322 lines
39 KiB
XML
322 lines
39 KiB
XML
<document id="300E9B1E834FEADB19AC8E39FC7EC771" ID-CLB-Dataset="34074" ID-DOI="http://dx.doi.org/10.3897/zookeys.568.7733" ID-GBIF-Dataset="92a3cb34-dc4e-4e01-b4e5-5ccc463ad0fb" ID-PMC="PMC4829672" ID-Pensoft-Pub="1313-2970-568-119" ID-PubMed="27103876" ID-ZooBank="DB611BCF653041B2A8887E1C26A51CE0" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-568-119" ModsDocOrigin="ZooKeys 568" ModsDocTitle="Cobitistakenoi sp. n. (Cypriniformes, Cobitidae): a new spined loach from Honshu Island, Japan" checkinTime="1456309275195" checkinUser="pensoft" docAuthor="Nakajima, Jun" docDate="2016" docId="CC888968EEB9DE4407DED2DB3B49C1C1" docLanguage="en" docName="ZooKeys 568: 119-128" docOrigin="ZooKeys 568" docSource="http://dx.doi.org/10.3897/zookeys.568.7733" docTitle="Cobitis takenoi Nakajima, 2016, sp. n." docType="treatment" docUuid="13895F3E-53D6-4C24-BE3B-E6DBDF7BEF07" docUuidSource="ZooBank" docVersion="7" lastPageNumber="125" masterDocId="6117B604FF87FF9BFFA5DB1A3810FFFA" masterDocTitle="Cobitistakenoi sp. n. (Cypriniformes, Cobitidae): a new spined loach from Honshu Island, Japan" masterLastPageNumber="128" masterPageNumber="119" pageNumber="121" updateTime="1701383209156" updateUser="ExternalLinkService">
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<mods:title id="3BE6FA454ACFF9FA02676539CB8C5CFD">Cobitistakenoi sp. n. (Cypriniformes, Cobitidae): a new spined loach from Honshu Island, Japan</mods:title>
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<mods:namePart id="FBDDBFFD95855B6E92362D9AEF284958">Nakajima, Jun</mods:namePart>
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<mods:date id="9866D4B819F90B0DDDC4F7B6EA690995">2016</mods:date>
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<mods:classification id="0C77793E248C5D7ECA452C512F2948F9">journal article</mods:classification>
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<mods:identifier id="4230E1033F4C283C29692095027C2E13" type="DOI">http://dx.doi.org/10.3897/zookeys.568.7733</mods:identifier>
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<treatment id="CC888968EEB9DE4407DED2DB3B49C1C1" ID-GBIF-Taxon="127890062" LSID="urn:lsid:zoobank.org:act:13895F3E-53D6-4C24-BE3B-E6DBDF7BEF07" httpUri="http://treatment.plazi.org/id/CC888968EEB9DE4407DED2DB3B49C1C1" lastPageId="6" lastPageNumber="125" pageId="2" pageNumber="121">
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<subSubSection id="7AAE9B2C82BE4E3047D7BFDFA5DB4D26" pageId="2" pageNumber="121" type="multiple">
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<paragraph id="04E1F218017D73D740985A034EC66D4D" pageId="2" pageNumber="121">Taxon classification Animalia Cypriniformes Cobitidae</paragraph>
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</subSubSection>
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<subSubSection id="87DDA3546F966FF5ED53FA7076228D35" pageId="2" pageNumber="121" type="nomenclature">
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<paragraph id="2F10220870D74DF933683148C5E3DB70" pageId="2" pageNumber="121">
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<taxonomicName id="BC7DBADF340CE765E0381572E0B6C3E8" LSID="http://zoobank.org/13895F3E-53D6-4C24-BE3B-E6DBDF7BEF07" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="2" pageNumber="121" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
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<taxonomicNameLabel id="4222274C15615DBD39048B15C31E1125" pageId="2" pageNumber="121">sp. n.</taxonomicNameLabel>
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Figs 3, 4, 5, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection id="778A802D4FBD98761A2BEBF93AC4CAAA" pageId="2" pageNumber="121" type="reference_group">
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<paragraph id="554F56676FE6BA82A58B1715315C8588" pageId="2" pageNumber="121">
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<taxonomicName id="B82602B4AAA5179334A4E431DE841568" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="2" pageNumber="121" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
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'Tango tetraploid
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<normalizedToken id="AD2022183DC62788B5DB8498EC0F37CA" originalValue="form’">form'</normalizedToken>
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of
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<taxonomicName id="ECEE3697AC28A1DB79D3765A10AE0383" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis striata" order="Cypriniformes" pageId="2" pageNumber="121" phylum="Chordata" rank="species" species="striata">Cobitis striata</taxonomicName>
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:
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<bibRefCitation id="9FD6EBB2A745BB0E587A95AC731E4386" author="Takeno, M" journalOrPublisher="Japanese Journal of Ichthyology" pageId="7" pageNumber="126" pagination="105 - 112" title="A distinctive allotetraploid spined loach population (genus Cobitis) from Tango District, Kyoto Prefecture, Japan." volume="57" year="2010">Takeno et al. 2010</bibRefCitation>
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: 108, fig. 2;
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<taxonomicName id="4F7FE91104C858828CEDAC0D27444E75" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis" order="Cypriniformes" pageId="2" pageNumber="121" phylum="Chordata" rank="genus">Cobitis</taxonomicName>
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sp. 5:
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<bibRefCitation id="2C98DF2DEB0C0ADAE8CC7BA380D37073" author="Nakajima, J" journalOrPublisher="Japanese Journal of Ichthyology" pageId="6" pageNumber="125" pagination="86 - 95" title="Proposal of standard Japanese names for Japanese loaches in the genus Cobitis (Cobitidae)." volume="59" year="2012">Nakajima et al. 2012</bibRefCitation>
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: 92, fig. 3e;
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<taxonomicName id="4C4CD86A8FAFE626D37DD22531E8D99C" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis" order="Cypriniformes" pageId="2" pageNumber="121" phylum="Chordata" rank="genus">Cobitis</taxonomicName>
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sp.:
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<bibRefCitation id="58339EF8AAB48F3278675594E64DD3D6" author="Hosoya, K" editor="Nakabo, T" journalOrPublisher="Tokai University Press, Hadano" pageId="6" pageNumber="125" pagination="328 - 334" title="Cobitidae." volumeTitle="Fishes of Japan with Pictorial Keys to the Species" year="2013">Hosoya 2013</bibRefCitation>
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: 331;
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<taxonomicName id="3FFB7BD47628893C2F090C59FA4F159E" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis" order="Cypriniformes" pageId="2" pageNumber="121" phylum="Chordata" rank="genus">Cobitis</taxonomicName>
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sp.:
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<bibRefCitation id="3A5CA03E00C7E1C4A07ACDD106D5E241" author="Kawase, S" editor="Hosoya, K" journalOrPublisher="Yama-Kei Publishers, Tokyo" pageId="6" pageNumber="125" title="Tango-suji-shima-dojyo." volumeTitle="Freshwater Fishes of Japan" year="2015">Kawase 2015</bibRefCitation>
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: 181.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5FAA9963303C2A51F781A3FD8A47307" pageId="2" pageNumber="121" type="type material">
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<paragraph id="3802E001E85D8DAAB236C38429AD59B4" pageId="2" pageNumber="121">Type materials.</paragraph>
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<paragraph id="7E75630BBE20B8D69FEEA26F82097BA9" pageId="2" pageNumber="121">Holotype: KPM-NI 31994, 60.4 mm standard length (SL), male, Tango District, Kyoto Prefecture, Honshu Island, Japan; collected by K. Tominaga on 17 Apr. 2010. Paratypes: 10 specimens, all from same locality as the holotype: TKPM-P 7363, 7364, 53.2-67.5 mm SL, male and female, same data as holotype; KPM-NI 31995-31999, 49.4-70.5 mm SL, 3 males and 2 females, collected by J. Nakajima on 12 Nov. 2010; KUN-P 45133, 57.7 mm SL, male, collected by K. Tominaga on 5 Jul. 2014; JNC 188, 189, 58.6-60.6 mm SL, 2 males, same data.</paragraph>
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</subSubSection>
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<subSubSection id="107B02BCB8F79CAC91952E7B56302E16" pageId="2" pageNumber="121" type="diagnosis">
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<paragraph id="1EECD80A1BE55AF2054F9F81BDFFB827" pageId="2" pageNumber="121">Diagnosis.</paragraph>
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<paragraph id="76FE160A2F9D467ED4931CAB44ADF2AE" pageId="2" pageNumber="121">Maxillary barbel short, more of the same eye diameter; lamina circularis in adult males simple and roundish; USP narrow; PMN 14; line L5 organised in 11-17 oblong or ovoid blotches out of spawning season, and lines L3 and L5 in adult male well-developed longitudinal obvious stripes during spawning season; upper and lower spot at caudal base not connected; tetraploid.</paragraph>
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</subSubSection>
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<subSubSection id="C815CCE6DB471508BDE9D1AFD29B6E57" lastPageId="3" lastPageNumber="122" pageId="2" pageNumber="121" type="description">
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<paragraph id="3DB397992FA863F923520347F20CC95C" pageId="2" pageNumber="121">Description.</paragraph>
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<paragraph id="BE956D6D45A359486E517BF9EFBE1DC0" lastPageId="3" lastPageNumber="122" pageId="2" pageNumber="121">
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Dorsal-fin rays iii, 7; anal-fin rays iii, 5; pectoral-fin rays i, 7-8; pelvic-fin rays ii, 6; caudal-fin rays 8+8. Body elongate, laterally compressed. Head and snout elongated. Interorbital space narrow, convex. Eye relatively large. Caudal peduncle relatively compressed. Mouth small, inferior, arched with fleshy lips; lower lip divided with 2 well-developed lobes; upper lip with transverse wrinkles on the surface. Barbels, 3 pairs, first on rostorum, second on maxilla, third on maxillomandibula; each barbel well-developed, length of maxillary barbel short, same as the eye diameter; the length of the rostral and maxillary barbels shorter than that of mandibular barbel (Fig. 4a). Lateral line short, reaching the pectoral-fin base. PMN 14. Very small cycloid scales on the trunk. Suborbital spine two-pronged and incurved; length of the outer spine one-third of that of the inner spine (Fig. 4b). First branched ray of the pectoral fin longer than rest (Fig. 4c); pectoral fin in adult males longer than that in females. USP narrow (Figs 4c, d, see also Fig. 6). Lamina circularis at the base of the pectoral fin in adult males simple and roundish (Fig. 4d). Dorsal-fin base equidistant from the base of the caudal fin and the top of the snout. Pelvic-fin origin below the second or third branched dorsal fin ray.
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<pageBreakToken id="61ED103B112E694FCA9AEA13986DB10C" pageId="3" pageNumber="122" start="start">Anal</pageBreakToken>
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fin not reaching the caudal-fin base. Margin of anal and dorsal fins slightly roundish. Caudal fin slightly roundish. Abdominal vertebrae 22 (21-23); caudal vertebrae 20 (19-21); total vertebrae 42 (40-44) (
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<bibRefCitation id="EBC2D078099E8AB1286407F994314E90" author="Takeno, M" journalOrPublisher="Japanese Journal of Ichthyology" pageId="7" pageNumber="126" pagination="105 - 112" title="A distinctive allotetraploid spined loach population (genus Cobitis) from Tango District, Kyoto Prefecture, Japan." volume="57" year="2010">Takeno et al. 2010</bibRefCitation>
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). Largest recorded specimens 65.5 mm SL in male and 84.9 mm SL in female (
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<bibRefCitation id="61739FF8C85E613AB75EA7B3A0CBF529" author="Takeno, M" journalOrPublisher="Japanese Journal of Ichthyology" pageId="7" pageNumber="126" pagination="105 - 112" title="A distinctive allotetraploid spined loach population (genus Cobitis) from Tango District, Kyoto Prefecture, Japan." volume="57" year="2010">Takeno et al. 2010</bibRefCitation>
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).
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</paragraph>
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<caption id="0761AC80FF0A28C64B49FAB62BCBC18B" pageId="3" pageNumber="122">
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<paragraph id="58BCFB1C65E769305B3AAF010DA13E0F" pageId="3" pageNumber="122">
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Figure 3. Male (a, c holotype, KPM-NI 31994, 60.4 mm SL) and female (b, d paratype, KPM-NI 31999, 70.5 mm SL) specimens of
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<taxonomicName id="CA6EBD2F83C8BC3E8C75BA7066918101" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="3" pageNumber="122" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
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sp. n. a, b Lateral view c, d dorsal view.
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</paragraph>
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</caption>
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<caption id="EE84CFAB9672ECF0E01739FBECDB19FF" pageId="3" pageNumber="122">
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<paragraph id="A049BD82AD50E76643F8FA01F46D0AEE" pageId="3" pageNumber="122">
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Figure 4.
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<taxonomicName id="EDC4AABA3AB36D13023DD640C1593074" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="3" pageNumber="122" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
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sp. n., KPM-NI 31994, holotype. a Mouth b right suborbital spine, lateral view (upper), and dorsal view (lower) c dorsal view of the pectoral
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<normalizedToken id="3E82720D18F5C6266970BE994E8AE2BE" originalValue="fin">fin</normalizedToken>
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d lamina circularis and upper segments of the
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<normalizedToken id="7E761808C9B0321FCD5443A2D716572F" originalValue="first">first</normalizedToken>
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branched soft ray (USP). Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption id="0D03B9305EDFAB6E25D15DAF879FE06C" pageId="3" pageNumber="122">
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<paragraph id="C179211B3CD99AD2B2578EFE1D8E8A98" pageId="3" pageNumber="122">
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Figure 5. Change in colouration of the
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<taxonomicName id="65066994381905711123DCFE763E26C8" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="3" pageNumber="122" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
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sp. n. adult male (paratype, KUN-P 45133, 57.7 mm SL). a Spawning season, 8 July 2014 b non-spawning season, 5 December 2014.
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</paragraph>
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</caption>
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<caption id="D2781779AA95295157CDA914E9F10711" pageId="3" pageNumber="122">
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<paragraph id="34A5F2E67313C9ED36A012F286307F8C" pageId="3" pageNumber="122">
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Figure 6. Narrow and broad types of upper segments of the first branched soft ray (USP). a Narrow type (
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<taxonomicName id="C5BA7BA70433CEC5467CA073935BFEA6" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis kaibarai" order="Cypriniformes" pageId="3" pageNumber="122" phylum="Chordata" rank="species" species="kaibarai">Cobitis kaibarai</taxonomicName>
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) b broad type (
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<taxonomicName id="E143FD504E1966BEB6C99BCE1ED98DFF" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis magnostriata" order="Cypriniformes" pageId="3" pageNumber="122" phylum="Chordata" rank="species" species="magnostriata">Cobitis magnostriata</taxonomicName>
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). Redrawn from
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<bibRefCitation id="A978FC2D99C834D4530A5D25744B4A61" author="Nakajima, J" journalOrPublisher="Zootaxa" pageId="6" pageNumber="125" pagination="103 - 130" title="Taxonomic study of the Cobitisstriata complex (Cypriniformes, Cobitidae) in Japan." volume="3586" year="2012">Nakajima (2012)</bibRefCitation>
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. Scale bars: 1 mm.
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</paragraph>
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</caption>
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<caption id="76E016D1B4ABE104C6E2C6F9B844FDF8" pageId="3" pageNumber="122">
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<paragraph id="A3BF38C6D045E5CA820E148262A28E54" pageId="3" pageNumber="122">
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Table 1. Counts and morphometric measurements of
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<taxonomicName id="8679F3371465D9D0C41BB3B7FFB400D8" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="3" pageNumber="122" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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<paragraph id="B95388203F8CAFA7971548C996B965BA" pageId="3" pageNumber="122">
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<table id="9C85AA377D46FAE69A4386ABCDB92A02" pageId="3" pageNumber="122">
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<tr id="992F51E9480224D0E2AF406BC4E00684" pageId="3" pageNumber="122">
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<th id="2E8A8C3982E387696FF88E553C060FD2" colspan="1" pageId="3" pageNumber="122" rowspan="2">Holotype</th>
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<th id="7EE14FF84BFC4EF7F395C51258AF8CAE" colspan="2" pageId="3" pageNumber="122" rowspan="1">Paratypes</th>
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</tr>
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<tr id="9CFEA99DED251FF845A584D17D1D6C91" pageId="3" pageNumber="122">
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<th id="01F9E588F2151DF550AC43729497F28E" colspan="1" pageId="3" pageNumber="122" rowspan="1">7 males</th>
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<th id="EC1210DDDEEB4DFB49E07127888EF5A9" colspan="1" pageId="3" pageNumber="122" rowspan="1">3 females</th>
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</tr>
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<tr id="6CAFB831E422CF1DD91E4F5D7B9B33BE" pageId="3" pageNumber="122">
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<td id="3683DCF1B0E771F5617C92EA0B655FE6" colspan="2" pageId="3" pageNumber="122" rowspan="1">SL (mm); mean (range)</td>
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</tr>
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<tr id="7CD35A93C4CD782DDB862A791A71D55A" pageId="3" pageNumber="122">
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<td id="74E63483E1F1850BF21CC246E2CE17EC" colspan="1" pageId="3" pageNumber="122" rowspan="5">Counts</td>
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</tr>
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<tr id="4A9185E48075080052F4DE1052A88AED" pageId="3" pageNumber="122">
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<td id="98EE0BFE6469AA3963BDA258F3C107AB" colspan="1" pageId="3" pageNumber="122" rowspan="7">In % SL; mean (range)</td>
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<td id="B8338486B711883DCCEDA5E2DEC73C65" colspan="1" pageId="3" pageNumber="122" rowspan="1">HL</td>
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</tr>
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<tr id="71AE589E449640838B2F647A7B9CFCCD" pageId="3" pageNumber="122">
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<td id="EAE44D99BF13201178712BFB0F70ED06" colspan="1" pageId="3" pageNumber="122" rowspan="1">LPP</td>
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</tr>
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<tr id="D8D83488280DF6247D9F9C9C59CC302C" pageId="3" pageNumber="122">
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<td id="4C70D188C14957D6DDF91580C17F035C" colspan="1" pageId="3" pageNumber="122" rowspan="1">LPA</td>
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</tr>
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<tr id="A90373D6260D5CC74D13A68DD26A6E41" pageId="3" pageNumber="122">
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<td id="9607F7A26DBE1DCFC07860830D33722A" colspan="1" pageId="3" pageNumber="122" rowspan="1">DCP</td>
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</tr>
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<tr id="FCE82AAC78E67EB22DEF1C5749DE6099" pageId="3" pageNumber="122">
|
||
<td id="63FECCA5C99E0BD4A76412D62E484262" colspan="1" pageId="3" pageNumber="122" rowspan="2">In % HL; mean (range)</td>
|
||
</tr>
|
||
<tr id="37A9A44C885AF4FA8577DD8879837844" pageId="3" pageNumber="122">
|
||
<td id="D293625896186E0691B0B9DA251EFE0F" colspan="2" pageId="3" pageNumber="122" rowspan="1">PMN</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="56114B21CCFBFEBA6150278A9D47F947" lastPageId="4" lastPageNumber="123" pageId="3" pageNumber="122" type="colouration">
|
||
<paragraph id="A3AFE013D03E8A8151D23B5690D97C4F" pageId="3" pageNumber="122">Colouration.</paragraph>
|
||
<paragraph id="08D601DA0C94193BD8023EF31A4DA713" lastPageId="4" lastPageNumber="123" pageId="3" pageNumber="122">
|
||
Body yellowish white with dark brown pigmentation in fresh. A clear streak running from the tip of the snout to the occiput, crossing to the eye. Upper part
|
||
<pageBreakToken id="6DA3445CC9E8AA0D12C5774342868E3E" pageId="4" pageNumber="123" start="start">of</pageBreakToken>
|
||
the head covered with amorphous spots; opercle and snout covered with amorphous patterns. Caudal and dorsal fins with 3-4 arcuate bars. Anal fin pigmented along fin rays. Upper spot at caudal base jet-black, size comparable to the eye diameter, lower spot at the caudal base relatively inconspicuous and small; upper and lower spots at the caudal base not connected. Male out of spawning season (Figs 3a, 5b). Body pigmentation organised in 1 dorsal and 4 lateral lines. Line L1 consisting of a series of 11-16, saddle or oval shaped blotches. Line L2 formed by a longitudinal jugged line or convex semicircular spots or chained small angular blotches, only present on dorsal part of body. Line L3 formed by a sharp longitudinal line or narrow dotted line, reaching to the post-dorsal body, with intermissive posterior part. Line L4 formed by narrow web like line or dots, reaching to dorsal body. Line L5 organised in 11-17 blotches from the upper part of the pectoral fin to the caudal-fin base; blotches roundish, frequently oblong or ovoid. Male in the spawning season (Fig. 5a). Line L4 not visible or formed by faint longitudinal line. Lines L3 and L5 well developed, forming longitudinal obvious stripes from the upper part of the pectoral-fin base to the caudal-fin base, often intermissive posterior part of L3. Female (Fig. 3b). Similar to males out of spawning season.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17D86D756F849931D27B29D61E9191EE" pageId="4" pageNumber="123" type="sexual dimorphism">
|
||
<paragraph id="52F357715B9761D2860B3AA409380CEE" pageId="4" pageNumber="123">Sexual dimorphism.</paragraph>
|
||
<paragraph id="8E6AB053EF24C4E4BE91FB18B9888570" pageId="4" pageNumber="123">Males having a roundish lamina circularis at the base of the pectoral fins; females do not. Generally, the body size of females larger than that of males. Lines L3 and L5 of adult males well developed, forming longitudinal obvious stripes during the spawning season; females do not.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F4EDAAF559E083AAE0353372EC854" pageId="5" pageNumber="124" type="ploidy">
|
||
<paragraph id="308E164C9D2AEF68AA812EBDEDC297EB" pageId="5" pageNumber="124">
|
||
<pageBreakToken id="C41AA2032499EDA99D4014D45FF2D73F" pageId="5" pageNumber="124" start="start">Ploidy</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B93831C0440614CC497597403B8E5AE3" pageId="5" pageNumber="124">
|
||
Tetraploid (
|
||
<bibRefCitation id="45AC8FC3CD288F2620E35E2CCF7FC844" author="Takeno, M" journalOrPublisher="Japanese Journal of Ichthyology" pageId="7" pageNumber="126" pagination="105 - 112" title="A distinctive allotetraploid spined loach population (genus Cobitis) from Tango District, Kyoto Prefecture, Japan." volume="57" year="2010">Takeno et al. 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6191E9A6E04B774C61065BE1249A2C7E" pageId="5" pageNumber="124" type="etymology">
|
||
<paragraph id="FC91D4F802F8BE141144B0AA3A5E33B6" pageId="5" pageNumber="124">Etymology.</paragraph>
|
||
<paragraph id="313CF8A47308BD861FB987F7289429F7" pageId="5" pageNumber="124">The specific name is dedicated to Mr. Makoto Takeno, the discoverer of this spined loach.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="478E1412970FC1E38DA5C7C326CE2E94" pageId="5" pageNumber="124" type="distribution">
|
||
<paragraph id="97915D19AD3411A7DC87AB44BAD1430D" pageId="5" pageNumber="124">Distribution.</paragraph>
|
||
<paragraph id="9D236DA8196A3D8D2CD5A1DA8EDF1E7B" pageId="5" pageNumber="124">Tango District, Kyoto prefecture, Honshu Island, Japan.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDE53A39F40AC1A1BF5B0B45B0E9EE7B" pageId="5" pageNumber="124" type="habitat and biology">
|
||
<paragraph id="0249CA09B9D84049CA3BAA1C3B9B6E9D" pageId="5" pageNumber="124">Habitat and biology.</paragraph>
|
||
<paragraph id="82C9AA143E69E05CD296E587AA5CFF62" pageId="5" pageNumber="124">This species inhabits sandy-mud bottoms of the middle and lower reaches of rivers (Fig. 2). Life histories are unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A3A53D041E5973BFBD333485922A5AC" pageId="5" pageNumber="124" type="mtdna cytb sequence">
|
||
<paragraph id="4E4D00B24B3D67E9FD9142F6F1351BC2" pageId="5" pageNumber="124">mtDNA cytb sequence.</paragraph>
|
||
<paragraph id="DF2DA374A7B4E02BB133C38BC3D05CCF" pageId="5" pageNumber="124">
|
||
AB533231-AB533234 (
|
||
<bibRefCitation id="6F43001EB0DB5CAF681CDC2F53A953E9" author="Takeno, M" journalOrPublisher="Japanese Journal of Ichthyology" pageId="7" pageNumber="126" pagination="105 - 112" title="A distinctive allotetraploid spined loach population (genus Cobitis) from Tango District, Kyoto Prefecture, Japan." volume="57" year="2010">Takeno et al. 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D41D1BD9B9E4DD317B88B1498E5C9798" pageId="5" pageNumber="124" type="japanese name">
|
||
<paragraph id="517E7D7C456671A5C7068ED476BB5AE5" pageId="5" pageNumber="124">Japanese name.</paragraph>
|
||
<paragraph id="FF89F7E56475896BFE9F90F640FFFCED" pageId="5" pageNumber="124">
|
||
<normalizedToken id="902A630DC032263CEF5B5CD39AABBF2D" originalValue="Tango-suji-shima-dojyô">Tango-suji-shima-dojyo</normalizedToken>
|
||
(
|
||
<bibRefCitation id="5D18F7176030AD7D1198E3DD21E0DFF6" author="Nakajima, J" journalOrPublisher="Japanese Journal of Ichthyology" pageId="6" pageNumber="125" pagination="86 - 95" title="Proposal of standard Japanese names for Japanese loaches in the genus Cobitis (Cobitidae)." volume="59" year="2012">Nakajima et al. 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5F3489110EA46EAD0AE1F3E251907105" lastPageId="6" lastPageNumber="125" pageId="5" pageNumber="124" type="comparison">
|
||
<paragraph id="8EB09B78741C90DA201D26EA039A8900" pageId="5" pageNumber="124">Comparison.</paragraph>
|
||
<paragraph id="04EA26F991D1BC35E058F81154A8904E" lastPageId="6" lastPageNumber="125" pageId="5" pageNumber="124">
|
||
This new species is distinguished from nine species of
|
||
<taxonomicName id="58AF417583003D80B35F8E956F240DFE" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="genus">Cobitis</taxonomicName>
|
||
in the Japanese archipelago (
|
||
<taxonomicName id="4D88D143EA081E88652DF5C39309F572" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis biwae" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="biwae">Cobitis biwae</taxonomicName>
|
||
,
|
||
<taxonomicName id="20A918555B9F98921DF9DEE0EC588B3A" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis striata" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="striata">Cobitis striata</taxonomicName>
|
||
,
|
||
<taxonomicName id="B9C50CBAADAD5EDD21D1658FB3C0670D" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis matsubarae" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="matsubarae">Cobitis matsubarae</taxonomicName>
|
||
,
|
||
<taxonomicName id="B39757CE501E7F4F49B7DC0A2D605259" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takatsuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="takatsuensis">Cobitis takatsuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="B42A129CB2FB9474C4593117D2AD97DE" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis shikokuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="shikokuensis">Cobitis shikokuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="725B7BCA8B79A6322FCE0C20F914C6E9" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis magnostriata" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="magnostriata">Cobitis magnostriata</taxonomicName>
|
||
,
|
||
<taxonomicName id="0FF17FBCB1C95ADF945C5BFAE5E1EF72" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis minamorii" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="minamorii">Cobitis minamorii</taxonomicName>
|
||
,
|
||
<taxonomicName id="032A45CD82B4DBB9DB44EB449A6D1A17" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis kaibarai" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="kaibarai">Cobitis kaibarai</taxonomicName>
|
||
and
|
||
<taxonomicName id="0AD535B6EFEE0FC49C8ECD73E390FF05" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis sakahoko" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="sakahoko">Cobitis sakahoko</taxonomicName>
|
||
) by a combination of the following character states: a short maxillary barbel equaling in length the eye diameter (vs. longer than the eye diameter in
|
||
<taxonomicName id="E7ED2810AA3A100C5ED8B78077523EBB" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis matsubarae" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="matsubarae">Cobitis matsubarae</taxonomicName>
|
||
,
|
||
<taxonomicName id="D7B9B5031048F84DA42F97332BE29FB8" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takatsuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="takatsuensis">Cobitis takatsuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="096CD414CD5E76F75EE4FC1E92D8140B" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis shikokuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="shikokuensis">Cobitis shikokuensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="8A4F401452C0BFA2A96D253183B089C4" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis sakahoko" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="sakahoko">Cobitis sakahoko</taxonomicName>
|
||
); a simple roundish lamina circularis (vs. beak-shaped or narrow in
|
||
<taxonomicName id="636B523C96E2C2DCDC43C5FA6090C94F" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis biwae" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="biwae">Cobitis biwae</taxonomicName>
|
||
; quite narrow in
|
||
<taxonomicName id="E3BB4762C553CE5F5CCBD6842050E4B0" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takatsuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="takatsuensis">Cobitis takatsuensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="51DA225ECA7DF8ACC03900DC68C7FA9E" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis shikokuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="shikokuensis">Cobitis shikokuensis</taxonomicName>
|
||
; rectangular with a neck in
|
||
<taxonomicName id="803E9E87E616FB081D857F3A4459508B" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis sakahoko" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="sakahoko">Cobitis sakahoko</taxonomicName>
|
||
); a narrow USP (vs. broad in
|
||
<taxonomicName id="FEF3E55EAF635CDCA9557DCC3DCD068B" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis matsubarae" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="matsubarae">Cobitis matsubarae</taxonomicName>
|
||
,
|
||
<taxonomicName id="E5D8FA56363B1380F6F9532AED67E81F" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takatsuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="takatsuensis">Cobitis takatsuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="72650FC22BC9F95C795D2268423E1CE3" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis shikokuensis" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="shikokuensis">Cobitis shikokuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="F3EE37EB3E1ABF7F1F0CB9CE12E5BC26" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis magnostriata" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="magnostriata">Cobitis magnostriata</taxonomicName>
|
||
and
|
||
<taxonomicName id="C1752546116A3C89242EC139C045865D" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis sakahoko" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="sakahoko">Cobitis sakahoko</taxonomicName>
|
||
); PMN 14 (vs. commonly 12 in
|
||
<taxonomicName id="33F3ECC982353C08758E8B17F5CF4783" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis minamorii" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="minamorii">Cobitis minamorii</taxonomicName>
|
||
; commonly 13 in
|
||
<taxonomicName id="E02CFD741B10312A3FA51B0ED1E2A555" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis striata" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="striata">Cobitis striata</taxonomicName>
|
||
and
|
||
<taxonomicName id="8B0ABFE9E93F6FE084A43542017C5C76" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis kaibarai" order="Cypriniformes" pageId="5" pageNumber="124" phylum="Chordata" rank="species" species="kaibarai">Cobitis kaibarai</taxonomicName>
|
||
); a L5 formed of blotches out of spawning
|
||
<pageBreakToken id="9374913B1B17CF5FD48789499A312F7C" pageId="6" pageNumber="125" start="start">season</pageBreakToken>
|
||
(vs. stripe-like in and out of spawning season in
|
||
<taxonomicName id="BFF39F936B24B00BF8D318F9901CF848" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takatsuensis" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="takatsuensis">Cobitis takatsuensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="2DA62D277A30125E9F1D298CA75136FB" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis magnostriata" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="magnostriata">Cobitis magnostriata</taxonomicName>
|
||
); both spots at caudal base obvious (vs. lower spot inconspicuous in
|
||
<taxonomicName id="CBCC7730B46E67985A8D0458B1A9036E" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis striata" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="striata">Cobitis striata</taxonomicName>
|
||
and
|
||
<taxonomicName id="776036EC1D15894462B8DE6361E53174" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis kaibarai" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="kaibarai">Cobitis kaibarai</taxonomicName>
|
||
); and ploidy tetraploid (vs. diploid in
|
||
<taxonomicName id="52B5BA59D151C557EF3E326FD7C7AB8E" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis striata" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="striata">Cobitis striata</taxonomicName>
|
||
,
|
||
<taxonomicName id="6A0C77BACD5BE573C299DCA2B8C69EB7" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takatsuensis" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="takatsuensis">Cobitis takatsuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="58B5FF46B662D289C1F6B4DBA206EB0E" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis shikokuensis" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="shikokuensis">Cobitis shikokuensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="29B27DF1942D37EA350A3AA13E959C61" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis minamorii" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="minamorii">Cobitis minamorii</taxonomicName>
|
||
and
|
||
<taxonomicName id="EEDE4463A586C8AD88F425C18CEA6DAB" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis kaibarai" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="kaibarai">Cobitis kaibarai</taxonomicName>
|
||
). These comparative data were summarised from
|
||
<bibRefCitation id="C999050FBA680A84B118EE8A1EAA9A61" author="Nakajima, J" journalOrPublisher="Ichthyological Research" pageId="6" pageNumber="125" pagination="68 - 78" title="Cobitissakahoko, a new species of spined loach (Cypriniformes: Cobitidae) from southern Kyushu Island, Japan." url="10.1007/s10228-015-0476-5" volume="63" year="2016">Nakajima and Suzawa (2016)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="51C66E3377E3A5D9775782435A14294C" pageId="6" pageNumber="125" type="remarks">
|
||
<paragraph id="13F32DC8D7DE04D69AD44D3D2534F55D" pageId="6" pageNumber="125">Remarks.</paragraph>
|
||
<paragraph id="5FF6BC69BEF2C8F2810CA0A85DB180B5" pageId="6" pageNumber="125">
|
||
Till date,
|
||
<taxonomicName id="623C3AEBA47F66ABD76CFCF92E62BA47" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis takenoi" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="takenoi">Cobitis takenoi</taxonomicName>
|
||
has only been found in one small river system, and the habitat is under threat from river improvement. In addition, some threatened freshwater fishes are captured and sold illegally in Japan (e.g.
|
||
<taxonomicName id="8313B25C283AFF5DEE1C180ACA55F8E0" family="Cobitidae" genus="Parabotia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parabotia curtus" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="curtus">Parabotia curtus</taxonomicName>
|
||
,
|
||
<bibRefCitation id="167BF5CE0DB238ED87AFDCF48A0FECD0" author="Watanabe, K" journalOrPublisher="The IUCN Red List of Threatened Species" pageId="7" pageNumber="126" title="Parabotiacurtus." url="10.2305/IUCN.UK.2015-4.RLTS.T11661A83606443.en" volume="2015" year="2015">Watanabe et al. 2015</bibRefCitation>
|
||
), and this new species is similarly at the risk of being commercially overfished for the ornamental fish market (
|
||
<bibRefCitation id="904F35B5C59409167DEA511CBEED76C3" author="Takeno, M" journalOrPublisher="Japanese Journal of Ichthyology" pageId="7" pageNumber="126" pagination="105 - 112" title="A distinctive allotetraploid spined loach population (genus Cobitis) from Tango District, Kyoto Prefecture, Japan." volume="57" year="2010">Takeno et al. 2010</bibRefCitation>
|
||
). Therefore, the species is ranked as a critically endangered species (CR) - as
|
||
<taxonomicName id="D376F22151180479EE6DA8ED0145180A" family="Cobitidae" genus="Cobitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cobitis" order="Cypriniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="genus">Cobitis</taxonomicName>
|
||
sp. - in the Japanese Red List (
|
||
<bibRefCitation id="D02BB32B3D182749E9CDAD62C6808FB1" author="Kitagawa, T" editor="Ministry, of the Environment" journalOrPublisher="GYOSEI Corporation, Tokyo" pageId="6" pageNumber="125" pagination="48 - 49" title="Tango-suji-shima-dojyo." volumeTitle="Red Data Book 2014 - Threatened Wildlife of Japan-, Volume 4, Pisces - Brackish and Fresh Water Fishes" year="2015">Kitagawa 2015</bibRefCitation>
|
||
). The distribution pattern, suitable habitat and life history of this species are not well-known. Basic biological investigations are required for its effective conservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |