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<document id="EBC6216C9FC71335AB1C852B6D6815AA" ID-DOI="10.11646/zootaxa.4816.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954419" ID-ZooBank="02EAEA41-160D-41FF-95E5-8F83E4A2BAD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595339832409" checkinUser="plazi" docAuthor="Johanson, Kjell Arne, Pham, Thai Hong, Malm, Tobias &amp; Sjöberg, Tin" docDate="2020" docId="03E74143FFEE0A05FF44320C73F6FE00" docLanguage="en" docName="zootaxa.4816.3.2.pdf" docOrigin="Zootaxa 4816 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oecetis porntipae Johanson &amp; Pham &amp; Malm &amp; Sjöberg 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="318" masterDocId="FFDE393BFFE80A02FFD336697039FFD3" masterDocTitle="Description of six new species of Oecetis (Trichoptera, Leptoceridae) from Vietnam" masterLastPageNumber="324" masterPageNumber="311" pageNumber="317" updateTime="1698783012615" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="AD3570043581FBE1177E69CA1CFF8738" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6D3CD80A86A30681C954BFD264BEF06">
<mods:title id="FD3A1EE4E322FD58F055AAAC4B813BCE">Description of six new species of Oecetis (Trichoptera, Leptoceridae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="F580B1B08386B5F0CA419A71151DFD5D" type="personal">
<mods:role id="CC6B3017AB20F8D2AF837ADBD66EABA0">
<mods:roleTerm id="EC0925B43DCBC76CA20174567A035CEB">Author</mods:roleTerm>
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<mods:namePart id="0E3828430FC12AD3D0BD39863FCB32A0">Johanson, Kjell Arne</mods:namePart>
<mods:affiliation id="5410D835BF6FDB5834E16AFB45EAE161">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden &amp; kjell. arne. johanson @ nrm. se; https: // orcid. org / 0000 - 0002 - 1893 - 3429</mods:affiliation>
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<mods:roleTerm id="47A571E333E129E285DC9FFADE6DA914">Author</mods:roleTerm>
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<mods:namePart id="295E4450141A545F0A4910D022A900FE">Pham, Thai Hong</mods:namePart>
<mods:affiliation id="685266286BA90B1102B8CEFE8C7578DC">Vietnam National Museum of Nature (VNMN) &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet St, Cau Giay, Hanoi, Vietnam. &amp; Faculty of Biology, Hanoi National University of Education, 136 Xuan Thuy, Cau Giay, Hanoi, Vietnam. &amp; phamthai @ vnmn. vast. vn; https: // orcid. org / 0000 - 0002 - 4763 - 3679</mods:affiliation>
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<mods:roleTerm id="1A6B3BD69701276B9AFFA66DB196D0A1">Author</mods:roleTerm>
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<mods:namePart id="5E9E975697C62089C03814C65CCE1194">Malm, Tobias</mods:namePart>
<mods:affiliation id="8F5DCE3474067EAFB37CE88B14BEAA1F">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden &amp; tobias. malm @ nrm. se; https: // orcid. org / 0000 - 0003 - 4610 - 4086</mods:affiliation>
</mods:name>
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<mods:roleTerm id="FD18A4C4CC7B6869CE8ED70A43A6672C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E9ADEE3AC315DC566C150C5C3D36B7B2">Sjöberg, Tin</mods:namePart>
<mods:affiliation id="3965451476BEE25A08A7DB850CCBD4F4">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden &amp; tin. sjoberg @ nrm. se; https: // orcid. org / 0000 - 0002 - 2813 - 9325</mods:affiliation>
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<mods:typeOfResource id="22EEC26544E399B9B3E0E5610296BD6C">text</mods:typeOfResource>
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<mods:title id="47279313FE1A9DD41FA95904E4C817D5">Zootaxa</mods:title>
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<mods:part id="24D83D38573D6E84C8BB4DB65BA80FAF">
<mods:date id="191880317735F1464DBED161A584AB85">2020</mods:date>
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<mods:number id="19DA4EEF16C44C2E3AF6FCCD2A4FCEDD">2020-07-17</mods:number>
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<mods:detail id="5E9F242FA2EA3024B8B51B33C27BDC37" type="volume">
<mods:number id="5B2607E1C7BC963909FFFE8AD0D2C9A8">4816</mods:number>
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<mods:classification id="BF67A43A70982D6EE17669152642E50F">journal article</mods:classification>
<mods:identifier id="3EB6EDFC0BA1C05DACB0F6F88B7DBC2F" type="DOI">10.11646/zootaxa.4816.3.2</mods:identifier>
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<treatment id="03E74143FFEE0A05FF44320C73F6FE00" LSID="urn:lsid:plazi:treatment:03E74143FFEE0A05FF44320C73F6FE00" httpUri="http://treatment.plazi.org/id/03E74143FFEE0A05FF44320C73F6FE00" lastPageId="7" lastPageNumber="318" pageId="6" pageNumber="317">
<subSubSection id="C354A3DEFFEE0A04FF44320C71F6FBAC" box="[151,463,1125,1151]" pageId="6" pageNumber="317" type="nomenclature">
<paragraph id="8BF1F055FFEE0A04FF44320C71F6FBAC" blockId="6.[151,463,1125,1187]" box="[151,463,1125,1151]" pageId="6" pageNumber="317">
<heading id="D0B94739FFEE0A04FF44320C71F6FBAC" bold="true" box="[151,463,1125,1151]" fontSize="11" level="1" pageId="6" pageNumber="317" reason="1">
<emphasis id="B93A2C47FFEE0A04FF44320C71F6FBAC" bold="true" box="[151,463,1125,1151]" pageId="6" pageNumber="317">
<taxonomicName id="4C4E8BD6FFEE0A04FF44320C7153FBAC" authority="Johanson &amp; Pham &amp; Malm &amp; Sjöberg, 2020" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[151,362,1125,1151]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="porntipae" status="sp. nov.">
<emphasis id="B93A2C47FFEE0A04FF44320C7153FBAC" bold="true" box="[151,362,1125,1151]" italics="true" pageId="6" pageNumber="317">Oecetis porntipae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209913CFFEE0A04FEA1320C71F6FBAC" box="[370,463,1125,1151]" pageId="6" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFEE0A04FF4432E0712CFB70" box="[151,277,1161,1187]" pageId="6" pageNumber="317" type="description">
<paragraph id="8BF1F055FFEE0A04FF4432E0712CFB70" blockId="6.[151,463,1125,1187]" box="[151,277,1161,1187]" pageId="6" pageNumber="317">
<figureCitation id="1375ECD0FFEE0A04FF4432E0712CFB70" box="[151,277,1161,1187]" captionStart="FIGURES 16–20" captionStartId="7.[151,264,1934,1959]" captionTargetBox="[160,1421,495,1897]" captionTargetId="figure@7.[151,1436,489,1909]" captionTargetPageId="7" captionText="FIGURES 16–20. Oecetis porntipae sp. nov., male holotype, genitalia. 16, left lateral; 17, dorsal; 18, ventral; 19, phallus, left lateral; 20, phallus, ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954427" httpUri="https://zenodo.org/record/3954427/files/figure.png" pageId="6" pageNumber="317">Figs 16–20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFEE0A04FF4432B974D4FA34" pageId="6" pageNumber="317" type="diagnosis">
<paragraph id="8BF1F055FFEE0A04FF4432B974D4FA34" blockId="6.[151,1437,1232,2015]" pageId="6" pageNumber="317">
<emphasis id="B93A2C47FFEE0A04FF4432B9712BFB38" bold="true" box="[151,274,1232,1259]" pageId="6" pageNumber="317">Diagnosis.</emphasis>
The species is distinguished from many of the Oriental and Palearctic species by the absence of honeycomb structure on pregenital tergites. In the genitalia, segment X is divided into a lower part and an upper part. The genitalia resemble those of
<taxonomicName id="4C4E8BD6FFEE0A04FE023370731AFAE0" authority="Malicky 2005" authorityName="Malicky" authorityYear="2005" box="[465,803,1304,1331]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="momos">
<emphasis id="B93A2C47FFEE0A04FE0233707245FAE0" box="[465,636,1305,1331]" italics="true" pageId="6" pageNumber="317">Oecetis momos</emphasis>
Malicky 2005
</taxonomicName>
and
<taxonomicName id="4C4E8BD6FFEE0A04FC8F337074F8FAE0" authority="Kimmins 1963" authorityName="Kimmins" authorityYear="1963" box="[860,1217,1305,1331]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="mekana">
<emphasis id="B93A2C47FFEE0A04FC8F33707429FAE0" box="[860,1040,1305,1331]" italics="true" pageId="6" pageNumber="317">Oecetis mekana</emphasis>
<bibRefCitation id="EFDF8DA4FFEE0A04FBCB337074F8FAE0" author="Kimmins, D. E." box="[1048,1217,1305,1331]" pageId="6" pageNumber="317" pagination="263 - 316" refId="ref6668" refString="Kimmins, D. E. (1963) On the Leptocerinae of the Indian sub-continent and north east Burma (Trichoptera). Bulletin of the British Museum (Natural History), Entomology, 14 (6), 263 - 316. https: // doi. org / 10.5962 / bhl. part. 8784" type="journal article" year="1963">
Kimmins
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in the shape of the superior appendages, coxopodites, and tergum X.
<taxonomicName id="4C4E8BD6FFEE0A04FD10335473B3FA84" authority="Johanson &amp; Pham &amp; Malm &amp; Sjöberg, 2020" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[707,906,1341,1367]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="porntipae" status="sp. nov.">
<emphasis id="B93A2C47FFEE0A04FD10335473B3FA84" box="[707,906,1341,1367]" italics="true" pageId="6" pageNumber="317">Oecetis porntipae</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A209913CFFEE0A04FC43335473D0FA84" box="[912,1001,1341,1367]" pageId="6" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="4C4E8BD6FFEE0A04FB3233547568FA84" authorityName="Malicky" authorityYear="2005" box="[1249,1361,1341,1367]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="momos">
<emphasis id="B93A2C47FFEE0A04FB32335474CCFA84" box="[1249,1269,1341,1367]" italics="true" pageId="6" pageNumber="317">O</emphasis>
.
<emphasis id="B93A2C47FFEE0A04FAD133577568FA84" box="[1282,1361,1342,1367]" italics="true" pageId="6" pageNumber="317">momos</emphasis>
</taxonomicName>
by the broader and more symmetrically narrowing coxopodites in lateral view, the larger and more pointed cone shaped posterior dorsolateral process of segment IX, and the shorter superior appendages. It is separated from
<taxonomicName id="4C4E8BD6FFEE0A04FAD033EC7543FA4C" authorityName="Kimmins" authorityYear="1963" box="[1283,1402,1413,1439]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="317" phylum="Arthropoda" rank="species" species="mekana">
<emphasis id="B93A2C47FFEE0A04FAD033EC752EFA4C" box="[1283,1303,1413,1439]" italics="true" pageId="6" pageNumber="317">O</emphasis>
.
<emphasis id="B93A2C47FFEE0A04FAF033EC7543FA4C" box="[1315,1402,1413,1439]" italics="true" pageId="6" pageNumber="317">mekana</emphasis>
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by the presence of a complete longitudinal apodeme on the ventrolateral part of each side of segment IX, the basodorsal lobe on each gonocoxite, and a prominent lower posterolateral process on each side of segment IX.
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFEE0A05FF1433997188FE5F" lastPageId="7" lastPageNumber="318" pageId="6" pageNumber="317" type="description">
<paragraph id="8BF1F055FFEE0A05FF14339970CFFECC" blockId="6.[151,1437,1232,2015]" lastBlockId="7.[151,1437,153,467]" lastPageId="7" lastPageNumber="318" pageId="6" pageNumber="317">
<emphasis id="B93A2C47FFEE0A04FF1433997199F9D8" bold="true" box="[199,416,1520,1547]" pageId="6" pageNumber="317">Description, male.</emphasis>
Length of each forewing
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(N = 1). Genitalia (
<figureCitation id="1375ECD0FFEE0A04FC2A33987440F9DF" box="[1017,1145,1521,1548]" captionStart="FIGURES 16–20" captionStartId="7.[151,264,1934,1959]" captionTargetBox="[160,1421,495,1897]" captionTargetId="figure@7.[151,1436,489,1909]" captionTargetPageId="7" captionText="FIGURES 16–20. Oecetis porntipae sp. nov., male holotype, genitalia. 16, left lateral; 17, dorsal; 18, ventral; 19, phallus, left lateral; 20, phallus, ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954427" httpUri="https://zenodo.org/record/3954427/files/figure.png" pageId="6" pageNumber="317">Figs 16–20</figureCitation>
) slightly longer than high in lateral view. Segment IX oriented almost vertically, anterior margin slightly convex from dorsum to venter; dorsal part strongly narrowing posteriorly, curving slightly posterad; acrotergite of segment IX subtriangular, apically blunt, about as wide as each superior appendage in dorsal view; dorsal process of segment IX minute, forming pair of semimembranous papillae; sternum with large semimembranous area covering bases of coxopodites in ventral view; lower posterolateral process on each side of segment IX large, rhomboid; each upper posterolateral process of segment IX broadly cone shaped, about three times longer than superior appendages; longitudinal lateral apodeme present on each side ventrolaterally on segment IX, running from anterior margin to ventral corner of lower posterolateral process of segment IX. Tergum X much longer than superior appendages; in lateral view dorsal margins of upper part and lower part almost straight, ventral margin below upper part expanded into pair of membranous lobes, upper part tuboid; in dorsal view lower part with wide base, upper part almost parallel-sided along its length, with apex narrowly rounded. Superior appendages very short, broadly rounded in dorsal and lateral views, without membranous areas. Each gonocoxite in lateral view with broad base tapering to blunt apex, curving slightly dorsad after mid-length, about two times as long as tergum X, unbranched, with basidorsal lobe scarcely evident before mid-length, dorsal margin slightly undulating, ventral margin concave at mid-length; in ventral view narrowing sharply before mid-length; curving mesad beyond base; without large setae. Phallus almost as long as coxopodites, with about five small sclerites pointing posterad; phallobase with lateral process along right side hooked mesad apically.
</paragraph>
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<emphasis id="B93A2C47FFEF0A05FF143740713DFE90" bold="true" box="[199,260,297,323]" pageId="7" pageNumber="318">Male</emphasis>
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<typeStatus id="54F54EF7FFEF0A05FEDD37407140FE90" box="[270,377,297,323]" pageId="7" pageNumber="318" type="holotype">holotype</typeStatus>
:
<collectingCountry id="F359B0C5FFEF0A05FE5A374171CAFE90" box="[393,499,296,323]" name="Vietnam" pageId="7" pageNumber="318">Vietnam</collectingCountry>
:
<collectingRegion id="498A3EB7FFEF0A05FDD1374072FDFE90" box="[514,708,296,323]" country="Vietnam" name="Dong Nai" pageId="7" pageNumber="318">Dong Nai Prov.</collectingRegion>
:
</emphasis>
<collectingCounty id="629088D9FFEF0A05FD003740739DFE90" box="[723,932,297,323]" pageId="7" pageNumber="318">Vinh Cuu District</collectingCounty>
,
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,
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stream,
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,
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,
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,
<collectingMethod id="520F8842FFEF0A05FDEC3724729CFEB4" box="[575,677,333,359]" pageId="7" pageNumber="318">light trap</collectingMethod>
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<collectingDate id="EFB42F7DFFEF0A05FD7F3724731AFEB4" box="[684,803,333,359]" pageId="7" pageNumber="318" value="2011-04-16">16.iv.2011</collectingDate>
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, loc#VN015, leg.
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&amp;
<collectorName id="26BB9583FFEF0A05FB7B372474CCFEB4" box="[1192,1269,333,359]" pageId="7" pageNumber="318">TT Du</collectorName>
, DNA voucher JG5 (
<collectionCode id="ED5F6890FFEF0A05FF0037187113FE58" box="[211,298,369,395]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="7" pageNumber="318">VNMN</collectionCode>
in alcohol)
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.
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFEF0A05FF1437FC73F6FE00" pageId="7" pageNumber="318" type="etymology">
<paragraph id="8BF1F055FFEF0A05FF1437FC73F6FE00" blockId="7.[151,1437,153,467]" pageId="7" pageNumber="318">
<emphasis id="B93A2C47FFEF0A05FF1437FC7177FE7C" bold="true" box="[199,334,405,431]" pageId="7" pageNumber="318">Etymology.</emphasis>
<emphasis id="B93A2C47FFEF0A05FE8737FF71FDFE7C" box="[340,452,406,431]" italics="true" pageId="7" pageNumber="318">Porntipae</emphasis>
, named for the trichopterologist Porntip Chantaramongkol, in recognition of her positive impact on the knowledge of the caddisfly diversity in the Oriental region.
</paragraph>
</subSubSection>
</treatment>
</document>
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