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<document id="D9296C66CBC86BFAE5CB9EACDA8D4313" ID-CLB-Dataset="31842" ID-DOI="10.11646/zootaxa.4284.1.1" ID-GBIF-Dataset="ac866645-a8b9-45a8-8c7d-d3aff74ede70" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010400" ID-ZooBank="1CD17B31-E3D0-4BFB-8B69-144D97949038" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507890517655" checkinUser="plazi" docAuthor="Kaczmarek, Łukasz" docDate="2017" docId="03ED740D32227E73FF6BFBDC6800FF43" docLanguage="en" docName="zootaxa.4284.1.1.pdf.imf" docOrigin="Zootaxa 4284 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paramacrobiotus areolatus Kaczmarek 2017, sensu lato" docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="FFD40C7532177E45FFFCFFEB6D1DFF96" masterDocTitle="Annotated zoogeography of non-marine Tardigrada. Part IV: Africa" masterLastPageNumber="74" masterPageNumber="1" pageNumber="54" updateTime="1698469609339" updateUser="ExternalLinkService">
<mods:mods id="D097D1351A7B0F1C3F27D27342419F91" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D20D3E03DFCBC30EDBFA61BB4AC6A607">
<mods:title id="FE6F1CA7F96E139BC6D2CD8779423868">Annotated zoogeography of non-marine Tardigrada. Part IV: Africa</mods:title>
</mods:titleInfo>
<mods:name id="E52E3B6E2585D3FD4CD5F01A4363EF37" type="personal">
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<mods:roleTerm id="2705CD0F60C146CBF9654FE68A910303">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9986E6E386F466D0DAE79381F81EC7E9">Kaczmarek, Łukasz</mods:namePart>
</mods:name>
<mods:typeOfResource id="5C7237982E7D9A3E7973B0D849F231F5">text</mods:typeOfResource>
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<mods:title id="4698A16B68F7E754B8108F87E341CE75">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6ADAFB687441DE3426B44292CB054F89">
<mods:date id="4C16794477DD8C65EEB24327DC25ED59">2017</mods:date>
<mods:detail id="9B78AFE1741F5AC8A804EF399AB2B4BF" type="pubDate">
<mods:number id="AE092D0D0B5CDA7E1AE41F7ED208C395">2017-06-28</mods:number>
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<mods:detail id="975BE801D95BA8609509AE4B5095DCBC" type="volume">
<mods:number id="E31C9907088CF6D5AA2E77AB25B9B253">4284</mods:number>
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<mods:classification id="8A28D3902AC574B5F33919BF328AFF54">journal article</mods:classification>
<mods:identifier id="4936476075DADFECA6FDD1FD64F5AEB4" type="CLB-Dataset">31842</mods:identifier>
<mods:identifier id="346B5632B9A9315C4F031FE6EDB714A7" type="DOI">10.11646/zootaxa.4284.1.1</mods:identifier>
<mods:identifier id="86FBB460C70DD15D7B935A8E26ADC15A" type="GBIF-Dataset">ac866645-a8b9-45a8-8c7d-d3aff74ede70</mods:identifier>
<mods:identifier id="7F95286393C0D0D983E96233B6336F53" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="874EF912497E9C642F3D6373D30A2D0C" type="Zenodo-Dep">1010400</mods:identifier>
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<treatment id="03ED740D32227E73FF6BFBDC6800FF43" ID-DOI="http://doi.org/10.5281/zenodo.6050227" ID-GBIF-Taxon="135435631" ID-Zenodo-Dep="6050227" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43" httpUri="http://treatment.plazi.org/id/03ED740D32227E73FF6BFBDC6800FF43" lastPageId="54" lastPageNumber="54" pageId="53" pageNumber="54">
<subSubSection id="C35E969032227E70FF6BFBDC6E64FBC6" box="[151,889,1079,1104]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="8BFBC51B32227E70FF6BFBDC6E64FBC6" blockId="53.[151,1436,1079,2037]" box="[151,889,1079,1104]" pageId="53" pageNumber="54">
<heading id="D0B3727732227E70FF6BFBDC6E64FBC6" bold="true" box="[151,889,1079,1104]" fontSize="10" level="2" pageId="53" pageNumber="54" reason="2">
<emphasis id="B930190932227E70FF6BFBDC6E64FBC6" bold="true" box="[151,889,1079,1104]" pageId="53" pageNumber="54">
156.
<taxonomicName id="4C44BE9832227E70FF33FBDC6FCCFBC6" ID-CoL="d787a5ff-484c-4a42-8638-c66ef3a8833b" authority="Murray, 1907c" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" box="[207,721,1079,1104]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" sensu="lato" species="areolatus">
<emphasis id="B930190932227E70FF33FBDC6F14FBC6" bold="true" box="[207,521,1079,1104]" italics="true" pageId="53" pageNumber="54">Paramacrobiotus areolatus</emphasis>
(
<bibRefCitation id="EFD5B8EA32227E70FDEBFBDC6FD4FBC6" author="Murray" box="[535,713,1079,1104]" pageId="53" pageNumber="54" refString="Murray, J. (1907 c) Arctic Tardigrada, collected by Wm. S. Bruce. TRansactions of the Royal Society of EdinbuRgh, 45, 669 - 681. https: // doi. org / 10.1017 / S 0080456800011789" type="journal article" year="1907" yearSuffix="c">Murray, 1907c</bibRefCitation>
)
</taxonomicName>
<emphasis id="B930190932227E70FD25FBD36E50FBC6" bold="true" box="[729,845,1079,1104]" italics="true" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A203A47232227E70FD25FBD36E50FBC6" box="[729,845,1079,1104]" pageId="53" pageNumber="54" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
[T]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E969032227E70FF3BFBB76EBCF96E" pageId="53" pageNumber="54" type="description">
<paragraph id="8BFBC51B32227E70FF3BFBB76EBBFBE4" blockId="53.[151,1436,1079,2037]" box="[199,934,1116,1138]" pageId="53" pageNumber="54">
<taxonomicName id="4C44BE9832227E70FF3BFBB76C55FBE4" authorityName="Radiatus Pilato, Binda &amp; Catanzaro" authorityYear="1991" box="[199,328,1116,1138]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="genus">
<emphasis id="B930190932227E70FF3BFBB76C55FBE4" box="[199,328,1116,1138]" italics="true" pageId="53" pageNumber="54">MacRobiotus</emphasis>
</taxonomicName>
<taxonomicName id="4C44BE9832227E70FEB1FBB76F20FBE4" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" box="[333,573,1116,1138]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="55" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="B930190932227E70FEB1FBB76CB1FBE4" box="[333,428,1116,1138]" italics="true" pageId="53" pageNumber="54">aReolatus</emphasis>
Murray, 1907
</taxonomicName>
(Binda 1984, Binda &amp; Pilato 1987)
</paragraph>
<paragraph id="8BFBC51B32227E70FF3BFB916E90FB06" blockId="53.[151,1436,1079,2037]" box="[199,909,1146,1168]" pageId="53" pageNumber="54">
<taxonomicName id="4C44BE9832227E70FF3BFB916C6AFB06" authorityName="Guidetti, Schill, Bertolani, Dandekar &amp; Wolf" authorityYear="2009" box="[199,375,1146,1168]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="genus">
<emphasis id="B930190932227E70FF3BFB916C6AFB06" box="[199,375,1146,1168]" italics="true" pageId="53" pageNumber="54">PaRamacRobiotus</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C44BE9832227E70FE62FB916F83FB06" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" box="[414,670,1146,1168]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="55" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="B930190932227E70FE62FB916CE0FB06" box="[414,509,1146,1168]" italics="true" pageId="53" pageNumber="54">aReolatus</emphasis>
(Murray, 1907)
</taxonomicName>
? (
<bibRefCitation id="EFD5B8EA32227E70FD49FB916E98FB06" author="Gasiorek" box="[693,901,1146,1168]" pageId="53" pageNumber="54" refString="Gasiorek, P., Morek, W., Stec, D., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688" type="book" year="2017">
Gąsiorek
<emphasis id="B930190932227E70FCE5FB906E51FB06" box="[793,844,1146,1168]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2017
</bibRefCitation>
)
</paragraph>
<paragraph id="8BFBC51B32227E70FF3BFB716F51FB39" blockId="53.[151,1436,1079,2037]" box="[199,588,1177,1199]" pageId="53" pageNumber="54">
<emphasis id="B930190932227E70FF3BFB716F51FB39" bold="true" box="[199,588,1177,1199]" pageId="53" pageNumber="54">
<taxonomicName id="4C44BE9832227E70FF3BFB716C59FB39" box="[199,324,1177,1199]" class="Insecta" family="Lycaenidae" genus="Terra" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="B930190932227E70FF3BFB716C59FB39" bold="true" box="[199,324,1177,1199]" italics="true" pageId="53" pageNumber="54">Terra typica</emphasis>
</taxonomicName>
: Svalbard (Spitsbergen)
</emphasis>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFB536DEDFB58" blockId="53.[151,1436,1079,2037]" box="[151,240,1208,1230]" pageId="53" pageNumber="54">
<emphasis id="B930190932227E70FF6BFB536DEDFB58" bold="true" box="[151,240,1208,1230]" pageId="53" pageNumber="54">Algeria:</emphasis>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFB3C699EFB7A" blockId="53.[151,1436,1079,2037]" box="[151,1155,1238,1260]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BFB3C699EFB7A" ID-GBIF-Occurrence="1671737429" box="[151,1155,1238,1260]" elevation="1400" latitude="36.666668" location="Pilato" longLatPrecision="1186" longitude="4.6" municipality="Akfadou Forest" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Tizi Ouzou Province">
•
<emphasis id="B930190932227E70FF3BFB3D6C9DFB7A" box="[199,384,1238,1260]" italics="true" pageId="53" pageNumber="54">
<geoCoordinate id="EE70A3DC32227E70FF3BFB3D6C00FB7A" box="[199,285,1238,1260]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="36.666668">36°40′N</geoCoordinate>
,
<geoCoordinate id="EE70A3DC32227E70FEDBFB3D6C67FB7A" box="[295,378,1238,1260]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="4.6">04°36′E</geoCoordinate>
;
</emphasis>
<emphasis id="B930190932227E70FE7BFB3D6F1AFB7A" bold="true" box="[391,519,1238,1260]" pageId="53" pageNumber="54">
<elevation id="0069222832227E70FE7BFB3D6F1EFB7A" box="[391,515,1238,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="53" pageNumber="54" unit="m" value="1400.0">
<quantity id="4CBC68FE32227E70FE7BFB3D6CC6FB7A" box="[391,475,1238,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="53" pageNumber="54" unit="m" value="1400.0">1,400 m</quantity>
asl
</elevation>
:
</emphasis>
<collectingRegion id="49800BF932227E70FDF2FB3D6FFEFB7A" box="[526,739,1238,1260]" country="Algeria" name="Tizi Ouzou" pageId="53" pageNumber="54">
Tizi
<collectingRegion id="49800BF932227E70FDC0FB3D6FFEFB7A" box="[572,739,1238,1260]" inferredBySuffix="true" pageId="53" pageNumber="54">Ouzou Province</collectingRegion>
</collectingRegion>
,
<collectingMunicipality id="6B9F5F6132227E70FD11FB3D6E93FB7A" box="[749,910,1238,1260]" pageId="53" pageNumber="54">Akfadou Forest</collectingMunicipality>
.
<emphasis id="B930190932227E70FC64FB3D699EFB7A" bold="true" box="[920,1155,1238,1260]" pageId="53" pageNumber="54">
Binda &amp;
<location id="8E9B93C032227E70FC06FB3D6924FB7A" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FC06FB3D6924FB7A" box="[1018,1081,1238,1260]" latitude="36.666668" longLatPrecision="1186" longitude="4.6" municipality="Akfadou Forest" name="Pilato" pageId="53" pageNumber="54" stateProvince="Tizi Ouzou Province">Pilato</location>
(1987)
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFB1E6C30FA9D" blockId="53.[151,1436,1079,2037]" box="[151,301,1269,1291]" pageId="53" pageNumber="54">
<emphasis id="B930190932227E70FF6BFB1E6C30FA9D" bold="true" box="[151,301,1269,1291]" pageId="53" pageNumber="54">Mozambique:</emphasis>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFAFE691DFABC" blockId="53.[151,1436,1079,2037]" box="[151,1024,1300,1322]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BFAFE691DFABC" ID-GBIF-Occurrence="1671737461" box="[151,1024,1300,1322]" latitude="-23.866667" location="Inhambene Province" longLatPrecision="1254" longitude="35.383335" municipality="Inhambene" pageId="53" pageNumber="54" specimenCount="1">
•
<emphasis id="B930190932227E70FF3BFAFF6CC9FABC" box="[199,468,1300,1322]" italics="true" pageId="53" pageNumber="54">
<geoCoordinate id="EE70A3DC32227E70FF3BFAFF6C04FABC" box="[199,281,1300,1322]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="-23.866667">23°52′S</geoCoordinate>
,
<geoCoordinate id="EE70A3DC32227E70FEDEFAFF6C6BFABC" box="[290,374,1300,1322]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="35.383335">35°23′E</geoCoordinate>
; 0 m asl:
</emphasis>
<location id="8E9B93C032227E70FE27FAFF6FB2FABC" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FE27FAFF6FB2FABC" box="[475,687,1300,1322]" latitude="-23.866667" longLatPrecision="1254" longitude="35.383335" municipality="Inhambene" name="Inhambene Province" pageId="53" pageNumber="54">Inhambene Province</location>
,
<collectingMunicipality id="6B9F5F6132227E70FD44FAFF6E36FABC" box="[696,811,1300,1322]" pageId="53" pageNumber="54">Inhambene</collectingMunicipality>
, moss.
<emphasis id="B930190932227E70FC8AFAFF691DFABC" bold="true" box="[886,1024,1300,1322]" pageId="53" pageNumber="54">Binda (1984)</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFAD96DF3FADE" blockId="53.[151,1436,1079,2037]" box="[151,238,1330,1352]" pageId="53" pageNumber="54">
<emphasis id="B930190932227E70FF6BFAD96DF3FADE" bold="true" box="[151,238,1330,1352]" pageId="53" pageNumber="54">Tunisia:</emphasis>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFAB96826FAF1" blockId="53.[151,1436,1079,2037]" box="[151,1339,1361,1383]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BFAB96826FAF1" ID-GBIF-Occurrence="1671737450" box="[151,1339,1361,1383]" elevation="120" latitude="37.179447" location="Raf Raf" longLatPrecision="19" longitude="10.253056" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Bizerte Governorate">
•
<emphasis id="B930190932227E70FF3BFABA6F55FAF1" bold="true" box="[199,584,1361,1383]" pageId="53" pageNumber="54">
<geoCoordinate id="EE70A3DC32227E70FF3BFABA6C58FAF1" box="[199,325,1361,1383]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="15" value="37.179447">37°10'46''N</geoCoordinate>
,
<geoCoordinate id="EE70A3DC32227E70FEADFABA6CD3FAF1" box="[337,462,1361,1383]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="15" value="10.253056">10°15'11''E</geoCoordinate>
;
<elevation id="0069222832227E70FE26FABA6F5EFAF1" box="[474,579,1361,1383]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="53" pageNumber="54" unit="m" value="120.0">
<quantity id="4CBC68FE32227E70FE26FABA6F01FAF1" box="[474,540,1361,1383]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="53" pageNumber="54" unit="m" value="120.0">120 m</quantity>
asl
</elevation>
:
</emphasis>
<collectingRegion id="49800BF932227E70FDB2FABA6E02FAF1" box="[590,799,1361,1383]" country="Tunisia" name="Bizerte" pageId="53" pageNumber="54">Bizerte Governorate</collectingRegion>
,
<location id="8E9B93C032227E70FCD5FABA6E66FAF1" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FCD5FABA6E66FAF1" box="[809,891,1361,1383]" latitude="37.179447" longLatPrecision="19" longitude="10.253056" name="Raf Raf" pageId="53" pageNumber="54" stateProvince="Bizerte Governorate">Raf Raf</location>
, forest, moss on soil.
<bibRefCitation id="EFD5B8EA32227E70FBAAFABA6826FAF1" box="[1110,1339,1361,1383]" pageId="53" pageNumber="54" refString="Gasiorek, P., Morek, W., Stec, D., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688" type="book">
<emphasis id="B930190932227E70FBAAFABA6826FAF1" bold="true" box="[1110,1339,1361,1383]" pageId="53" pageNumber="54">
Gąsiorek
<emphasis id="B930190932227E70FB43FAB969ECFAF1" bold="true" box="[1215,1265,1361,1383]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</emphasis>
</bibRefCitation>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BFA9A682EFA10" blockId="53.[151,1436,1079,2037]" box="[151,1331,1392,1414]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BFA9A682EFA10" ID-GBIF-Occurrence="1671737422" box="[151,1331,1392,1414]" elevation="67" latitude="36.89917" location="Gammarth" longLatPrecision="19" longitude="10.305278" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Tunis Governorate">
•
<emphasis id="B930190932227E70FF3BFA9B6F20FA10" bold="true" box="[199,573,1392,1414]" pageId="53" pageNumber="54">
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,
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Gąsiorek
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Gąsiorek
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Gąsiorek
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M'tir, forest, moss on soil.
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Gąsiorek
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M'tir, forest, moss on soil.
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Gąsiorek
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Gąsiorek
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asl
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:
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M'tir, forest, moss on soil.
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<emphasis id="B930190932227E70FB7AF9AD6877F9CA" bold="true" box="[1158,1386,1606,1628]" pageId="53" pageNumber="54">
Gąsiorek
<emphasis id="B930190932227E70FB12F9AC683DF9CA" bold="true" box="[1262,1312,1606,1628]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</emphasis>
</bibRefCitation>
</materialsCitation>
</paragraph>
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•
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<geoCoordinate id="EE70A3DC32227E70FF3BF98E6C58F9ED" box="[199,325,1637,1659]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="15" value="36.72222">36°43'20''N</geoCoordinate>
,
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;
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asl
</elevation>
:
</emphasis>
<collectingRegion id="49800BF932227E70FDACF98E6E2AF9ED" box="[592,823,1637,1659]" country="Tunisia" name="Jendouba" pageId="53" pageNumber="54">Jendouba Governorate</collectingRegion>
,
<location id="8E9B93C032227E70FCBDF98E6E6CF9ED" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FCBDF98E6E6CF9ED" box="[833,881,1637,1659]" latitude="36.72222" longLatPrecision="19" longitude="8.682778" name="Beni" pageId="53" pageNumber="54" stateProvince="Jendouba Governorate">Beni</location>
M'tir, forest, moss on soil.
<bibRefCitation id="EFD5B8EA32227E70FB7AF98E6877F9ED" box="[1158,1386,1637,1659]" pageId="53" pageNumber="54" refString="Gasiorek, P., Morek, W., Stec, D., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688" type="book">
<emphasis id="B930190932227E70FB7AF98E6877F9ED" bold="true" box="[1158,1386,1637,1659]" pageId="53" pageNumber="54">
Gąsiorek
<emphasis id="B930190932227E70FB12F98D683DF9ED" bold="true" box="[1262,1312,1637,1659]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</emphasis>
</bibRefCitation>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BF96E6877F90C" blockId="53.[151,1436,1079,2037]" box="[151,1386,1668,1690]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BF96E6877F90C" ID-GBIF-Occurrence="1671737460" box="[151,1386,1668,1690]" elevation="474" latitude="36.721947" location="Beni" longLatPrecision="19" longitude="8.724999" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Jendouba Governorate">
•
<emphasis id="B930190932227E70FF3BF96F6F54F90C" bold="true" box="[199,585,1668,1690]" pageId="53" pageNumber="54">
<geoCoordinate id="EE70A3DC32227E70FF3BF96F6C58F90C" box="[199,325,1668,1690]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="15" value="36.721947">36°43'19''N</geoCoordinate>
,
<geoCoordinate id="EE70A3DC32227E70FEADF96F6CCDF90C" box="[337,464,1668,1690]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="15" value="8.724999">08°43'30''E</geoCoordinate>
;
<elevation id="0069222832227E70FE20F96F6F58F90C" box="[476,581,1668,1690]" metricMagnitude="2" metricUnit="m" metricValue="4.74" pageId="53" pageNumber="54" unit="m" value="474.0">
<quantity id="4CBC68FE32227E70FE20F96F6F00F90C" box="[476,541,1668,1690]" metricMagnitude="2" metricUnit="m" metricValue="4.74" pageId="53" pageNumber="54" unit="m" value="474.0">474 m</quantity>
asl
</elevation>
:
</emphasis>
<collectingRegion id="49800BF932227E70FDACF96F6E2AF90C" box="[592,823,1668,1690]" country="Tunisia" name="Jendouba" pageId="53" pageNumber="54">Jendouba Governorate</collectingRegion>
,
<location id="8E9B93C032227E70FCBDF96F6E6CF90C" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FCBDF96F6E6CF90C" box="[833,881,1668,1690]" latitude="36.721947" longLatPrecision="19" longitude="8.724999" name="Beni" pageId="53" pageNumber="54" stateProvince="Jendouba Governorate">Beni</location>
M'tir, forest, moss on soil.
<bibRefCitation id="EFD5B8EA32227E70FB7AF96F6877F90C" box="[1158,1386,1668,1690]" pageId="53" pageNumber="54" refString="Gasiorek, P., Morek, W., Stec, D., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688" type="book">
<emphasis id="B930190932227E70FB7AF96F6877F90C" bold="true" box="[1158,1386,1668,1690]" pageId="53" pageNumber="54">
Gąsiorek
<emphasis id="B930190932227E70FB12F96F683DF90C" bold="true" box="[1262,1312,1668,1690]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</emphasis>
</bibRefCitation>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BF9486877F92E" blockId="53.[151,1436,1079,2037]" box="[151,1386,1698,1720]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BF9486877F92E" ID-GBIF-Occurrence="1671737477" box="[151,1386,1698,1720]" elevation="371" latitude="36.713333" location="Beni" longLatPrecision="19" longitude="8.724999" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Jendouba Governorate">
•
<emphasis id="B930190932227E70FF3BF9496F54F92E" bold="true" box="[199,585,1698,1720]" pageId="53" pageNumber="54">
<geoCoordinate id="EE70A3DC32227E70FF3BF9496C58F92E" box="[199,325,1698,1720]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="15" value="36.713333">36°42'48''N</geoCoordinate>
,
<geoCoordinate id="EE70A3DC32227E70FEADF9496CCDF92E" box="[337,464,1698,1720]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="15" value="8.724999">08°43'30''E</geoCoordinate>
;
<elevation id="0069222832227E70FE20F9496F58F92E" box="[476,581,1698,1720]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="53" pageNumber="54" unit="m" value="371.0">
<quantity id="4CBC68FE32227E70FE20F9496F00F92E" box="[476,541,1698,1720]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="53" pageNumber="54" unit="m" value="371.0">371 m</quantity>
asl
</elevation>
:
</emphasis>
<collectingRegion id="49800BF932227E70FDACF9496E2AF92E" box="[592,823,1698,1720]" country="Tunisia" name="Jendouba" pageId="53" pageNumber="54">Jendouba Governorate</collectingRegion>
,
<location id="8E9B93C032227E70FCBDF9496E6CF92E" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FCBDF9496E6CF92E" box="[833,881,1698,1720]" latitude="36.713333" longLatPrecision="19" longitude="8.724999" name="Beni" pageId="53" pageNumber="54" stateProvince="Jendouba Governorate">Beni</location>
M'tir, forest, moss on soil.
<bibRefCitation id="EFD5B8EA32227E70FB7AF9496877F92E" box="[1158,1386,1698,1720]" pageId="53" pageNumber="54" refString="Gasiorek, P., Morek, W., Stec, D., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688" type="book">
<emphasis id="B930190932227E70FB7AF9496877F92E" bold="true" box="[1158,1386,1698,1720]" pageId="53" pageNumber="54">
Gąsiorek
<emphasis id="B930190932227E70FB12F948683DF92E" bold="true" box="[1262,1312,1698,1720]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</emphasis>
</bibRefCitation>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BF9296877F941" blockId="53.[151,1436,1079,2037]" box="[151,1386,1729,1751]" pageId="53" pageNumber="54">
<materialsCitation id="3B2CCF4632227E70FF6BF9296877F941" ID-GBIF-Occurrence="1671737491" box="[151,1386,1729,1751]" elevation="462" latitude="36.702778" location="Beni" longLatPrecision="19" longitude="8.679444" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Jendouba Governorate">
•
<emphasis id="B930190932227E70FF3BF92A6F54F941" bold="true" box="[199,585,1729,1751]" pageId="53" pageNumber="54">
<geoCoordinate id="EE70A3DC32227E70FF3BF92A6C58F941" box="[199,325,1729,1751]" direction="north" orientation="latitude" pageId="53" pageNumber="54" precision="15" value="36.702778">36°42'10''N</geoCoordinate>
,
<geoCoordinate id="EE70A3DC32227E70FEADF92A6CCDF941" box="[337,464,1729,1751]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="15" value="8.679444">08°40'46''E</geoCoordinate>
;
<elevation id="0069222832227E70FE20F92A6F58F941" box="[476,581,1729,1751]" metricMagnitude="2" metricUnit="m" metricValue="4.62" pageId="53" pageNumber="54" unit="m" value="462.0">
<quantity id="4CBC68FE32227E70FE20F92A6F00F941" box="[476,541,1729,1751]" metricMagnitude="2" metricUnit="m" metricValue="4.62" pageId="53" pageNumber="54" unit="m" value="462.0">462 m</quantity>
asl
</elevation>
:
</emphasis>
<collectingRegion id="49800BF932227E70FDACF92A6E2AF941" box="[592,823,1729,1751]" country="Tunisia" name="Jendouba" pageId="53" pageNumber="54">Jendouba Governorate</collectingRegion>
,
<location id="8E9B93C032227E70FCBDF92A6E6CF941" LSID="urn:lsid:plazi:treatment:03ED740D32227E73FF6BFBDC6800FF43:8E9B93C032227E70FCBDF92A6E6CF941" box="[833,881,1729,1751]" latitude="36.702778" longLatPrecision="19" longitude="8.679444" name="Beni" pageId="53" pageNumber="54" stateProvince="Jendouba Governorate">Beni</location>
M'tir, forest, moss on soil.
<bibRefCitation id="EFD5B8EA32227E70FB7AF92A6877F941" box="[1158,1386,1729,1751]" pageId="53" pageNumber="54" refString="Gasiorek, P., Morek, W., Stec, D., Marnissi, J. &amp; Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688" type="book">
<emphasis id="B930190932227E70FB7AF92A6877F941" bold="true" box="[1158,1386,1729,1751]" pageId="53" pageNumber="54">
Gąsiorek
<emphasis id="B930190932227E70FB12F929683DF941" bold="true" box="[1262,1312,1729,1751]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2017)
</emphasis>
</bibRefCitation>
</materialsCitation>
</paragraph>
<paragraph id="8BFBC51B32227E70FF6BF9346EBCF96E" blockId="53.[151,1436,1079,2037]" box="[151,929,1759,1784]" pageId="53" pageNumber="54">
<emphasis id="B930190932227E70FF6BF9346C7BF96E" bold="true" box="[151,358,1759,1784]" pageId="53" pageNumber="54">Record numbers.</emphasis>
<collectingCountry id="F353858B32227E70FE90F9346CD8F96E" box="[364,453,1759,1784]" name="Algeria" pageId="53" pageNumber="54">Algeria</collectingCountry>
: 1,
<collectingCountry id="F353858B32227E70FE17F9346F9EF96E" box="[491,643,1759,1784]" name="Mozambique" pageId="53" pageNumber="54">Mozambique</collectingCountry>
: 1,
<collectingCountry id="F353858B32227E70FD56F9346E1FF96E" box="[682,770,1759,1784]" name="Tunisia" pageId="53" pageNumber="54">Tunisia</collectingCountry>
: 13;
<emphasis id="B930190932227E70FCCAF9346EBCF96E" bold="true" box="[822,929,1759,1784]" pageId="53" pageNumber="54">total: 15.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35E969032227E73FF6BF8EF6800FF43" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="discussion">
<paragraph id="8BFBC51B32227E73FF6BF8EF6800FF43" blockId="53.[151,1436,1079,2037]" lastBlockId="54.[199,1436,151,213]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
<emphasis id="B930190932227E70FF6BF8EF6C16F88B" bold="true" box="[151,267,1796,1821]" pageId="53" pageNumber="54">Remarks.</emphasis>
The genus
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<emphasis id="B930190932227E70FE6CF8EF6F4EF88B" box="[400,595,1796,1821]" italics="true" pageId="53" pageNumber="54">Paramacrobiotus</emphasis>
</taxonomicName>
(until recently a species-group within
<taxonomicName id="4C44BE9832227E70FBF6F8EF6985F88B" authorityName="Radiatus Pilato, Binda &amp; Catanzaro" authorityYear="1991" box="[1034,1176,1796,1821]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="genus">
<emphasis id="B930190932227E70FBF6F8EF6985F88B" box="[1034,1176,1796,1821]" italics="true" pageId="53" pageNumber="54">Macrobiotus</emphasis>
</taxonomicName>
) has been divided into
<taxonomicName id="4C44BE9832227E70FF3BF8CC6C3BF8D6" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1911" box="[199,294,1831,1856]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" species="richtersi">
<emphasis id="B930190932227E70FF3BF8CC6C3BF8D6" box="[199,294,1831,1856]" italics="true" pageId="53" pageNumber="54">richtersi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C44BE9832227E70FE90F8CC6CC8F8D6" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" box="[364,469,1831,1856]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="B930190932227E70FE90F8CC6CC8F8D6" box="[364,469,1831,1856]" italics="true" pageId="53" pageNumber="54">areolatus</emphasis>
</taxonomicName>
(with and without macroplacoid, respectively) species groups.
<taxonomicName id="4C44BE9832227E70FB25F8CC6C2DF8F3" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="B930190932227E70FB25F8CC6C2DF8F3" italics="true" pageId="53" pageNumber="54">Paramacrobiotus areolatus</emphasis>
</taxonomicName>
, once considered cosmopolitan (
<bibRefCitation id="EFD5B8EA32227E70FD4CF8A76E46F8F2" author="McInnes" box="[688,859,1868,1892]" pageId="53" pageNumber="54" refString="McInnes, S. J. (1994) Zoogeographic distribution of terrestrial / freshwater tardigrades from current literature. JouRnal of NatuRal HistoRy, 28, 257 - 352. https: // doi. org / 10.1080 / 00222939400770131" type="journal article" year="1994">McInnes 1994</bibRefCitation>
), is now recognised as the nominal species for a group of very similar taxa that require careful taxonomic examination of adults and egg morphology for correct identification. A few new (including African) species of the
<taxonomicName id="4C44BE9832227E70FC09F87F6943F83B" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" box="[1013,1118,1940,1965]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="53" pageNumber="54" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="B930190932227E70FC09F87F6943F83B" box="[1013,1118,1940,1965]" italics="true" pageId="53" pageNumber="54">areolatus</emphasis>
</taxonomicName>
group (e.g.
<bibRefCitation id="EFD5B8EA32227E70FB0AF87F6C8AF846" author="Michalczyk" pageId="53" pageNumber="54" refString="Michalczyk, L. &amp; Kaczmarek, L. (2006 b) MacRobiotus huzioRi, a new species of Tardigrada (Eutardigrada: Macrobiotidae) from Costa Rica (Central America). Zootaxa, 1169, 47 - 59." type="journal article" year="2006" yearSuffix="b">Michalczyk &amp; Kaczmarek 2006b</bibRefCitation>
,
<bibRefCitation id="EFD5B8EA32227E70FE5FF8536F6AF846" author="Degma" box="[419,631,1975,2000]" pageId="53" pageNumber="54" refString="Degma, P., Michalczyk, L. &amp; Kaczmarek, L. (2008) MacRobiotus deRkai, a new species of Tardigrada (Eutardigrada, Macrobiotidae, huzoRi group) from the Colombian Andes (South America). Zootaxa, 1731, 1 - 23." type="journal article" year="2008">
Degma
<emphasis id="B930190932227E70FE00F8526F28F846" box="[508,565,1975,2000]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2008
</bibRefCitation>
,
<bibRefCitation id="EFD5B8EA32227E70FD78F85C6E8AF846" author="Kaczmarek" box="[644,919,1975,2000]" pageId="53" pageNumber="54" refString="Kaczmarek, L., Cytan, J., Zawierucha, K., Diduszko, D. &amp; Michalczyk, L. (2014 b) Tardigrades from Peru (South America), with descriptions of three new species of Parachela. Zootaxa, 3790 (2), 357 - 379. https: // doi. org / 10.11646 / zootaxa. 3790.2.5" type="journal article" year="2014" yearSuffix="b">
Kaczmarek
<emphasis id="B930190932227E70FCF7F8526E59F846" box="[779,836,1975,2000]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2014b
</bibRefCitation>
). Both
<bibRefCitation id="EFD5B8EA32227E70FC15F85C6984F846" author="Richters" box="[1001,1177,1975,2000]" pageId="53" pageNumber="54" refString="Richters, F. (1908) Moosbewohner. Wissenschaftliche ERgebnisse deR Schwedischen SudpolaR-Expedition (1901 - 1903), Stockholm, 6, 1 - 16." type="journal article" year="1908">Richters (1908)</bibRefCitation>
and Binda (1984) predate the taxonomic changes and failed to find eggs, essential for correct species identification. This, in conjunction with
<taxonomicName id="4C44BE9832217E73FE77FF726F12FF26" authorityName="Kaczmarek" authorityYear="2017" baseAuthorityName="Murray" baseAuthorityYear="1907" box="[395,527,151,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="54" pageNumber="55" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="B930190932217E73FE77FF726F12FF26" box="[395,527,151,176]" italics="true" pageId="54" pageNumber="55">P. areolatus</emphasis>
</taxonomicName>
being originally described from the Arctic, implies all African records should be treated with caution until relevant examples are re-examined or new African material is confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>
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