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<document id="5125D8C28C7C5982FD46E7238D1C67C0" ID-DOI="10.11646/zootaxa.5323.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8204101" ID-ZooBank="0ABC4CFF-9D2A-449A-B7C2-CFE16DF16004" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690887012857" checkinUser="plazi" docAuthor="Soares, Matheus M. M., Runyon, Justin B., Capellari, Renato S. &amp; Ale-Rocha, Rosaly" docDate="2023" docId="03E62055FF930015FF7BF8B2FA463182" docLanguage="en" docName="zootaxa.5323.2.1.pdf" docOrigin="Zootaxa 5323 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5323.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraclius arcuatus" docType="key" docVersion="4" lastPageNumber="154" masterDocId="FFDF582DFF910016FFECFFBDFFDB3247" masterDocTitle="Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species" masterLastPageNumber="182" masterPageNumber="151" pageNumber="153" updateTime="1691081561564" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FBAB0DB903F4AB578EAE8F6647B06AD6">Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species</mods:title>
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<mods:namePart id="60DA99C3CAF6798BC9FBFC475B62D87A">Soares, Matheus M. M.</mods:namePart>
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<mods:namePart id="91B9103B9AFB1C11DAD9F41E9E0A615B">Runyon, Justin B.</mods:namePart>
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<mods:affiliation id="ACB879C6CCCD58DBB536BD253468CA76">Rocky Mountain Research Station, USDA Forest Service, 1648 S. 7 th Avenue, Bozeman, Montana 59717, USA. &amp; justin. runyon @ usda. gov; https: // orcid. org / 0000 - 0002 - 0271 - 0511</mods:affiliation>
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<mods:namePart id="15145A858B336EDFF78B7D30EB9070B7">Capellari, Renato S.</mods:namePart>
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<mods:affiliation id="77C7B816C0D7576BCF42A12D0916A843">Instituto Federal do Triângulo Mineiro, Campus Uberaba, Rua João Batista Ribeiro 4000, Distrito Industrial II, Uberaba, Minas Gerais 38064 - 790, Brazil. &amp; rscapellari @ gmail. com; https: // orcid. org / 0000 - 0002 - 8410 - 9235</mods:affiliation>
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<mods:affiliation id="A3FEB1EAADB1CD01503954125D29575B">Coordenação de Biodiversidade (COBIO, INPA), Av. André Araújo, 2936, Petrópolis, CEP 69067 - 375, Manaus, Amazonas, Brazil. &amp; alerocha @ inpa. gov. br; https: // orcid. org / 0000 - 0001 - 9874 - 9770</mods:affiliation>
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Key to species of the
<taxonomicName id="4C4FEAC0FF930014FE71F8B2FDDC356E" ID-CoL="6TNMC" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[413,519,1807,1833]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FE71F8B2FDDC356E" bold="true" box="[413,519,1807,1833]" italics="true" pageId="2" pageNumber="153">arcuatus</emphasis>
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-group of
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<emphasis id="B93B4D51FF930014FD6EF8B3FD2D356E" bold="true" box="[642,758,1806,1833]" italics="true" pageId="2" pageNumber="153">Paraclius</emphasis>
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(males)
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<emphasis id="B93B4D51FF930014FF7BF8EEFF78352F" bold="true" box="[151,163,1875,1896]" pageId="2" pageNumber="153">1</emphasis>
Antenna wholly dark brown to black (
<emphasis id="B93B4D51FF930014FDBEF8E9FDAE352E" box="[594,629,1876,1897]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FD97F8E9FD1C352E" box="[635,711,1876,1898]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="2" pageNumber="153">Figs 1E</figureCitation>
,
<figureCitation id="13748DC6FF930014FD3EF8E9FD35352E" box="[722,750,1876,1897]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">2E</figureCitation>
,
<figureCitation id="13748DC6FF930014FD15F8E9FCCC352E" box="[761,791,1876,1897]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">8E</figureCitation>
); face gradually narrowing below (funnel-shaped), narrowest point slightly wider than mid ocellus (
<emphasis id="B93B4D51FF930014FDFAF8CFFDE235C0" box="[534,569,1906,1927]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FDD2F8CCFD5035C1" box="[574,651,1905,1927]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="2" pageNumber="153">Figs 1D</figureCitation>
,
<figureCitation id="13748DC6FF930014FD79F8CCFD6835C1" box="[661,691,1905,1926]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">2C</figureCitation>
,
<figureCitation id="13748DC6FF930014FD50F8CCFD0135C1" box="[700,730,1905,1926]" captionStart="FIGURE 7" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[158,1430,771,1915]" captionTargetId="figure-209@11.[158,1430,771,1915]" captionTargetPageId="11" captionText="FIGURE 7. Paraclius angustipennis Van Duzee, 1929, male holotype (NMNH). AB. Habitus, lateral and dorsal views, respectively; C. Abdominal segments 5 and 6; D. Labels; EF. Head, anterior and anterolateral views, respectively. Images B, C, E originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204125" httpUri="https://zenodo.org/record/8204125/files/figure.png" pageId="2" pageNumber="153">7E</figureCitation>
); clypeus short, about 1/4 as long as face (
<emphasis id="B93B4D51FF930014FB87F8CFFB5535C0" box="[1131,1166,1906,1927]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FB7FF8CCFB3A35C1" box="[1171,1249,1905,1927]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="2" pageNumber="153">Figs 1D</figureCitation>
,
<figureCitation id="13748DC6FF930014FB07F8CCFAD335C1" box="[1259,1288,1905,1926]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">2C</figureCitation>
,
<figureCitation id="13748DC6FF930014FAFEF8CCFAF435C1" box="[1298,1327,1905,1926]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">8C</figureCitation>
,
<figureCitation id="13748DC6FF930014FAD5F8CCFAB835C1" box="[1337,1379,1905,1926]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="2" pageNumber="153">11C</figureCitation>
); anal lobe barely developed (
<emphasis id="B93B4D51FF930014FE29F832FE3335E3" box="[453,488,1935,1956]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FE02F833FDE335E4" box="[494,568,1934,1956]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="2" pageNumber="153">Figs 2F</figureCitation>
,
<figureCitation id="13748DC6FF930014FDAEF833FDBC35E4" box="[578,615,1934,1955]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="2" pageNumber="153">11F</figureCitation>
,
<figureCitation id="13748DC6FF930014FD9DF833FD5635E4" box="[625,653,1934,1955]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">8F</figureCitation>
); tergite 6 either with short or long and strong lateral setae (
<emphasis id="B93B4D51FF930014FB3EF832FB2E35E3" box="[1234,1269,1935,1956]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FB17F833FA9135E4" box="[1275,1354,1934,1956]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="2" pageNumber="153">Figs 1H</figureCitation>
,
<figureCitation id="13748DC6FF930014FAB9F833FAAF35E4" box="[1365,1396,1934,1955]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="2" pageNumber="153">8D</figureCitation>
,
<figureCitation id="13748DC6FF930014FA93F833FA4335E4" box="[1407,1432,1934,1955]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9F</figureCitation>
,
<figureCitation id="13748DC6FF930014FF0EF811FED13586" box="[226,266,1964,1985]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="2" pageNumber="153">11E</figureCitation>
); apicoventral epandrial lobe not developed (
<emphasis id="B93B4D51FF930014FD51F811FD3B3586" box="[701,736,1964,1985]" italics="true" pageId="2" pageNumber="153">e.g.</emphasis>
<figureCitation id="13748DC6FF930014FD09F811FCEF3586" box="[741,820,1964,1986]" captionStart="FIGURE 3" captionStartId="6.[151,250,1458,1482]" captionTargetBox="[151,1436,181,1435]" captionTargetId="figure-20@6.[151,1436,181,1435]" captionTargetPageId="6" captionText="FIGURE 3. Paraclius arcuatus (Loew, 1861), identified male specimens from Honduras (CNC). A. Abdomen, ventral view; B. Tergite 7 and Sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Internal appendages, left lateral view; E. Hypandrium, ventral view; F. Dorsal lobe of surstylus, outer view; G. Ventral lobe of surstylus, outer view. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct br = proctiger brush; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204111" httpUri="https://zenodo.org/record/8204111/files/figure.png" pageId="2" pageNumber="153">Figs 3G</figureCitation>
,
<figureCitation id="13748DC6FF930014FCD2F811FC803586" box="[830,859,1964,1985]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="2" pageNumber="153">9E</figureCitation>
,
<figureCitation id="13748DC6FF930014FC89F811FC543586" box="[869,911,1964,1985]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="2" pageNumber="153">12D</figureCitation>
)...................
<taxonomicName id="4C4FEAC0FF930014FB96F811FB163586" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[1146,1229,1964,1985]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF930014FB96F811FB163586" box="[1146,1229,1964,1985]" italics="true" pageId="2" pageNumber="153">arcuatus</emphasis>
</taxonomicName>
-group of
<taxonomicName id="4C4FEAC0FF930014FAC1F811FA513586" authorityName="Loew" authorityYear="1864" box="[1325,1418,1964,1985]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B93B4D51FF930014FAC1F811FA513586" box="[1325,1418,1964,1985]" italics="true" pageId="2" pageNumber="153">Paraclius</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF930014FA7CF816FA473587" bold="true" box="[1424,1436,1963,1984]" pageId="2" pageNumber="153">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF09143FF930015FF7BF874FA47328D" blockId="2.[151,1437,1875,2015]" lastBlockId="3.[151,1437,151,965]" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153">
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- Antenna usually orangish-yellow, with only apical 1/2 to apical 1/3 of postpedicel brown to dark brown; face various, distinctly wider than mid ocellus at narrowest point; clypeus usually 1/2 to 1/3 as long as face; anal lobe normally developed; tergite 6 bare; apicoventral epandrial developed................................................... other
<taxonomicName id="4C4FEAC0FF920015FB14FF08FA8E328D" authorityName="Loew" authorityYear="1864" box="[1272,1365,181,202]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B93B4D51FF920015FB14FF08FA8E328D" box="[1272,1365,181,202]" italics="true" pageId="3" pageNumber="154">Paraclius</emphasis>
</taxonomicName>
groups
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BB89E6FF920015FF7BFF6FFA473378" pageId="3" pageNumber="154">
<paragraph id="8BF09143FF920015FF7BFF6FFA473343" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFF6FFA473343" pageId="3" pageNumber="154">
<emphasis id="B93B4D51FF920015FF7BFF6FFF7832A0" bold="true" box="[151,163,210,231]" pageId="3" pageNumber="154">2</emphasis>
Postpedicel subrectangular, pointed dorsally at apex (
<figureCitation id="13748DC6FF920015FD33FF6FFCF732A0" box="[735,812,210,232]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Figs 1E</figureCitation>
,
<figureCitation id="13748DC6FF920015FCDAFF6FFC8932A0" box="[822,850,210,231]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="3" pageNumber="154">2E</figureCitation>
,
<figureCitation id="13748DC6FF920015FCB0FF6FFC5C32A0" box="[860,903,210,231]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">11D</figureCitation>
); femur I wholly yellow (
<figureCitation id="13748DC6FF920015FB97FF6FFB1132A0" box="[1147,1226,210,232]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Figs 1A</figureCitation>
,
<figureCitation id="13748DC6FF920015FB38FF6FFB2F32A0" box="[1236,1268,210,231]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="3" pageNumber="154">2A</figureCitation>
,
<figureCitation id="13748DC6FF920015FB11FF6FFAC732AF" box="[1277,1308,210,232]" captionStart="FIGURE 6" captionStartId="10.[151,250,1628,1652]" captionTargetBox="[151,1436,1029,1604]" captionTargetId="figure-375@10.[151,1436,1029,1604]" captionTargetPageId="10" captionText="FIGURE 6. Paraclius femoratus Aldrich, 1901, female syntype (NMNH). AB. Habitus, lateral and dorsal views, respectively. C. Scutellum, dorsal view (arrow pointing to the apical pair of scutellar setae); D. Labels. Images B and C originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204119" httpUri="https://zenodo.org/record/8204119/files/figure.png" pageId="3" pageNumber="154">6A</figureCitation>
,
<figureCitation id="13748DC6FF920015FACAFF6FFA8A32A0" box="[1318,1361,210,231]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">11A</figureCitation>
); CuAx ratio: 0.6 or greater (
<figureCitation id="13748DC6FF920015FE44FF52FE353342" box="[424,494,239,261]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Figs 1I</figureCitation>
,
<figureCitation id="13748DC6FF920015FE14FF4DFDC93342" box="[504,530,240,261]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="3" pageNumber="154">2F</figureCitation>
,
<figureCitation id="13748DC6FF920015FDF0FF4DFD9A3342" box="[540,577,240,261]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">11F</figureCitation>
).....................................................................
<emphasis id="B93B4D51FF920015FA7CFF52FA473343" bold="true" box="[1424,1436,239,260]" pageId="3" pageNumber="154">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF09143FF920015FF7BFEB0FA473378" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFEB0FA473378" pageId="3" pageNumber="154">
<emphasis id="B93B4D51FF920015FF7BFEB0FF443365" bold="true" box="[151,159,269,290]" pageId="3" pageNumber="154">-</emphasis>
Postpedicel various, not pointed dorsally at apex (
<figureCitation id="13748DC6FF920015FD2AFEB0FCCF3365" box="[710,788,269,291]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="3" pageNumber="154">Figs 8E</figureCitation>
,
<figureCitation id="13748DC6FF920015FCF3FEB0FCB83365" box="[799,867,269,291]" captionStart="FIGURE 15" captionStartId="22.[151,250,1709,1733]" captionTargetBox="[151,1436,489,1685]" captionTargetId="figure-152@22.[151,1436,489,1685]" captionTargetPageId="22" captionText="FIGURE 15. Paraclius pumilio Loew, 1872, AE identified male specimens from USA, Texas (CNC), F male from USA, Montana (MTEC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Plumule, lateral view; D. Left antenna, outer view (male from Texas); E. Left wing; F. Right antenna, inner view. Image F originally lacked scale bar." figureDoi="http://doi.org/10.5281/zenodo.8204147" httpUri="https://zenodo.org/record/8204147/files/figure.png" pageId="3" pageNumber="154">15D, F</figureCitation>
,
<figureCitation id="13748DC6FF920015FC81FEB0FC4D3365" box="[877,918,269,290]" captionStart="FIGURE 19" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[177,1411,597,1917]" captionTargetId="figure-128@25.[177,1411,597,1917]" captionTargetPageId="25" captionText="FIGURE 19. Paraclius xibun sp. nov., AE male holotype (CNC), FG male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Hypopygium, lateral view; E. Male abdominal segments 5 and 6; F. Left antenna, inner view; G. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204163" httpUri="https://zenodo.org/record/8204163/files/figure.png" pageId="3" pageNumber="154">19F</figureCitation>
); femur I mostly brown to dark brown (e.g.,
<figureCitation id="13748DC6FF920015FAA5FEB0FA433365" box="[1353,1432,269,291]" captionStart="FIGURE 7" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[158,1430,771,1915]" captionTargetId="figure-209@11.[158,1430,771,1915]" captionTargetPageId="11" captionText="FIGURE 7. Paraclius angustipennis Van Duzee, 1929, male holotype (NMNH). AB. Habitus, lateral and dorsal views, respectively; C. Abdominal segments 5 and 6; D. Labels; EF. Head, anterior and anterolateral views, respectively. Images B, C, E originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204125" httpUri="https://zenodo.org/record/8204125/files/figure.png" pageId="3" pageNumber="154">Figs 7A</figureCitation>
,
<figureCitation id="13748DC6FF920015FF0EFE97FEDA3378" box="[226,257,298,319]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="3" pageNumber="154">8A</figureCitation>
,
<figureCitation id="13748DC6FF920015FEE7FE97FEE33378" box="[267,312,298,319]" captionStart="FIGURE 14" captionStartId="20.[151,250,1211,1235]" captionTargetBox="[151,1436,181,1188]" captionTargetId="figure-20@20.[151,1436,181,1188]" captionTargetPageId="20" captionText="FIGURE 14. Paraclius pumilio Loew, 1872, male syntype (MCZ). AB. Habitus, lateral and dorsal views respectively; C. Head, anterior view; D. Left wing; E. Labels." figureDoi="http://doi.org/10.5281/zenodo.8204145" httpUri="https://zenodo.org/record/8204145/files/figure.png" pageId="3" pageNumber="154">14A</figureCitation>
); CuAx ratio: 0.30.5 (e.g.,
<figureCitation id="13748DC6FF920015FDAAFE97FD4F3378" box="[582,660,298,320]" captionStart="FIGURE 7" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[158,1430,771,1915]" captionTargetId="figure-209@11.[158,1430,771,1915]" captionTargetPageId="11" captionText="FIGURE 7. Paraclius angustipennis Van Duzee, 1929, male holotype (NMNH). AB. Habitus, lateral and dorsal views, respectively; C. Abdominal segments 5 and 6; D. Labels; EF. Head, anterior and anterolateral views, respectively. Images B, C, E originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204125" httpUri="https://zenodo.org/record/8204125/files/figure.png" pageId="3" pageNumber="154">Figs 7A</figureCitation>
,
<figureCitation id="13748DC6FF920015FD72FE97FD633378" box="[670,696,298,319]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="3" pageNumber="154">8F</figureCitation>
,
<figureCitation id="13748DC6FF920015FD2EFE97FD323307" box="[706,745,298,320]" captionStart="FIGURE 15" captionStartId="22.[151,250,1709,1733]" captionTargetBox="[151,1436,489,1685]" captionTargetId="figure-152@22.[151,1436,489,1685]" captionTargetPageId="22" captionText="FIGURE 15. Paraclius pumilio Loew, 1872, AE identified male specimens from USA, Texas (CNC), F male from USA, Montana (MTEC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Plumule, lateral view; D. Left antenna, outer view (male from Texas); E. Left wing; F. Right antenna, inner view. Image F originally lacked scale bar." figureDoi="http://doi.org/10.5281/zenodo.8204147" httpUri="https://zenodo.org/record/8204147/files/figure.png" pageId="3" pageNumber="154">15E</figureCitation>
).......................................................
<emphasis id="B93B4D51FF920015FA7CFE97FA473378" bold="true" box="[1424,1436,298,319]" pageId="3" pageNumber="154">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BB89E6FF920015FF7BFEFAFA4733F3" pageId="3" pageNumber="154">
<paragraph id="8BF09143FF920015FF7BFEFAFA47333C" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFEFAFA47333C" pageId="3" pageNumber="154">
<emphasis id="B93B4D51FF920015FF7BFEFAFF78331B" bold="true" box="[151,163,327,348]" pageId="3" pageNumber="154">3</emphasis>
Scutellum with 1 pair of short apical setae (
<figureCitation id="13748DC6FF920015FD6DFEFAFD16331A" box="[641,717,327,349]" captionStart="FIGURE 6" captionStartId="10.[151,250,1628,1652]" captionTargetBox="[151,1436,1029,1604]" captionTargetId="figure-375@10.[151,1436,1029,1604]" captionTargetPageId="10" captionText="FIGURE 6. Paraclius femoratus Aldrich, 1901, female syntype (NMNH). AB. Habitus, lateral and dorsal views, respectively. C. Scutellum, dorsal view (arrow pointing to the apical pair of scutellar setae); D. Labels. Images B and C originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204119" httpUri="https://zenodo.org/record/8204119/files/figure.png" pageId="3" pageNumber="154">Fig. 6C</figureCitation>
); femur II yellow with apical 1/3 to 1/2 brown to dark brown (
<figureCitation id="13748DC6FF920015FAF3FEFAFAB4331A" box="[1311,1391,327,349]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="3" pageNumber="154">Figs 2A</figureCitation>
,
<figureCitation id="13748DC6FF920015FA95FEF5FA43331A" box="[1401,1432,328,349]" captionStart="FIGURE 4" captionStartId="9.[151,250,582,606]" captionTargetBox="[151,1436,181,558]" captionTargetId="figure-18@9.[151,1436,181,558]" captionTargetPageId="9" captionText="FIGURE 4. Paraclius arcuatus (Loew, 1861), identified female specimen from Mexico (CNC). A. Habitus, lateral view; B. Head, anterior view; C. Left antenna, inner view." figureDoi="http://doi.org/10.5281/zenodo.8204113" httpUri="https://zenodo.org/record/8204113/files/figure.png" pageId="3" pageNumber="154">4A</figureCitation>
,
<figureCitation id="13748DC6FF920015FF0EFED8FED9333C" box="[226,258,357,379]" captionStart="FIGURE 6" captionStartId="10.[151,250,1628,1652]" captionTargetBox="[151,1436,1029,1604]" captionTargetId="figure-375@10.[151,1436,1029,1604]" captionTargetPageId="10" captionText="FIGURE 6. Paraclius femoratus Aldrich, 1901, female syntype (NMNH). AB. Habitus, lateral and dorsal views, respectively. C. Scutellum, dorsal view (arrow pointing to the apical pair of scutellar setae); D. Labels. Images B and C originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204119" httpUri="https://zenodo.org/record/8204119/files/figure.png" pageId="3" pageNumber="154">6A</figureCitation>
); male tergite 6 bare (
<figureCitation id="13748DC6FF920015FE38FED8FDC4333D" box="[468,543,357,379]" captionStart="FIGURE 2" captionStartId="5.[151,250,1753,1777]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-18@5.[151,1436,181,1730]" captionTargetPageId="5" captionText="FIGURE 2. Paraclius arcuatus (Loew, 1861), A, D, E, F male specimen from Honduras (CNC), B, C male specimen from Mexico (NMNH). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D.Abdomen and hypopyium, lateral view; E. Left antenna, inner view; F. Left wing.Abbreviations: dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R2+3 = second branch of radius; R = third branch of radius." figureDoi="http://doi.org/10.5281/zenodo.8204107" httpUri="https://zenodo.org/record/8204107/files/figure.png" pageId="3" pageNumber="154">Fig. 2D</figureCitation>
)...............
<taxonomicName id="4C4FEAC0FF920015FD0FFED8FC45333D" authority="(Loew)" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[739,926,357,378]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="B93B4D51FF920015FD0FFED8FD29333D" box="[739,754,357,378]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FD12FED8FC8A333D" box="[766,849,357,378]" italics="true" pageId="3" pageNumber="154">arcuatus</emphasis>
(Loew)
</taxonomicName>
[Widespread in Central America,
<collectingCountry id="F358D1D3FF920015FB0AFED8FAC2333C" box="[1254,1305,357,379]" name="Cuba" pageId="3" pageNumber="154">Cuba</collectingCountry>
and
<collectingCountry id="F358D1D3FF920015FAABFED8FA4C333C" box="[1351,1431,357,379]" name="Jamaica" pageId="3" pageNumber="154">Jamaica</collectingCountry>
]
</keyLead>
</paragraph>
<paragraph id="8BF09143FF920015FF7BFE3FFA4733F3" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFE3FFA4733F3" pageId="3" pageNumber="154">
<emphasis id="B93B4D51FF920015FF7BFE3FFF4433D0" bold="true" box="[151,159,386,407]" pageId="3" pageNumber="154">-</emphasis>
Scutellum without pair of apical short setae (
<figureCitation id="13748DC6FF920015FD78FE3FFD3933D0" box="[660,738,386,408]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Figs 1B</figureCitation>
,
<figureCitation id="13748DC6FF920015FD01FE3FFCCC33D0" box="[749,791,386,407]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">11B</figureCitation>
); femur II yellow (
<figureCitation id="13748DC6FF920015FC22FE3FFBC433D0" box="[974,1055,386,408]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Figs 1A</figureCitation>
,
<figureCitation id="13748DC6FF920015FBC6FE3FFB8E33D0" box="[1066,1109,386,407]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">11A</figureCitation>
); lateral margins of male tergite 6 with strong setae (
<figureCitation id="13748DC6FF920015FE7DFE22FE3A33F2" box="[401,481,415,437]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Figs 1H</figureCitation>
,
<figureCitation id="13748DC6FF920015FE07FE1DFDCA33F2" box="[491,529,416,437]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">11E</figureCitation>
).........................................................................
<emphasis id="B93B4D51FF920015FA7CFE22FA4733F3" bold="true" box="[1424,1436,415,436]" pageId="3" pageNumber="154">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BB89E6FF920015FF7BFE00FA47300F" pageId="3" pageNumber="154">
<paragraph id="8BF09143FF920015FF7BFE00FA4733B7" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFE00FA4733B7" pageId="3" pageNumber="154">
<emphasis id="B93B4D51FF920015FF7BFE00FF783395" bold="true" box="[151,163,445,466]" pageId="3" pageNumber="154">4</emphasis>
Wing wholly brownish (
<figureCitation id="13748DC6FF920015FE3DFE00FDFC3395" box="[465,551,445,467]" captionStart="FIGURE 11" captionStartId="17.[151,250,1597,1621]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURE 11. Paraclius brooksi sp. nov., AE male holotype (CNC), F male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; CD. Head anterior and lateral views, respectively; E. Abdomen and hypopygium, lateral view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204135" httpUri="https://zenodo.org/record/8204135/files/figure.png" pageId="3" pageNumber="154">Fig. 11F</figureCitation>
); basal apicoventral epandrial seta about 2X longer than apical seta, and both shorter than ventral lobe of surstylus (
<figureCitation id="13748DC6FF920015FE3AFE67FD9233A8" box="[470,585,474,496]" captionStart="FIGURE12" captionStartId="18.[151,250,1647,1671]" captionTargetBox="[151,1436,237,1624]" captionTargetId="figure-42@18.[151,1436,237,1624]" captionTargetPageId="18" captionText="FIGURE12.Paraclius brooksisp.nov., male paratype (CNC).A. Abdomen, ventralview;B. Cerci, dorsalview;C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium; E. Internal appendages; F. Hypandrium and phallus, ventral view; G. Tergite 7, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204139" httpUri="https://zenodo.org/record/8204139/files/figure.png" pageId="3" pageNumber="154">Fig. 12C, D</figureCitation>
)..................
<taxonomicName id="4C4FEAC0FF920015FCC3FE66FC4B33A8" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[815,912,474,496]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="brooksi" status="sp. nov.">
<emphasis id="B93B4D51FF920015FCC3FE66FCE533B7" box="[815,830,475,496]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FCA5FE67FC4B33A8" box="[841,912,474,495]" italics="true" pageId="3" pageNumber="154">brooksi</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF920015FC7AFE67FC3833A8" bold="true" box="[918,995,474,495]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="A208F02AFF920015FC7AFE67FC3833A8" box="[918,995,474,495]" pageId="3" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
[
<collectingCountry id="F358D1D3FF920015FC02FE67FBE133B7" box="[1006,1082,474,496]" name="Mexico" pageId="3" pageNumber="154">Mexico</collectingCountry>
(
<collectingRegion id="498B5FA1FF920015FBABFE67FB4D33B7" box="[1095,1174,474,496]" country="Mexico" name="Chiapas" pageId="3" pageNumber="154">Chiapas</collectingRegion>
and
<collectingRegion id="498B5FA1FF920015FB28FE67FAD533A8" box="[1220,1294,474,495]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="154">Oaxaca</collectingRegion>
) and
<collectingCountry id="F358D1D3FF920015FAAFFE67FA4C33B7" box="[1347,1431,474,496]" name="Grenada" pageId="3" pageNumber="154">Grenada</collectingCountry>
]
</keyLead>
</paragraph>
<paragraph id="8BF09143FF920015FF7BFE45FA47300F" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFE45FA47300F" pageId="3" pageNumber="154">
- Wing wholly grayish (
<figureCitation id="13748DC6FF920015FE52FE4AFDDA304A" box="[446,513,503,525]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Fig. 1I</figureCitation>
); basal apicoventral epandrial seta as long as apical seta, and both longer than ventral lobe of surstylus (
<figureCitation id="13748DC6FF920015FEA8FDA8FE54306D" box="[324,399,533,555]" captionStart="FIGURE 1" captionStartId="4.[151,250,1581,1605]" captionTargetBox="[151,1436,306,1555]" captionTargetId="figure-87@4.[151,1436,306,1555]" captionTargetPageId="4" captionText="FIGURE 1. Paraclius angusticauda Van Duzee, 1933, male holotype (AMNH). AB. Habitus, lateral and dorsal views respectively; C. Labels; DE. Head, anterior and lateral views, respectively; FH. Hypopygium, lateral view; I. Right wing. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dm-m = discal medial crossvein; M1 = first branch of media; M4 = fourth branch of media; R 4+5 = third branch of radius. Images originally lacked scale bars. Photographs by Agnieszka Pierwola, AMNH. " figureDoi="http://doi.org/10.5281/zenodo.8204105" httpUri="https://zenodo.org/record/8204105/files/figure.png" pageId="3" pageNumber="154">Fig. 1G</figureCitation>
)..............................................................................................
<taxonomicName id="4C4FEAC0FF920015FEBEFD8EFD433000" authority="Van Duzee, 1933" authorityName="Van Duzee" authorityYear="1933" box="[338,664,562,584]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="angusticauda">
<emphasis id="B93B4D51FF920015FEBEFD8EFEBA300F" box="[338,353,563,584]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FE81FD8FFE353000" box="[365,494,562,583]" italics="true" pageId="3" pageNumber="154">angusticauda</emphasis>
<bibRefCitation id="EFDEECB2FF920015FE18FD8EFD433000" author="Van Duzee, M. C." box="[500,664,562,584]" pageId="3" pageNumber="154" pagination="1 - 27" refId="ref22526" refString="Van Duzee, M. C. (1933) New Dolichopidae from North America with notes on several described species. American Museum Novitates, 599, 1 - 27, 52 figs." type="journal article" year="1933">Van Duzee, 1933</bibRefCitation>
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[
<collectingCountry id="F358D1D3FF920015FD48FD8FFCCF300F" box="[676,788,562,584]" name="Guatemala" pageId="3" pageNumber="154">Guatemala</collectingCountry>
];
<taxonomicName id="4C4FEAC0FF920015FCCFFD8EFB9D3000" authority="Van Duzee, 1930" authorityName="Van Duzee" authorityYear="1930" box="[803,1094,562,584]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B93B4D51FF920015FCCFFD8EFCE9300F" box="[803,818,563,584]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FCD2FD8FFC403000" box="[830,923,562,583]" italics="true" pageId="3" pageNumber="154">elongatus</emphasis>
<bibRefCitation id="EFDEECB2FF920015FC4DFD8EFB9D3000" author="Van Duzee, M. C." box="[929,1094,562,584]" pageId="3" pageNumber="154" pagination="53 - 55" refId="ref22492" refString="Van Duzee, M. C. (1930) New species of Dolichopodidae from North America (Diptera). Entomological News, 41, 53 - 55 + 70 - 73." type="journal article" year="1930">Van Duzee, 1930</bibRefCitation>
</taxonomicName>
[
<collectingCountry id="F358D1D3FF920015FBBDFD8FFA4C300F" box="[1105,1431,562,584]" name="Saint Vincent and the Grenadines" pageId="3" pageNumber="154">Saint Vincent and the Grenadines</collectingCountry>
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<paragraph id="8BF09143FF920015FF7BFDEDFA4730E7" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
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<emphasis id="B93B4D51FF920015FF7BFDEDFF783022" bold="true" box="[151,163,592,613]" pageId="3" pageNumber="154">5</emphasis>
IIt 1 wholly dark brown (
<figureCitation id="13748DC6FF920015FE24FDF2FDCC3022" box="[456,535,591,613]" captionStart="FIGURE 7" captionStartId="11.[151,250,1939,1963]" captionTargetBox="[158,1430,771,1915]" captionTargetId="figure-209@11.[158,1430,771,1915]" captionTargetPageId="11" captionText="FIGURE 7. Paraclius angustipennis Van Duzee, 1929, male holotype (NMNH). AB. Habitus, lateral and dorsal views, respectively; C. Abdominal segments 5 and 6; D. Labels; EF. Head, anterior and anterolateral views, respectively. Images B, C, E originally lacked scale bars. Photographs by Alyssa Seemann, NMNH." figureDoi="http://doi.org/10.5281/zenodo.8204125" httpUri="https://zenodo.org/record/8204125/files/figure.png" pageId="3" pageNumber="154">Figs 7A</figureCitation>
,
<figureCitation id="13748DC6FF920015FDCDFDEDFD993022" box="[545,578,592,613]" captionStart="FIGURE 8" captionStartId="12.[151,250,1973,1997]" captionTargetBox="[151,1436,489,1950]" captionTargetId="figure-189@12.[151,1436,489,1950]" captionTargetPageId="12" captionText="FIGURE 8. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Abdomen, lateral view; E. Left antenna, outer view; F. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204127" httpUri="https://zenodo.org/record/8204127/files/figure.png" pageId="3" pageNumber="154">8A</figureCitation>
); phallus with fringe of spines at apical 1/2, not reaching the apex of phallus (
<figureCitation id="13748DC6FF920015FAC4FDF2FA4A3022" box="[1320,1425,591,613]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="3" pageNumber="154">Fig. 9BD</figureCitation>
); ventral lobe of surstylus short, gently curved ventrally, about 1/3 as long as basal apicoventral epandrial seta (
<figureCitation id="13748DC6FF920015FB13FDD0FA5A30C5" box="[1279,1409,621,643]" captionStart="FIGURE 9" captionStartId="14.[151,250,1639,1663]" captionTargetBox="[151,1436,181,1615]" captionTargetId="figure-20@14.[151,1436,181,1615]" captionTargetPageId="14" captionText="FIGURE 9. Paraclius angustipennis Van Duzee, 1929, identified male specimen from Mexico (CNC). A. Abdomen, ventral view; B. Hypopygium, left lateral view; C. Internal appendages, left lateral view; D. Apical part of phallus; E. Apical edge of epandrium; F. Tergite 6, lateral view; G. Right cercus, outer view; H. Left ventral surstylus, outer view; I. Hypandrium, ventral view; JK. Surstyli, left and right lateral views, respectively. Abbreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; ph pr = phallus process; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204131" httpUri="https://zenodo.org/record/8204131/files/figure.png" pageId="3" pageNumber="154">Fig. 9B, C, H</figureCitation>
).....................................................
<taxonomicName id="4C4FEAC0FF920015FCB0FD36FB7D30D8" authority="Van Duzee, 1929" authorityName="Van Duzee" authorityYear="1929" box="[860,1190,650,672]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="angustipennis">
<emphasis id="B93B4D51FF920015FCB0FD36FCB030E7" box="[860,875,651,672]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FC9AFD36FC2030E7" box="[886,1019,651,672]" italics="true" pageId="3" pageNumber="154">angustipennis</emphasis>
<bibRefCitation id="EFDEECB2FF920015FBEEFD36FB7D30D8" author="Van Duzee, M. C." box="[1026,1190,650,672]" pageId="3" pageNumber="154" pagination="1 - 64" refId="ref22434" refString="Van Duzee, M. C. (1929) Tropical American Diptera or two-winged flies of the family Dolichopodidae from Central and South America. Proceedings of the United States of National Museum, 74 (10), 1 - 64. https: // doi. org / 10.5479 / si. 00963801.74 - 2755.1" type="journal article" year="1929">Van Duzee, 1929</bibRefCitation>
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[
<collectingCountry id="F358D1D3FF920015FB5EFD37FAC630E7" box="[1202,1309,650,672]" name="Guatemala" pageId="3" pageNumber="154">Guatemala</collectingCountry>
and
<collectingCountry id="F358D1D3FF920015FAA7FD37FA4C30E7" box="[1355,1431,650,672]" name="Mexico" pageId="3" pageNumber="154">Mexico</collectingCountry>
]
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</paragraph>
<paragraph id="8BF09143FF920015FF7BFD15FA4730B0" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
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<emphasis id="B93B4D51FF920015FF7BFD15FF4430FA" bold="true" box="[151,159,680,701]" pageId="3" pageNumber="154">-</emphasis>
IIt 1 yellow on basal 1/2 and brownish on apical 1/2 (
<figureCitation id="13748DC6FF920015FD0DFD1AFCE030FA" box="[737,827,679,701]" captionStart="FIGURE 19" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[177,1411,597,1917]" captionTargetId="figure-128@25.[177,1411,597,1917]" captionTargetPageId="25" captionText="FIGURE 19. Paraclius xibun sp. nov., AE male holotype (CNC), FG male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Hypopygium, lateral view; E. Male abdominal segments 5 and 6; F. Left antenna, inner view; G. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204163" httpUri="https://zenodo.org/record/8204163/files/figure.png" pageId="3" pageNumber="154">Fig. 19A</figureCitation>
); phallus plain, lacking spines (
<figureCitation id="13748DC6FF920015FB81FD1AFB1330FA" box="[1133,1224,679,701]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="3" pageNumber="154">Figs 16E</figureCitation>
,
<figureCitation id="13748DC6FF920015FB3EFD15FB2630FA" box="[1234,1277,680,701]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="3" pageNumber="154">20E</figureCitation>
); ventral lobe of surstylus long, gently to strongly curved ventrally, about 1/2 as long or slightly shorter than basal apicoventral epandrial seta (
<figureCitation id="13748DC6FF920015FF05FD5FFE8530BF" box="[233,350,738,760]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="3" pageNumber="154">Figs 16CE</figureCitation>
,
<figureCitation id="13748DC6FF920015FE84FD5FFE4830B0" box="[360,403,738,759]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="3" pageNumber="154">20D</figureCitation>
)....................................................................................
<emphasis id="B93B4D51FF920015FA7CFD5FFA4730B0" bold="true" box="[1424,1436,738,759]" pageId="3" pageNumber="154">6</emphasis>
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<keyStep id="30BB89E6FF920015FF7BFD42FA463182" pageId="3" pageNumber="154">
<paragraph id="8BF09143FF920015FF7BFD42FA47312A" blockId="3.[151,1437,151,965]" pageId="3" pageNumber="154">
<keyLead id="30BE3276FF920015FF7BFD42FA47312A" pageId="3" pageNumber="154">
<emphasis id="B93B4D51FF920015FF7BFD42FF783153" bold="true" box="[151,163,767,788]" pageId="3" pageNumber="154">6</emphasis>
Tergite 6 with row of 34 strong lateral setae increasing in length towards posterior margin (
<figureCitation id="13748DC6FF920015FB94FD42FB123152" box="[1144,1225,767,789]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="3" pageNumber="154">Fig. 16I</figureCitation>
); sternite 5 U-shaped anteriorly (
<figureCitation id="13748DC6FF920015FEBDFCA0FE743174" box="[337,431,797,819]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="3" pageNumber="154">Fig. 16A</figureCitation>
); ventral lobe of surstylus strongly curved ventrally (
<figureCitation id="13748DC6FF920015FC3DFCA0FB923174" box="[977,1097,797,819]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="3" pageNumber="154">Fig. 16CE</figureCitation>
); dorsal lobe of surstylus weakly sclerotized, short, subrectangular, with 1 slender apicoventral seta, as long as dorsal lobe of surstylus, and 2 strong setae, 1 at apical margin and 1 subapical at dorsal edge (
<figureCitation id="13748DC6FF920015FD74FCEAFD37312A" box="[664,748,855,877]" captionStart="FIGURE 16" captionStartId="23.[151,250,1794,1818]" captionTargetBox="[272,1315,381,1770]" captionTargetId="figure-119@23.[272,1315,381,1770]" captionTargetPageId="23" captionText="FIGURE 16. Paraclius pumilio Loew, 1872, identified male specimen from Texas (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Dorsal lobe of surstylus, outer view; G. Hypandrium, ventral view; H. Cercus, dorsal view; I. Tergite 6, lateral view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; hypd pr = hypandrial process; pgt = postgonite; ph = phallus; prct = proctiger; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204149" httpUri="https://zenodo.org/record/8204149/files/figure.png" pageId="3" pageNumber="154">Fig. 16F</figureCitation>
)...............
<taxonomicName id="4C4FEAC0FF920015FC41FCE5FB95312A" authority="Loew" authorityName="Loew" authorityYear="1872" box="[941,1102,855,877]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B93B4D51FF920015FC41FCE5FC67312A" box="[941,956,856,877]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FC24FCEAFBCA312B" box="[968,1041,855,876]" italics="true" pageId="3" pageNumber="154">pumilio</emphasis>
Loew
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[Widespread in Mexico and
<collectingCountry id="F358D1D3FF920015FA88FCE5FA4C312A" box="[1380,1431,856,877]" name="United States of America" pageId="3" pageNumber="154">USA</collectingCountry>
]
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<keyLead id="30BE3276FF920015FF7BFCC8FA463182" pageId="3" pageNumber="154">
- Tergite 6 with two rows of 78 short lateral setae (
<figureCitation id="13748DC6FF920015FD57FCC8FCCB31CD" box="[699,784,885,907]" captionStart="FIGURE 19" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[177,1411,597,1917]" captionTargetId="figure-128@25.[177,1411,597,1917]" captionTargetPageId="25" captionText="FIGURE 19. Paraclius xibun sp. nov., AE male holotype (CNC), FG male paratype (CNC). A. Habitus, lateral view; B. Thorax, dorsal view; C. Head, anterior view; D. Hypopygium, lateral view; E. Male abdominal segments 5 and 6; F. Left antenna, inner view; G. Left wing." figureDoi="http://doi.org/10.5281/zenodo.8204163" httpUri="https://zenodo.org/record/8204163/files/figure.png" pageId="3" pageNumber="154">Fig. 19E</figureCitation>
); sternite 5 divided in two narrow hemitergites anteriorly (
<figureCitation id="13748DC6FF920015FAD4FCC8FA4A31CD" box="[1336,1425,885,907]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="3" pageNumber="154">Fig. 20A</figureCitation>
); ventral lobe of surstylus gently curved ventrally (
<figureCitation id="13748DC6FF920015FD50FC2FFCEA31E0" box="[700,817,914,936]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="3" pageNumber="154">Fig. 20C, D</figureCitation>
); dorsal lobe of surstylus suboval, with dorsal row of short setae at basal 1/2, ending in 1 strong seta at middle and with 1 strong seta at apex (
<figureCitation id="13748DC6FF920015FC29FC12FBC63182" box="[965,1053,943,965]" captionStart="FIGURE 20" captionStartId="26.[151,250,1827,1851]" captionTargetBox="[222,1366,525,1804]" captionTargetId="figure-203@26.[222,1366,525,1804]" captionTargetPageId="26" captionText="FIGURE 20. Paraclius xibun sp. nov., male paratype (CNC). A. Abdomen, ventral view; B. Tergite 7 and sternite 8, dorsal view; C. Hypopygium, left lateral view; D. Surstyli and apical edge of epandrium, outer view; E. Internal appendages, left lateral view; F. Hypandrium, ventral view; G. Cercus, dorsal view. Abreviations: a apv s = apical apicoventral epandrial seta; b apv s = basal apicoventral epandrial seta; cerc = cercus; dsur = dorsal lobe of surstylus; ej apod = ejaculatory apodeme; epand = epandrium; hypd = hypandrium; pgt = postgonite; ph = phallus; sp = sperm pump; st = sternite; tg = tergite; vsur = ventral lobe of surstylus." figureDoi="http://doi.org/10.5281/zenodo.8204167" httpUri="https://zenodo.org/record/8204167/files/figure.png" pageId="3" pageNumber="154">Fig. 20D</figureCitation>
)...........
<taxonomicName id="4C4FEAC0FF920015FB4AFC0DFB2D3183" authority="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha, 2023" authorityName="Soares &amp; Runyon &amp; Capellari &amp; Ale-Rocha" authorityYear="2023" box="[1190,1270,943,965]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="xibun" status="sp. nov.">
<emphasis id="B93B4D51FF920015FB4AFC0DFB6E3182" box="[1190,1205,944,965]" italics="true" pageId="3" pageNumber="154">P</emphasis>
.
<emphasis id="B93B4D51FF920015FB2DFC12FB2D3183" box="[1217,1270,943,964]" italics="true" pageId="3" pageNumber="154">xibun</emphasis>
</taxonomicName>
<emphasis id="B93B4D51FF920015FB10FC0DFA923182" bold="true" box="[1276,1353,944,965]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="A208F02AFF920015FB10FC0DFA923182" box="[1276,1353,944,965]" pageId="3" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
[
<collectingCountry id="F358D1D3FF920015FAB9FC12FA4C3182" box="[1365,1431,943,965]" name="Belize" pageId="3" pageNumber="154">Belize</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>