321 lines
48 KiB
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321 lines
48 KiB
XML
<document id="339485FD86E2A61139DF545B9629C6F8" ID-CLB-Dataset="5669" ID-DOI="10.5852/ejt.2021.758.1423" ID-GBIF-Dataset="de9c0c06-0e7d-44a6-a582-d41a346233f6" ID-ISSN="2118-9773" ID-Zenodo-Dep="5075824" ID-ZooBank="01BBC12C-E715-4393-A9F6-6EA85CB1289F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625579995989" checkinUser="felipe" docAuthor="Wesener, Thomas & Sagorny, Christina" docDate="2021" docId="03BF4B62FFA5E0291C55FEF3875CF9F0" docLanguage="en" docName="ejt-758_1-48.pdf" docOrigin="European Journal of Taxonomy 758 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.758.1423" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zoosphaerium voahangy Wesener & Sagorny 2021, sp. nov." docType="treatment" docUuid="11B9C806-A111-48FC-AC36-035385125232" docUuidSource="ZooBank" docVersion="7" lastPageNumber="30" masterDocId="FF86331AFFBEE0341E06FFFB8261FFB5" masterDocTitle="Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="28" updateTime="1699025899143" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="53DB7D280BAE82ED91936A8DA8051CBE">Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)</mods:title>
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<mods:namePart id="6F8C2DA487725C8AAC42C2A5584F3D4C">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="D41F094F43D438891113AD3B6F0598AB">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="A2C85C7CAD348A7F8779A283D4DD0379" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:namePart id="E3D1CC377A5BCD9D560E6225C481F1C1">Sagorny, Christina</mods:namePart>
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<mods:affiliation id="8238263C93EC6614079EB37F568A2A00">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="EFB422796E90D2403D8796A8CFB9B860" type="email">csagorny@evolution.uni-bonn.de</mods:nameIdentifier>
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<mods:title id="EF61EE65BEBC1D2D69756D68ADF9DC34">European Journal of Taxonomy</mods:title>
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<mods:date id="EB0D23A237F3BBF26CDC3A5FAC658C09">2021</mods:date>
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<treatment id="03BF4B62FFA5E0291C55FEF3875CF9F0" ID-DOI="http://doi.org/10.5281/zenodo.5076208" ID-GBIF-Taxon="183392513" ID-Zenodo-Dep="5076208" LSID="urn:lsid:zoobank.org:act:11B9C806-A111-48FC-AC36-035385125232" httpUri="http://treatment.plazi.org/id/03BF4B62FFA5E0291C55FEF3875CF9F0" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
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<subSubSection id="C30CA9FFFFA5E02F1C55FEF38180FE96" box="[595,993,264,291]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BA9FA74FFA5E02F1C55FEF38180FE96" blockId="27.[377,1210,264,362]" box="[595,993,264,291]" pageId="27" pageNumber="28">
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<heading id="D0E14D18FFA5E02F1C55FEF38180FE96" box="[595,993,264,291]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="28" reason="2">
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<taxonomicName id="4C1681F7FFA5E02F1C55FEF3811EFE96" ID-CoL="9BDGT" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[595,895,264,291]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
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<emphasis id="B9622666FFA5E02F1C55FEF3811EFE96" bold="true" box="[595,895,264,291]" italics="true" pageId="27" pageNumber="28">Zoosphaerium voahangy</emphasis>
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<taxonomicNameLabel id="A2519B1DFFA5E02F1D81FEF28180FE96" box="[903,993,265,291]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<uuid id="FFB0C0A1FFA5E02F1C90FED786DBFEF2" box="[662,1210,300,327]" pageId="27" pageNumber="28">11B9C806-A111-48FC-AC36-035385125232</uuid>
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</uri>
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<paragraph id="8BA9FA74FFA5E02F1CBEFEB4811DFEDC" blockId="27.[377,1210,264,362]" box="[696,892,335,362]" pageId="27" pageNumber="28">
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<figureCitation id="132DE6F1FFA5E02F1CBEFEB48174FEDC" box="[696,789,335,361]" captionStart="Fig" captionStartId="9.[189,232,1764,1790]" captionTargetBox="[198,1390,265,1713]" captionTargetId="figure-92@9.[189,1398,265,1719]" captionTargetPageId="9" captionText="Fig. 2. Habitus photographs of the seven new species of Zoosphaerium, lateral view. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1).D. Z. beanka sp. nov., holotype (FMNH-INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). F. Z. masoala sp. nov., holotype (FMNH-INS 3195651). G. Z. spinopiligerum sp. nov., holotype (FMNH-INS 3119885). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="27" pageNumber="28">Figs 2E</figureCitation>
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,
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<geoCoordinate id="EE229CB3FFA5E02F1D27FEB48100FEDF" box="[801,865,335,362]" degrees="3, 5" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="5555" value="3.5">
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<figureCitation id="132DE6F1FFA5E02F1D27FEB48150FEDC" box="[801,817,335,361]" captionStart="Fig" captionStartId="10.[189,232,1891,1917]" captionTargetBox="[228,1372,266,1870]" captionTargetId="figure-87@10.[202,1385,239,1881]" captionTargetPageId="10" captionText="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." figureDoi="http://doi.org/10.5281/zenodo.5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="27" pageNumber="28">3</figureCitation>
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,
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<figureCitation id="132DE6F1FFA5E02F1D38FEB48100FEDF" box="[830,865,335,362]" captionStart="Fig" captionStartId="12.[189,232,1891,1917]" captionTargetBox="[270,1317,266,1874]" captionTargetId="figure-78@12.[270,1317,266,1875]" captionTargetPageId="12" captionText="Fig. 5. Scanning electron micrographs of the endoterga of midbody rings. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1). D. Z. beanka sp. nov., holotype (FMNH- INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="27" pageNumber="28">5E</figureCitation>
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,
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<figureCitation id="132DE6F1FFA5E02F1D6BFEB4811DFEDC" box="[877,892,335,361]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="27" pageNumber="28">9</figureCitation>
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</paragraph>
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<subSubSection id="C30CA9FFFFA5E02F1EBBFE6B834DFD65" pageId="27" pageNumber="28" type="diagnosis">
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<paragraph id="8BA9FA74FFA5E02F1EBBFE6B8357FE1F" blockId="27.[189,310,400,426]" box="[189,310,400,426]" pageId="27" pageNumber="28">
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<heading id="D0E14D18FFA5E02F1EBBFE6B8357FE1F" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
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<emphasis id="B9622666FFA5E02F1EBBFE6B8357FE1F" bold="true" box="[189,310,400,426]" pageId="27" pageNumber="28">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9FA74FFA5E02F1EBBFE44834DFD65" blockId="27.[187,1399,447,720]" pageId="27" pageNumber="28">
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The position of the locking carinae close to the last pleurite identifies
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<taxonomicName id="4C1681F7FFA5E02F1DD1FE3B8607FE6C" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[983,1126,447,473]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
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<emphasis id="B9622666FFA5E02F1DD1FE3B8607FE6C" box="[983,1126,447,473]" italics="true" pageId="27" pageNumber="28">Z. voahangy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFA5E02F1A6DFE3B86A5FE6F" box="[1131,1220,448,474]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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as a member of the
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<taxonomicName id="4C1681F7FFA5E02F1EEEFE1883EAFE49" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1902" box="[232,395,482,508]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="platylabum">
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<emphasis id="B9622666FFA5E02F1EEEFE1883EAFE49" box="[232,395,482,508]" italics="true" pageId="27" pageNumber="28">Z. platylabum</emphasis>
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</taxonomicName>
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species-group. The uniquely slender and elongated posterior telopods in combination with the presence of a single stridulation rib on the male harp and more than 35 apical cones on the antennae, are characters only shared with
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<taxonomicName id="4C1681F7FFA5E02F1CC0FDD1865FFDF6" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[710,1086,553,579]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="broelemanni">
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<emphasis id="B9622666FFA5E02F1CC0FDD1811CFDF6" box="[710,893,553,579]" italics="true" pageId="27" pageNumber="28">Z. broelemanni</emphasis>
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Wesener, 2009
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</taxonomicName>
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.
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<taxonomicName id="4C1681F7FFA5E02F1A49FDD28717FDF6" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[1103,1398,553,579]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
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<emphasis id="B9622666FFA5E02F1A49FDD28717FDF6" box="[1103,1398,553,579]" italics="true" pageId="27" pageNumber="28">Zoosphaerium voahangy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2519B1DFFA5E02F1EBBFDB6837AFDD2" box="[189,283,589,615]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C1681F7FFA5E02F1FC6FDB68017FDD3" authorityName="Wesener" authorityYear="2009" box="[448,630,588,614]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="broelemanni">
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<emphasis id="B9622666FFA5E02F1FC6FDB68017FDD3" box="[448,630,588,614]" italics="true" pageId="27" pageNumber="28">Z. broelemanni</emphasis>
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</taxonomicName>
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in the differently shaped anterior telopods, spination and teeth number of the movable finger of the posterior telopod (see key), as well as in characters of the endotergum, which has two dense rows of long marginal bristles in the former, but a single row of short bristles in the latter.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30CA9FFFFA5E02F1EBBFD0C8330FCD6" pageId="27" pageNumber="28" type="etymology">
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<paragraph id="8BA9FA74FFA5E02F1EBBFD0C8325FCA4" blockId="27.[189,324,759,785]" box="[189,324,759,785]" pageId="27" pageNumber="28">
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<heading id="D0E14D18FFA5E02F1EBBFD0C8325FCA4" bold="true" box="[189,324,759,785]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
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<emphasis id="B9622666FFA5E02F1EBBFD0C8325FCA4" bold="true" box="[189,324,759,785]" pageId="27" pageNumber="28">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9FA74FFA5E02F1EBBFCDD8330FCD6" blockId="27.[189,1399,806,867]" pageId="27" pageNumber="28">‘Voahangy’, noun in apposition, to honor the great collector and Malagasy scientist Voahangy Soarimalala.</paragraph>
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</subSubSection>
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<subSubSection id="C30CA9FFFFA5E02F1EBBFC718753FB2F" pageId="27" pageNumber="28" type="materials_examined">
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<paragraph id="8BA9FA74FFA5E02F1EBBFC7183CBFC11" blockId="27.[189,426,906,932]" box="[189,426,906,932]" pageId="27" pageNumber="28">
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<emphasis id="B9622666FFA5E02F1EBBFC7183CBFC11" bold="true" box="[189,426,906,932]" pageId="27" pageNumber="28">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9FA74FFA5E02F1EECFC428300FB88" blockId="27.[189,1399,953,1085]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B7EF029FFA5E02F1EECFC428300FB88" ID-GBIF-Occurrence="3314642351" collectingDate="2004-12-08" collectionCode="FMNH-" collectorName="V. Soarimalala" country="Madagascar" elevation="120" latitude="-14.31" location="Analalava" longLatPrecision="128" longitude="47.915" pageId="27" pageNumber="28" specimenCode="INS 3119921" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="holotype">
|
||
<emphasis id="B9622666FFA5E02F1EECFC42833DFC66" bold="true" box="[234,348,953,979]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54AD44D6FFA5E02F1EECFC42833DFC66" box="[234,348,953,979]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F301BAE4FFA5E02F1EBBFC2783EFFC43" box="[189,398,988,1014]" name="Madagascar" pageId="27" pageNumber="28">MADAGASCAR</collectingCountry>
|
||
–
|
||
<emphasis id="B9622666FFA5E02F1FBDFC27802BFC43" bold="true" box="[443,586,988,1014]" pageId="27" pageNumber="28">
|
||
<collectingRegion id="49D23496FFA5E02F1FBDFC27802BFC43" box="[443,586,988,1014]" country="Madagascar" name="Mahajanga" pageId="27" pageNumber="28">Mahajanga</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D1031FDFFA5E02F1C74FC2680EAFC42" box="[626,651,989,1015]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
;
|
||
<location id="8EC9ACAFFFA5E02F1C98FC278179FC43" LSID="urn:lsid:plazi:treatment:03BF4B62FFA5E0291C55FEF3875CF9F0:8EC9ACAFFFA5E02F1C98FC278179FC43" box="[670,792,988,1014]" country="Madagascar" latitude="-14.31" longLatPrecision="128" longitude="47.915" name="Analalava" pageId="27" pageNumber="28" stateProvince="Mahajanga">Analalava</location>
|
||
,
|
||
<locationDeviation id="9A1BE356FFA5E02F1D2DFC2786B8FC43" box="[811,1241,988,1015]" location="Ambinda Village" pageId="27" pageNumber="28">
|
||
<quantity id="4CEE5791FFA5E02F1D2DFC2781E5FC43" box="[811,900,988,1015]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="27" pageNumber="28" unit="km" value="5.5">5.5 km</quantity>
|
||
NNE of Ambinda Village
|
||
</locationDeviation>
|
||
;
|
||
<geoCoordinate id="EE229CB3FFA5E02F1AEBFC218710FC43" box="[1261,1393,986,1014]" degrees="14" direction="south" minutes="18.6" orientation="latitude" pageId="27" pageNumber="28" precision="92" value="-14.31">14°18.6′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE229CB3FFA5E02F1EBBFC05835AFBAF" box="[189,315,1022,1050]" degrees="47" direction="east" minutes="54.9" orientation="longitude" pageId="27" pageNumber="28" precision="92" value="47.915">47°54.9′ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CEE5791FFA5E02F1F77FC0483DFFBAF" box="[369,446,1023,1050]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="27" pageNumber="28" unit="m" value="120.0">
|
||
<elevation id="003B1D47FFA5E02F1F77FC0483DFFBAF" box="[369,446,1023,1050]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="27" pageNumber="28" unit="m" value="120.0">120 m</elevation>
|
||
</quantity>
|
||
; dry deciduous forest;
|
||
<date id="FFA8DCB4FFA5E02F1CC3FC048134FBAC" box="[709,853,1023,1050]" pageId="27" pageNumber="28" value="2004-12-08">
|
||
<collectingDate id="EFEC255CFFA5E02F1CC3FC048134FBAC" box="[709,853,1023,1050]" pageId="27" pageNumber="28" value="2004-12-08">8 Dec. 2004</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26E39FA2FFA5E02F1D59FBFB866BFBAC" box="[863,1034,1023,1050]" pageId="27" pageNumber="28">V. Soarimalala</collectorName>
|
||
leg.; coll. no. VS-508;
|
||
<collectionCode id="ED0762B1FFA5E02F1B1EFBFB8716FBAF" box="[1304,1399,1024,1050]" pageId="27" pageNumber="28">FMNH-</collectionCode>
|
||
<specimenCode id="DBB0520FFFA5E02F1EBBFBD8833CFB88" box="[189,349,1059,1085]" collectionCode="FMNH-" pageId="27" pageNumber="28">INS 3119921</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EECFBA78753FB2F" blockId="27.[189,1330,1116,1178]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="3B7EF029FFA5E02F1EECFBA7874CFB2F" ID-GBIF-Occurrence="3314642302" collectingDate="2004-12-08" collectionCode="FMNH-" collectorName="V. Soarimalala" country="Madagascar" elevation="120" latitude="-14.31" location="Mahajanga" longLatPrecision="128" longitude="47.915" pageId="27" pageNumber="28" specimenCode="FMNH-INS 3119919" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga" typeStatus="paratype">
|
||
<emphasis id="B9622666FFA5E02F1EECFBA7833DFBC3" bold="true" box="[234,348,1116,1142]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54AD44D6FFA5E02F1EECFBA7833DFBC3" box="[234,348,1116,1142]" pageId="27" pageNumber="28" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F301BAE4FFA5E02F1EBBFB7B83EFFB2F" box="[189,398,1152,1178]" name="Madagascar" pageId="27" pageNumber="28">MADAGASCAR</collectingCountry>
|
||
–
|
||
<emphasis id="B9622666FFA5E02F1FADFB84805BFB2C" bold="true" box="[427,570,1151,1177]" pageId="27" pageNumber="28">
|
||
<collectingRegion id="49D23496FFA5E02F1FADFB84805BFB2C" box="[427,570,1151,1177]" country="Madagascar" name="Mahajanga" pageId="27" pageNumber="28">Mahajanga</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D1031FDFFA5E02F1C55FB7B800DFB2F" box="[595,620,1152,1178]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="DBB0520FFFA5E02F1A29FB7B874CFB2F" box="[1071,1325,1152,1178]" collectionCode="FMNH-INS" pageId="27" pageNumber="28">FMNH-INS 3119919</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFA5E0291EBBFB3A8358FA3A" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBFB3A832FFB6E" blockId="27.[189,334,1217,1243]" box="[189,334,1217,1243]" pageId="27" pageNumber="28">
|
||
<heading id="D0E14D18FFA5E02F1EBBFB3A832FFB6E" bold="true" box="[189,334,1217,1243]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B9622666FFA5E02F1EBBFB3A832FFB6E" bold="true" box="[189,334,1217,1243]" pageId="27" pageNumber="28">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBFB0B866EFA9B" blockId="27.[188,1399,1263,1326]" pageId="27" pageNumber="28">
|
||
BODY LENGTH. Male (
|
||
<typeStatus id="54AD44D6FFA5E02F1FBDFB148048FABF" box="[443,553,1263,1290]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
): length
|
||
<quantity id="4CEE5791FFA5E02F1C8DFB148094FABF" box="[651,757,1263,1290]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="27" pageNumber="28" unit="mm" value="26.3">26.3 mm</quantity>
|
||
, width of thoracic shield
|
||
<quantity id="4CEE5791FFA5E02F1A11FB1486E3FABF" box="[1047,1154,1263,1290]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="27" pageNumber="28" unit="mm" value="16.1">16.1 mm</quantity>
|
||
, of tergite 7
|
||
<quantity id="4CEE5791FFA5E02F1B09FB148716FABF" box="[1295,1399,1263,1290]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="27" pageNumber="28" unit="mm" value="16.7">16.7 mm</quantity>
|
||
(widest), height of thoracic shield
|
||
<quantity id="4CEE5791FFA5E02F1C48FAE880CAFA98" box="[590,683,1299,1325]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="27" pageNumber="28" unit="mm" value="8.9">8.9 mm</quantity>
|
||
, of tergite 7
|
||
<quantity id="4CEE5791FFA5E02F1D38FAE881F9FA98" box="[830,920,1299,1325]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="27" pageNumber="28" unit="mm" value="9.3">9.3 mm</quantity>
|
||
(highest).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBFAAF872AFA27" blockId="27.[189,1398,1364,1426]" pageId="27" pageNumber="28">
|
||
COLORATION. Head, collum, thoracic shield and tergites light brown with dark posterior margins. Anal shield entirely light brown (
|
||
<figureCitation id="132DE6F1FFA5E02F1C02FA8C8001FA24" box="[516,608,1399,1425]" captionStart="Fig" captionStartId="9.[189,232,1764,1790]" captionTargetBox="[198,1390,265,1713]" captionTargetId="figure-92@9.[189,1398,265,1719]" captionTargetPageId="9" captionText="Fig. 2. Habitus photographs of the seven new species of Zoosphaerium, lateral view. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1).D. Z. beanka sp. nov., holotype (FMNH-INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). F. Z. masoala sp. nov., holotype (FMNH-INS 3195651). G. Z. spinopiligerum sp. nov., holotype (FMNH-INS 3119885). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5075828" httpUri="https://zenodo.org/record/5075828/files/figure.png" pageId="27" pageNumber="28">Fig. 2E</figureCitation>
|
||
). Antennae dark green, legs basally light brown, apically green.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBFA428369F9AF" blockId="27.[189,1398,1465,1562]" pageId="27" pageNumber="28">HEAD. Eyes with more than 65 ommatidia. Antennae short, protruding back to leg 5. Length of antennomeres 1=2 =3=4 =5<6. A shallow groove present on antennomere 1, disc with 48/49 apical cones.</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBF9BA8169F9EE" blockId="27.[189,776,1601,1627]" box="[189,776,1601,1627]" pageId="27" pageNumber="28">GNATHOCHILARIUM AND MANDIBLE. Not dissected.</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBF9798097F90A" blockId="27.[189,1399,1666,1728]" pageId="27" pageNumber="28">
|
||
STIGMATIC PLATES. First stigmatic plate triangular, apex sharply edged, pointed, turned towards coxa. Hairs most abundant on apical margin (
|
||
