159 lines
18 KiB
XML
159 lines
18 KiB
XML
<document id="C57E39A372CB2DFCEEA9AEE42E9E9D76" ID-DOI="10.5281/zenodo.10667704" ID-ISSN="1407-8619" ID-Zenodo-Dep="10667704" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708024650417" checkinUser="felipe" docAuthor="Fanti, Fabrizio" docDate="2021" docId="CC0016512A0DFFE7FC5891626ADBB4DD" docLanguage="en" docName="BalticJColeopterol.21.1.1-9.pdf" docOrigin="Baltic Journal of Coleopterology 21 (1)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.1:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="1" docTitle="Malthodes (Libertimalthodes) headsi Fanti 2021, sp." docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="30396E292A0FFFE4FFCC947D6E5CB04A" masterDocTitle="Two new Malthodes and a new Podistra from Eocene Baltic amber (Coleoptera, Cantharidae)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="1" updateTime="1708024956114" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="BAB2E0874B2FC0F50485B0068CCF6EB1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1B7439DDFFED50D31BF372BA88CC5D76">
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<mods:title id="2F73B5CC2006FAF6EE204F6AD119A78D">Two new Malthodes and a new Podistra from Eocene Baltic amber (Coleoptera, Cantharidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="BC2D60932FD0EA24DEBEBBAA0BEB3ABF" type="personal">
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<mods:role id="2316C43902E6F83A8985121563F30084">
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<mods:roleTerm id="34C7F6E804BD05346ABDDB2DE433C1AE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="445638982E391F358D113AA4A5537F75">Fanti, Fabrizio</mods:namePart>
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<mods:typeOfResource id="961F6DEC10DE94A27B178380DD54B07F">text</mods:typeOfResource>
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<mods:titleInfo id="9DF2AFB3AF039E18A12290E60101DC2C">
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<mods:title id="29C285B8560E60473B4D7491406659E6">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="ADC324E648DDF18EF920DA9535264615">
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<mods:date id="AB7D0272A990B90DE84F6571ABDF2C9A">2021</mods:date>
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<mods:detail id="888A73BBCB8C791ECE05605B0AB83F90" type="pubDate">
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<mods:number id="DF8ABC4FB4EEEBDA7BE3BBBFA354E1F9">2021-09-30</mods:number>
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</mods:detail>
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<mods:detail id="101E3D6BAB9B6E78F53E69F72E33B83A" type="volume">
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<mods:number id="637C62399AFB03B94156FFFBE85B9035">21</mods:number>
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</mods:detail>
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<mods:detail id="7B685F1D195C02DAEB7C6D232AB81426" type="issue">
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<mods:number id="E9AD7DD4D6763DCD9E4D3438388BC48E">1</mods:number>
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<mods:extent id="BE5D7B3B46AA46A9FB84CF5A70CE26DE" unit="page">
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<mods:start id="93766EF270DBC192CDFECF9C04716F9B">1</mods:start>
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<mods:end id="1CB4D046DF0C5D94F802D844D1C93039">9</mods:end>
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<mods:classification id="8AA96F00D41C24280BDECD0C0BA6B9E9">journal article</mods:classification>
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<mods:identifier id="2068A21713434C56516369E6E0D4F803" type="DOI">10.5281/zenodo.10667704</mods:identifier>
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<mods:identifier id="3357815DB83CA3C016E2020E6F0D7DEB" type="ISSN">1407-8619</mods:identifier>
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<mods:identifier id="6CFABA99D931ED0090AB5378EA9C6577" type="Zenodo-Dep">10667704</mods:identifier>
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</mods:mods>
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<treatment id="CC0016512A0DFFE7FC5891626ADBB4DD" LSID="urn:lsid:plazi:treatment:CC0016512A0DFFE7FC5891626ADBB4DD" httpUri="http://treatment.plazi.org/id/CC0016512A0DFFE7FC5891626ADBB4DD" lastPageId="3" pageId="2">
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<subSubSection id="0CB3F4CC2A0DFFE6FC5891626DE0B51D" pageId="2" type="nomenclature">
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<paragraph id="4416A7472A0DFFE6FC5891626B0CB57D" blockId="2.[916,1360,1311,1399]" box="[916,1360,1311,1335]" pageId="2">
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<heading id="1F5E102B2A0DFFE6FC5891626B0CB57D" bold="true" box="[916,1360,1311,1335]" fontSize="10" level="2" pageId="2" reason="4">
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<emphasis id="76DD7B552A0DFFE6FC5891626B0CB57D" bold="true" box="[916,1360,1311,1335]" pageId="2">
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<taxonomicName id="83A9DCC42A0DFFE6FC5891626B70B57D" authority="Fanti, 2021" authorityName="Fanti" authorityYear="2021" box="[916,1324,1311,1335]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="headsi" status="sp." subGenus="Libertimalthodes">
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<emphasis id="76DD7B552A0DFFE6FC5891626A56B57D" bold="true" box="[916,1034,1311,1335]" italics="true" pageId="2">Malthodes</emphasis>
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(
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<emphasis id="76DD7B552A0DFFE6FBD491626A8AB57D" bold="true" box="[1048,1238,1311,1335]" italics="true" pageId="2">Libertimalthodes</emphasis>
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)
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<emphasis id="76DD7B552A0DFFE6FB2891626B70B57D" bold="true" box="[1252,1324,1311,1335]" italics="true" pageId="2">headsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6DEEC62E2A0DFFE6FAFD91626B0CB57D" box="[1329,1360,1311,1335]" pageId="2">sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4416A7472A0DFFE6FC5891426DE0B51D" blockId="2.[916,1360,1311,1399]" box="[916,956,1343,1367]" pageId="2">
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<emphasis id="76DD7B552A0DFFE6FC5891426DE0B51D" bold="true" box="[916,956,1343,1367]" pageId="2">nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="0CB3F4CC2A0DFFE6FC5891226A50B53D" box="[916,1036,1375,1399]" pageId="2" type="description">
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<paragraph id="4416A7472A0DFFE6FC5891226A50B53D" blockId="2.[916,1360,1311,1399]" box="[916,1036,1375,1399]" pageId="2">
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(
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<figureCitation id="DC92BBC22A0DFFE6FC5091226A58B53D" box="[924,1028,1375,1399]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[303,343,620,644]" captionStartId-1="4.[303,342,1699,1723]" captionTargetBox-1="[595,1087,1346,1636]" captionTargetId-1="figure-170@4.[584,1119,1311,1655]" captionTargetPageId-1="4" captionText-0="Fig. 3. Malthodes (Libertimalthodes) headsi sp. nov. Head completely exposed, rounded, wide, INHSP-18001 in Baltic amber. Holotype, dorso-lateral with shallow punctation and short setae. view, bar = 1.0 mm Eyes large, rounded, convex, inserted laterally and in the upper part of the head. Mandibles falciform, pointed apically. Maxillary palpi 4-segmented, first palpomere short and robust, second elongated, third shorter than second, fourth globular and distally pointed.Labial palpi 3-segmented; last palpomere globular and pointed.Antennae 11-segmented, filiform, inserted close to the eyes, strongly pubescent with many long setae, not particularly long and almost reaching the half of elytra; scape club-shaped, very robust; antennomere II short, about 1.9 times shorter than scape; antennomeres III–X Fig. 4. Malthodes (Libertimalthodes) headsi sp. nov. sub-equal and filiform; antennomere XI INHSP-18001 in Baltic amber. Holotype, detail of legs, elongated and rounded apically. bar = 0.2 mm Pronotum transverse, slightly narrower" captionText-1="Fig. 5. Malthodes (Libertimalthodes) headsi sp. nov. INHSP-18001 in Baltic amber. Drawing of last urites (lateral view)" figureDoi-1="http://doi.org/10.5281/zenodo.10667706" httpUri-1="https://zenodo.org/record/10667706/files/figure.png" pageId="2">Figs. 3 - 5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0CB3F4CC2A0DFFE7FC5891E26D47B0EE" lastPageId="3" pageId="2" type="materials_examined">
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<paragraph id="4416A7472A0DFFE6FC5891E26B68B59D" blockId="2.[916,1360,1439,1495]" pageId="2">
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<emphasis id="76DD7B552A0DFFE6FC5891E26A5CB5FD" bold="true" box="[916,1024,1439,1463]" pageId="2">
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<typeStatus id="9B1219E52A0DFFE6FC5891E26DA7B5FD" box="[916,1019,1439,1463]" pageId="2" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male, in Baltic amber, deposited under accession number INHSP-18001.
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</paragraph>
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<paragraph id="4416A7472A0DFFE6FC5891826B74B67D" blockId="2.[916,1360,1535,1592]" pageId="2">
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<emphasis id="76DD7B552A0DFFE6FC5891826A6DB65D" bold="true" box="[916,1073,1535,1559]" pageId="2">Type locality.</emphasis>
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<collectingCountry id="3CBEE7D72A0DFFE6FBF791826AD6B65D" box="[1083,1162,1535,1559]" name="Poland" pageId="2">Poland</collectingCountry>
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, Baltic Sea coast, Gdańsk city area, Wisła River estuary.
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</paragraph>
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<caption id="10D6F7CF2A0DFFE6FEE3921C6A1AB6FD" pageId="2" startId="2.[303,343,1633,1657]" targetType="figure">
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<paragraph id="4416A7472A0DFFE6FEE3921C6A1AB6FD" blockId="2.[303,1360,1631,1721]" pageId="2">
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Fig. 2.
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<taxonomicName id="83A9DCC42A0DFFE6FE48921C6D59B633" authority="Fanti, 2021" authorityName="Fanti" authorityYear="2021" box="[388,773,1633,1657]" class="Insecta" family="Cantharidae" genus="Podistra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="guthriei" status="sp. nov." subGenus="Pseudoabsidia">
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<emphasis id="76DD7B552A0DFFE6FE48921C6FBFB633" box="[388,483,1633,1657]" italics="true" pageId="2">Podistra</emphasis>
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(
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<emphasis id="76DD7B552A0DFFE6FE34921C6CC5B633" box="[504,665,1633,1657]" italics="true" pageId="2">Pseudoabsidia</emphasis>
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)
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<emphasis id="76DD7B552A0DFFE6FD61921C6D59B633" box="[685,773,1633,1657]" italics="true" pageId="2">guthriei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6DEEC62E2A0DFFE6FCDC921F6D35B633" box="[784,873,1634,1657]" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="76DD7B552A0DFFE6FC5892226A69B63D" bold="true" box="[916,1077,1631,1655]" pageId="2">Type horizon.</emphasis>
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Middle Eocene: Lutetian INHSP-18000 in Baltic amber. Holotype, detail of elytral (47.8–41.2 Mya) to late Eocene: Priabonian apex and last tergites, bar = 0.2 mm (37.8–33.9 Mya).
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</paragraph>
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</caption>
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<paragraph id="4416A7472A0CFFE7FD0194EF6D47B0EE" blockId="3.[717,795,146,164]" box="[717,795,146,164]" pageId="3">
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<emphasis id="76DD7B552A0CFFE7FD0194EF6D47B0EE" box="[717,795,146,164]" italics="true" pageId="3">Fanti F.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="0CB3F4CC2A0CFFE7FF2F94C26F9FB1B3" pageId="3" type="diagnosis">
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<paragraph id="4416A7472A0CFFE7FF2F94C26F9FB1B3" blockId="3.[227,738,191,505]" pageId="3">
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<emphasis id="76DD7B552A0CFFE7FF2F94C26FABB09D" bold="true" box="[227,503,191,215]" pageId="3">Differential diagnosis.</emphasis>
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The last maxillary palpomere globular and distally pointed, the long elytra and the last urites little differentiated, allow this taxon to be abscribed to the genus
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<taxonomicName id="83A9DCC42A0CFFE7FF2F95426F0AB11D" authorityName="Kiesenwetter" authorityYear="1852" box="[227,342,319,343]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="genus">
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<emphasis id="76DD7B552A0CFFE7FF2F95426F0AB11D" box="[227,342,319,343]" italics="true" pageId="3">Malthodes</emphasis>
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</taxonomicName>
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and its subgenus
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<taxonomicName id="83A9DCC42A0CFFE7FDEA95426CBCB11D" authorityName="Kupryjanowicz & Fanti" authorityYear="2019" box="[550,736,319,343]" pageId="3" rank="subGenus" subGenus="Libertimalthodes">
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<emphasis id="76DD7B552A0CFFE7FDEA95426CBCB11D" box="[550,736,319,343]" italics="true" pageId="3">Libertimalthodes</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="2038DAB62A0CFFE7FF2795226C70B13D" author="Fanti F. & Michalski A. R." box="[235,556,351,375]" pageId="3" pagination="127 - 132" refId="ref4043" refString="Fanti F., Michalski A. R. 2018. An unusual fossil Malthodes with long elytra (Insecta Coleoptera Cantharidae). Giornale italiano di Entomologia 15 (63): 127 - 132." type="journal article" year="2018">Fanti & Michalski 2018</bibRefCitation>
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;
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<bibRefCitation id="2038DAB62A0CFFE7FDF295226C87B13D" author="Fanti F." box="[574,731,351,375]" pageId="3" pagination="65 - 68" refId="ref3849" refString="Fanti F. 2019 c. New fossil Malthodes Kiesenwetter, 1852 of the subgenus Libertimalthodes Kupryjanowicz & Fanti, 2019 from Baltic amber (Coleoptera: Cantharidae). Palaeodiversity 12: 65 - 68." type="journal article" year="2019">Fanti 2019c</bibRefCitation>
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;
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<bibRefCitation id="2038DAB62A0CFFE7FF2F95026C72B1DD" author="Kupryjanowicz J. & Fanti F." box="[227,558,383,407]" pageId="3" pagination="1 - 13" refId="ref4570" refString="Kupryjanowicz J., Fanti F. 2019. New subgenus and three new species of soldier beetles from the Eocene of Baltic amber. Palaeontologia Electronica 22.2.22 A: 1 - 13." type="book chapter" year="2019">Kupryjanowicz & Fanti 2019</bibRefCitation>
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). The species is easily distinguishable by the last sternite elongated, concave in the centre with at the sides a lobe bent upwards.
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</paragraph>
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</subSubSection>
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<subSubSection id="0CB3F4CC2A0CFFE7FF2F966A6D9FB2DD" pageId="3" type="description">
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<paragraph id="4416A7472A0CFFE7FF2F966A6F71B2E5" blockId="3.[227,738,535,687]" pageId="3">
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<emphasis id="76DD7B552A0CFFE7FF2F966A6F2CB265" bold="true" box="[227,368,535,559]" pageId="3">Description.</emphasis>
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Adult, winged, slender. Male defined on the basis of the last sternite elongated and with the sides apically bent upwards. Body length: about 4.0 mm, elytra 3.0 mm. Entirely dark brown.
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</paragraph>
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<paragraph id="4416A7472A0CFFE7FF2F96AA6AFEB21D" blockId="3.[227,738,727,1743]" lastBlockId="3.[774,1285,191,599]" pageId="3">
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Head not completely exposed, rounded, wide, almost smooth. Eyes large, rounded, convex, inserted laterally and in the upper part of the head; interocular dorsal distance about 3.1 times greater than eye diameter. Mandibles not well visible. Maxillary palpi 4-segmented; first palpomere short and robust, second elongated, third shorter than second, fourth globular and distally pointed. Labial palpi 3-segmented; last palpomere globular and pointed. Antennae presumably 11-segmented, filiform, inserted close to the eyes, slightly pubescent with numerous short setae; scape club-shaped, very robust and enlarged apically; antennomere II short, about 2.1 times shorter than scape; antennomeres III–
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<collectionCode id="22B83F822A0CFFE7FD4090E56CF4B4E5" box="[652,680,1176,1199]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3">VI</collectionCode>
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subequal, filiform and very elongated. Pronotum transverse, narrower than head; surface almost flat, equipped with very short setae, apical margin slightly rounded and bordered; basal margin straight and bordered; sides straight and slightly bordered, propleura rounded and wide. Scutellum triangular. Elytra wider than pronotum, parallel-sided, rounded apically, very elongate, surpassing the last abdominal segments, strongly wrinkled, with semi-recumbent short setae. Hind wings transparent, almost completely covered by the elytra. Legs relatively short, rather robust; coxae robust and very elongated; trochanters elongated and rounded apically; femora robust, subcylindrical, slightly curved, equipped with many setae; tibiae shorter than femora especially the metatibiae, cylindrical, thin, without spurs at apex and equipped with setae; tarsal formula 5-5- 5; tarsomeres slender; tarsomere I elongated; tarsomere II about 1.3 times shorter than I; tarsomere III slightly shorter than II and sturdier; tarsomere IV strongly bilobed, with curved lobes; tarsomere
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<collectionCode id="22B83F822A0CFFE7FC7B951D6D96B13D" box="[951,970,352,375]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" type="Museum">V</collectionCode>
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elongated and flat; claws simple, without lobes or teeth. Metasternum squadrate and robust; abdominal segments transverse, short, with sparse setae; last urotergite and penultimate urosternite very large and wide; last urosternite obovate, elongate, concave in the middle, with two short lobes strongly bent upwards at apex (
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<figureCitation id="DC92BBC22A0CFFE7FB9896426AC8B21D" box="[1108,1172,575,599]" captionStart="Fig" captionStartId="4.[303,342,1699,1723]" captionTargetBox="[595,1087,1346,1636]" captionTargetId="figure-170@4.[584,1119,1311,1655]" captionTargetPageId="4" captionText="Fig. 5. Malthodes (Libertimalthodes) headsi sp. nov. INHSP-18001 in Baltic amber. Drawing of last urites (lateral view)" figureDoi="http://doi.org/10.5281/zenodo.10667706" httpUri="https://zenodo.org/record/10667706/files/figure.png" pageId="3">Fig. 5</figureCitation>
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).
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</paragraph>
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<paragraph id="4416A7472A0CFFE7FCCA96026D9FB2DD" blockId="3.[774,963,639,663]" box="[774,963,639,663]" pageId="3">Female unknown.</paragraph>
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</subSubSection>
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<subSubSection id="0CB3F4CC2A0CFFE7FCCA96C26DD9B3DD" pageId="3" type="etymology">
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<paragraph id="4416A7472A0CFFE7FCCA96C26A87B37D" blockId="3.[774,1285,703,823]" pageId="3">
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<emphasis id="76DD7B552A0CFFE7FCCA96C26DD4B29D" bold="true" box="[774,904,703,727]" pageId="3">Etymology.</emphasis>
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Named in honour of my friend, the great palaeontologist and entomologist Sam W. Heads (
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<collectingRegion id="866D69A52A0CFFE7FC9A96826DF9B35D" box="[854,933,767,791]" country="United States of America" name="Illinois" pageId="3">Illinois</collectingRegion>
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||
Natural History Survey, University of
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<collectingRegion id="866D69A52A0CFFE7FC8797626DCBB37D" box="[843,919,799,823]" country="United States of America" name="Illinois" pageId="3">Illinois</collectingRegion>
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at Urbana-Champaign,
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<collectingCountry id="3CBEE7D72A0CFFE7FB5A97626A93B37D" box="[1174,1231,799,823]" name="United States of America" pageId="3">USA</collectingCountry>
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).
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</paragraph>
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<paragraph id="4416A7472A0CFFE7FCCA97226DD9B3DD" blockId="3.[774,1285,863,919]" pageId="3">
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<emphasis id="76DD7B552A0CFFE7FCCA97226DF7B33D" bold="true" box="[774,939,863,887]" pageId="3">Syninclusions.</emphasis>
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Very few botanical remains and air bubbles.
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</paragraph>
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</subSubSection>
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<subSubSection id="0CB3F4CC2A0CFFE7FCCA97C26ADBB4DD" pageId="3" type="discussion">
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<paragraph id="4416A7472A0CFFE7FCCA97C26ADBB4DD" blockId="3.[774,1285,959,1175]" pageId="3">
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<emphasis id="76DD7B552A0CFFE7FCCA97C26D29B39D" bold="true" box="[774,885,959,983]" pageId="3">Remarks.</emphasis>
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The amber piece is transparent, measures 35 x
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<quantity id="83510AA22A0CFFE7FCA097A26DEAB3BD" box="[876,950,991,1015]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" unit="mm" value="11.0">11 mm</quantity>
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and shows the apex much thinner. The right metatibia is detached between the femur and tibia but still present; the right antenna is preserved up to the second antennomere and part of the third, and the left antenna is preserved up to the sixth antennomere.
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</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |