="46.[173,660,1336,1730]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
Interantennal groove broadly and moderately deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 510 at least 2 times longer than wide; antennomeres 411 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with Lshaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3
<superScript id="7C4B0C26FFDDBC69FDDDF985FDABC807" attach="left" box="[519,534,1571,1585]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
palpomere 1.61.8 times longer than palpomere 2.
<emphasis id="B94A7D7CFFDDBC69FDF0F9E5FDC0C86E" bold="true" box="[554,637,1603,1624]" pageId="45" pageNumber="46">Thorax:</emphasis>
Pronotum moderately wide (Fig. 16F), length to width ratio = 0.630.71; ratio of pronotal width to head width = 1.451.70; maximum width subequal to elytral width at humeral angles; punctures on dorsal surface separated by distance slightly less than to slightly more than diameter of a puncture; lateral margins explanate posterad lateral foveae, narrowly explanate anterad lateral foveae, narrowing to not at all explanate at apex; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.22.6 times longer than pronotum; apical margins broadly convex. Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along entire midline of mesosternum (
<figureCitation id="1305BDEBFFDDBC69FC72F9DEFBBDC8BB" box="[936,1024,1656,1677]" captionStart="Fig" captionStartId="25.[709,740,1185,1204]" captionTargetBox="[689,1169,195,1168]" captionTargetId="figure-245@25.[688,1170,194,1169]" captionTargetPageId="25" captionText="Fig. 20. Phlaeopterus species, mesosterna, ventral oblique view. A) P. longipennis, B) P. obsoletus, C) P. olympicus, D) P. frosti, E) P. occidentalis, F) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102871" httpUri="https://zenodo.org/record/5102871/files/figure.png" pageId="45" pageNumber="46">Fig. 20E</figureCitation>
).
<emphasis id="B94A7D7CFFDDBC69FBCBF9DEFBF7C8BB" bold="true" box="[1041,1098,1656,1677]" pageId="45" pageNumber="46">Legs:</emphasis>
Apices of all tibia without pubescence, length of subglabrous apex of metatibia as ratio to metatibial length in males = 7.628.0, in females = 8.619.4. Metatrochanter without tooth on apical margin.
<emphasis id="B94A7D7CFFDEBC6AFED3FAC8FEC8CBB5" bold="true" box="[265,373,1390,1411]" pageId="46" pageNumber="47">Abdomen:</emphasis>
Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI with wing-folding spicules.
<emphasis id="B94A7D7CFFDEBC6AFF77FA18FEA5CBE5" bold="true" box="[173,280,1470,1491]" pageId="46" pageNumber="47">Aedeagus:</emphasis>
Length
<quantity id="4CC60C8BFFDEBC6AFEA8FA18FDBBCBE5" box="[370,518,1469,1491]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" metricValueMax="1.5" metricValueMin="1.24" pageId="46" pageNumber="47" unit="mm" value="1.37" valueMax="1.5" valueMin="1.24">1.241.50 mm</quantity>
. Median lobe irregularly narrowing from base to abruptly truncate apex. Parameres diverging from base to apical fourth that strongly converges towards apex of median lobe (
<figureCitation id="1305BDEBFFDEBC6AFEF4F98EFE2DC80B" box="[302,400,1576,1597]" captionStart="Fig" captionStartId="28.[197,228,1491,1510]" captionTargetBox="[177,1188,196,1463]" captionTargetId="figure-70@28.[175,1190,194,1475]" captionTargetPageId="28" captionText="Fig. 23. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. bakerensis, B) P. smetanai, C) P. smetanai, D) P. occidentalis." figureDoi="http://doi.org/10.5281/zenodo.5102877" httpUri="https://zenodo.org/record/5102877/files/figure.png" pageId="46" pageNumber="47">Figs. 23D</figureCitation>
,
<figureCitation id="1305BDEBFFDEBC6AFE42F98EFE78C80B" box="[408,453,1576,1597]" captionStart="Fig" captionStartId="29.[175,206,1486,1505]" captionTargetBox="[156,1167,196,1468]" captionTargetId="figure-90@29.[154,1169,194,1470]" captionTargetPageId="29" captionText="Fig. 24. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. occidentalis, B) P. olympicus, C) P. loganensis, D) P. fusconiger." figureDoi="http://doi.org/10.5281/zenodo.5102879" httpUri="https://zenodo.org/record/5102879/files/figure.png" pageId="46" pageNumber="47">24A</figureCitation>
). Internal sack lightly sclerotized; with subapical transverse fold; apical 2/3 covered in microspinules except for smooth area of transverse fold.
</paragraph>
</subSubSection>
<caption id="DF41F1E6FFDEBC6AFF1FFB0BFDBFCB3E" ID-DOI="http://doi.org/10.5281/zenodo.5102911" ID-Zenodo-Dep="5102911" httpUri="https://zenodo.org/record/5102911/files/figure.png" pageId="46" pageNumber="47" startId="46.[197,228,1197,1216]" targetBox="[176,656,195,1180]" targetPageId="46">
<paragraph id="8B81A16EFFDEBC6AFF1FFB0BFDBFCB3E" blockId="46.[173,660,1197,1288]" pageId="46" pageNumber="47">
<emphasis id="B94A7D7CFFDEBC6AFF1FFB0BFEB3CAF6" bold="true" box="[197,270,1197,1216]" pageId="46" pageNumber="47">Fig. 42.</emphasis>
Heads, ventral view, of A)
<taxonomicName id="4C3EDAEDFFDEBC6AFDF6FB0BFD29CAF6" box="[556,660,1197,1216]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFDEBC6AFDF6FB0BFDD3CAF6" box="[556,622,1197,1216]" italics="true" pageId="46" pageNumber="47">Unamis</emphasis>
sp.
</taxonomicName>
undescribed, B)
<taxonomicName id="4C3EDAEDFFDEBC6AFEBBFB63FE6ACAEE" box="[353,471,1221,1240]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFDEBC6AFEBBFB63FE1ECAEE" box="[353,419,1221,1240]" italics="true" pageId="46" pageNumber="47">Unamis</emphasis>
sp.
</taxonomicName>
, C)
<taxonomicName id="4C3EDAEDFFDEBC6AFDF9FB63FEAFCAC6" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFDEBC6AFDF9FB63FEAFCAC6" italics="true" pageId="46" pageNumber="47">Phlaeopterus lagrandeuri</emphasis>
</taxonomicName>
, D)
<taxonomicName id="4C3EDAEDFFDEBC6AFE9AFB7BFE4ACAC6" authorityName="Hatch" authorityYear="1957" box="[320,503,1245,1264]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="houkae">
<emphasis id="B94A7D7CFFDEBC6AFE9AFB7BFE4ACAC6" box="[320,503,1245,1264]" italics="true" pageId="46" pageNumber="47">Phlaeopterus houkae</emphasis>
</taxonomicName>
, E)
<taxonomicName id="4C3EDAEDFFDEBC6AFDF9FB7BFEB2CB3E" authorityName="Mullen &amp; Campbell &amp; Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B94A7D7CFFDEBC6AFDF9FB7BFEB2CB3E" italics="true" pageId="46" pageNumber="47">Phlaeopterus longipennis</emphasis>
</taxonomicName>
, F)
<taxonomicName id="4C3EDAEDFFDEBC6AFEEEFB53FE40CB3E" authorityName="Mullen and Campbell" authorityYear="2018" box="[308,509,1269,1288]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B94A7D7CFFDEBC6AFEEEFB53FE40CB3E" box="[308,509,1269,1288]" italics="true" pageId="46" pageNumber="47">Phlaeopterus elongatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C324F2E5FFDEBC6AFF1FF934FC8ACF9D" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8B81A16EFFDEBC6AFF1FF934FC8ACF9D" blockId="46.[173,660,1336,1730]" lastBlockId="46.[707,1194,194,1730]" pageId="46" pageNumber="47">
<emphasis id="B94A7D7CFFDEBC6AFF1FF934FEC8C891" bold="true" box="[197,373,1682,1703]" pageId="46" pageNumber="47">Type Specimens.</emphasis>
<materialsCitation id="3B56AB33FFDEBC6AFE5EF934FC08CF3A" collectingDate="1979-08-03" collectorName="Mt. Rainier N. P. &amp; J. M. &amp; B. A. Campbell" location="Mt. Rainier" municipality="West Side Rd." pageId="46" pageNumber="47" specimenCount="2" typeStatus="holotype">
<typeStatus id="54851FCCFFDEBC6AFE5EF934FE61C891" box="[388,476,1682,1703]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
male (UAMObs: Ento:235826) and
<typeStatus id="54851FCCFFDEBC6AFE86F90BFE15C8F4" box="[348,424,1709,1730]" pageId="46" pageNumber="47" type="allotype">allotype</typeStatus>
female (UAMObs:Ento: 235827) labeled as follows: WASH.,
<collectorName id="26CBC4B8FFDEBC6AFBE2FF64FD4BCEC7" pageId="46" pageNumber="47">
<location id="8EE1F7B5FFDEBC6AFBE2FF64FB15CEE1" LSID="urn:lsid:plazi:treatment:03971078FFDCBC6AFD3FFAEAFBD1C8F4:8EE1F7B5FFDEBC6AFBE2FF64FB15CEE1" box="[1080,1192,194,215]" municipality="West Side Rd." name="Mt. Rainier" pageId="46" pageNumber="47">Mt. Rainier</location>
N. P.
</collectorName>
, end of
<collectingMunicipality id="6BE53B14FFDEBC6AFC91FF7AFC61CEC7" box="[843,988,220,241]" pageId="46" pageNumber="47">West Side Rd.</collectingMunicipality>
,
<date id="FF8087AEFFDEBC6AFC32FF7AFBE3CEC7" box="[1000,1118,220,241]" pageId="46" pageNumber="47" value="1979-08-03">
<collectingDate id="EFC47E46FFDEBC6AFC32FF7AFBE3CEC7" box="[1000,1118,220,241]" pageId="46" pageNumber="47" value="1979-08-03">3.VIII.1979</collectingDate>
</date>
, 3700,
<collectorName id="26CBC4B8FFDEBC6AFD19FF51FC08CF3A" box="[707,949,247,268]" pageId="46" pageNumber="47">J. M. &amp; B. A. Campbell</collectorName>
</materialsCitation>
/
<typeStatus id="54851FCCFFDEBC6AFC66FF51FB81CF3A" box="[956,1084,247,268]" pageId="46" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
³ (or AL- LOTYPE
<specimenCount id="9D386AE7FFDEBC6AFCF0FEB2FC8ACF10" box="[810,823,276,294]" pageId="46" pageNumber="47" type="female"></specimenCount>
)
<taxonomicName id="4C3EDAEDFFDEBC6AFC93FEB7FBFBCF10" authorityName="Mullen and Campbell" authorityYear="2018" box="[841,1094,273,294]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentalis">Phlaeopterus occidentalis</taxonomicName>
desig. L. J. Mullen and J. M. Campbell CNC No. 18466 (red label). Both specimens are in the CNC, Ottawa.
<typeStatus id="54851FCCFFDEBC6AFD19FEC7FC9CCF40" box="[707,801,353,374]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
(
<emphasis id="B94A7D7CFFDEBC6AFCEAFEC7FC81CF40" box="[816,828,353,374]" italics="true" pageId="46" pageNumber="47">n</emphasis>
= 232) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, ROM, USNM, and UCRC.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFDEBC6AFD01FE17FC93CD89" pageId="46" pageNumber="47" type="distribution">
<paragraph id="8B81A16EFFDEBC6AFD01FE17FBD7CCAD" blockId="46.[707,1194,194,1730]" pageId="46" pageNumber="47">
<emphasis id="B94A7D7CFFDEBC6AFD01FE17FCD8CFF0" bold="true" box="[731,869,433,454]" pageId="46" pageNumber="47">Distribution.</emphasis>
<taxonomicName id="4C3EDAEDFFDEBC6AFCACFE17FB35CFF0" authorityName="Mullen and Campbell" authorityYear="2018" box="[886,1160,433,454]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B94A7D7CFFDEBC6AFCACFE17FB35CFF0" box="[886,1160,433,454]" italics="true" pageId="46" pageNumber="47">Phlaeopterus occidentalis</emphasis>
</taxonomicName>
is known from southern
<collectingRegion id="49FA6F8CFFDEBC6AFC6AFE6AFBB2CFD7" box="[944,1039,460,481]" country="United States of America" name="California" pageId="46" pageNumber="47">California</collectingRegion>
north through
<collectingRegion id="49FA6F8CFFDEBC6AFD19FE40FCB0CFCD" box="[707,781,486,507]" country="United States of America" name="Oregon" pageId="46" pageNumber="47">Oregon</collectingRegion>
,
<collectingRegion id="49FA6F8CFFDEBC6AFCFAFE40FC29CFCD" box="[800,916,486,507]" country="United States of America" name="Washington" pageId="46" pageNumber="47">Washington</collectingRegion>
, and
<collectingRegion id="49FA6F8CFFDEBC6AFC03FE40FB3ACFCD" box="[985,1159,486,507]" country="Canada" name="British Columbia" pageId="46" pageNumber="47">British Columbia</collectingRegion>
to southeastern
<collectingRegion id="49FA6F8CFFDEBC6AFC9AFDA7FC3FCC20" box="[832,898,513,534]" country="United States of America" name="Alaska" pageId="46" pageNumber="47">Alaska</collectingRegion>
(
<figureCitation id="1305BDEBFFDEBC6AFC55FDA7FC5BCC20" box="[911,998,513,534]" captionStart="Fig" captionStartId="19.[175,206,1027,1046]" captionText="Fig. 13. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. obsoletus, B) P. occidentalis. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="46" pageNumber="47">Fig. 13B</figureCitation>
). A single specimen (UAM:Ento:332755) was collected from Great Basin National Park,
<collectingRegion id="49FA6F8CFFDEBC6AFC7EFD90FC4DCC7D" box="[932,1008,566,587]" country="United States of America" name="Nevada" pageId="46" pageNumber="47">Nevada</collectingRegion>
. We attempted to re-collect this species from the 1968 locality (UAMObs:Ento:233463) near Haines,
<collectingRegion id="49FA6F8CFFDEBC6AFB9EFDCDFB35CCB6" box="[1092,1160,619,640]" country="United States of America" name="Alaska" pageId="46" pageNumber="47">Alaska</collectingRegion>
, in the summer of 2014 but were unsuccessful.
</paragraph>
<paragraph id="8B81A16EFFDEBC6AFD01FD06FC93CD89" blockId="46.[707,1194,194,1730]" pageId="46" pageNumber="47">
<emphasis id="B94A7D7CFFDEBC6AFD01FD06FCF7CC83" bold="true" box="[731,842,672,693]" pageId="46" pageNumber="47">Bionomics.</emphasis>
Adults of this species have been collected at
<quantity id="4CC60C8BFFDEBC6AFC9FFD1DFC6DCCE6" box="[837,976,698,720]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="3.2" metricValueMin="0.3" pageId="46" pageNumber="47" unit="m" value="1750.0" valueMax="3200.0" valueMin="300.0">3003,200 m</quantity>
, a broader elevation range than any other species in the genus. In
<collectingRegion id="49FA6F8CFFDEBC6AFD19FD56FCCBCD33" box="[707,886,752,773]" country="Canada" name="British Columbia" pageId="46" pageNumber="47">British Columbia</collectingRegion>
and
<collectingRegion id="49FA6F8CFFDEBC6AFC6BFD56FC47CD33" box="[945,1018,752,773]" country="United States of America" name="Alaska" pageId="46" pageNumber="47">Alaska</collectingRegion>
, the species has only been collected at
<quantity id="4CC60C8BFFDEBC6AFC78FCADFB95CD16" box="[930,1064,778,800]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.8" metricValueMin="0.3" pageId="46" pageNumber="47" unit="m" value="1050.0" valueMax="1800.0" valueMin="300.0">3001,800 m</quantity>
. Adults have been collected during MayOctober in
<collectingRegion id="49FA6F8CFFDEBC6AFBA4FC83FCBACD63" country="United States of America" name="California" pageId="46" pageNumber="47">California</collectingRegion>
and
<collectingRegion id="49FA6F8CFFDEBC6AFC91FCE6FC24CD63" box="[843,921,832,853]" country="United States of America" name="Oregon" pageId="46" pageNumber="47">Oregon</collectingRegion>
, but only during June August in
<collectingRegion id="49FA6F8CFFDEBC6AFCF7FCFCFC18CD59" box="[813,933,858,879]" country="United States of America" name="Washington" pageId="46" pageNumber="47">Washington</collectingRegion>
and
<collectingRegion id="49FA6F8CFFDEBC6AFC02FCFCFB9FCD59" box="[984,1058,858,879]" country="United States of America" name="Alaska" pageId="46" pageNumber="47">Alaska</collectingRegion>
. This species has been collected under rocks and in moss in the splash-zone at the edges of small streams or waterfalls.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFDEBC6AFD01FC63FCB7C8BA" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8B81A16EFFDEBC6AFD01FC63FC89CBB5" blockId="46.[707,1194,194,1730]" pageId="46" pageNumber="47">
<emphasis id="B94A7D7CFFDEBC6AFD01FC63FC80CDEC" bold="true" box="[731,829,965,986]" pageId="46" pageNumber="47">Remarks.</emphasis>
Specimens from Vancouver Island vary from those of the mainland by having the base of the internal sac of the aedeagus broader and the apex of the internal sac concave. The aedeagus of the Vancouver Island (
<figureCitation id="1305BDEBFFDEBC6AFC75FB89FBB0CA72" box="[943,1037,1071,1092]" captionStart="Fig" captionStartId="29.[175,206,1486,1505]" captionTargetBox="[156,1167,196,1468]" captionTargetId="figure-90@29.[154,1169,194,1470]" captionTargetPageId="29" captionText="Fig. 24. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. occidentalis, B) P. olympicus, C) P. loganensis, D) P. fusconiger." figureDoi="http://doi.org/10.5281/zenodo.5102879" httpUri="https://zenodo.org/record/5102879/files/figure.png" pageId="46" pageNumber="47">Fig. 24A</figureCitation>
) and mainland (
<figureCitation id="1305BDEBFFDEBC6AFD10FBEFFC95CA68" box="[714,808,1097,1118]" captionStart="Fig" captionStartId="28.[197,228,1491,1510]" captionTargetBox="[177,1188,196,1463]" captionTargetId="figure-70@28.[175,1190,194,1475]" captionTargetPageId="28" captionText="Fig. 23. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. bakerensis, B) P. smetanai, C) P. smetanai, D) P. occidentalis." figureDoi="http://doi.org/10.5281/zenodo.5102877" httpUri="https://zenodo.org/record/5102877/files/figure.png" pageId="46" pageNumber="47">Fig. 23D</figureCitation>
) populations are illustrated. There is also a slight difference in the length of the posterior tibiae
<emphasis id="B94A7D7CFFDEBC6AFCE4FBD9FCEFCAA2" box="[830,850,1151,1172]" italics="true" pageId="46" pageNumber="47">vs</emphasis>
. posterior tarsus and in the length of the antennomeres approaching the apex. Furthermore, the mainland population shows geographic variation in the aedeagus, with the aedeagus becoming slightly longer and the apex of the median lobe progressively more narrowly triangular moving from south to north. We deem these differences insufficient to warrant splitting these populations into multiple species or subspecies.
</paragraph>
<paragraph id="8B81A16EFFDEBC6AFD01FA2EFCB7C8BA" blockId="46.[707,1194,194,1730]" pageId="46" pageNumber="47">
<bibRefCitation id="EFAFDC9FFFDEBC6AFD01FA2EFCDCCBAB" author="Casey" box="[731,865,1416,1437]" pageId="46" pageNumber="47" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey (1893)</bibRefCitation>
erroneously referred to a specimen of
<taxonomicName id="4C3EDAEDFFDEBC6AFCC1FA05FC0FCB8E" authorityName="Mullen and Campbell" authorityYear="2018" box="[795,946,1443,1464]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="occidentalis">
<emphasis id="B94A7D7CFFDEBC6AFCC1FA05FC0FCB8E" box="[795,946,1443,1464]" italics="true" pageId="46" pageNumber="47">P. occidentalis</emphasis>
</taxonomicName>
as
<taxonomicName id="4C3EDAEDFFDEBC6AFC3BFA05FBE2CB8E" baseAuthorityName="Fauvel" baseAuthorityYear="1878" box="[993,1119,1443,1464]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cavicollis">
<emphasis id="B94A7D7CFFDEBC6AFC3BFA05FBE2CB8E" box="[993,1119,1443,1464]" italics="true" pageId="46" pageNumber="47">T. cavicollis</emphasis>
</taxonomicName>
in his treatment of the latter species, likely because the type series of the latter species was mixed. This error was fixed by a
<typeStatus id="54851FCCFFDEBC6AFC4FFA54FC4DC831" box="[917,1008,1522,1543]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
designation for
<emphasis id="B94A7D7CFFDEBC6AFB4DFA54FC94C814" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="4C3EDAEDFFDEBC6AFB4DFA54FC9BC814" baseAuthorityName="Fauvel" baseAuthorityYear="1878" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cavicollis">P. cavicollis</taxonomicName>
.
</emphasis>
The specimen examined by
<bibRefCitation id="EFAFDC9FFFDEBC6AFBB7F9ABFCB9C80B" author="Casey" pageId="46" pageNumber="47" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey (1893)</bibRefCitation>
is labeled as follows: Vanc./
<specimenCount id="9D386AE7FFDEBC6AFB9EF98CFBECC80A" box="[1092,1105,1578,1596]" pageId="46" pageNumber="47" type="female"></specimenCount>
/CASEY bequest 1925/
<taxonomicName id="4C3EDAEDFFDEBC6AFC95F9E4FC60C861" authority="Fvl." authorityName="Fvl." box="[847,989,1602,1623]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="cavicollis">cavicollis Fvl.</taxonomicName>
/May be from original type series. see Casey, Col. notices
<date id="FF8087AEFFDEBC6AFB8CF9FBFB18C844" box="[1110,1189,1629,1650]" pageId="46" pageNumber="47" value="1893-05">V. 1893</date>
, p. 402.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFDEBC6AFD01F934FBD1C8F4" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8B81A16EFFDEBC6AFD01F934FBD1C8F4" blockId="46.[707,1194,194,1730]" pageId="46" pageNumber="47">
<emphasis id="B94A7D7CFFDEBC6AFD01F934FCF0C891" bold="true" box="[731,845,1682,1703]" pageId="46" pageNumber="47">Etymology.</emphasis>
This species is named for its wide distribution in northwestern North America.
</paragraph>
</subSubSection>
</treatment>
</document>