<figureCitation id="132DE6F1FFA5E02F1C8AF95E8089F90A" box="[652,744,1701,1727]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="27" pageNumber="28">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBF91C8607F891" blockId="27.[189,1398,1767,1828]" pageId="27" pageNumber="28">PLEURITES. First pleurite extending posteriorly in a well-rounded process. Second pleurite also with a well-rounded process, but not projecting. Remaining pleurites broadly rounded.</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBF8B081FCF8D0" blockId="27.[189,925,1867,1894]" box="[189,925,1867,1894]" pageId="27" pageNumber="28">COLLUM. Margins with a few short hairs, central part glabrous.</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E02F1EBBF8768107F812" blockId="27.[189,870,1933,1959]" box="[189,870,1933,1959]" pageId="27" pageNumber="28">THORACIC SHIELD. Surface like those of tergites, glabrous.</paragraph>
|
||
<paragraph id="8BA9FA74FFA5E0291EBBF83586F6FEDF" blockId="27.[189,1333,1998,2025]" lastBlockId="29.[189,1399,265,362]" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
|
||
TERGITES. Smooth and glabrous, no small pits present. Paratergites projecting slightly posteriorly. ENDOTERGUM. Inner section with numerous short spines and isolated bristles. Between marginal ridge and inner area a single row of sparse, circular cuticular impressions. Externally two dense rows of marginal bristles. Bristles scaly, the longest reaching above tergite margin (
|
||
<figureCitation id="132DE6F1FFA3E0291A29FEB486E8FEDF" box="[1071,1161,335,362]" captionStart="Fig" captionStartId="12.[189,232,1891,1917]" captionTargetBox="[270,1317,266,1874]" captionTargetId="figure-78@12.[270,1317,266,1875]" captionTargetPageId="12" captionText="Fig. 5. Scanning electron micrographs of the endoterga of midbody rings. A. Zoosphaerium nigrum sp. nov., holotype (CASENT 9068289). B. Z. silens sp. nov., holotype (FMNH-INS 3119887). C. Z. ambatovaky sp. nov., holotype (CASENT 9068294/1). D. Z. beanka sp. nov., holotype (FMNH- INS 3119897). E. Z. voahangy sp. nov., holotype (FMNH-INS 3119921). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5075834" httpUri="https://zenodo.org/record/5075834/files/figure.png" pageId="29" pageNumber="30">Fig. 5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF69AAFCFFA2E0281EBBF9018716F85D" ID-DOI="http://doi.org/10.5281/zenodo.5075842" ID-Zenodo-Dep="5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1786,1812]" targetBox="[243,1356,268,1770]" targetPageId="28">
|
||
<paragraph id="8BA9FA74FFA2E0281EBBF9018716F85D" blockId="28.[188,1399,1786,2024]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9622666FFA2E0281EBBF9018371F8A1" bold="true" box="[189,272,1786,1812]" pageId="28" pageNumber="29">Fig. 9.</emphasis>
|
||
<taxonomicName id="4C1681F7FFA2E0281F1AF9018025F8A1" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[284,580,1786,1812]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
|
||
<emphasis id="B9622666FFA2E0281F1AF9018025F8A1" box="[284,580,1786,1812]" italics="true" pageId="28" pageNumber="29">Zoosphaerium voahangy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFA2E0281C57F90080D2F8A0" box="[593,691,1787,1813]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (FMNH-INS 3119921).
|
||
<emphasis id="B9622666FFA2E0281A59F9018615F8A1" bold="true" box="[1119,1140,1786,1812]" pageId="28" pageNumber="29">A</emphasis>
|
||
. left leg 9, posterior view.
|
||
<emphasis id="B9622666FFA2E0281F05F8E58376F88D" bold="true" box="[259,279,1822,1848]" pageId="28" pageNumber="29">B</emphasis>
|
||
. Left coxa 2 with gonopore, posterior view.
|
||
<emphasis id="B9622666FFA2E0281D27F8E68157F882" bold="true" box="[801,822,1821,1847]" pageId="28" pageNumber="29">C</emphasis>
|
||
. Left coxa 1 with stigmatic plate, posterior view.
|
||
<emphasis id="B9622666FFA2E0281EBBF8BA82B3F8EE" bold="true" box="[189,210,1857,1883]" pageId="28" pageNumber="29">D</emphasis>
|
||
. Left anterior gonopod, anterior view.
|
||
<emphasis id="B9622666FFA2E0281C96F8BA80C5F8EE" bold="true" box="[656,676,1857,1883]" pageId="28" pageNumber="29">E</emphasis>
|
||
. Left anterior gonopod, lateral view.
|
||
<emphasis id="B9622666FFA2E0281A56F8BA8603F8EE" bold="true" box="[1104,1122,1857,1883]" pageId="28" pageNumber="29">F</emphasis>
|
||
. Left anterior gonopod, posterior view.
|
||
<emphasis id="B9622666FFA2E0281F7EF89F83EEF8CB" bold="true" box="[376,399,1892,1918]" pageId="28" pageNumber="29">G</emphasis>
|
||
. Right posterior telopod, posterior view.
|
||
<emphasis id="B9622666FFA2E0281D85F89F81FBF8CB" bold="true" box="[899,922,1892,1918]" pageId="28" pageNumber="29">H</emphasis>
|
||
. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFE6A830EFE44" blockId="29.[189,1399,401,497]" pageId="29" pageNumber="30">ANAL SHIELD. Rounded, neither bell-shaped nor tapered, glabrous. Ventral side with two black locking carinae, located closely to laterotergites. Anterior carina small and posterior carina 2–3 times as long as anterior carina.</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFDE286EEFDCC" blockId="29.[189,1399,536,634]" pageId="29" pageNumber="30">
|
||
LEGS. Leg 1 with 4 ventral spines, leg 2 with 5 or 6, leg 3 with 7. First two leg pairs without an apical spine, spine present on leg 3. Leg pairs 4–21 with 9 or 10 ventral spines and an apical spine. On leg 9, femur 2.1 and tarsus 5.1 times as long as wide. All podomeres with setae (
|
||
<figureCitation id="132DE6F1FFA3E0291A25FDA486E2FDCC" box="[1059,1155,607,633]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EECFD5A806AFD0E" blockId="29.[189,1398,673,769]" box="[234,523,673,699]" pageId="29" pageNumber="30">
|
||
<emphasis id="B9622666FFA3E0291EECFD5A806AFD0E" bold="true" box="[234,523,673,699]" pageId="29" pageNumber="30">Male sexual characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFD3F8652FCB4" blockId="29.[189,1398,673,769]" pageId="29" pageNumber="30">
|
||
GONOPORE. Covered with a single undivided, apically rounded, membranous, basally sclerotized plate covering ¼ of surface of coxa. Gonopore located basally on joint (
|
||
<figureCitation id="132DE6F1FFA3E0291DCEFD1C8644FCB4" box="[968,1061,743,769]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFCD383C4FBA0" blockId="29.[186,1400,808,1045]" pageId="29" pageNumber="30">
|
||
ANTERIOR TELOPODS. Harp with single stridulation rib located mesally, long and well developed (
|
||
<figureCitation id="132DE6F1FFA3E0291B0BFCD38708FCF7" box="[1293,1385,808,834]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9D</figureCitation>
|
||
). First podomere 1.1 times as wide as long. Second podomere process lobe-like and slightly curved with rounded edges, protruding up to half of third podomere height, apically with sclerotized spots (
|
||
<figureCitation id="132DE6F1FFA3E0291B16FC948708FC3C" box="[1296,1385,879,905]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9F</figureCitation>
|
||
). Third podomere long, almost as long as first; cavity mesally with numerous sclerotized spots and three thin, sclerotized spines located apically (
|
||
<figureCitation id="132DE6F1FFA3E0291CA6FC4E817DFC7A" box="[672,796,949,975]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9E–F</figureCitation>
|
||
); one sclerotized spot located close to tip, laterally with a row of ca 11 crenulated teeth (
|
||
<figureCitation id="132DE6F1FFA3E0291C85FC238081FC47" box="[643,736,984,1010]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9E</figureCitation>
|
||
). Podomeres 1–3 covered on both sides with numerous long, isolated hairs.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFBC683C5FA9F" blockId="29.[189,1399,1085,1322]" pageId="29" pageNumber="30">
|
||
POSTERIOR TELOPODS. Movable finger slender and long, with a well-rounded tip, not tapering towards tip; hollowed-out inner margin with 3 or 4 sclerotized spines located apically, posterior aspect with ca 25 small sclerotized teeth (
|
||
<figureCitation id="132DE6F1FFA3E0291C20FB7880E6FB28" box="[550,647,1155,1181]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9G</figureCitation>
|
||
). Movable finger 4 times as long as wide, slightly longer than fixed finger. Fixed finger straight, only apically curved towards movable finger, not tapering, more slender than third podomere. Fixed finger at margin with numerous sclerotized spots (
|
||
<figureCitation id="132DE6F1FFA3E0291ACFFB318748FB51" box="[1225,1321,1226,1252]" captionStart="Fig" captionStartId="28.[189,232,1786,1812]" captionTargetBox="[243,1356,268,1770]" captionTargetId="figure-142@28.[226,1362,258,1778]" captionTargetPageId="28" captionText="Fig. 9. Zoosphaerium voahangy sp. nov., holotype (FMNH-INS 3119921). A. left leg 9, posterior view. B. Left coxa 2 with gonopore, posterior view. C. Left coxa 1 with stigmatic plate, posterior view. D. Left anterior gonopod, anterior view. E. Left anterior gonopod, lateral view. F. Left anterior gonopod, posterior view. G. Right posterior telopod, posterior view. H. Right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cx= coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-ps = sclerotized spots; sr = stridulation rib; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. Roman numerals indicate telopoditomeres. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5075842" httpUri="https://zenodo.org/record/5075842/files/figure.png" pageId="29" pageNumber="30">Fig. 9H</figureCitation>
|
||
). First podomere in posterior view almost glabrous, podomeres 2 and 3 covered on both sides with numerous long, isolated hairs.
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EECFAA98324FAD9" blockId="29.[189,325,1362,1423]" box="[234,325,1362,1388]" pageId="29" pageNumber="30">
|
||
<emphasis id="B9622666FFA3E0291EECFAA98324FAD9" bold="true" box="[234,325,1362,1388]" pageId="29" pageNumber="30">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFA8E8358FA3A" blockId="29.[189,325,1362,1423]" box="[189,313,1397,1423]" pageId="29" pageNumber="30">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CA9FFFFA3E0291EBBFA4D875CF9F0" pageId="29" pageNumber="30" type="distribution">
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFA4D8390FA65" blockId="29.[189,497,1462,1488]" box="[189,497,1462,1488]" pageId="29" pageNumber="30">
|
||
<heading id="D0E14D18FFA3E0291EBBFA4D8390FA65" bold="true" box="[189,497,1462,1488]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis id="B9622666FFA3E0291EBBFA4D8390FA65" bold="true" box="[189,497,1462,1488]" pageId="29" pageNumber="30">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA9FA74FFA3E0291EBBFA1E875CF9F0" blockId="29.[189,1399,1509,1606]" pageId="29" pageNumber="30">
|
||
The only known habitat of
|
||
<taxonomicName id="4C1681F7FFA3E0291FFDFA1D80EDFA4A" authority="Wesener & Sagorny, 2021" authorityName="Wesener & Sagorny" authorityYear="2021" box="[507,652,1509,1535]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="voahangy" status="sp. nov.">
|
||
<emphasis id="B9622666FFA3E0291FFDFA1D80EDFA4A" box="[507,652,1509,1535]" italics="true" pageId="29" pageNumber="30">Z. voahangy</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2519B1DFFA3E0291C92FA1E8090FA4A" box="[660,753,1509,1535]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, the forest of Analalava (
|
||
<figureCitation id="132DE6F1FFA3E0291A11FA1E863CFA4A" box="[1047,1117,1509,1535]" captionStart="Fig" captionStartId="10.[189,232,1891,1917]" captionTargetBox="[228,1372,266,1870]" captionTargetId="figure-87@10.[202,1385,239,1881]" captionTargetPageId="10" captionText="Fig. 3. Distribution map of the newly described species of Zoosphaerium. Vegetation color marks modified from Moat & Smith 2007. 1 = Z. nigrum sp. nov.; 2 = Z. silens sp. nov.; 3 = Z. ambatovaky sp. nov.; 4 = Z. beanka sp. nov.; 5 = Z. voahangy sp. nov.; 6 = Z. masoala sp. nov.; 7 = Z. spinopiligerum sp. nov. Numbers followed by ‘a’ mark additional, non-type localities." figureDoi="http://doi.org/10.5281/zenodo.5075830" httpUri="https://zenodo.org/record/5075830/files/figure.png" pageId="29" pageNumber="30">Fig. 3</figureCitation>
|
||
), is a small (
|
||
<quantity id="4CEE5791FFA3E0291AF5FA1E874EFA4A" box="[1267,1327,1509,1535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="km" value="2.0">2 km</quantity>
|
||
south to north,
|
||
<quantity id="4CEE5791FFA3E0291F2FF9F38315F997" box="[297,372,1544,1571]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="29" pageNumber="30" unit="m" value="750.0">750 m</quantity>
|
||
east to west) fragment of disturbed and partly degraded lowland rainforest, isolated by
|
||
<quantity id="4CEE5791FFA3E0291EB8F9D08366F9F0" box="[190,263,1579,1605]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="km" value="30.0">30 km</quantity>
|
||
from Betampona, the only other remaining lowland rainforest fragment in a
|
||
<quantity id="4CEE5791FFA3E0291A88F9D08687F9F0" box="[1166,1254,1579,1605]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="km" value="100.0">100 km</quantity>
|
||
radius.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |