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<document id="710730D9A7A8AE6C48822F0FE3CEAEFB" ID-CLB-Dataset="29928" ID-DOI="10.11646/zootaxa.4433.1.2" ID-GBIF-Dataset="af2ba010-3b23-4b91-8e35-dfeadcb442cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1287093" ID-ZooBank="27EA8C94-C2E1-4C6C-B2EB-B40BC4759ED8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528732629291" checkinUser="plazi" docAuthor="Trivedi, Jigneshkumar N., Vachhrajani, Kauresh D. &amp; Ng, Peter K. L." docDate="2018" docId="03E2FE21FFF3FFA1FF75FB39CA95FA5A" docLanguage="en" docName="zootaxa.4433.1.2.pdf" docOrigin="Zootaxa 4433 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcotheres placunae" docType="treatment" docVersion="8" masterDocId="FFDB8659FFF2FFA6FFE2FFBACE30FFBF" masterDocTitle="Redescription of Arcotheres placunae (Hornell &amp; Southwell, 1909) (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India and Pakistan" updateTime="1698548289357" updateUser="ExternalLinkService">
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<mods:title id="B8758E6B327013DC2EBA6249D04584B3">Redescription of Arcotheres placunae (Hornell &amp; Southwell, 1909) (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India and Pakistan</mods:title>
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<mods:namePart id="0AF7E533F7E200FAF76EF01755D5F4A2">Trivedi, Jigneshkumar N.</mods:namePart>
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<mods:namePart id="B076DC50A9C9C6C7B614A9E258338773">Vachhrajani, Kauresh D.</mods:namePart>
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<mods:namePart id="313BEA7EBC618BBA4D7959B8A0CE00F5">Ng, Peter K. L.</mods:namePart>
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<mods:date id="97117AADAF07CAD0442890EE0DC406BB">2018</mods:date>
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<treatment id="03E2FE21FFF3FFA1FF75FB39CA95FA5A" ID-DOI="http://doi.org/10.5281/zenodo.5983966" ID-GBIF-Taxon="145939535" ID-Zenodo-Dep="5983966" LSID="urn:lsid:plazi:treatment:03E2FE21FFF3FFA1FF75FB39CA95FA5A" httpUri="http://treatment.plazi.org/id/03E2FE21FFF3FFA1FF75FB39CA95FA5A" lastPageId="7" pageId="1">
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<paragraph id="8BF44F37FFF3FFA7FF75FB39CCC7FB21" blockId="1.[151,759,1155,1215]" box="[151,759,1155,1182]" pageId="1">
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<taxonomicName id="4C4B34B4FFF3FFA7FF75FB39CCC7FB21" ID-CoL="67PXW" authority="(Hornell &amp; Southwell, 1909)" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[151,759,1155,1182]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF3FFA7FF75FB39CCC7FB21" bold="true" box="[151,759,1155,1182]" pageId="1">
<emphasis id="B93F9325FFF3FFA7FF75FB39CFA1FB22" bold="true" box="[151,401,1155,1181]" italics="true" pageId="1">Arcotheres placunae</emphasis>
(
<bibRefCitation id="EFDA32C6FFF3FFA7FE42FB3ECCDFFB21" box="[416,751,1156,1182]" pageId="1" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell, 1909</bibRefCitation>
)
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</paragraph>
<paragraph id="8BF44F37FFF3FFA7FF75FB1CCF22FB00" blockId="1.[151,759,1155,1215]" box="[151,274,1190,1215]" pageId="1">
(
<figureCitation id="137053B2FFF3FFA7FF7DFB1CCF39FB00" box="[159,265,1190,1215]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1733,1755]" captionStartId-1="3.[151,250,1566,1588]" captionStartId-2="5.[151,250,1947,1969]" captionStartId-3="6.[151,250,1609,1631]" captionTargetBox-0="[151,1436,483,1708]" captionTargetBox-1="[160,1417,315,1521]" captionTargetBox-2="[162,1428,816,1872]" captionTargetBox-3="[153,1410,211,1503]" captionTargetId-0="figure@2.[151,1436,483,1708]" captionTargetId-1="figure@3.[151,1436,301,1545]" captionTargetId-2="figure@5.[151,1436,787,1900]" captionTargetId-3="figure@6.[151,1436,193,1587]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." captionText-1="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." captionText-2="FIGURE 3. Arcotheres placunae (Hornell &amp; Southwell, 1909). AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded)." captionText-3="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri-0="https://zenodo.org/record/1287095/files/figure.png" httpUri-1="https://zenodo.org/record/1287097/files/figure.png" httpUri-2="https://zenodo.org/record/1287099/files/figure.png" httpUri-3="https://zenodo.org/record/1287101/files/figure.png" pageId="1">Figs. 14</figureCitation>
)
</paragraph>
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<paragraph id="8BF44F37FFF3FFA7FF75FB54CC97FADF" blockId="1.[151,1436,1262,1438]" pageId="1">
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<taxonomicName id="4C4B34B4FFF3FFA7FF75FB54CC82FABB" ID-CoL="4HZ8Q" authority="Hornell &amp; Southwell 1909: 99" authorityName="Hornell &amp; Southwell" authorityPageNumber="99" authorityYear="1909" box="[151,690,1262,1284]" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF3FFA7FF75FB54CF44FABB" box="[151,372,1262,1284]" italics="true" pageId="1">Pinnotheres placunae</emphasis>
<treatmentCitation id="0AEA6926FFF3FFA7FE98FB54CC82FABB" author="" box="[378,690,1262,1284]" page="99" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FE98FB54CCBCFABB" box="[378,652,1262,1284]" pageId="1" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
: 99
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</taxonomicName>
, 100, pl. 1, figs. 110;
<treatmentCitation id="0AEA6926FFF3FFA7FC7EFB54CA76FABB" author="" box="[924,1094,1262,1284]" page="250" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FC7EFB54CA24FABB" box="[924,1044,1262,1284]" pageId="1" refId="ref6674" refString="Tesch, J. J. (1918) Goneplacidae and Pinnotheridae. The Decapoda Brachyura of the Siboga-Expedition. II. Siboga Expeditie Monografie, 39 c (84), 149 - 295, pls. 7 - 18." type="journal article">Tesch 1918</bibRefCitation>
: 250
</treatmentCitation>
, 251 (list);
<treatmentCitation id="0AEA6926FFF3FFA7FB59FB54CBA8FABB" author="" box="[1211,1432,1262,1284]" page="503" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FB59FB54CB56FABB" box="[1211,1382,1262,1284]" pageId="1" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar 1957a</bibRefCitation>
: 503
</treatmentCitation>
, 504, pl. 12, figs. ak;
<treatmentCitation id="0AEA6926FFF3FFA7FE59FAB7CCA4FA9C" author="" box="[443,660,1293,1315]" page="41" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FE59FAB7CC5CFA9C" box="[443,620,1293,1315]" pageId="1" refId="ref5738" refString="Chhapgar, B. F. (1957 b) Marine crabs of Bombay State. Contribution No. 1 of the Taraporevala Marine Biological Station Marine Biological Station. Department of Fisheries, Bombay, 89 pp." type="book">Chhapgar 1957b</bibRefCitation>
: 41
</treatmentCitation>
, 42, 66, 74;
<treatmentCitation id="0AEA6926FFF3FFA7FCC6FAB7CDC1FA9C" author="" box="[804,1009,1293,1315]" page="37" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FCC6FAB7CDF8FA9C" box="[804,968,1293,1315]" pageId="1" refId="ref5777" refString="Chhapgar, B. F. (1959) On the breeding habits and larval stages of some crabs of Bombay. Records of the Indian Museum, 54, 33 - 52." type="journal article">Chhapgar 1959</bibRefCitation>
: 37
</treatmentCitation>
;
<treatmentCitation id="0AEA6926FFF3FFA7FBE3FAB7CB6AFA9C" author="" box="[1025,1370,1293,1315]" page="1179" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FBE3FAB7CB28FA9C" box="[1025,1304,1293,1315]" pageId="1" refId="ref6525" refString="Silas, E. G. &amp; Algarswami, K. (1967) On an instance of parasitisati on by the pea crab (Pinnotheres sp.) on the backwater clam Meretrix casta (Chemnitz) from India with a review of the work on the systematics, ecology, biology and ethology of the pea-crabs of the genus Pinnotheres Latreille. In: Proceeding of Symposium in Crustacea. Pt. III. Marine Biological Association of India, Cochin, Kerala, pp. 1161 - 1227," type="book chapter">Silas &amp; Algarswami 1967</bibRefCitation>
: 1179
</treatmentCitation>
, 1207 (discussion &amp; list);
<treatmentCitation id="0AEA6926FFF3FFA7FE72FA96CC48FAFD" author="" box="[400,632,1324,1346]" page="455" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FE72FA96CC79FAFD" box="[400,585,1324,1346]" pageId="1" refId="ref5811" refString="Devi, S. L. &amp; Rao, K. H. (1986) Some aspects of biology of the pea crab Pinnotheres placunae Hornell &amp; Southwell. Proceeding of the Indian National Science Academy, Part B, 52 (4), 455 - 459." type="journal article">Devi &amp; Rao 1986</bibRefCitation>
: 455
</treatmentCitation>
459 (discussion);
<treatmentCitation id="0AEA6926FFF3FFA7FCD9FA96CA15FAFD" author="" box="[827,1061,1324,1346]" page="81" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FCD9FA96CA31FAFD" box="[827,1025,1324,1346]" pageId="1" refId="ref6459" refString="Schmitt, W. L., McCain, J. C. &amp; Davidson, E. S. (1973) Decapoda. I. Brachyura. I. Fam. Pinnotheridae. In: Gruner, H. E. &amp; Holthuis, L. B. (Eds.), Crustaceorum Catalogus, 3, pp. 1 - 160." type="journal article">
Schmitt
<emphasis id="B93F9325FFF3FFA7FC73FA97CDF5FAFD" box="[913,965,1324,1346]" italics="true" pageId="1">et al.</emphasis>
1973
</bibRefCitation>
: 81
</treatmentCitation>
(discussion);
<treatmentCitation id="0AEA6926FFF3FFA7FB57FA96CEDFFADF" author="" page="189" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FB57FA96CBA9FAFD" box="[1205,1433,1324,1346]" pageId="1" refId="ref6716" refString="Yeragi, S. G. &amp; Yeragi, S. S. (1997) A study on Pinnotheres crabs associated with newly recorded Soleteuiiva violacea (Lam.) and Placuna placenta (Linn.) Journal of Marine Biological Association of India, 39 (1 - 2), 188 - 193." type="journal article">Yeragi &amp; Yeragi 1997</bibRefCitation>
: 189
</treatmentCitation>
, fig. 1;
<treatmentCitation id="0AEA6926FFF3FFA7FEDEFAF0CC5AFADF" author="" box="[316,618,1354,1376]" page="52" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FEDEFAF0CC77FADF" box="[316,583,1354,1376]" pageId="1" refId="ref6617" refString="Subba Rao, N. V. &amp; Sastry, D. R. K. (2005) Fauna of Marine National Park, Gulf of Kachchh (Gujarat). An overview. In: Conservation Area Series. Vol. 23. Zoological Survey India, Kolkata, pp. 1 - 79." type="book">Subba Rao &amp; Sastry 2005</bibRefCitation>
: 52
</treatmentCitation>
(list).
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</paragraph>
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<taxonomicName id="4C4B34B4FFF3FFA7FF75FAD3CC42FAC0" ID-CoL="67PXW" authority=", Ahyong &amp; Ng 2007: 191" authorityName="Ahyong &amp; Ng" authorityPageNumber="191" authorityYear="2007" box="[151,626,1385,1407]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF3FFA7FF75FAD3CF57FAC0" box="[151,359,1385,1407]" italics="true" pageId="1">Arcotheres placunae</emphasis>
,
<treatmentCitation id="0AEA6926FFF3FFA7FE91FAD3CC42FAC0" author="" box="[371,626,1385,1407]" page="191" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FE91FAD3CC71FAC0" box="[371,577,1385,1407]" pageId="1" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: 191
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, 205 (discussion); Ng
<emphasis id="B93F9325FFF3FFA7FCB6FAD0CDB7FAC0" box="[852,903,1385,1407]" italics="true" pageId="1">et al.</emphasis>
2008: 264 (list);
<treatmentCitation id="0AEA6926FFF3FFA7FBD4FAD3CB32FAC0" author="" box="[1078,1282,1385,1407]" page="384" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FBD4FAD3CAE3FAC0" box="[1078,1235,1385,1407]" pageId="1" refId="ref6384" refString="Pati, S. K., Sahu, K. C., Swain, D., Baliarsingh, S. K. &amp; Sharma, R. M. (2012) Marine crabs (Decapoda: Anomura and Brachyura). In: Fauna of Maharashtra. State Fauna Series. Vol. 20. Part 2. Zoological Survey of India, Kolkata, pp. 381 - 385." type="book chapter">
Pati
<emphasis id="B93F9325FFF3FFA7FB81FAD0CAA6FAC0" box="[1123,1174,1385,1407]" italics="true" pageId="1">et al.</emphasis>
2012
</bibRefCitation>
: 384
</treatmentCitation>
(list); Ng
<emphasis id="B93F9325FFF3FFA7FA8BFAD0CBA6FAC0" box="[1385,1430,1385,1407]" italics="true" pageId="1">et al</emphasis>
. 2017: 1092, 1094 (discussion);
<treatmentCitation id="0AEA6926FFF3FFA7FDE5FA32CD13FA21" author="" box="[519,803,1416,1438]" page="7" pageId="1">
<bibRefCitation id="EFDA32C6FFF3FFA7FDE5FA32CD3BFA21" box="[519,779,1416,1438]" pageId="1" refId="ref6080" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography. Zootaxa, 3374, 1 - 67." type="journal article">Naderloo &amp; Türkay 2012</bibRefCitation>
: 7
</treatmentCitation>
, 55 (discussion).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3511CBCFFF3FFA7FF75FA70CF0AF8BC" pageId="1" type="materials_examined">
<paragraph id="8BF44F37FFF3FFA7FF75FA70CC6DF904" blockId="1.[151,1436,1482,2011]" pageId="1">
<materialsCitation id="3B23456AFFF3FFA7FF75FA70CEFEF9F0" ID-GBIF-Occurrence="1848743986" collectingDate="2015-01-06" collectionCode="ZRC" collectorName="J. Trivedi" country="India" latitude="22.452223" location="Material" longLatPrecision="21" longitude="69.13583" municipality="Balapur Bay" pageId="1" specimenCode="ZRC 2017.1293" specimenCount="2" specimenCount-male="2" stateProvince="Gujarat" typeStatus="neotype">
<emphasis id="B93F9325FFF3FFA7FF75FA70CFE8FA5C" bold="true" box="[151,472,1482,1507]" pageId="1">
<location id="8E9419ECFFF3FFA7FF75FA70CECFFA5C" LSID="urn:lsid:plazi:treatment:03E2FE21FFF3FFA1FF75FB39CA95FA5A:8E9419ECFFF3FFA7FF75FA70CECFFA5C" box="[151,255,1482,1507]" country="India" latitude="22.452223" longLatPrecision="21" longitude="69.13583" municipality="Balapur Bay" name="Material" pageId="1" stateProvince="Gujarat">Material</location>
examined.
<collectingCountry id="F35C0FA7FFF3FFA7FE6DFA70CFE3FA5C" box="[399,467,1482,1507]" name="India" pageId="1">India</collectingCountry>
:
</emphasis>
<typeStatus id="54F0F195FFF3FFA7FE01FA71CC72FA5C" box="[483,578,1483,1507]" pageId="1" type="neotype">Neotype</typeStatus>
(here designated), female (CW 11.0 mm, CL
<quantity id="4CB3E2D2FFF3FFA7FB81FA71CA8EFA5C" box="[1123,1214,1483,1507]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="1" unit="mm" value="8.1">8.1 mm</quantity>
) (
<specimenCode id="DBEDE74CFFF3FFA7FB3BFA71CBA5FA5C" box="[1241,1429,1483,1507]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1">ZRC 2017.1293</specimenCode>
)
<collectingMunicipality id="6B90D54DFFF3FFA7FF75FA54CF15F9B8" box="[151,293,1518,1543]" pageId="1">Balapur Bay</collectingMunicipality>
(
<geoCoordinate id="EE7F29F0FFF3FFA7FED4FA54CF8FF9B8" box="[310,447,1518,1543]" direction="north" orientation="latitude" pageId="1" precision="15" value="22.452223">22°2708″N</geoCoordinate>
,
<geoCoordinate id="EE7F29F0FFF3FFA7FE29FA54CC61F9B8" box="[459,593,1518,1543]" direction="east" orientation="longitude" pageId="1" precision="15" value="69.13583">69°0809″E</geoCoordinate>
), muddy shore, in
<taxonomicName id="4C4B34B4FFF3FFA7FCC4FA54CA96F9B8" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[806,1190,1518,1543]" class="Bivalvia" family="Placunidae" genus="Placuna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="1" phylum="Mollusca" rank="species" species="placenta">
<emphasis id="B93F9325FFF3FFA7FCC4FA54CDDCF9B9" box="[806,1004,1518,1542]" italics="true" pageId="1">Placuna placenta</emphasis>
Linnaeus, 1758
</taxonomicName>
, Beyt Dwarka Island, Gulf of Kachchh,
<collectingRegion id="498F81D5FFF3FFA7FE83F9A8CF88F994" box="[353,440,1554,1579]" country="India" name="Gujarat" pageId="1">Gujarat</collectingRegion>
, coll.
<collectorName id="26BE2AE1FFF3FFA7FE1EF9A9CC5AF994" box="[508,618,1554,1579]" pageId="1">J. Trivedi</collectorName>
,
<collectingDate id="EFB1901FFFF3FFA7FD96F9A8CD13F994" box="[628,803,1554,1579]" pageId="1" value="2015-01-06">
<date id="FFF569F7FFF3FFA7FD96F9A8CD13F994" box="[628,803,1554,1579]" pageId="1" value="2015-01-06">6 January 2015</date>
</collectingDate>
;
<specimenCount id="9D4D84BEFFF3FFA7FCCDF9A9CDB5F994" box="[815,901,1554,1579]" pageId="1" type="male">2 males</specimenCount>
(CW
<quantity id="4CB3E2D2FFF3FFA7FC28F9A8CA12F995" box="[970,1058,1554,1579]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="1" unit="mm" value="6.4">6.4 mm</quantity>
, CL
<quantity id="4CB3E2D2FFF3FFA7FBBBF9A8CA82F995" box="[1113,1202,1554,1579]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" unit="mm" value="5.8">5.8 mm</quantity>
; CW
<quantity id="4CB3E2D2FFF3FFA7FB09F9A8CB72F995" box="[1259,1346,1554,1579]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" unit="mm" value="6.8">6.8 mm</quantity>
, CL
<quantity id="4CB3E2D2FFF3FFA7FA9BF9A8CEF7F9F0" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="1" unit="mm" value="5.9">5.9 mm</quantity>
)
</materialsCitation>
,
<materialsCitation id="3B23456AFFF3FFA7FF39F98CCAE7F9F0" ID-GBIF-Occurrence="1848743983" box="[219,1239,1590,1615]" collectingDate="2015-01-06" collectionCode="ZRC" collectorName="J. Trivedi" country="India" latitude="22.452223" location="Material" longLatPrecision="21" longitude="69.13583" municipality="Balapur Bay" pageId="1" specimenCount="5" specimenCount-female="5" stateProvince="Gujarat" typeStatus="neotype">
<specimenCount id="9D4D84BEFFF3FFA7FF39F98CCF79F9F0" box="[219,329,1590,1615]" pageId="1" type="female">5 females</specimenCount>
(CW
<quantity id="4CB3E2D2FFF3FFA7FE73F98CCC19F9F1" box="[401,553,1590,1615]" metricMagnitude="-3" metricUnit="m" metricValue="8.55" metricValueMax="10.6" metricValueMin="6.5" pageId="1" unit="mm" value="8.55" valueMax="10.6" valueMin="6.5">6.510.6 mm</quantity>
, CL
<quantity id="4CB3E2D2FFF3FFA7FD81F98CCCDEF9F0" box="[611,750,1590,1615]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.7" metricValueMin="5.4" pageId="1" unit="mm" value="6.55" valueMax="7.7" valueMin="5.4">5.47.7 mm</quantity>
) (ZLARCR100), same data as neotype
</materialsCitation>
;
<materialsCitation id="3B23456AFFF3FFA7FB06F98DCD00F9CC" ID-GBIF-Occurrence="1848744173" collectingDate="2015-01-06" collectionCode="ZRC" collectorName="J. Trivedi" country="India" latitude="22.452223" location="Material" longLatPrecision="21" longitude="69.13583" municipality="Balapur Bay" pageId="1" specimenCode="ZRC 2017.1065" specimenCount="1" specimenCount-male="1" stateProvince="Gujarat" typeStatus="neotype">
<specimenCount id="9D4D84BEFFF3FFA7FB06F98DCB01F9F0" box="[1252,1329,1590,1615]" pageId="1" type="male">1 male</specimenCount>
(CW 7.0 mm, CL
<quantity id="4CB3E2D2FFF3FFA7FF1FF9E0CF65F9CC" box="[253,341,1626,1651]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" unit="mm" value="5.8">5.8 mm</quantity>
) (
<specimenCode id="DBEDE74CFFF3FFA7FE8FF9E1CC19F9CC" box="[365,553,1626,1651]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1">ZRC 2017.1065</specimenCode>
), same data as neotype
</materialsCitation>
.
<materialsCitation id="3B23456AFFF3FFA7FCDEF9E0CC6DF904" ID-GBIF-Occurrence="1848743985" collectingDate="1998-11-04" collectingDateMax="1999-10-25" collectingDateMin="1998-11-04" collectionCode="ZRC" collectorName="Q. Kazmi" country="Pakistan" latitude="24.763334" location="Korangi Creek" longLatPrecision="21" longitude="67.17472" municipality="Karachi" pageId="1" specimenCode="ZRC 2017.1008" specimenCount="10" specimenCount-female="10" typeStatus="neotype">
<emphasis id="B93F9325FFF3FFA7FCDEF9E0CD9EF9CC" bold="true" box="[828,942,1626,1651]" pageId="1">
<collectingCountry id="F35C0FA7FFF3FFA7FCDEF9E0CD99F9CC" box="[828,937,1626,1651]" name="Pakistan" pageId="1">Pakistan</collectingCountry>
:
</emphasis>
<specimenCount id="9D4D84BEFFF3FFA7FC57F9E1CA00F9CC" box="[949,1072,1626,1651]" pageId="1" type="female">10 females</specimenCount>
(CW 9.213.0 mm, CL
<quantity id="4CB3E2D2FFF3FFA7FAAAF9E0CEF4F928" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.7" metricValueMin="6.9" pageId="1" unit="mm" value="8.3" valueMax="9.7" valueMin="6.9">6.99.7 mm</quantity>
) (
<specimenCode id="DBEDE74CFFF3FFA7FF38F9C5CFA5F928" box="[218,405,1663,1687]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1">ZRC 2017.1008</specimenCode>
),
<location id="8E9419ECFFF3FFA7FE45F9C4CC7FF928" LSID="urn:lsid:plazi:treatment:03E2FE21FFF3FFA1FF75FB39CA95FA5A:8E9419ECFFF3FFA7FE45F9C4CC7FF928" box="[423,591,1662,1687]" country="Pakistan" latitude="24.763334" longLatPrecision="21" longitude="67.17472" municipality="Karachi" name="Korangi Creek" pageId="1">Korangi Creek</location>
,
<collectingMunicipality id="6B90D54DFFF3FFA7FDB8F9C4CC83F928" box="[602,691,1662,1687]" pageId="1">Karachi</collectingMunicipality>
(
<geoCoordinate id="EE7F29F0FFF3FFA7FD23F9C4CD7AF928" box="[705,842,1662,1687]" direction="north" orientation="latitude" pageId="1" precision="15" value="24.763334">24°4548″N</geoCoordinate>
,
<geoCoordinate id="EE7F29F0FFF3FFA7FCB7F9C4CDEBF928" box="[853,987,1662,1687]" direction="east" orientation="longitude" pageId="1" precision="15" value="67.17472">67°1029″E</geoCoordinate>
), in
<taxonomicName id="4C4B34B4FFF3FFA7FBE9F9C4CAFFF929" authorityName="Linnaeus" authorityYear="1758" box="[1035,1231,1662,1686]" class="Bivalvia" family="Placunidae" genus="Placuna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="1" phylum="Mollusca" rank="species" species="placenta">
<emphasis id="B93F9325FFF3FFA7FBE9F9C4CAFFF929" box="[1035,1231,1662,1686]" italics="true" pageId="1">Placuna placenta</emphasis>
</taxonomicName>
, coll.
<collectorName id="26BE2AE1FFF3FFA7FAF7F9C5CBB1F928" box="[1301,1409,1662,1687]" pageId="1">Q. Kazmi</collectorName>
<emphasis id="B93F9325FFF3FFA7FA6AF9C5CE84F904" italics="true" pageId="1">et al.</emphasis>
,
<collectingDate id="EFB1901FFFF3FFA7FF20F919CFBBF904" box="[194,395,1698,1723]" pageId="1" value="1998-11-04">
<date id="FFF569F7FFF3FFA7FF20F919CFBBF904" box="[194,395,1698,1723]" pageId="1" value="1998-11-04">4 November 1998</date>
</collectingDate>
<collectingDate id="EFB1901FFFF3FFA7FE78F918CC69F904" box="[410,601,1698,1723]" pageId="1" value="1999-10-25">
<date id="FFF569F7FFF3FFA7FE78F918CC69F904" box="[410,601,1698,1723]" pageId="1" value="1999-10-25">25 October 1999</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF44F37FFF3FFA7FF25F97CCF0AF8BC" blockId="1.[151,1436,1482,2011]" pageId="1">
<materialsCitation id="3B23456AFFF3FFA7FF25F97CCF0AF8BC" ID-GBIF-Occurrence="1848743984" country="India" county="Rann of Kutch" latitude="22.452223" location="Balapur Bay" longLatPrecision="21" longitude="69.13583" municipality="Gulf of Kachchh" pageId="1" specimenCount="1" stateProvince="Gujarat" typeStatus="holotype">
<emphasis id="B93F9325FFF3FFA7FF25F97CCF6FF960" bold="true" box="[199,351,1734,1759]" pageId="1">
<typeStatus id="54F0F195FFF3FFA7FF25F97CCF32F960" box="[199,258,1734,1759]" pageId="1">Type</typeStatus>
locality
</emphasis>
.
<location id="8E9419ECFFF3FFA7FE8DF97CCFCEF960" LSID="urn:lsid:plazi:treatment:03E2FE21FFF3FFA1FF75FB39CA95FA5A:8E9419ECFFF3FFA7FE8DF97CCFCEF960" box="[367,510,1734,1759]" country="India" county="Rann of Kutch" latitude="22.452223" longLatPrecision="21" longitude="69.13583" municipality="Gulf of Kachchh" name="Balapur Bay" pageId="1" stateProvince="Gujarat">Balapur Bay</location>
(
<geoCoordinate id="EE7F29F0FFF3FFA7FDEDF97CCC91F961" box="[527,673,1734,1759]" direction="north" orientation="latitude" pageId="1" precision="15" value="22.452223">22°2708″ N</geoCoordinate>
,
<geoCoordinate id="EE7F29F0FFF3FFA7FD4CF97CCD0CF960" box="[686,828,1734,1759]" direction="east" orientation="longitude" pageId="1" precision="15" value="69.13583">69°0809″ E</geoCoordinate>
),
<location id="8E9419ECFFF3FFA7FCB0F97DCDDBF960" LSID="urn:lsid:plazi:treatment:03E2FE21FFF3FFA1FF75FB39CA95FA5A:8E9419ECFFF3FFA7FCB0F97DCDDBF960" box="[850,1003,1734,1759]" country="India" county="Rann of Kutch" latitude="22.452223" longLatPrecision="21" longitude="69.13583" municipality="Gulf of Kachchh" name="Beyt Dwarka" pageId="1" stateProvince="Gujarat">Beyt Dwarka</location>
,
<collectingMunicipality id="6B90D54DFFF3FFA7FC1AF97CCA89F960" box="[1016,1209,1734,1759]" pageId="1">Gulf of Kachchh</collectingMunicipality>
(=
<collectingCounty id="629537BBFFF3FFA7FB01F97DCBA0F960" box="[1251,1424,1734,1759]" pageId="1">Rann of Kutch</collectingCounty>
),
<collectingRegion id="498F81D5FFF3FFA7FF75F950CEDEF8BC" box="[151,238,1770,1795]" country="India" name="Gujarat" pageId="1">Gujarat</collectingRegion>
,
<collectingCountry id="F35C0FA7FFF3FFA7FF1BF950CF06F8BC" box="[249,310,1770,1795]" name="India" pageId="1">India</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3511CBCFFF3FFA2FF25F8B4CC3BFCE3" lastPageId="4" pageId="1" type="description">
<paragraph id="8BF44F37FFF3FFA4FF25F8B4CCCAFE13" blockId="1.[151,1436,1482,2011]" lastBlockId="2.[151,1437,151,429]" lastPageId="2" pageId="1">
<emphasis id="B93F9325FFF3FFA7FF25F8B4CF61F898" bold="true" box="[199,337,1806,1831]" pageId="1">Description</emphasis>
. Female: carapace subhexagonal (
<figureCitation id="137053B2FFF3FFA7FD03F8B4CDB1F899" box="[737,897,1806,1831]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="1">Figs.1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF3FFA7FC72F8B5CD85F898" box="[912,949,1807,1831]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="1">2A</figureCitation>
), slightly wider than long, regions clearly defined with prominent shallow cervical and cardiac grooves, anterior margin flat, clearly defined, front not projecting but demarcated by emargination above each orbit, posterior margin concave. Eyes small, pigmented, not visible in dorsal view; orbits oval. Antennules folded obliquely (
<figureCitation id="137053B2FFF3FFA7FC6BF8C0CDD5F82C" box="[905,997,1914,1939]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="1">Fig. 2B</figureCitation>
), antennular fossa slightly larger than orbit; antennae with 6 articles, basal article with large round urinal article, fused with epistome, distal articles setose, reaching beyond inner edge of orbit. Epistome and endostome separated, margin of first sternite emarginated. MXP3 obliquely placed in buccal cavity, outer surface with few scattered setae on mesial margin (
<figureCitation id="137053B2FFF0FFA4FF7DFF06CEC5FF6B" box="[159,245,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="2">Fig. 2F</figureCitation>
), inner margin with long scattered setae (
<figureCitation id="137053B2FFF0FFA4FD24FF06CD10FF6B" box="[710,800,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="2">Fig. 2G</figureCitation>
); propodus about 2.2 times as long as high, stout, longer than carpus, dorsal margin with fringe of long setae, anterior margin rounded with long setae; dactylus digitiform distally setose, inserted medially in notch on ventral margin of propodus, distally not overreaching tip of propodus but overreaching anteromesial angle of merus; ischiomerus completely fused, 2.6 times as long as wide, lateral margin convex without setae, mesial margin slightly angular at the widest point forming antero mesial angle, inner margin with long setae. Exopod slightly longer than half length of ischiomerus, with shallow median furrow, flagellum with 2 articles before fine setae (
<figureCitation id="137053B2FFF0FFA4FD95FE2ECCDDFE13" box="[631,749,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="2">Fig. 2F, G</figureCitation>
).
</paragraph>
<caption id="DF341FBFFFF0FFA4FF75F97FCB55F8A7" httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="2" startId="2.[151,250,1733,1755]" targetBox="[151,1436,483,1708]" targetPageId="2">
<paragraph id="8BF44F37FFF0FFA4FF75F97FCB55F8A7" blockId="2.[151,1436,1732,1816]" pageId="2">
<emphasis id="B93F9325FFF0FFA4FF75F97FCF25F965" bold="true" box="[151,277,1733,1755]" pageId="2">FIGURE 1.</emphasis>
<taxonomicName id="4C4B34B4FFF0FFA4FEFFF97ECD16F965" authority="(Hornell &amp; Southwell, 1909)" authorityName="Hornell &amp; Southwell" authorityYear="1909" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[285,806,1732,1754]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF0FFA4FEFFF97ECFDFF965" box="[285,495,1732,1754]" italics="true" pageId="2">Arcotheres placunae</emphasis>
(Hornell &amp; Southwell, 1909)
</taxonomicName>
. A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view.
</paragraph>
</caption>
<paragraph id="8BF44F37FFF0FFA5FF25F8FACFE6FF47" blockId="2.[151,1436,1855,2024]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" pageId="2">
P1 (cheliped) stouter than P2P5 (
<figureCitation id="137053B2FFF0FFA4FDB0F885CCDFF8E7" box="[594,751,1855,1880]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="2">Fig. 1AD, F</figureCitation>
), merus slightly longer than carpus, chela relatively slender (
<figureCitation id="137053B2FFF0FFA4FF7DF8D9CF2AF8C3" box="[159,282,1891,1916]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="2">Fig. 2D, E</figureCitation>
), ventral margin straight, outer margin slightly curved, palm about 1.8 times as long as high, dactylus and pollex slightly shorter than dorsal margin of palm, longer than greatest height of palm, palm relatively stout, dactylus dorsal antero distal margin with patch of long setae, cutting edge with 1 large proximal tooth, ends in sharp claw, covered with setae. Pollex proximal margin with patch of long setae and distal margin with row of small setae present medially visible on inner surface (
<figureCitation id="137053B2FFF1FFA5FD13FF2DCD77FF0F" box="[753,839,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="3">Fig. 2E</figureCitation>
), outer surface smooth (
<figureCitation id="137053B2FFF1FFA5FBB4FF2DCA80FF0F" box="[1110,1200,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="3">Fig. 2D</figureCitation>
), cutting edge with 2 large proximal teeth followed by undulating margin ending with sharp claw, covered with setae, ventral margin with long setae (
<figureCitation id="137053B2FFF1FFA5FEB2FF65CFF9FF47" box="[336,457,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="3">Fig. 2D, E</figureCitation>
).
</paragraph>
<caption id="DF341FBFFFF1FFA5FF75F9A4CF68F9CD" httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="3" startId="3.[151,250,1566,1588]" targetBox="[160,1417,315,1521]" targetPageId="3">
<paragraph id="8BF44F37FFF1FFA5FF75F9A4CF68F9CD" blockId="3.[151,1436,1566,1650]" pageId="3">
<emphasis id="B93F9325FFF1FFA5FF75F9A4CF22F98B" bold="true" box="[151,274,1566,1588]" pageId="3">FIGURE 2.</emphasis>
<taxonomicName id="4C4B34B4FFF1FFA5FEFBF9A4CD26F98B" authority="(Hornell &amp; Southwell, 1909)" authorityName="Hornell &amp; Southwell" authorityYear="1909" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[281,790,1566,1588]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF1FFA5FEFBF9A4CFD8F98B" box="[281,488,1566,1588]" italics="true" pageId="3">Arcotheres placunae</emphasis>
(Hornell &amp; Southwell, 1909)
</taxonomicName>
, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view.
</paragraph>
</caption>
<paragraph id="8BF44F37FFF1FFA5FF25F924CA65F868" blockId="3.[151,1437,1694,2007]" pageId="3">
Ambulatory legs almost symmetrical in length except P4 (
<figureCitation id="137053B2FFF1FFA5FCB7F924CDE2F908" box="[853,978,1694,1719]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="3">Fig. 4AH</figureCitation>
); P2, P3 similar, merus, carpus glabrous, propodus longer than carpus with scattered setae on distodorsal and distoventral margin, dactylus shorter than carpus and propodus, hooked at tip, ventral margin covered with short setae, outer surface with scattered setae, P2 dactylus dorsal margin with few short setae; P4 markedly asymmetrical, longer P4 stouter than P2 and P3, carpus outer margin slightly curved, dactylus sickle-like, slightly smaller than propodus, ends in small curved tip, longer than dactyli of P2 and P3, dorsal, ventral and outer margin with short setae. P5 more slender than P2P4; segments similar in shape than those of P2 and P3; except dactylus, longer than those of P2P4, dorsal, ventral and outer margins with short setae, tip with row of small spines, ending in sharp curved spine (
<figureCitation id="137053B2FFF1FFA5FBB6F820CAA7F80C" box="[1108,1175,1946,1971]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="3">Fig. 4</figureCitation>
H'). Relative lengths of ambulatory legs P4&gt;P3&gt;P2&gt;P5 and dactyli of longer ambulatory legs P5&gt;P4&gt;P3&gt;P2.
</paragraph>
<paragraph id="8BF44F37FFF6FFA2FF25FF2DCAA5FF6A" blockId="4.[151,1437,151,2013]" pageId="4">
Pleon wide, ovate (
<figureCitation id="137053B2FFF6FFA2FE41FF2DCC36FF0F" box="[419,518,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="4">Figs. 1B</figureCitation>
,
<figureCitation id="137053B2FFF6FFA2FDF6FF22CC08FF0F" box="[532,568,152,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="4">2C</figureCitation>
), with 6 free somites and telson, covering bases of ambulatory legs; pleonal somites increasing in length distally, widening from somite 15, decreasing towards telson.
</paragraph>
<paragraph id="8BF44F37FFF6FFA2FF25FF65CF7BFDA7" blockId="4.[151,1437,151,2013]" pageId="4">
Male: carapace subhexagonal (
<figureCitation id="137053B2FFF6FFA2FDC0FF65CCB5FF47" box="[546,645,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="4">Figs. 1D</figureCitation>
,
<figureCitation id="137053B2FFF6FFA2FD70FF5ACC87FF47" box="[658,695,224,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[162,1428,816,1872]" captionTargetId="figure@5.[151,1436,787,1900]" captionTargetPageId="5" captionText="FIGURE 3. Arcotheres placunae (Hornell &amp; Southwell, 1909). AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded)." httpUri="https://zenodo.org/record/1287099/files/figure.png" pageId="4">3A</figureCitation>
), wider than long, regions clearly defined with prominent shallow cervical and cardiac grooves, front broadly triangular, projecting anteriorly, emarginated, with flat anterior margin, eyes visible in dorsal view. MXP3 as in female. Chela slightly stouter than in female (
<figureCitation id="137053B2FFF6FFA2FB61FE9DCB3CFE80" box="[1155,1292,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="4">Figs. 1D, E</figureCitation>
,
<figureCitation id="137053B2FFF6FFA2FAF8FE92CB0BFEFF" box="[1306,1339,296,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[162,1428,816,1872]" captionTargetId="figure@5.[151,1436,787,1900]" captionTargetPageId="5" captionText="FIGURE 3. Arcotheres placunae (Hornell &amp; Southwell, 1909). AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded)." httpUri="https://zenodo.org/record/1287099/files/figure.png" pageId="4">3C</figureCitation>
) margin slightly curved, ventral margin straight, outer margin covered with short scattered setae. Dactylus slightly shorter than dorsal margin of palm, few scattered setae present on distodorsal margin, outer surface with short scattered setae, ending in sharp claw, cutting edge denticulate with 1 large tooth proximally, covered with short setae. Pollex cutting edge with 2 large teeth separated by deep notch proximally followed by denticulate ridge ending before reaching distal margin, ending in sharp claw, ventral margin with fringe of short setae ending on mid-ventral margin of palm.
</paragraph>
<paragraph id="8BF44F37FFF6FFA2FF25FD9ECB00FD72" blockId="4.[151,1437,151,2013]" pageId="4">
P2P5 (
<figureCitation id="137053B2FFF6FFA2FEC2FD9ECFBCFD83" box="[288,396,548,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="4">
Fig.
<date id="FFF569F7FFF6FFA2FEB6FD9ECF5BFD83" box="[340,363,548,572]" pageId="4">4I</date>
L
</figureCitation>
) more slender than cheliped; merus with fringe of short setae on dorsal and ventral margins; carpus with strongly curved outer margin; propodus longer than carpus with curved outer margin; dactylus dorsal and ventral margins with short setae. P3 and P4 with fringe of long swimming setae on outer face of carpus and propodus increasing in size distally; tip of P5 dactylus with row of small spines ending in sharp spine (
<figureCitation id="137053B2FFF6FFA2FAD0FD35CB46FD17" box="[1330,1398,655,680]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="4">Fig. 4</figureCitation>
L'). Relative lengths of ambulatory legs P4&gt;P3&gt;P2&gt;P5 and dactyli of longer ambulatory legs P5&gt;P4&gt;P3&gt;P2.
</paragraph>
<paragraph id="8BF44F37FFF6FFA2FF25FD6DCC3BFCE3" blockId="4.[151,1437,151,2013]" pageId="4">
Pleon (
<figureCitation id="137053B2FFF6FFA2FEFBFD6DCF41FD4F" box="[281,369,727,752]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[162,1428,816,1872]" captionTargetId="figure@5.[151,1436,787,1900]" captionTargetPageId="5" captionText="FIGURE 3. Arcotheres placunae (Hornell &amp; Southwell, 1909). AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded)." httpUri="https://zenodo.org/record/1287099/files/figure.png" pageId="4">Fig. 3B</figureCitation>
) broadly triangular, with six free somites and telson, widest at somite 3, decreasing towards evenly rounded telson, lateral and dorsal margins with short setae. G1(
<figureCitation id="137053B2FFF6FFA2FC44FD46CDCEFCAB" box="[934,1022,764,789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[162,1428,816,1872]" captionTargetId="figure@5.[151,1436,787,1900]" captionTargetPageId="5" captionText="FIGURE 3. Arcotheres placunae (Hornell &amp; Southwell, 1909). AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded)." httpUri="https://zenodo.org/record/1287099/files/figure.png" pageId="4">Fig. 3D</figureCitation>
) long, curved, with short angular tip, shallow furrow running parallel to dorsal margin, dorsal and ventral surface with long setae; G2 stout with blunt tip, exopod elongate (
<figureCitation id="137053B2FFF6FFA2FE68FCFECFCFFCE3" box="[394,511,836,861]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[162,1428,816,1872]" captionTargetId="figure@5.[151,1436,787,1900]" captionTargetPageId="5" captionText="FIGURE 3. Arcotheres placunae (Hornell &amp; Southwell, 1909). AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded)." httpUri="https://zenodo.org/record/1287099/files/figure.png" pageId="4">Fig. 3E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3511CBCFFF6FFA2FF25FCDDCA5DFB1F" pageId="4" type="distribution">
<paragraph id="8BF44F37FFF6FFA2FF25FCDDCA5DFB1F" blockId="4.[151,1437,151,2013]" pageId="4">
<emphasis id="B93F9325FFF6FFA2FF25FCDDCF50FC3F" bold="true" box="[199,352,871,896]" pageId="4">Distribution.</emphasis>
In
<collectingCountry id="F35C0FA7FFF6FFA2FE68FCDDCFF6FC3F" box="[394,454,871,896]" name="India" pageId="4">India</collectingCountry>
, the species has been recorded from various localities:
<collectingRegion id="498F81D5FFF6FFA2FBA7FCDDCAA9FC3F" box="[1093,1177,871,896]" country="India" name="Gujarat" pageId="4">Gujarat</collectingRegion>
(
<bibRefCitation id="EFDA32C6FFF6FFA2FB48FCDDCEE3FC1B" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
;
<bibRefCitation id="EFDA32C6FFF6FFA2FF3DFC36CC3DFC1A" box="[223,525,908,933]" pageId="4" refId="ref6617" refString="Subba Rao, N. V. &amp; Sastry, D. R. K. (2005) Fauna of Marine National Park, Gulf of Kachchh (Gujarat). An overview. In: Conservation Area Series. Vol. 23. Zoological Survey India, Kolkata, pp. 1 - 79." type="book">Subba Rao &amp; Sastry 2005</bibRefCitation>
),
<collectingRegion id="498F81D5FFF6FFA2FDC0FC36CC9EFC1A" box="[546,686,908,933]" country="India" name="Maharashtra" pageId="4">Maharashtra</collectingRegion>
(
<bibRefCitation id="EFDA32C6FFF6FFA2FD5DFC36CD4DFC1A" box="[703,893,908,933]" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar 1957a</bibRefCitation>
,
<bibRefCitation id="EFDA32C6FFF6FFA2FC68FC36CDABFC1A" box="[906,923,908,933]" pageId="4" refId="ref5738" refString="Chhapgar, B. F. (1957 b) Marine crabs of Bombay State. Contribution No. 1 of the Taraporevala Marine Biological Station Marine Biological Station. Department of Fisheries, Bombay, 89 pp." type="book">b</bibRefCitation>
;
<bibRefCitation id="EFDA32C6FFF6FFA2FC4BFC36CA94FC1B" box="[937,1188,908,933]" pageId="4" refId="ref6716" refString="Yeragi, S. G. &amp; Yeragi, S. S. (1997) A study on Pinnotheres crabs associated with newly recorded Soleteuiiva violacea (Lam.) and Placuna placenta (Linn.) Journal of Marine Biological Association of India, 39 (1 - 2), 188 - 193." type="journal article">Yeragi &amp; Yeragi 1997</bibRefCitation>
) and
<collectingRegion id="498F81D5FFF6FFA2FB04FC36CBACFC1A" box="[1254,1436,908,933]" country="India" name="Andhra Pradesh" pageId="4">Andhra Pradesh</collectingRegion>
(
<bibRefCitation id="EFDA32C6FFF6FFA2FF7CFC15CF5BFC77" box="[158,363,943,968]" pageId="4" refId="ref6121" refString="Narasimham, K. A. (1984) Biology of windowpane oyster Placenta placenta (Linnaeus) in Kakinada Bay. Indian Journal of Fisheries, 31, 272 - 284." type="journal article">Narasimham1984</bibRefCitation>
;
<bibRefCitation id="EFDA32C6FFF6FFA2FE9BFC15CC7BFC77" box="[377,587,943,968]" pageId="4" refId="ref5811" refString="Devi, S. L. &amp; Rao, K. H. (1986) Some aspects of biology of the pea crab Pinnotheres placunae Hornell &amp; Southwell. Proceeding of the Indian National Science Academy, Part B, 52 (4), 455 - 459." type="journal article">Devi &amp; Rao 1986</bibRefCitation>
). It is also present in
<collectingCountry id="F35C0FA7FFF6FFA2FCA8FC15CD99FC77" box="[842,937,943,968]" name="Pakistan" pageId="4">Pakistan</collectingCountry>
(present study). In describing
<taxonomicName id="4C4B34B4FFF6FFA2FAEEFC0ACFE7FC53" authority=", Hornell &amp; Southwell (1909)" authorityName=", Hornell &amp; Southwell" authorityYear="1909" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF6FFA2FAEEFC0ACBA4FC77" box="[1292,1428,944,968]" italics="true" pageId="4">A. placunae</emphasis>
,
<bibRefCitation id="EFDA32C6FFF6FFA2FF75FC6ECFE7FC53" box="[151,471,980,1005]" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell (1909)</bibRefCitation>
</taxonomicName>
reported that they had on hand
<specimenCount id="9D4D84BEFFF6FFA2FCA1FC6ECDAEFC52" box="[835,926,980,1005]" pageId="4" type="male">31 male</specimenCount>
specimens and three female specimens from Balapur Bay in the Gulf of Kachchh. They, however, also noted that “
<emphasis id="B93F9325FFF6FFA2FC22FC42CA1AFBAF" box="[960,1066,1016,1040]" italics="true" pageId="4">Placunae</emphasis>
from
<collectingCountry id="F35C0FA7FFF6FFA2FB91FC4DCAF4FBAF" box="[1139,1220,1015,1040]" name="Sri Lanka" pageId="4">Ceylon</collectingCountry>
rarely contain this commensal. One large female was, however, taken by one of us some years ago from a large
<taxonomicName id="4C4B34B4FFF6FFA2FB0DFBA6CB7DFB8B" box="[1263,1357,1052,1076]" class="Bivalvia" family="Placunidae" genus="Placuna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="4" phylum="Mollusca" rank="genus">
<emphasis id="B93F9325FFF6FFA2FB0DFBA6CB7DFB8B" box="[1263,1357,1052,1076]" italics="true" pageId="4">Placuna</emphasis>
</taxonomicName>
fished at Tampalakam Bay, proving the geographical range to extend from the Gulf of Kutch to
<collectingCountry id="F35C0FA7FFF6FFA2FB4CFB85CB34FBE7" box="[1198,1284,1087,1112]" name="Sri Lanka" pageId="4">Ceylon</collectingCountry>
.” (
<bibRefCitation id="EFDA32C6FFF6FFA2FAC6FB85CFB8FBC3" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909: 101</bibRefCitation>
). This record from
<collectingCountry id="F35C0FA7FFF6FFA2FD89FBDECCEBFBC2" box="[619,731,1124,1149]" name="Sri Lanka" pageId="4">Sri Lanka</collectingCountry>
(=
<collectingCountry id="F35C0FA7FFF6FFA2FCE5FBDECD6AFBC2" box="[775,858,1124,1149]" name="Sri Lanka" pageId="4">Ceylon</collectingCountry>
) will need to be confirmed in any case with fresh collections. There is no indication, however, if this specimen was retained or preserved.
</paragraph>
</subSubSection>
<subSubSection id="C3511CBCFFF6FFA2FF25FB16CDCDF8F2" pageId="4" type="discussion">
<paragraph id="8BF44F37FFF6FFA2FF25FB16CFA7FA8F" blockId="4.[151,1437,151,2013]" pageId="4">
<emphasis id="B93F9325FFF6FFA2FF25FB16CF04FB7A" bold="true" box="[199,308,1196,1221]" pageId="4">Remarks</emphasis>
.
<bibRefCitation id="EFDA32C6FFF6FFA2FEA0FB16CC87FB7B" box="[322,695,1196,1221]" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell (1909: 100)</bibRefCitation>
described the female P4 and P5 dactyli of
<taxonomicName id="4C4B34B4FFF6FFA2FB75FB16CB4DFB7B" authorityName=", Hornell &amp; Southwell" authorityYear="1909" box="[1175,1405,1196,1220]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF6FFA2FB75FB16CB4DFB7B" box="[1175,1405,1196,1220]" italics="true" pageId="4">Arcotheres placunae</emphasis>
</taxonomicName>
as one and a half times longer than those of the P2 and P3, but this observation does not match with their figures of a female specimen, which show P2P4 symmetrical and with all the dactyli subequal in length (
<bibRefCitation id="EFDA32C6FFF6FFA2FB52FB4ECFBBFA8F" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909: pl. 1 figs. 310</bibRefCitation>
).
</paragraph>
<paragraph id="8BF44F37FFF6FFA2FF25FA86CAD7F993" blockId="4.[151,1437,151,2013]" pageId="4">
<bibRefCitation id="EFDA32C6FFF6FFA2FF25FA86CFA2FAEA" box="[199,402,1340,1365]" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar (1957a)</bibRefCitation>
also recorded
<taxonomicName id="4C4B34B4FFF6FFA2FDD4FA86CC8CFAEB" authorityName=", Hornell &amp; Southwell" authorityYear="1909" box="[566,700,1340,1364]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF6FFA2FDD4FA86CC8CFAEB" box="[566,700,1340,1364]" italics="true" pageId="4">A. placunae</emphasis>
</taxonomicName>
from
<collectingCountry id="F35C0FA7FFF6FFA2FCE3FA86CD53FAEA" box="[769,867,1340,1365]" name="India" pageId="4">Mumbai</collectingCountry>
,
<collectingRegion id="498F81D5FFF6FFA2FC8CFA86CDCAFAEA" box="[878,1018,1340,1365]" country="India" name="Maharashtra" pageId="4">Maharashtra</collectingRegion>
state,
<collectingCountry id="F35C0FA7FFF6FFA2FBA1FA86CAB0FAEA" box="[1091,1152,1340,1365]" name="India" pageId="4">India</collectingCountry>
. The figure of the female specimen in
<bibRefCitation id="EFDA32C6FFF6FFA2FEC7FAE5CFC1FAC7" box="[293,497,1375,1400]" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar (1957a)</bibRefCitation>
is imprecise in many ways and also does not match his description. His figure (see
<bibRefCitation id="EFDA32C6FFF6FFA2FF75FA3ECF6BFA22" box="[151,347,1412,1437]" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar 1957a</bibRefCitation>
: pl. 12 fig. a) shows the right P2P4 as longer than those on the left but based on the measurements of P2P
<quantity id="4CB3E2D2FFF6FFA2FE42FA12CFFEFA7F" box="[416,462,1447,1472]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" unit="in" value="4.0">4 in</quantity>
his description, the left P2P4 should be longer than those on the right.
<bibRefCitation id="EFDA32C6FFF6FFA2FACDFA1DCEDEFA5A" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar (1957a)</bibRefCitation>
also described the P4 and P5 dactyli being one and a half times longer than those of the P2 and P3, but his figure (
<bibRefCitation id="EFDA32C6FFF6FFA2FF05FA55CF95F9B7" box="[231,421,1519,1544]" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar 1957a</bibRefCitation>
: pl. 12 fig. a) shows the left P5 dactylus to be shorter than those on the left P2P4; while on the right side, the P3 dactylus is longer than those of P2, P4 and P5 (
<bibRefCitation id="EFDA32C6FFF6FFA2FC77F9AECA64F992" box="[917,1108,1556,1581]" pageId="4" refId="ref5696" refString="Chhapgar, B. F. (1957 a) On the marine crabs (Decapoda: Brachyura) of Bombay State, part 2. Journal of Bombay Natural History Society, 54 (3), 503 - 549." type="journal article">Chhapgar 1957a</bibRefCitation>
:pl. 12 fig. a).
</paragraph>
<paragraph id="8BF44F37FFF6FFA2FF25F98DCDCDF8F2" blockId="4.[151,1437,151,2013]" pageId="4">
The specimens examined in the present study generally agree with the description and figures of the female carapace, male pleon and some pereiopods provided by
<bibRefCitation id="EFDA32C6FFF6FFA2FCF2F9E6CA62F9CB" box="[784,1106,1628,1653]" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell (1909)</bibRefCitation>
.
<bibRefCitation id="EFDA32C6FFF6FFA2FBBFF9E6CBACF9CB" box="[1117,1436,1628,1653]" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell (1909)</bibRefCitation>
mentioned in the description that the dactyli of the female P4 and P5 are 1.5 times longer than those of P2 and P3 but the artist must have simplified the drawings by making all the dactyli of P2P5 equal in length. The dactyli of P2P5 of the female specimens examined in the present study, however, match the description provided by
<bibRefCitation id="EFDA32C6FFF6FFA2FAA5F97DCF4BF8BB" pageId="4" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell (1909)</bibRefCitation>
.
<taxonomicName id="4C4B34B4FFF6FFA2FE64F956CC5DF8BB" authorityName=", Hornell &amp; Southwell" authorityYear="1909" box="[390,621,1772,1796]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF6FFA2FE64F956CC5DF8BB" box="[390,621,1772,1796]" italics="true" pageId="4">Arcotheres placunae</emphasis>
</taxonomicName>
closely resembles species such as
<taxonomicName id="4C4B34B4FFF6FFA2FC16F956CB28F8BA" authority="(Burger 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1012,1304,1772,1797]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF6FFA2FC16F956CA42F8BB" box="[1012,1138,1772,1796]" italics="true" pageId="4">A. latifrons</emphasis>
(
<bibRefCitation id="EFDA32C6FFF6FFA2FB62F956CB20F8BA" box="[1152,1296,1772,1797]" pageId="4" refId="ref5578" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger 1895</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4B34B4FFF6FFA2FAADF956CFB2F897" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF6FFA2FAADF956CBACF8BB" box="[1359,1436,1772,1796]" italics="true" pageId="4">A. rayi</emphasis>
<bibRefCitation id="EFDA32C6FFF6FFA2FF75F8B5CFB2F897" box="[151,386,1807,1833]" pageId="4" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng, 2007</bibRefCitation>
</taxonomicName>
(both from
<collectingRegion id="498F81D5FFF6FFA2FDEFF8B5CC64F897" box="[525,596,1807,1832]" country="Philippines" name="Bohol" pageId="4">Bohol</collectingRegion>
,
<collectingCountry id="F35C0FA7FFF6FFA2FD80F8B5CCD2F897" box="[610,738,1807,1832]" name="Philippines" pageId="4">Philippines</collectingCountry>
) in having a subtrapezoidal carapace, a flat anterior carapace margin and having the MXP3 dactylus not reaching the apex of the propodus.
</paragraph>
</subSubSection>
<subSubSection id="C3511CBCFFF6FFA1FF25F8E2CAFFFBAF" lastPageId="7" pageId="4" type="reference_group">
<paragraph id="8BF44F37FFF6FFA3FF25F8E2CCEAFD72" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,717]" lastPageId="5" pageId="4">
<taxonomicName id="4C4B34B4FFF6FFA2FF25F8E2CF81F8CF" ID-CoL="67PXW" authorityName=", Hornell &amp; Southwell" authorityYear="1909" box="[199,433,1880,1904]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF6FFA2FF25F8E2CF81F8CF" box="[199,433,1880,1904]" italics="true" pageId="4">Arcotheres placunae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4B34B4FFF6FFA2FDAFF8E2CCFEF8CF" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[589,718,1880,1904]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF6FFA2FDAFF8E2CCFEF8CF" box="[589,718,1880,1904]" italics="true" pageId="4">A. latifrons</emphasis>
</taxonomicName>
in the following features: 1) the female carapace is 1.4 times wider than long (
<figureCitation id="137053B2FFF6FFA2FEB4F8C6CFC7F82B" box="[342,503,1916,1941]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="4">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF6FFA2FDE0F8C6CC16F82B" box="[514,550,1916,1940]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="4">2A</figureCitation>
) (vs. carapace 1.1 time wider than long in
<emphasis id="B93F9325FFF6FFA2FBE7F8C6CABAF82B" box="[1029,1162,1916,1940]" italics="true" pageId="4">
<taxonomicName id="4C4B34B4FFF6FFA2FBE7F8C6CAB5F82B" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1029,1157,1916,1940]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="latifrons">A. latifrons</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF6FFA2FB56F8C6CBA8F82B" box="[1204,1432,1916,1941]" pageId="4" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5A); 2) the eyes are present but not visible in dorsal view (
<figureCitation id="137053B2FFF6FFA2FCBCF81ACA31F807" box="[862,1025,1952,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="4">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF6FFA2FBEFF81ACA01F807" box="[1037,1073,1952,1976]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="4">2A</figureCitation>
) (vs. eyes absent in
<emphasis id="B93F9325FFF6FFA2FAF5F81ACBACF807" box="[1303,1436,1952,1976]" italics="true" pageId="4">
<taxonomicName id="4C4B34B4FFF6FFA2FAF5F81ACBA8F807" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1303,1432,1952,1976]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="latifrons">A. latifrons</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF6FFA2FF5FF87ECF98F863" box="[189,424,1988,2013]" pageId="4" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5A); 3) the carapace posterior margin is strongly concave (
<figureCitation id="137053B2FFF6FFA2FB68F87ECB00F863" box="[1162,1328,1988,2013]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="4">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF6FFA2FADCF87ECB52F863" box="[1342,1378,1988,2012]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="4">2A</figureCitation>
) (vs. moderately concave in
<taxonomicName id="4C4B34B4FFF7FFA3FE7FFF22CC2BFF0F" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[413,539,152,176]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FE7FFF22CC2BFF0F" box="[413,539,152,176]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FDADFF2DCD04FF0F" box="[591,820,151,176]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5A); 4) the antennae reach beyond the inner edge of the orbit (
<figureCitation id="137053B2FFF7FFA3FEC3FF06CF46FF6B" box="[289,374,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2B</figureCitation>
) (vs. do not reach the inner edge of the orbit in
<taxonomicName id="4C4B34B4FFF7FFA3FC65FF06CA33FF6B" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[903,1027,188,212]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FC65FF06CA33FF6B" box="[903,1027,188,212]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FBD6FF06CB26FF6B" box="[1076,1302,188,213]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5B); 5) the MXP3 propodus is 2.2 times as long as high (
<figureCitation id="137053B2FFF7FFA3FD5CFF65CD01FF47" box="[702,817,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2F, G</figureCitation>
) (vs. 3 times as long as high in
<taxonomicName id="4C4B34B4FFF7FFA3FB71FF5ACB20FF47" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1171,1296,224,248]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FB71FF5ACB20FF47" box="[1171,1296,224,248]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FAA0FF65CF28FEA3" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5D); 6) the anterior margin of the MXP3 propodus is rounded with long setae (
<figureCitation id="137053B2FFF7FFA3FB22FEBECB04FEA3" box="[1216,1332,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2F, G</figureCitation>
) (vs. less rounded with short setae in
<taxonomicName id="4C4B34B4FFF7FFA3FE2CFE92CC7BFEFF" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[462,587,296,320]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FE2CFE92CC7BFEFF" box="[462,587,296,320]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FD9FFE9DCD52FEFF" box="[637,866,295,320]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5D); 7) the MXP3 ischiomerus is 2.6 times as long as wide (
<figureCitation id="137053B2FFF7FFA3FED7FEF6CF99FEDB" box="[309,425,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2F, G</figureCitation>
) (vs. less than 2 times as long as wide in
<taxonomicName id="4C4B34B4FFF7FFA3FC9FFEF6CDCAFEDB" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[893,1018,332,356]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FC9FFEF6CDCAFEDB" box="[893,1018,332,356]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FBCFFEF6CB21FEDB" box="[1069,1297,332,357]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5D); 8) the mesial margin of the MXP3 ischiomerus is slightly angular at its widest point (
<figureCitation id="137053B2FFF7FFA3FBC8FED5CAACFE37" box="[1066,1180,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2F, G</figureCitation>
) (vs. distinctly angular in
<taxonomicName id="4C4B34B4FFF7FFA3FF57FE2ECF04FE13" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[181,308,404,428]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FF57FE2ECF04FE13" box="[181,308,404,428]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FE8BFE2ECC62FE13" box="[361,594,404,429]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5D); 9) the inner and outer margins of the MXP3 exopod are almost straight (
<figureCitation id="137053B2FFF7FFA3FF1FFE0DCF64FE6F" box="[253,340,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2F</figureCitation>
) (vs. inner and outer margin convex in
<taxonomicName id="4C4B34B4FFF7FFA3FCCAFE02CD99FE6F" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[808,937,440,464]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FCCAFE02CD99FE6F" box="[808,937,440,464]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FC03FE0DCAFFFE6F" box="[993,1231,439,464]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5D); 10) the cheliped dactylus is slightly shorter than the dorsal length of the palm (
<figureCitation id="137053B2FFF7FFA3FC24FE66CA70FE4B" box="[966,1088,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="5">Fig. 2D, E</figureCitation>
) (vs. dactylus about half palm length in
<taxonomicName id="4C4B34B4FFF7FFA3FEE6FDBACFB3FDA7" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[260,387,512,536]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FEE6FDBACFB3FDA7" box="[260,387,512,536]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FE5AFE45CC91FDA7" box="[440,673,511,536]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5C); 11) the P2 and P3 dactyli have short setae on the dorsal, ventral and lateral margins (
<figureCitation id="137053B2FFF7FFA3FE33FD9ECCB7FD83" box="[465,647,548,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="5">Fig. 4A, B, E, F</figureCitation>
) (vs. dactyli glabrous in
<taxonomicName id="4C4B34B4FFF7FFA3FC7CFD9ECA2BFD83" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[926,1051,548,572]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FC7CFD9ECA2BFD83" box="[926,1051,548,572]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FBAFFD9ECB00FD83" box="[1101,1328,548,573]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5A); 12) the P4 dactylus has short setae on the dorsal, ventral and lateral margins (
<figureCitation id="137053B2FFF7FFA3FBDEFDFDCA8DFDDF" box="[1084,1213,583,608]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="5">Fig. 4C, G</figureCitation>
) (vs. few setae on distoventral margin in
<taxonomicName id="4C4B34B4FFF7FFA3FE77FDD6CC23FD3B" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[405,531,620,644]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FE77FDD6CC23FD3B" box="[405,531,620,644]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FDA7FDD6CD19FD3B" box="[581,809,620,645]" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5A); and 13) the P5 dactylus has short setae on the dorsal, ventral and lateral margins (
<figureCitation id="137053B2FFF7FFA3FDC2FD35CCA8FD17" box="[544,664,655,680]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="5">Fig. 4D, H</figureCitation>
) (vs. ventral margin with short setae in
<taxonomicName id="4C4B34B4FFF7FFA3FBB2FD2ACAFDFD17" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1104,1229,656,680]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FBB2FD2ACAFDFD17" box="[1104,1229,656,680]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF7FFA3FB1FFD35CEE4FD73" pageId="5" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 5A). Males of
<taxonomicName id="4C4B34B4FFF7FFA3FE4CFD0ECC1BFD73" ID-CoL="67Q9X" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[430,555,692,716]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93F9325FFF7FFA3FE4CFD0ECC1BFD73" box="[430,555,692,716]" italics="true" pageId="5">A. latifrons</emphasis>
</taxonomicName>
are not known.
</paragraph>
<caption id="DF341FBFFFF7FFA3FF75F821CC99F851" httpUri="https://zenodo.org/record/1287099/files/figure.png" pageId="5" startId="5.[151,250,1947,1969]" targetBox="[162,1428,816,1872]" targetPageId="5">
<paragraph id="8BF44F37FFF7FFA3FF75F821CC99F851" blockId="5.[151,1436,1947,2030]" pageId="5">
<emphasis id="B93F9325FFF7FFA3FF75F821CF23F80F" bold="true" box="[151,275,1947,1969]" pageId="5">FIGURE 3.</emphasis>
<taxonomicName id="4C4B34B4FFF7FFA3FEFBF821CD29F80E" ID-CoL="67PXW" authority="(Hornell &amp; Southwell, 1909)" authorityName="Hornell &amp; Southwell" authorityYear="1909" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[281,793,1947,1969]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF7FFA3FEFBF821CFDAF80E" box="[281,490,1947,1969]" italics="true" pageId="5">Arcotheres placunae</emphasis>
(Hornell &amp; Southwell, 1909)
</taxonomicName>
. AE, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, male, ZRC 2017.1065 (CW 7.0 mm, CL 5.8 mm), India. A, dorsal view; B, pleon; C, right chela, outer view; D, G1; E, G2; F, left G2 with exopod (lateral view, setae denuded).
</paragraph>
</caption>
<caption id="DF341FBFFFF4FFA0FF75F9F3CBA2F923" httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="6" startId="6.[151,250,1609,1631]" targetBox="[153,1410,211,1503]" targetPageId="6">
<paragraph id="8BF44F37FFF4FFA0FF75F9F3CBA2F923" blockId="6.[151,1436,1609,1692]" pageId="6">
<emphasis id="B93F9325FFF4FFA0FF75F9F3CF24F9E1" bold="true" box="[151,276,1609,1631]" pageId="6">FIGURE 4.</emphasis>
<taxonomicName id="4C4B34B4FFF4FFA0FEFFF9F3CD16F9E0" ID-CoL="67PXW" authority="(Hornell &amp; Southwell, 1909)" authorityName="Hornell &amp; Southwell" authorityYear="1909" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[285,806,1609,1631]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF4FFA0FEFFF9F3CFC0F9E0" box="[285,496,1609,1631]" italics="true" pageId="6">Arcotheres placunae</emphasis>
(Hornell &amp; Southwell, 1909)
</taxonomicName>
. AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5.
</paragraph>
</caption>
<paragraph id="8BF44F37FFF4FFA1FF25F97ECCCAFE6F" blockId="6.[151,1436,1731,2008]" lastBlockId="7.[151,1437,151,1509]" lastPageId="7" pageId="6">
<taxonomicName id="4C4B34B4FFF4FFA0FF25F97ECF9EF963" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[199,430,1732,1756]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF4FFA0FF25F97ECF9EF963" box="[199,430,1732,1756]" italics="true" pageId="6">Arcotheres placunae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4B34B4FFF4FFA0FDA6F97ECCA1F964" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[580,657,1732,1755]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF4FFA0FDA6F97ECCA1F964" box="[580,657,1732,1755]" italics="true" pageId="6">A. rayi</emphasis>
</taxonomicName>
in following features: 1) the female carapace is 1.4 times wider than long (
<figureCitation id="137053B2FFF4FFA0FF3BF95DCF4CF940" box="[217,380,1767,1792]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="6">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF4FFA0FE6AF952CF9CF8BF" box="[392,428,1768,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="6">2A</figureCitation>
) (vs. carapace 1.3 times wider than long in
<taxonomicName id="4C4B34B4FFF4FFA0FC7EF952CDD9F940" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[924,1001,1768,1791]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF4FFA0FC7EF952CDD9F940" box="[924,1001,1768,1791]" italics="true" pageId="6">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF4FFA0FBFFF95DCB32F8BF" box="[1053,1282,1767,1792]" pageId="6" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11A); 2) the carapace posterior margin is strongly concave (
<figureCitation id="137053B2FFF4FFA0FD37F8B1CD46F89C" box="[725,886,1803,1828]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="6">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF4FFA0FC60F8B6CD96F89B" box="[898,934,1804,1828]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="6">2A</figureCitation>
) (vs. slightly concave in
<taxonomicName id="4C4B34B4FFF4FFA0FB20F8B6CB3EF89C" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1218,1294,1804,1827]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF4FFA0FB20F8B6CB3EF89C" box="[1218,1294,1804,1827]" italics="true" pageId="6">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF4FFA0FAA0F8B1CF10F8F7" pageId="6" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11A); 3) the antennae clearly reach beyond the inner edge of the orbit (
<figureCitation id="137053B2FFF4FFA0FB4AF895CB31F8F7" box="[1192,1281,1839,1864]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="6">Fig. 2B</figureCitation>
) (vs. slightly reaching beyond the inner edge of the orbit in
<taxonomicName id="4C4B34B4FFF4FFA0FD3FF8EECD00F8D4" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[733,816,1876,1899]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF4FFA0FD3FF8EECD00F8D4" box="[733,816,1876,1899]" italics="true" pageId="6">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF4FFA0FC92F8E9CA59F8D3" box="[880,1129,1875,1900]" pageId="6" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11B); 4) the MXP3 ishchiomerus is 2.6 times as long as wide (
<figureCitation id="137053B2FFF4FFA0FD9EF8CDCCC0F82F" box="[636,752,1911,1936]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="6">Fig. 2F, G</figureCitation>
) (vs. about 2 times as long as wide in
<taxonomicName id="4C4B34B4FFF4FFA0FB46F8C2CAC1F830" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1188,1265,1912,1935]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF4FFA0FB46F8C2CAC1F830" box="[1188,1265,1912,1935]" italics="true" pageId="6">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF4FFA0FAC6F8CDCF30F80B" pageId="6" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11E); 5) the MXP3 exopod has the inner and outer margins almost straight with a clear median shallow furrow (
<figureCitation id="137053B2FFF4FFA0FEB1F805CF97F867" box="[339,423,1983,2008]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="6">Fig. 2F</figureCitation>
) (vs. inner and outer margin convex without median furrow in
<taxonomicName id="4C4B34B4FFF4FFA0FB9AF87ACAF5F868" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1144,1221,1984,2007]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF4FFA0FB9AF87ACAF5F868" box="[1144,1221,1984,2007]" italics="true" pageId="6">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF4FFA1FB1BF805CEE4FF0F" lastPageId="7" pageId="6" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11E); 6) the palm is about 1.8 times as long as high (
<figureCitation id="137053B2FFF5FFA1FCBDFF2DCDE7FF0F" box="[863,983,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2D, E</figureCitation>
) (vs. about 3 times as long as high in
<taxonomicName id="4C4B34B4FFF5FFA1FA66FF22CEF4FF6B" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FA66FF22CEF4FF6B" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FF18FF06CFD5FF6B" box="[250,485,188,213]" pageId="7" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11D); 7) the cheliped dactylus is slightly shorter than the dorsal length of the palm with a tuft of short setae on the distodorsal margin (
<figureCitation id="137053B2FFF5FFA1FCFCFF65CDA5FF47" box="[798,917,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2D, E</figureCitation>
) (vs. dactylus about half palm length with the distodorsal margin glabrous in
<taxonomicName id="4C4B34B4FFF5FFA1FE1AFEBECC76FEA3" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[504,582,260,284]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FE1AFEBECC76FEA3" box="[504,582,260,284]" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FD99FEBECD52FEA3" box="[635,866,260,285]" pageId="7" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11D); 8) the ventral margin of the pollex has short setae (
<figureCitation id="137053B2FFF5FFA1FEFEFE9DCFA1FEFF" box="[284,401,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2D, E</figureCitation>
) (vs. glabrous in
<taxonomicName id="4C4B34B4FFF5FFA1FDB1FE92CCAEFE80" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[595,670,296,319]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FDB1FE92CCAEFE80" box="[595,670,296,319]" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FD2DFE9DCD80FEFF" box="[719,944,295,320]" pageId="7" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11D); 9) the order of relative lengths of the ambulatory dactyli is P5&gt;P4&gt;P3&gt;P2 (
<figureCitation id="137053B2FFF5FFA1FD93FEF6CCDEFEDB" box="[625,750,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="7">Fig. 4AH</figureCitation>
) (vs. P4&gt;P5&gt;P3=P
<quantity id="4CB3E2D2FFF5FFA1FC2AFEF6CDC5FEDA" box="[968,1013,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C4B34B4FFF5FFA1FC1FFEF6CA7BFEDB" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1021,1099,332,356]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FC1FFEF6CA7BFEDB" box="[1021,1099,332,356]" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FB63FEF6CB5AFEDB" box="[1153,1386,332,357]" pageId="7" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11A); 10) the fifth pleonal somite is the widest (
<figureCitation id="137053B2FFF5FFA1FD52FED5CD36FE37" box="[688,774,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2C</figureCitation>
) (vs. fourth pleonal somite widest in
<taxonomicName id="4C4B34B4FFF5FFA1FB4AFECACAC3FE38" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1192,1267,368,391]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FB4AFECACAC3FE38" box="[1192,1267,368,391]" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FAC7FED5CECDFE13" pageId="7" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11C); and 11) the lower margin of the telson is concave (
<figureCitation id="137053B2FFF5FFA1FC54FE2ECA3CFE13" box="[950,1036,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2C</figureCitation>
) (vs. convex in
<taxonomicName id="4C4B34B4FFF5FFA1FB21FE2ECB3FFE13" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1219,1295,404,428]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FB21FE2ECB3FFE13" box="[1219,1295,404,428]" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
, cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FAA0FE2ECF29FE6F" pageId="7" refId="ref5519" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 11C). Males of
<taxonomicName id="4C4B34B4FFF5FFA1FE1DFE02CC7BFE70" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[511,587,440,463]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B93F9325FFF5FFA1FE1DFE02CC7BFE70" box="[511,587,440,463]" italics="true" pageId="7">A. rayi</emphasis>
</taxonomicName>
are not known.
</paragraph>
<paragraph id="8BF44F37FFF5FFA1FF25FE66CAFFFBAF" blockId="7.[151,1437,151,1509]" pageId="7">
Comparing specimens of
<taxonomicName id="4C4B34B4FFF5FFA1FE12FE66CC6EFE4B" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[496,606,476,500]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">
<emphasis id="B93F9325FFF5FFA1FE12FE66CC6EFE4B" box="[496,606,476,500]" italics="true" pageId="7">A. tivelae</emphasis>
</taxonomicName>
with
<taxonomicName id="4C4B34B4FFF5FFA1FD46FE66CD1DFE4B" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[676,813,476,500]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF5FFA1FD46FE66CD1DFE4B" box="[676,813,476,500]" italics="true" pageId="7">A. placunae</emphasis>
</taxonomicName>
, the following differences are obvious: 1) the female carapace is 1.4 times wider than long (
<figureCitation id="137053B2FFF5FFA1FDADFE45CCC4FDA8" box="[591,756,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="7">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF5FFA1FCE2FDBACD14FDA7" box="[768,804,512,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">2A</figureCitation>
) (vs.1.2 times wider than long in
<emphasis id="B93F9325FFF5FFA1FB45FDBACB29FDA7" box="[1191,1305,512,536]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FB45FDBACB24FDA7" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[1191,1300,512,536]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FAA4FE45CEE3FD82" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4a;
<bibRefCitation id="EFDA32C6FFF5FFA1FED4FD9ECFD2FD83" box="[310,482,548,573]" pageId="7" refId="ref6040" refString="Naderloo, R. (2017) Atlas of Crabs of the Persian Gulf. Springer, Cham, 440 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book">Naderloo 2017</bibRefCitation>
: fig. 38.2); 2) the carapace posterior margin is strongly concave (
<figureCitation id="137053B2FFF5FFA1FB20FD9ECB55FD83" box="[1218,1381,548,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="7">Figs. 1A, C, F</figureCitation>
,
<figureCitation id="137053B2FFF5FFA1FA93FD9ECBA4FD83" box="[1393,1428,548,572]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">2A</figureCitation>
) (vs.slightly convex in
<emphasis id="B93F9325FFF5FFA1FE70FDF2CC34FDDF" box="[402,516,584,608]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FE70FDF2CFCFFDDF" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[402,511,584,608]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FDD2FDFDCCFAFDDF" box="[560,714,583,608]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4a;
<bibRefCitation id="EFDA32C6FFF5FFA1FCCCFDFDCDEBFDDF" box="[814,987,583,608]" pageId="7" refId="ref6040" refString="Naderloo, R. (2017) Atlas of Crabs of the Persian Gulf. Springer, Cham, 440 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book">Naderloo 2017</bibRefCitation>
: fig. 38.2); 3) the MXP3 ischiomerus is 2.6 times as long as wide (
<figureCitation id="137053B2FFF5FFA1FE29FDD6CC72FD3B" box="[459,578,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2F, G</figureCitation>
) (vs. 2.2 two times as long as wide in
<emphasis id="B93F9325FFF5FFA1FBE3FDD6CA44FD3B" box="[1025,1140,620,644]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FBE3FDD6CA5FFD3B" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[1025,1135,620,644]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FB40FDD6CB0DFD3A" box="[1186,1341,620,645]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4b;
<bibRefCitation id="EFDA32C6FFF5FFA1FF75FD35CF72FD17" box="[151,322,655,680]" pageId="7" refId="ref6040" refString="Naderloo, R. (2017) Atlas of Crabs of the Persian Gulf. Springer, Cham, 440 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book">Naderloo 2017</bibRefCitation>
: fig. 38.5); 4) the cheliped dactylus is slightly shorter than the dorsal length of the palm (
<figureCitation id="137053B2FFF5FFA1FAFCFD35CBA4FD17" box="[1310,1428,655,680]" captionStart="FIGURE 2" captionStartId="3.[151,250,1566,1588]" captionTargetBox="[160,1417,315,1521]" captionTargetId="figure@3.[151,1436,301,1545]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres placunae (Hornell &amp; Southwell, 1909), female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India. A, dorsal view; B, frontal view; C, pleon; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view; G, left MXP3, outer view." httpUri="https://zenodo.org/record/1287097/files/figure.png" pageId="7">Fig. 2D, E</figureCitation>
) (vs. dactylus about half palm length in
<emphasis id="B93F9325FFF5FFA1FDA5FD0ECC87FD73" box="[583,695,692,716]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FDA5FD0ECC83FD73" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[583,691,692,716]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FD03FD0ECD49FD72" box="[737,889,692,717]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4b); 5) the P4 merus is 1.3 times longer than that of P5 (
<figureCitation id="137053B2FFF5FFA1FEF6FD6DCFFDFD4F" box="[276,461,727,752]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="7">Fig. 4C, D, G, H</figureCitation>
) (vs. P4 merus 1.7 times longer than that of P
<quantity id="4CB3E2D2FFF5FFA1FC30FD6DCDCDFD4F" box="[978,1021,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="7" unit="in" value="5.0">5 in</quantity>
<emphasis id="B93F9325FFF5FFA1FBE1FD62CA44FD4F" box="[1027,1140,728,752]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FBE1FD62CA40FD4F" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[1027,1136,728,752]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FB7DFD6DCB08FD4F" box="[1183,1336,727,752]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4a); 6) the P4 dactylus is 1.2 times longer than the P3 dactylus (
<figureCitation id="137053B2FFF5FFA1FCCBFD46CDEBFCAB" box="[809,987,764,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="7">Fig. 4B, C, F, G</figureCitation>
) (vs. P4 dactylus 2 times longer than P3 dactylus in
<emphasis id="B93F9325FFF5FFA1FEF5FC9ACFB7FC87" box="[279,391,800,824]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FEF5FC9ACFB3FC87" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[279,387,800,824]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FE53FCA5CC79FC87" box="[433,585,799,824]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4d, e); 7) the P5 dactylus is 0.84 times longer than the P4 dactylus (
<figureCitation id="137053B2FFF5FFA1FA8DFCA5CF2BFCE3" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="7">Fig. 4C, D, G, H</figureCitation>
) (vs. P5 dactylus 0.75 times longer than P4 dactylus in
<emphasis id="B93F9325FFF5FFA1FC67FCFECDC5FCE3" box="[901,1013,836,860]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FC67FCFECDC1FCE3" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[901,1009,836,860]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FBFDFCFECA88FCE2" box="[1055,1208,836,861]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4e, f); 8) the P5 dactylus dorsodistal half is covered with short setae (
<figureCitation id="137053B2FFF5FFA1FD0DFCDDCD5BFC3F" box="[751,875,871,896]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="7">Fig. 4D, H</figureCitation>
) (vs. glabrous in
<emphasis id="B93F9325FFF5FFA1FBD0FCD2CA94FC3F" box="[1074,1188,872,896]" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FBD0FCD2CA90FC3F" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[1074,1184,872,896]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FB33FCDDCB5AFC3F" box="[1233,1386,871,896]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4f); and 9) the P2P5 merus is glabrous (
<figureCitation id="137053B2FFF5FFA1FD94FC36CCC7FC1B" box="[630,759,908,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1609,1631]" captionTargetBox="[153,1410,211,1503]" captionTargetId="figure@6.[151,1436,193,1587]" captionTargetPageId="6" captionText="FIGURE 4. Arcotheres placunae (Hornell &amp; Southwell, 1909). AH, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; IL, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India. AD, female left P2P5 outer view; EH, female right P2P4 outer view; H', tip of dactylus of female right P5; IL, male left P2P5, outer view; L', tip of dactylus of male right P5." httpUri="https://zenodo.org/record/1287101/files/figure.png" pageId="7">Fig. 4AH</figureCitation>
)(vs. P4 and P5 merus with setae on proximal margin in
<emphasis id="B93F9325FFF5FFA1FA66FC36CED8FC77" italics="true" pageId="7">
<taxonomicName id="4C4B34B4FFF5FFA1FA66FC36CED4FC77" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">A. tivelae</taxonomicName>
,
</emphasis>
cf.
<bibRefCitation id="EFDA32C6FFF5FFA1FEC6FC15CFF5FC77" box="[292,453,943,968]" pageId="7" refId="ref5861" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 4a). Males of
<taxonomicName id="4C4B34B4FFF5FFA1FD23FC0ACD04FC77" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[705,820,944,968]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">
<emphasis id="B93F9325FFF5FFA1FD23FC0ACD04FC77" box="[705,820,944,968]" italics="true" pageId="7">A. tivelae</emphasis>
</taxonomicName>
are not known. Considering the above mentioned morphological differences between
<taxonomicName id="4C4B34B4FFF5FFA1FDD3FC6ECC8AFC53" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[561,698,980,1004]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF5FFA1FDD3FC6ECC8AFC53" box="[561,698,980,1004]" italics="true" pageId="7">A. placunae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4B34B4FFF5FFA1FD14FC6ECD53FC53" ID-CoL="67PXV" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[758,867,980,1004]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="tivelae">
<emphasis id="B93F9325FFF5FFA1FD14FC6ECD53FC53" box="[758,867,980,1004]" italics="true" pageId="7">A. tivelae</emphasis>
</taxonomicName>
and that the two species have different host (see Saeedi &amp;Ardalan 2010;
<bibRefCitation id="EFDA32C6FFF5FFA1FE44FC4DCC62FBAF" box="[422,594,1015,1040]" pageId="7" refId="ref6040" refString="Naderloo, R. (2017) Atlas of Crabs of the Persian Gulf. Springer, Cham, 440 pp. https: // doi. org / 10.1007 / 978 - 3 - 319 - 49374 - 9" type="book">Naderloo 2017</bibRefCitation>
), there is no doubt about the distinctness of both species.
</paragraph>
</subSubSection>
<subSubSection id="C3511CBCFFF5FFA1FF25FBA6CA95FA5A" pageId="7" type="description">
<paragraph id="8BF44F37FFF5FFA1FF25FBA6CF7FFA8F" blockId="7.[151,1437,151,1509]" pageId="7">
As discussed earlier, the
<typeStatus id="54F0F195FFF5FFA1FE3EFBA7CC27FB8A" box="[476,535,1053,1077]" pageId="7">types</typeStatus>
of
<taxonomicName id="4C4B34B4FFF5FFA1FDD9FBA6CA3AFB8B" authority="(Hornell &amp; Southwell, 1909)" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[571,1034,1052,1077]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF5FFA1FDD9FBA6CCF1FB8B" box="[571,705,1052,1076]" italics="true" pageId="7">A. placunae</emphasis>
(
<bibRefCitation id="EFDA32C6FFF5FFA1FD2DFBA6CA32FB8B" box="[719,1026,1052,1077]" pageId="7" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell, 1909</bibRefCitation>
)
</taxonomicName>
are almost certainly lost. In view of the uncertainty of what the species actually was until recently (see above) and the discrepancies in the descriptions and figures by the original authors, it is better to designate one of the recent specimens from the original
<typeStatus id="54F0F195FFF5FFA1FA8EFBDFCBACFBC2" box="[1388,1436,1125,1149]" pageId="7">type</typeStatus>
locality (Gulf of Kachchh) as the
<typeStatus id="54F0F195FFF5FFA1FDFAFB33CC41FB1E" box="[536,625,1161,1185]" pageId="7" type="neotype">neotype</typeStatus>
of the species. This will not only stabilize the taxonomy of the species but also be useful in distinguishing other allied species in subsequent taxonomic studies of the
<taxonomicName id="4C4B34B4FFF5FFA1FB2FFB16CB5CFB7A" authorityName="De Haan" authorityYear="1833" box="[1229,1388,1196,1221]" class="Malacostraca" family="Pinnotheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
. We here designate a female specimen (CW 11.0 mm, CL
<quantity id="4CB3E2D2FFF5FFA1FD18FB6ACD64FB57" box="[762,852,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="7" unit="mm" value="8.1">8.1 mm</quantity>
; ZRC 2017.1293) (
<figureCitation id="137053B2FFF5FFA1FBD2FB75CAB6FB57" box="[1072,1158,1231,1256]" captionStart="FIGURE 1" captionStartId="2.[151,250,1733,1755]" captionTargetBox="[151,1436,483,1708]" captionTargetId="figure@2.[151,1436,483,1708]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres placunae (Hornell &amp; Southwell, 1909). A, B, female, ZL-AR-CR-100 (CW 10.6 mm, CL 7.7 mm), India; C, neotype female, ZRC 2017.1293 (CW 11.0 mm, CL 8.1 mm), India; D, E, male, ZL-AR-CR-100 (CW 6.8 mm, CL 5.9 mm), India; F, female, ZRC 2017.1008 (CW 11.5 mm, CL 7.9 mm), Pakistan. A, C, D, F, dorsal view; B, E, ventral view." httpUri="https://zenodo.org/record/1287095/files/figure.png" pageId="7">Fig. 1C</figureCitation>
) collected from Balapur bay in Beyt Dwarka Island, Gulf of Kachchh,
<collectingRegion id="498F81D5FFF5FFA1FD4EFB4ECD33FAB2" box="[684,771,1268,1293]" country="India" name="Gujarat" pageId="7">Gujarat</collectingRegion>
,
<collectingCountry id="F35C0FA7FFF5FFA1FCF2FB4ECD7CFAB2" box="[784,844,1268,1293]" name="India" pageId="7">India</collectingCountry>
, as the
<typeStatus id="54F0F195FFF5FFA1FC44FB4FCDCFFAB2" box="[934,1023,1269,1293]" pageId="7" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C4B34B4FFF5FFA1FBCAFB4ECF7AFA8F" authority="Hornell &amp; Southwell 1909" authorityName="Hornell &amp; Southwell" authorityYear="1909" class="Malacostraca" family="Pinnotheridae" genus="Pinnotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="B93F9325FFF5FFA1FBCAFB4ECB2EFAB3" box="[1064,1310,1268,1292]" italics="true" pageId="7">Pinnotheres placunae</emphasis>
<bibRefCitation id="EFDA32C6FFF5FFA1FAC5FB4ECF7AFA8F" pageId="7" refId="ref5922" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF44F37FFF5FFA1FF25FA86CA95FA5A" blockId="7.[151,1437,151,1509]" pageId="7">
Hornell &amp; Southwells (1909) material is unusual because it is composed almost entirely of males—in almost all studies; female specimens invariably outnumber male ones, with males often not even found. Even more unusual was his report that he found two
<taxonomicName id="4C4B34B4FFF5FFA1FD67FA3ECCD2FA23" box="[645,738,1412,1436]" class="Bivalvia" family="Placunidae" genus="Placuna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pectinida" pageId="7" phylum="Mollusca" rank="genus">
<emphasis id="B93F9325FFF5FFA1FD67FA3ECCD2FA23" box="[645,738,1412,1436]" italics="true" pageId="7">Placuna</emphasis>
</taxonomicName>
shells that had 11 and
<specimenCount id="9D4D84BEFFF5FFA1FC18FA3ECA67FA22" box="[1018,1111,1412,1437]" pageId="7" type="male">12 male</specimenCount>
specimens each—normally, bivalves only have single adult individuals or heterosexual pairs. It suggests that many of their specimens may perhaps be juveniles which may have just only recently metamorphosed in the bivalve hosts.
</paragraph>
</subSubSection>
</treatment>
</document>