241 lines
31 KiB
XML
241 lines
31 KiB
XML
<document id="807336C9E7A9F0EB0A5D0CA1EA53FBA3" ID-DOI="10.11646/zootaxa.4790.1.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="3887813" ID-ZooBank="E9496778-7D5E-4D55-BF7D-BC820CCBC021" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591774886097" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03A12F692105AE014A9EFF14EE30B09C" docLanguage="en" docName="zootaxa.4790.1.10.pdf" docOrigin="Zootaxa 4790 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clidicus mawarensis Jałoszyński 2020, sp. n." docType="treatment" docVersion="3" lastPageNumber="191" masterDocId="FF9857112107AE054A09FFF5EF18B43F" masterDocTitle="A new species of Clidicus in East Malaysia (Coleoptera, Staphylinidae Scydmaeninae)" masterLastPageNumber="192" masterPageNumber="187" pageNumber="189" updateTime="1700276526832" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="206B6FE3F98AE36B22E7B3385FBBE7FB">
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<mods:title id="75695D9FC2E14F5F2AE0363E9750C777">A new species of Clidicus in East Malaysia (Coleoptera, Staphylinidae Scydmaeninae)</mods:title>
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<mods:name id="8822B6FA796C81382CC3ED943335930A" type="personal">
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<mods:roleTerm id="92780D561E405583229B2F538D48C640">Author</mods:roleTerm>
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<mods:namePart id="A3943615C72B5A00C778775A9F7DF88C">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="A79F63419E5B89F1B8E86B057BB19B7D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F93FB7D5693DC34CF28FCAEC4F0DC0B4">
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<mods:date id="77CBEB7137781DC2780EE61004B75958">2020</mods:date>
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<mods:detail id="CA13DA619DA889C555D508C5F01554C3" type="pubDate">
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<mods:number id="7C1DA404353166749B3A6064D274359A">2020-06-10</mods:number>
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<mods:number id="406D233FCBD5165E996EA715F282A504">4790</mods:number>
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<mods:detail id="A157A6C6CD5A474F93355F92179E0626" type="issue">
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<mods:number id="E74D9C9E1F78BA6C62C218FD00D07C43">1</mods:number>
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<mods:start id="5CCC6ADD6E4CE685608CC99A28FEF410">187</mods:start>
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<mods:classification id="F0E079AECBDA842F248D59D671C62FD2">journal article</mods:classification>
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<mods:identifier id="7F399A5ED59C889C0C287B6587047C84" type="DOI">10.11646/zootaxa.4790.1.12</mods:identifier>
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<mods:identifier id="AE8EC00C49FD5C826C06C7A6FF85EFC0" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C312CDF42105AE074A9EFF14EEC3B4C3" box="[151,475,225,252]" pageId="2" pageNumber="189" type="nomenclature">
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<paragraph id="8BB79E7F2105AE074A9EFF14EEC3B4C3" blockId="2.[151,475,225,287]" box="[151,475,225,252]" pageId="2" pageNumber="189">
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<heading id="D0FF29132105AE074A9EFF14EEC3B4C3" bold="true" box="[151,475,225,252]" fontSize="11" level="1" pageId="2" pageNumber="189" reason="1">
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<emphasis id="B97C426D2105AE074A9EFF14EEC3B4C3" bold="true" box="[151,475,225,252]" pageId="2" pageNumber="189">
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<taxonomicName id="4C08E5FC2105AE074A9EFF14EE89B4C4" ID-CoL="8XXT8" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,401,225,252]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="mawarensis" status="sp. nov.">
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<emphasis id="B97C426D2105AE074A9EFF14EE89B4C4" bold="true" box="[151,401,225,252]" italics="true" pageId="2" pageNumber="189">Clidicus mawarensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24FFF162105AE074B90FF17EEC3B4C3" box="[409,475,226,252]" pageId="2" pageNumber="189" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C312CDF42105AE074A9EFEF0EE01B520" box="[151,281,261,287]" pageId="2" pageNumber="189" type="description">
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<paragraph id="8BB79E7F2105AE074A9EFEF0EE01B520" blockId="2.[151,475,225,287]" box="[151,281,261,287]" pageId="2" pageNumber="189">
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(
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<figureCitation id="133382FA2105AE074A96FEF0EE09B520" box="[159,273,261,287]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1962,1987]" captionTargetBox="[163,1424,849,1938]" captionTargetId="figure@3.[151,1436,849,1938]" captionTargetPageId="3" captionText="FIGURES 7–12. Clidicus mawarensis sp. n. apical portion of median lobe in dorsal (7–8), ventral (9–10) and lateral (11–12) views. Abbreviations: cp, copulatory piece; fl, flagellum; gon, gonopore; hlp, hook-like projection." figureDoi="http://doi.org/10.5281/zenodo.3887819" httpUri="https://zenodo.org/record/3887819/files/figure.png" pageId="2" pageNumber="189">Figs 7–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C312CDF42105AE074A9EFEB8EDBBB590" pageId="2" pageNumber="189" type="materials_examined">
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<paragraph id="8BB79E7F2105AE074A9EFEB8EDBBB590" blockId="2.[151,1436,333,612]" pageId="2" pageNumber="189">
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<emphasis id="B97C426D2105AE074A9EFEB8EE7DB557" bold="true" box="[151,357,333,360]" pageId="2" pageNumber="189">Material studied.</emphasis>
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<materialsCitation id="3B6094222105AE074B65FEB8EDBBB590" collectionCode="NMPC" collectorName="P. Hlavac" country="Malaysia" location="Patau village" pageId="2" pageNumber="189" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
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<emphasis id="B97C426D2105AE074B65FEB8ED0CB557" bold="true" box="[364,532,333,360]" pageId="2" pageNumber="189">
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<typeStatus id="54B320DD2105AE074B65FEB8ED17B557" box="[364,527,333,360]" pageId="2" pageNumber="189" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D0E55F62105AE074815FEBBED29B557" box="[540,561,334,360]" pageId="2" pageNumber="189" type="male">♂</specimenCount>
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:
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<emphasis id="B97C426D2105AE074848FEB8EC49B557" bold="true" box="[577,849,333,360]" pageId="2" pageNumber="189">
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<collectingCountry id="F31FDEEF2105AE074848FEB8EDCDB557" box="[577,725,333,360]" name="Malaysia" pageId="2" pageNumber="189">MALAYSIA</collectingCountry>
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(
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<collectingRegion id="49CC509D2105AE0748ECFEB8EC51B557" box="[741,841,333,360]" country="Malaysia" name="Sabah" pageId="2" pageNumber="189">SABAH</collectingRegion>
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)
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</emphasis>
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, with two labels: “MALAYSIA-Sabah / Mawar wa- terfall / near
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<location id="8ED7C8A42105AE074B2EFE87EEA0B5B4" LSID="urn:lsid:plazi:treatment:03A12F692105AE014A9EFF14EE30B09C:8ED7C8A42105AE074B2EFE87EEA0B5B4" box="[295,440,369,396]" country="Malaysia" name="Patau village" pageId="2" pageNumber="189" stateProvince="Sabah">Patau village</location>
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/ 305.-31.5.98,
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<collectorName id="26FDFBA92105AE074863FE84EDD7B5B4" box="[618,719,369,395]" pageId="2" pageNumber="189">P.Hlaváč</collectorName>
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” [white, printed], “
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<taxonomicName id="4C08E5FC2105AE0749A1FE87EB3EB5B3" authorityName="Laporte de Castelnau" authorityYear="1832" box="[936,1062,370,396]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="genus">
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<emphasis id="B97C426D2105AE0749A1FE87EB3EB5B3" box="[936,1062,370,396]" italics="true" pageId="2" pageNumber="189">CLIDICUS</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C08E5FC2105AE074E35FE87EBD9B5B4" authorityName="Jałoszyński" authorityYear="2020" box="[1084,1217,370,395]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="mawarensis">
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<emphasis id="B97C426D2105AE074E35FE87EBD9B5B4" box="[1084,1217,370,395]" italics="true" pageId="2" pageNumber="189">mawarensis</emphasis>
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</taxonomicName>
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m. / P.
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<bibRefCitation id="EF99E38E2105AE074F1DFE84EFD7B590" author="Jaloszynski, P." pageId="2" pageNumber="189" pagination="87 - 94" refId="ref2019" refString="Jaloszynski, P. (2020) Clidicus minilankanus sp. n., with notes on remaining Sri Lankan Clidicus species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4718 (1), 87 - 94. https: // doi. org / 10.11646 / zootaxa. 4718.1.7" type="journal article" year="2020">Jałoszyński, 2020</bibRefCitation>
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/
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<typeStatus id="54B320DD2105AE074AECFE60EE93B590" box="[229,395,405,431]" pageId="2" pageNumber="189" type="holotype">HOLOTYPUS</typeStatus>
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” [red, printed] (
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<collectionCode id="ED1906BA2105AE074848FE60ED8FB590" box="[577,663,405,431]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="189">NMPC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C312CDF42105AE074ACEFE4CED84B65B" pageId="2" pageNumber="189" type="diagnosis">
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<paragraph id="8BB79E7F2105AE074ACEFE4CED84B65B" blockId="2.[151,1436,333,612]" pageId="2" pageNumber="189">
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<emphasis id="B97C426D2105AE074ACEFE4CEE5AB5EB" bold="true" box="[199,322,441,468]" pageId="2" pageNumber="189">Diagnosis.</emphasis>
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Body large, BL>
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<quantity id="4CF0339A2105AE074800FE4CED78B5EB" box="[521,608,441,468]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="189" unit="mm" value="6.5">6.5 mm</quantity>
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; punctures on pronotum separated by distances subequal to their diameters; most flagellomeres elongate; aedeagus stout and massive, with short, slender and slightly asymmetrical parameres (one slightly longer than the other), with broad apex of median lobe and complex apical structures, with a pair of lateral apical hook-like projections at sides of gonopore-bearing median projection and with ventral subtrapezoidal shield-like plate with deeply emarginate apex.
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</paragraph>
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</subSubSection>
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||
<caption id="DF77CEF72105AE074A9EF843EBDFB3F0" ID-DOI="http://doi.org/10.5281/zenodo.3887817" ID-Zenodo-Dep="3887817" box="[151,1223,1974,1999]" httpUri="https://zenodo.org/record/3887817/files/figure.png" pageId="2" pageNumber="189" startId="2.[151,264,1974,1999]" targetBox="[163,1425,646,1935]" targetPageId="2">
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<paragraph id="8BB79E7F2105AE074A9EF843EBDFB3F0" blockId="2.[151,1223,1974,1999]" box="[151,1223,1974,1999]" pageId="2" pageNumber="189">
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<emphasis id="B97C426D2105AE074A9EF843EE23B3F0" bold="true" box="[151,315,1974,1999]" pageId="2" pageNumber="189">FIGURES 4–6.</emphasis>
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<taxonomicName id="4C08E5FC2105AE074B48F842ED0CB3F0" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[321,532,1975,1999]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="mawarensis" status="sp. nov.">
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<emphasis id="B97C426D2105AE074B48F842ED0CB3F0" box="[321,532,1975,1999]" italics="true" pageId="2" pageNumber="189">Clidicus mawarensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24FFF162105AE074813F84DED49B3F0" box="[538,593,1976,1999]" pageId="2" pageNumber="189" rank="species">sp. n.</taxonomicNameLabel>
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Aedeagus in dorsal (4) and lateral (5) views; sperm pump (6).
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</paragraph>
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</caption>
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<subSubSection id="C312CDF42104AE014ACEFF6DEEBAB6CC" lastPageId="4" lastPageNumber="191" pageId="3" pageNumber="190" type="description">
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<paragraph id="8BB79E7F2104AE064ACEFF6DEEA7B4C4" blockId="3.[151,1437,152,827]" pageId="3" pageNumber="190">
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<emphasis id="B97C426D2104AE064ACEFF6DEE41B48C" bold="true" box="[199,345,152,179]" pageId="3" pageNumber="190">Description.</emphasis>
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Body of male (
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<figureCitation id="133382FA2104AE06480EFF6CED5FB48C" box="[519,583,153,179]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1964,1989]" captionTargetBox="[166,1433,744,1929]" captionTargetId="figure@1.[151,1436,741,1938]" captionTargetPageId="1" captionText="FIGURES 1–3. Clidicus mawarensis sp. n. Habitus of male (1); head of male in dorsal view (2), apex of male terminalia in ventral view (3). Abbreviation: st9, sternite IX." figureDoi="http://doi.org/10.5281/zenodo.3887815" httpUri="https://zenodo.org/record/3887815/files/figure.png" pageId="3" pageNumber="190">Fig. 1</figureCitation>
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) elongate and slender, head flattened, prothorax and elytra strongly convex; legs and antennae long and slender, BL
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<quantity id="4CF0339A2104AE064847FF49EDACB4E8" box="[590,692,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" pageId="3" pageNumber="190" unit="mm" value="6.75">6.75 mm</quantity>
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; cuticle glossy, pigmentation uniformly reddish brown, setae indistinctly lighter than cuticle.
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</paragraph>
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<paragraph id="8BB79E7F2104AE064ACEFEF0ED27B6B8" blockId="3.[151,1437,152,827]" pageId="3" pageNumber="190">
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Head (
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<figureCitation id="133382FA2104AE064B18FEF0EE4BB520" box="[273,339,261,287]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1964,1989]" captionTargetBox="[166,1433,744,1929]" captionTargetId="figure@1.[151,1436,741,1938]" captionTargetPageId="1" captionText="FIGURES 1–3. Clidicus mawarensis sp. n. Habitus of male (1); head of male in dorsal view (2), apex of male terminalia in ventral view (3). Abbreviation: st9, sternite IX." figureDoi="http://doi.org/10.5281/zenodo.3887815" httpUri="https://zenodo.org/record/3887815/files/figure.png" pageId="3" pageNumber="190">Fig. 2</figureCitation>
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) broadest distinctly behind middle of tempora, HL
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<quantity id="4CF0339A2104AE064983FEF0ECF7B520" box="[906,1007,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="3" pageNumber="190" unit="mm" value="1.13">1.13 mm</quantity>
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, HW
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<quantity id="4CF0339A2104AE064E27FEF1EB8CB520" box="[1070,1172,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="190" unit="mm" value="1.5">1.50 mm</quantity>
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; tempora in dorsal view nearly 4 times as long as eyes, nearly straight in slightly more than anterior half and weakly divergent posterad, posteriorly strongly and evenly rounded; vertex with a deep subtrapezoidal posteromedian impression and a pair of indistinct posterior pits marking sites where dorsal tentorial arms are fused with dorsal wall of head. Sides of vertex convex, frons near each antennal insertion slightly impressed, entire frons and vertex with a narrow median longitudinal groove, supraantennal tubercles small but distinctly elevated; clypeus transverse and convex, demarcated from frons by a deep and distinct frontoclypeal groove; compound eyes small, nearly round, strongly convex but in dorsal view not projecting from head silhouette, finely faceted. Punctures on vertex and frons sparse and fine, inconspicuous; supraantennal tubercles virtually impunctate. Setae moderately long, sparse, erect. Antennae slender, scape slightly longer than head, antennomeres II–VIII elongate (VIII indistinctly so), IX–X as long as broad, XI indistinctly transverse; AnL
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<quantity id="4CF0339A2104AE064BDDFD98ED23B6B8" box="[468,571,621,647]" metricMagnitude="-3" metricUnit="m" metricValue="3.63" pageId="3" pageNumber="190" unit="mm" value="3.63">3.63 mm</quantity>
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.
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||
</paragraph>
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<paragraph id="8BB79E7F2104AE064ACEFD64EC26B694" blockId="3.[151,1437,152,827]" box="[199,830,657,683]" pageId="3" pageNumber="190">Labrum with shallow subtriangular anteromedian notch.</paragraph>
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<paragraph id="8BB79E7F2104AE064ACEFD40EA93B704" blockId="3.[151,1437,152,827]" pageId="3" pageNumber="190">
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Pronotum with subglobose disc and short posterior ‘collar’, broadest near anterior third; PL
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||
<quantity id="4CF0339A2104AE064EC3FD40EA28B6F0" box="[1226,1328,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="3" pageNumber="190" unit="mm" value="1.63">1.63 mm</quantity>
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, PW
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<quantity id="4CF0339A2104AE064F62FD41EFDEB6CC" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="190" unit="mm" value="1.55">1.55 mm</quantity>
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; ‘collar’ demarcated by a narrow transverse groove with twelve pits increasing in diameter toward sides. Punc- tures on disc denser than those on head, in median area separated by spaces subequal to diameters of punctures; setae moderately long, dense, and suberect. Cuticle between punctures on pronotal disc as glossy as that on head.
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</paragraph>
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<caption id="DF77CEF72104AE064A9EF85FEB9AB3D8" ID-DOI="http://doi.org/10.5281/zenodo.3887819" ID-Zenodo-Dep="3887819" httpUri="https://zenodo.org/record/3887819/files/figure.png" pageId="3" pageNumber="190" startId="3.[151,264,1962,1987]" targetBox="[163,1424,849,1938]" targetPageId="3">
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||
<paragraph id="8BB79E7F2104AE064A9EF85FEB9AB3D8" blockId="3.[151,1437,1962,2024]" pageId="3" pageNumber="190">
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<emphasis id="B97C426D2104AE064A9EF85FEE50B3FC" bold="true" box="[151,328,1962,1987]" pageId="3" pageNumber="190">FIGURES 7–12.</emphasis>
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<taxonomicName id="4C08E5FC2104AE064B46F85EED3BB3FC" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[335,547,1963,1987]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="mawarensis" status="sp. nov.">
|
||
<emphasis id="B97C426D2104AE064B46F85EED3BB3FC" box="[335,547,1963,1987]" italics="true" pageId="3" pageNumber="190">Clidicus mawarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24FFF162104AE064823F859ED7AB3FC" box="[554,610,1964,1987]" pageId="3" pageNumber="190" rank="species">sp. n.</taxonomicNameLabel>
|
||
apical portion of median lobe in dorsal (7–8), ventral (9–10) and lateral (11–12) views. Abbreviations: cp, copulatory piece; fl, flagellum; gon, gonopore; hlp, hook-like projection.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB79E7F2103AE014ACEFF6CEC44B57C" blockId="4.[151,1437,152,1187]" pageId="4" pageNumber="191">
|
||
Elytra together oval and strongly elongate, broadest slightly in front of middle, EL 4.00 mm, EW
|
||
<quantity id="4CF0339A2103AE014F19FF6DEA6EB48C" box="[1296,1398,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="4" pageNumber="191" unit="mm" value="2.15">2.15 mm</quantity>
|
||
, EI 1.86; each elytron with five dorsal and one lateral rows of large and deep, densely distributed pits, cuticle between pits glossy, between rows with fine and sparse setiferous punctures; setae longer and slightly thicker than those on head and pronotum, moderately dense, suberect. Apex of each elytron obliquely truncated, with lateral corner form- ing short denticle; adsutural corner weakly projecting posterad.
|
||
</paragraph>
|
||
<paragraph id="8BB79E7F2103AE014ACEFEB8EB55B558" blockId="4.[151,1437,152,1187]" box="[199,1101,333,359]" pageId="4" pageNumber="191">Hind wings present, longer than elytra but narrow and apparently not functional.</paragraph>
|
||
<paragraph id="8BB79E7F2103AE014ACEFE84ED4BB5B4" blockId="4.[151,1437,152,1187]" box="[199,595,369,395]" pageId="4" pageNumber="191">Legs long and slender, unmodified.</paragraph>
|
||
<paragraph id="8BB79E7F2103AE014ACEFE60EFFBB5EC" blockId="4.[151,1437,152,1187]" pageId="4" pageNumber="191">
|
||
Abdominal sternite IX (
|
||
<figureCitation id="133382FA2103AE014BDCFE60ED01B590" box="[469,537,405,431]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1964,1989]" captionTargetBox="[166,1433,744,1929]" captionTargetId="figure@1.[151,1436,741,1938]" captionTargetPageId="1" captionText="FIGURES 1–3. Clidicus mawarensis sp. n. Habitus of male (1); head of male in dorsal view (2), apex of male terminalia in ventral view (3). Abbreviation: st9, sternite IX." figureDoi="http://doi.org/10.5281/zenodo.3887815" httpUri="https://zenodo.org/record/3887815/files/figure.png" pageId="4" pageNumber="191">Fig. 3</figureCitation>
|
||
) with broadly V-shaped subapical transverse carina, area behind carina densely setose.
|
||
</paragraph>
|
||
<paragraph id="8BB79E7F2103AE014ACEFE28EBB4B6F0" blockId="4.[151,1437,152,1187]" pageId="4" pageNumber="191">
|
||
Aedeagus (
|
||
<figureCitation id="133382FA2103AE014B4FFE28EEB4B5C8" box="[326,428,476,503]" captionStart="FIGURES 4–6" captionStartId="2.[151,264,1974,1999]" captionTargetBox="[163,1425,646,1935]" captionTargetId="figure@2.[151,1436,633,1949]" captionTargetPageId="2" captionText="FIGURES 4–6. Clidicus mawarensis sp. n. Aedeagus in dorsal (4) and lateral (5) views; sperm pump (6)." figureDoi="http://doi.org/10.5281/zenodo.3887817" httpUri="https://zenodo.org/record/3887817/files/figure.png" pageId="4" pageNumber="191">Figs 4–5</figureCitation>
|
||
) relatively stout, in lateral view weakly curved, AeL
|
||
<quantity id="4CF0339A2103AE014E06FE28EB60B5C8" box="[1039,1144,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="4" pageNumber="191" unit="mm" value="1.33">1.33 mm</quantity>
|
||
; parameres short, slightly asymmetrical, one slightly longer than the other, their apical portions projecting beyond apex of median lobe nearly parallel; flagellum differentiated from ductus ejaculatorius by a rapid and strong broadening of walls near distal fourth of median lobe. Apex of median lobe (between parameres;
|
||
<figureCitation id="133382FA2103AE014976FDBCECE8B65C" box="[895,1008,585,611]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1962,1987]" captionTargetBox="[163,1424,849,1938]" captionTargetId="figure@3.[151,1436,849,1938]" captionTargetPageId="3" captionText="FIGURES 7–12. Clidicus mawarensis sp. n. apical portion of median lobe in dorsal (7–8), ventral (9–10) and lateral (11–12) views. Abbreviations: cp, copulatory piece; fl, flagellum; gon, gonopore; hlp, hook-like projection." figureDoi="http://doi.org/10.5281/zenodo.3887819" httpUri="https://zenodo.org/record/3887819/files/figure.png" pageId="4" pageNumber="191">Figs 7–12</figureCitation>
|
||
) broad and with median rounded projection with broad gonopore opening ventrally, flanked by a pair of robust tooth-like projections directed ventrad, ventral apical region of median lobe with shield-like plate with deeply emarginate distal margin. Sperm pump (
|
||
<figureCitation id="133382FA2103AE014F66FD64EFBEB6EF" captionStart="FIGURES 4–6" captionStartId="2.[151,264,1974,1999]" captionTargetBox="[163,1425,646,1935]" captionTargetId="figure@2.[151,1436,633,1949]" captionTargetPageId="2" captionText="FIGURES 4–6. Clidicus mawarensis sp. n. Aedeagus in dorsal (4) and lateral (5) views; sperm pump (6)." figureDoi="http://doi.org/10.5281/zenodo.3887817" httpUri="https://zenodo.org/record/3887817/files/figure.png" pageId="4" pageNumber="191">Fig. 6</figureCitation>
|
||
)
|
||
<quantity id="4CF0339A2103AE014ABFFD40EE02B6F0" box="[182,282,693,719]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="191" unit="mm" value="0.8">0.80 mm</quantity>
|
||
in length, with two large ‘funnels’, distal funnel much shorter than proximal one.
|
||
</paragraph>
|
||
<paragraph id="8BB79E7F2103AE014ACEFD2CEEBAB6CC" blockId="4.[151,1437,152,1187]" box="[199,418,729,755]" pageId="4" pageNumber="191">
|
||
<emphasis id="B97C426D2103AE014ACEFD2CEE3DB6CC" bold="true" box="[199,293,729,755]" pageId="4" pageNumber="191">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312CDF42103AE014ACEFD09ED36B728" box="[199,558,764,791]" pageId="4" pageNumber="191" type="distribution">
|
||
<paragraph id="8BB79E7F2103AE014ACEFD09ED36B728" blockId="4.[151,1437,152,1187]" box="[199,558,764,791]" pageId="4" pageNumber="191">
|
||
<emphasis id="B97C426D2103AE014ACEFD09EE79B728" bold="true" box="[199,353,764,791]" pageId="4" pageNumber="191">Distribution.</emphasis>
|
||
Northern Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312CDF42103AE014ACEFCD4ED4FB704" box="[199,599,801,827]" pageId="4" pageNumber="191" type="etymology">
|
||
<paragraph id="8BB79E7F2103AE014ACEFCD4ED4FB704" blockId="4.[151,1437,152,1187]" box="[199,599,801,827]" pageId="4" pageNumber="191">
|
||
<emphasis id="B97C426D2103AE014ACEFCD4EE56B704" bold="true" box="[199,334,801,827]" pageId="4" pageNumber="191">Etymology.</emphasis>
|
||
After Mawar waterfall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312CDF42103AE014ACEFCB0EE30B09C" pageId="4" pageNumber="191" type="discussion">
|
||
<paragraph id="8BB79E7F2103AE014ACEFCB0EE30B09C" blockId="4.[151,1437,152,1187]" pageId="4" pageNumber="191">
|
||
<emphasis id="B97C426D2103AE014ACEFCB0EE23B760" bold="true" box="[199,315,837,863]" pageId="4" pageNumber="191">Remarks.</emphasis>
|
||
<taxonomicName id="4C08E5FC2103AE014B4AFCB0ED36B760" authorityName="Jałoszyński" authorityYear="2020" box="[323,558,837,863]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="mawarensis">
|
||
<emphasis id="B97C426D2103AE014B4AFCB0ED36B760" box="[323,558,837,863]" italics="true" pageId="4" pageNumber="191">Clidicus mawarensis</emphasis>
|
||
</taxonomicName>
|
||
has large-bodied adults, which are externally similar to several congeners inhabiting SE Asia. Examination of the aedeagus is necessary to identify the new species. The male copulatory organ of
|
||
<taxonomicName id="4C08E5FC2103AE014ABEFC78EE47B798" authorityName="Jałoszyński" authorityYear="2020" box="[183,351,909,935]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="mawarensis">
|
||
<emphasis id="B97C426D2103AE014ABEFC78EE47B798" box="[183,351,909,935]" italics="true" pageId="4" pageNumber="191">C. mawarensis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to that of
|
||
<taxonomicName id="4C08E5FC2103AE014883FC78ECCCB798" authority="Reitter, 1887" authorityName="Reitter" authorityYear="1887" box="[650,980,909,935]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="ganglbaueri">
|
||
<emphasis id="B97C426D2103AE014883FC78EC2EB798" box="[650,822,909,935]" italics="true" pageId="4" pageNumber="191">C. ganglbaueri</emphasis>
|
||
<bibRefCitation id="EF99E38E2103AE014937FC78ECCCB798" author="Reitter, E." box="[830,980,909,935]" pageId="4" pageNumber="191" pagination="64" refId="ref2332" refString="Reitter, E. (1887) Ubersicht der bekannten Arten der Coleopteren-Gattung Clidicus Casteln. Wiener Entomologische Zeitung, 6, 64. https: // doi. org / 10.5962 / bhl. part. 17733" type="journal article" year="1887">Reitter, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
, known to occur in
|
||
<collectingCountry id="F31FDEEF2103AE014EB3FC78EA3FB798" box="[1210,1319,909,935]" name="Indonesia" pageId="4" pageNumber="191">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49CC509D2103AE014F31FC78EA73B798" box="[1336,1387,909,935]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="191">Java</collectingRegion>
|
||
and
|
||
<collectingRegion id="49CC509D2103AE014A9EFC44EFE2B7F4" box="[151,250,945,971]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="191">Sumatra</collectingRegion>
|
||
). In both species the median lobe is relatively massive and stout, the parameres are slightly asymmetrical, and the sperm pump has a similar shape. However, in
|
||
<taxonomicName id="4C08E5FC2103AE0148E2FC20EC88B7D0" authorityName="Jałoszyński" authorityYear="2020" box="[747,912,981,1007]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="mawarensis">
|
||
<emphasis id="B97C426D2103AE0148E2FC20EC88B7D0" box="[747,912,981,1007]" italics="true" pageId="4" pageNumber="191">C. mawarensis</emphasis>
|
||
</taxonomicName>
|
||
the entire aedeagus is much slenderer in lateral view; the median lobe is broadest in the subapical region (in sub-basal region in
|
||
<taxonomicName id="4C08E5FC2103AE014E28FC0CEBD2B02C" authorityName="Reitter" authorityYear="1887" box="[1057,1226,1017,1043]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="ganglbaueri">
|
||
<emphasis id="B97C426D2103AE014E28FC0CEBD2B02C" box="[1057,1226,1017,1043]" italics="true" pageId="4" pageNumber="191">C. ganglbaueri</emphasis>
|
||
</taxonomicName>
|
||
); the subapical region bears a small shield-like ventral plate with pointed distolateral corners (in
|
||
<taxonomicName id="4C08E5FC2103AE014E03FBE8EBACB008" authorityName="Reitter" authorityYear="1887" box="[1034,1204,1053,1079]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="ganglbaueri">
|
||
<emphasis id="B97C426D2103AE014E03FBE8EBACB008" box="[1034,1204,1053,1079]" italics="true" pageId="4" pageNumber="191">C. ganglbaueri</emphasis>
|
||
</taxonomicName>
|
||
an elongated ventral plate with rounded distolateral corners); and the parameres are distinctly shorter in relation to the median lobe and constricted (long and gradually narrowing in
|
||
<taxonomicName id="4C08E5FC2103AE014884FB90EC2EB040" authorityName="Reitter" authorityYear="1887" box="[653,822,1125,1151]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="ganglbaueri">
|
||
<emphasis id="B97C426D2103AE014884FB90EC2EB040" box="[653,822,1125,1151]" italics="true" pageId="4" pageNumber="191">C. ganglbaueri</emphasis>
|
||
</taxonomicName>
|
||
). Moreover, the elytra of males are more elongate in
|
||
<emphasis id="B97C426D2103AE014F8AFB90EE30B09C" italics="true" pageId="4" pageNumber="191">
|
||
<taxonomicName id="4C08E5FC2103AE014F8AFB90EE3CB09C" authorityName="Reitter" authorityYear="1887" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="ganglbaueri">C. ganglbaueri</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |