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<document id="E490035E05C56C40063A152BCF628826" ID-DOI="10.11646/zootaxa.5397.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10468563" ID-ZooBank="C55178A5-DABB-4615-8443-59B388F99A36" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704706602791" checkinUser="plazi" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2024" docId="03DC9030FFB6FFB86EFAFCD3FC30E577" docLanguage="en" docName="zootaxa.5397.2.2.pdf" docOrigin="Zootaxa 5397 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5397.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea indistincta Dunin 1991" docType="treatment" docVersion="2" lastPageNumber="212" masterDocId="FFE5E848FFA7FFA96E6DFFB4FFFBE051" masterDocTitle="New data on Dysdera Latreille, 1804 and Harpactea Bristowe, 1939 (Araneae: Dysderidae) of the Caucasus, with new species and records" masterLastPageNumber="217" masterPageNumber="195" pageNumber="212" updateTime="1704707340498" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="93C0A63968EF69D8B37308C29A32A551">New data on Dysdera Latreille, 1804 and Harpactea Bristowe, 1939 (Araneae: Dysderidae) of the Caucasus, with new species and records</mods:title>
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<mods:namePart id="9CA0DEF617DA6282E6904BCCBDD67C85">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="70E426DC3ECE9A442E4B52B208DA8DBD">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku 20500, Finland.</mods:affiliation>
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<mods:namePart id="6AF467D5328CF3081398199A41682FAC">Marusik, Yuri M.</mods:namePart>
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<mods:affiliation id="D270C36B85AB9215C5DAB3516DE3C9BC">Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa. &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Institute for Biological Problems of the North, Portovaya Str. 18, Magadan 685000, Russia.</mods:affiliation>
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<subSubSection id="C36F72ADFFB6FFB86EFAFCD3FDC7E3D1" box="[151,572,870,897]" pageId="17" pageNumber="212" type="nomenclature">
<paragraph id="8BCA2126FFB6FFB86EFAFCD3FDC7E3D1" blockId="17.[151,572,870,933]" box="[151,572,870,897]" pageId="17" pageNumber="212">
<heading id="D082964AFFB6FFB86EFAFCD3FDC7E3D1" bold="true" box="[151,572,870,897]" fontSize="11" level="1" pageId="17" pageNumber="212" reason="1">
<taxonomicName id="4C755AA5FFB6FFB86EFAFCD3FDC7E3D1" ID-CoL="3JRTF" authority="Dunin, 1991" authorityName="Dunin" authorityYear="1991" box="[151,572,870,897]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="indistincta">
<emphasis id="B901FD34FFB6FFB86EFAFCD3FDC7E3D1" bold="true" box="[151,572,870,897]" pageId="17" pageNumber="212">
<emphasis id="B901FD34FFB6FFB86EFAFCD3FE66E3D0" bold="true" box="[151,413,870,897]" italics="true" pageId="17" pageNumber="212">Harpactea indistincta</emphasis>
Dunin, 1991
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F72ADFFB6FFB86EFAFC3FFEE7E3F4" box="[151,284,907,933]" pageId="17" pageNumber="212" type="description">
<paragraph id="8BCA2126FFB6FFB86EFAFC3FFEE7E3F4" blockId="17.[151,572,870,933]" box="[151,284,907,933]" pageId="17" pageNumber="212">
<figureCitation id="134E3DA3FFB6FFB86EFAFC3FFEE7E3F4" box="[151,284,907,933]" captionStart="FIGURE 18" captionStartId="18.[152,255,1943,1967]" captionTargetBox="[212,1373,181,1916]" captionTargetId="figure-32@18.[205,1382,181,1919]" captionTargetPageId="18" captionText="FIGURE 18. Males of Harpactea indistincta (AC), H. dunini sp. n. (DF), and H. modesta (GI), left bulbs. A, D, G prolateral B, E, H retrolateral C, F, I ventral. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10468603" httpUri="https://zenodo.org/record/10468603/files/figure.png" pageId="17" pageNumber="212">Fig. 18AC</figureCitation>
</paragraph>
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<subSubSection id="C36F72ADFFB6FFB86EFAFC64FD3BE459" pageId="17" pageNumber="212" type="reference_group">
<paragraph id="8BCA2126FFB6FFB86EFAFC64FD24E3BB" blockId="17.[151,735,975,1032]" box="[151,735,975,1002]" pageId="17" pageNumber="212">
<treatmentCitationGroup id="AB650608FFB6FFB86EFAFC64FD24E3BB" box="[151,735,975,1002]" pageId="17" pageNumber="212">
<treatmentCitation id="0AD40737FFB6FFB86EFAFC64FDD4E3B6" author="Dunin, P. M." box="[151,559,975,999]" page="83" pageId="17" pageNumber="212" year="1991">
<taxonomicName id="4C755AA5FFB6FFB86EFAFC64FDD4E3B6" ID-CoL="3JRTF" authority="Dunin, 1991 b: 83" authorityName="Dunin" authorityPageNumber="83" authorityYear="1991" box="[151,559,975,999]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="indistincta">
<emphasis id="B901FD34FFB6FFB86EFAFC64FE8EE3B6" box="[151,373,975,999]" italics="true" pageId="17" pageNumber="212">Harpactea indistincta</emphasis>
<bibRefCitation id="EFE45CD7FFB6FFB86F16FC7BFDD4E3B6" author="Dunin, P. M." box="[379,559,975,999]" pageId="17" pageNumber="212" pagination="76 - 86" refId="ref10009" refString="Dunin, P. M. (1991 b) Spiders of genus Harpactea in the fauna of Azerbaijan (Aranei, Haplogynae, Dysderidae). Zoologicheskii Zhurnal, 70 (9), 76 - 86. [in Russian]" type="journal article" year="1991">Dunin, 1991b: 83</bibRefCitation>
</taxonomicName>
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, fig. 3.79 (
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).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA2126FFB6FFB86EFAFC5BFD3BE459" blockId="17.[151,735,975,1032]" box="[151,704,1006,1032]" pageId="17" pageNumber="212">
<treatmentCitationGroup id="AB650608FFB6FFB86EFAFC5BFD3BE459" box="[151,704,1006,1032]" pageId="17" pageNumber="212">
<taxonomicName id="4C755AA5FFB6FFB86EFAFC5BFE8EE457" ID-CoL="3JRTF" authorityName="Dunin" authorityYear="1991" box="[151,373,1006,1030]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="212" phylum="Arthropoda" rank="species" species="indistincta">
<emphasis id="B901FD34FFB6FFB86EFAFC5BFE8EE457" box="[151,373,1006,1030]" italics="true" pageId="17" pageNumber="212">Harpactea indistincta</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AD40737FFB6FFB86FEFFC5AFDCBE457" author="Dunin, P. M." box="[386,560,1006,1030]" page="71" pageId="17" pageNumber="212" year="1992">
<bibRefCitation id="EFE45CD7FFB6FFB86FEFFC5AFDCBE457" author="Dunin, P. M." box="[386,560,1006,1030]" pageId="17" pageNumber="212" pagination="35 - 76" refId="ref10097" refString="Dunin, P. M. (1992 b) The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta, 1 (3), 35 - 76. [in Russian]" type="journal article" year="1992">Dunin 1992b: 71</bibRefCitation>
</treatmentCitation>
, fig. 17 (
<specimenCount id="9D73EAAFFFB6FFB86CE2FC5BFD54E459" box="[655,687,1007,1032]" count="3" pageId="17" pageNumber="212" type="female">³♀</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36F72ADFFB6FFB86EFAFB80FC66E423" pageId="17" pageNumber="212" type="materials_examined">
<paragraph id="8BCA2126FFB6FFB86EFAFB80FC66E423" blockId="17.[151,1437,1076,1318]" pageId="17" pageNumber="212">
<emphasis id="B901FD34FFB6FFB86EFAFB80FEFDE41F" bold="true" box="[151,262,1076,1102]" pageId="17" pageNumber="212">Material.</emphasis>
<materialsCitation id="3B1D2B7BFFB6FFB86F62FB80FCB8E41F" box="[271,835,1076,1104]" collectionCode="ZMMU" country="Azerbaijan" location="Quba District" pageId="17" pageNumber="212" specimenCount="11" specimenCount-female="11" stateProvince="Quba">
<collectingCountry id="F36261B6FFB6FFB86F62FB80FE3BE41F" box="[271,448,1076,1102]" name="Azerbaijan" pageId="17" pageNumber="212">AZERBAIJAN</collectingCountry>
:
<emphasis id="B901FD34FFB6FFB86FA0FB80FD91E41F" box="[461,618,1076,1102]" italics="true" pageId="17" pageNumber="212">
<collectingRegion id="49B1EFC4FFB6FFB86FA0FB80FD91E41F" box="[461,618,1076,1102]" country="Azerbaijan" name="Quba" pageId="17" pageNumber="212">Quba District</collectingRegion>
</emphasis>
: 2³
<specimenCount id="9D73EAAFFFB6FFB86CF4FB81FD35E401" box="[665,718,1077,1104]" count="11" pageId="17" pageNumber="212" type="female">11♀</specimenCount>
(
<collectionCode id="ED64B9E3FFB6FFB86CB3FB80FCC0E41F" box="[734,827,1076,1102]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="17" pageNumber="212" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B1D2B7BFFB6FFB86D3DFB80FEAAE423" collectionCode="ZMMU" collectorName="N. Snegovaya" country="Azerbaijan" latitude="41.35" location="Quba" longLatPrecision="1156" longitude="48.5" pageId="17" pageNumber="212" specimenCount="6" specimenCount-female="6" stateProvince="Quba">
<collectingRegion id="49B1EFC4FFB6FFB86D3DFB80FC74E41F" box="[848,911,1076,1102]" country="Azerbaijan" name="Quba" pageId="17" pageNumber="212">Quba</collectingRegion>
, [
<geoCoordinate id="EE4147E1FFB6FFB86DC9FB80FBF9E41F" box="[932,1026,1076,1102]" degrees="41" direction="north" minutes="21" orientation="latitude" pageId="17" pageNumber="212" precision="925" value="41.35">41°21'N</geoCoordinate>
<geoCoordinate id="EE4147E1FFB6FFB86A61FB80FB92E41F" box="[1036,1129,1076,1102]" degrees="48" direction="east" minutes="30" orientation="longitude" pageId="17" pageNumber="212" precision="925" value="48.5">48°30'E</geoCoordinate>
], 04.2010 (
<collectorName id="268044F0FFB6FFB86A86FB80FA6BE41F" box="[1259,1424,1076,1102]" pageId="17" pageNumber="212">N. Snegovaya</collectorName>
); 1³
<specimenCount id="9D73EAAFFFB6FFB86EDFFBEDFF26E425" box="[178,221,1113,1140]" count="6" pageId="17" pageNumber="212" type="female">6♀</specimenCount>
(
<collectionCode id="ED64B9E3FFB6FFB86E81FBECFEB2E423" box="[236,329,1112,1138]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="17" pageNumber="212" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B1D2B7BFFB6FFB86F31FBECFC61E423" box="[348,922,1112,1138]" collectorName="N. Snegovaya" country="Azerbaijan" latitude="41.35" location="Quba" longLatPrecision="1156" longitude="48.5" pageId="17" pageNumber="212" specimenCount="1" stateProvince="Quba">
<collectingRegion id="49B1EFC4FFB6FFB86F31FBECFE60E423" box="[348,411,1112,1138]" country="Azerbaijan" name="Quba" pageId="17" pageNumber="212">Quba</collectingRegion>
, [
<geoCoordinate id="EE4147E1FFB6FFB86FC3FBECFDF7E423" box="[430,524,1112,1138]" degrees="41" direction="north" minutes="21" orientation="latitude" pageId="17" pageNumber="212" precision="925" value="41.35">41°21'N</geoCoordinate>
<geoCoordinate id="EE4147E1FFB6FFB86C7EFBECFD8AE423" box="[531,625,1112,1138]" degrees="48" direction="east" minutes="30" orientation="longitude" pageId="17" pageNumber="212" precision="925" value="48.5">48°30'E</geoCoordinate>
], 06.2010 (
<collectorName id="268044F0FFB6FFB86C82FBECFC69E423" box="[751,914,1112,1138]" pageId="17" pageNumber="212">N. Snegovaya</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F72ADFFB6FFB86EAAFBC8FB3DE553" pageId="17" pageNumber="212" type="discussion">
<paragraph id="8BCA2126FFB6FFB86EAAFBC8FB3DE553" blockId="17.[151,1437,1076,1318]" pageId="17" pageNumber="212">
<emphasis id="B901FD34FFB6FFB86EAAFBC8FEB4E4C7" bold="true" box="[199,335,1148,1174]" pageId="17" pageNumber="212">Comments.</emphasis>
This species has only been treated in the two publications mentioned above and was previously known solely from two neighbouring localities in northeastern
<collectingCountry id="F36261B6FFB6FFB86D3CFB14FC37E4EB" box="[849,972,1184,1210]" name="Azerbaijan" pageId="17" pageNumber="212">Azerbaijan</collectingCountry>
and the adjacent southern Dagestan (
<bibRefCitation id="EFE45CD7FFB6FFB86B04FB14FF2FE48F" author="Otto, S." pageId="17" pageNumber="212" refId="ref10910" refString="Otto, S. (2022) Caucasian Spiders. A faunistic database on the spiders of the Caucasus. Version February 2022. Available from: https: // caucasus-spiders. info / (accessed 23 May 2023)" type="url" year="2022">Otto 2022</bibRefCitation>
). The examined material was collected from the
<typeStatus id="54CE9F84FFB6FFB86C93FB71FCD5E48E" box="[766,814,1221,1247]" pageId="17" pageNumber="212">type</typeStatus>
locality. It inhabits oak-hornbeam forests at elevations ranging from
<quantity id="4C8D8CC3FFB6FFB86F5CFB5CFE3DE552" box="[305,454,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="17" pageNumber="212" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600 to 800 m</quantity>
and can be found within leaf litter or beneath stones (
<bibRefCitation id="EFE45CD7FFB6FFB86A4DFB5CFB42E553" author="Dunin, P. M." box="[1056,1209,1256,1282]" pageId="17" pageNumber="212" pagination="76 - 86" refId="ref10009" refString="Dunin, P. M. (1991 b) Spiders of genus Harpactea in the fauna of Azerbaijan (Aranei, Haplogynae, Dysderidae). Zoologicheskii Zhurnal, 70 (9), 76 - 86. [in Russian]" type="journal article" year="1991">Dunin 1991b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F72ADFFB6FFB86EAAFAB8FC30E577" box="[199,971,1292,1318]" pageId="17" pageNumber="212" type="distribution">
<paragraph id="8BCA2126FFB6FFB86EAAFAB8FC30E577" blockId="17.[151,1437,1076,1318]" box="[199,971,1292,1318]" pageId="17" pageNumber="212">
<emphasis id="B901FD34FFB6FFB86EAAFAB8FE9AE577" bold="true" box="[199,353,1292,1318]" pageId="17" pageNumber="212">Distribution.</emphasis>
Caucasus:
<collectingCountry id="F36261B6FFB6FFB86FB3FAB8FDA2E577" box="[478,601,1292,1318]" name="Azerbaijan" pageId="17" pageNumber="212">Azerbaijan</collectingCountry>
and North Caucasus (Dagestan).
</paragraph>
</subSubSection>
</treatment>
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