treatments-xml/data/03/DC/6C/03DC6C76CC234C30F7DFFC89FBF2FD1C.xml
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<document id="30C8D18EBAE3C9C511334FF04272B5C2" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC234C30F7DFFC89FBF2FD1C" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana tubiraensis Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="158874DE-1356-42D8-A039-F7A62D93F8AF" docUuidSource="ZooBank" docVersion="3" lastPageNumber="21" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="19" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="861ACC29AF7B455CFAFFFC4A5603A94C">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:namePart id="A4B959CA45B1D2CB54C429BAEAB0FEB1">Lehmann, Ingo</mods:namePart>
<mods:nameIdentifier id="B43DD12F3E060E110C490D987CC401DD" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:namePart id="771A0EF3540647C9EEF41302BF4AD508">Zahiri, Reza</mods:namePart>
<mods:nameIdentifier id="9D583734049B4B89B4BFFDB2490B1643" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
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<mods:namePart id="939EB48D5D386A3361F724EB5866DCA2">Husemann, Martin</mods:namePart>
<mods:nameIdentifier id="226ED8A0767D14DC2DDB31C224A4F3BB" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="2E13F77C9841E0225FE7561A9EA6713A">Zootaxa</mods:title>
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<mods:date id="8F7B1C92BB24AE3668F626659BD68CE3">2023</mods:date>
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<treatment id="03DC6C76CC234C30F7DFFC89FBF2FD1C" LSID="urn:lsid:zoobank.org:act:158874DE-1356-42D8-A039-F7A62D93F8AF" httpUri="http://treatment.plazi.org/id/03DC6C76CC234C30F7DFFC89FBF2FD1C" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="C36F8EEBCC234C36F7DFFC89FC04FC34" box="[151,922,870,897]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BCADD60CC234C36F7DFFC89FC04FC34" blockId="18.[151,961,870,969]" box="[151,922,870,897]" pageId="18" pageNumber="19">
<heading id="D0826A0CCC234C36F7DFFC89FC04FC34" bold="true" box="[151,922,870,897]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="B9010172CC234C36F7DFFC89FC04FC34" bold="true" box="[151,922,870,897]" pageId="18" pageNumber="19">
<taxonomicName id="4C75A6E3CC234C36F7DFFC89FCABFC34" ID-CoL="B8R8B" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,821,870,897]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tubiraensis" status="sp. nov.">
<emphasis id="B9010172CC234C36F7DFFC89FE37FC34" bold="true" box="[151,425,870,897]" italics="true" pageId="18" pageNumber="19">Lukeniana tubiraensis</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC234C36F474FC88FC04FC34" box="[828,922,871,897]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC234C36F7DFFC64FACFFBA4" pageId="18" pageNumber="19" type="description">
<paragraph id="8BCADD60CC234C36F7DFFC64FC5FFC7C" blockId="18.[151,961,870,969]" pageId="18" pageNumber="19">
<figureCitation id="134EC1E5CC234C36F7DFFC64FF76FC10" box="[151,232,907,933]" captionStart="FIGURE 3" captionStartId="19.[151,250,1753,1777]" captionTargetBox="[151,1436,381,1729]" captionTargetId="figure-103@19.[151,1436,381,1729]" captionTargetPageId="19" captionText="FIGURE 3. a. Lukeniana enaiposha, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha; b. L. obliqualinea, male, Kenya, County Nairobi, Nairobi; c. L. obliqualinea, female, Kenya, County Nairobi, Loresho Ridge; d. L. raymondrevellii, holotype, male, Namibia, Otjozondjupa Region, Ozombanda; e. L. rajaeii, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole; f. L. tubiraensis, holotype, male, Tanzania, Kigoma Region, Tubira Forest." figureDoi="http://doi.org/10.5281/zenodo.7840792" httpUri="https://zenodo.org/record/7840792/files/figure.png" pageId="18" pageNumber="19">Figs 3f</figureCitation>
,
<figureCitation id="134EC1E5CC234C36F7BCFC64FE8FFC10" box="[244,273,907,933]" captionStart="FIGURE 9" captionStartId="59.[151,250,1714,1738]" captionTargetBox="[235,1352,188,1685]" captionTargetId="figure-19@59.[223,1364,181,1690]" captionTargetPageId="59" captionText="FIGURE 9. Wing venation of: a. Zambezia darrelplowesi, female, holotype (Zambezia = subgroup G1 in Lehmann 2019b); b. Lukeniana obliqualinea, female, Kenya, County Nairobi, Ngong (Lukeniana = subgroup G3 in Lehmann 2019b); c. Zambezia madambae, male, holotype; d. Z. diredaouaensis, male, paratype; e. Lukeniana tubiraensis, male, holotype; f. Metarbelodes umtaliana, male, holotype (Metarbelodes = subgroup G2 in Lehmann 2019b). The areole (blue arrows) is a possible plesiomorphy (in Lehmann 2019b, p. 24), the continuous CuP fold (red arrows), partly sclerotized, is a very rare apomorphy (in Lehmann 2019b, p. 25) among species of Metarbelidae. It has been found only in species close to or to the South of the Zambezi River. In the same areas an intact CuP on the forewing occurs only in basal species (as defined by Lehmann 2019b, pp. 24 and 25) of Metarbelidae (Lehmann 2019b). The complete loss of the CuP in the forewing is another apomorphy (Lehmann 2019b, p. 322) that occurs in the majority of species of Metarbelidae in theAfrotropical Region and Oriental Region (seeFigure 39).Drawings by I.L." figureDoi="http://doi.org/10.5281/zenodo.7840807" httpUri="https://zenodo.org/record/7840807/files/figure.png" pageId="18" pageNumber="19">9e</figureCitation>
,
<figureCitation id="134EC1E5CC234C36F654FC64FEDAFC10" box="[284,324,907,933]" captionStart="FIGURE 13" captionStartId="64.[151,250,1283,1307]" captionTargetBox="[163,1424,181,1249]" captionTargetId="figure-28@64.[151,1436,166,1259]" captionTargetPageId="64" captionText="FIGURE 13. a. Lukeniana enaiposha, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha; b. L. obliqualinea, male, Kenya, County Nairobi, Nairobi; c. L. raymondrevellii, holotype, male, Namibia, Otjozondjupa Region, Ozombanda; d. L. rajaeii, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole; e. L. tubiraensis, holotype, male, Tanzania, Kigoma Region, Tubira Forest; f. L. lutztoepferi, holotype, male, Kenya, County Nairobi, Ololua Forest." figureDoi="http://doi.org/10.5281/zenodo.7840820" httpUri="https://zenodo.org/record/7840820/files/figure.png" pageId="18" pageNumber="19">13e</figureCitation>
<uri id="FFE4D162CC234C36F7DFFC41FC5FFC7C" box="[151,961,942,969]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BCADD60CC234C36F7DFFC18FACFFBA4" blockId="18.[151,1361,1015,1041]" box="[151,1361,1015,1041]" pageId="18" pageNumber="19">
<materialsCitation id="3B1DD73DCC234C36F7DFFC18FAD3FBA4" box="[151,1357,1015,1041]" collectionCode="NHMO" country="Tanzania" location="University" municipality="Natural History Museum" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Oslo" typeStatus="holotype">
<typeStatus id="54CE63C2CC234C36F7DFFC18FF51FBA4" box="[151,207,1015,1041]" pageId="18" pageNumber="19">Type</typeStatus>
locality and repository:
<collectingCountry id="F3629DF0CC234C36F6A8FC18FDD6FBA4" box="[480,584,1015,1041]" name="Tanzania" pageId="18" pageNumber="19">Tanzania</collectingCountry>
, the
<collectingMunicipality id="6BAE471ACC234C36F534FC18FC09FBA4" box="[636,919,1015,1041]" pageId="18" pageNumber="19">Natural History Museum</collectingMunicipality>
,
<location id="8EAA8BBBCC234C36F4EAFC18FB86FBA4" LSID="urn:lsid:plazi:treatment:03DC6C76CC234C30F7DFFC89FBF2FD1C:8EAA8BBBCC234C36F4EAFC18FB86FBA4" box="[930,1048,1015,1041]" country="Tanzania" municipality="Natural History Museum" name="University" pageId="18" pageNumber="19" stateProvince="Oslo">University</location>
of
<collectingRegion id="49B11382CC234C36F375FC18FBEBFBA4" box="[1085,1141,1015,1041]" country="Norway" name="Oslo" pageId="18" pageNumber="19">Oslo</collectingRegion>
,
<collectingCountry id="F3629DF0CC234C36F3C8FC18FB44FBA4" box="[1152,1242,1015,1041]" name="Norway" pageId="18" pageNumber="19">Norway</collectingCountry>
(
<collectionCode id="ED6445A5CC234C36F3A1FC18FADBFBA4" box="[1257,1349,1015,1041]" country="Norway" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="18" pageNumber="19" type="Museum">NHMO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC234C36F7DFFBD1FE64FB5C" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BCADD60CC234C36F7DFFBD1FE64FB5C" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F7DFFBD1FE1CFBED" bold="true" box="[151,386,1086,1112]" pageId="18" pageNumber="19">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC234C36F6C3FBD0FE9DFB14" collectingDate="1989-07-28" collectionCode="NHMO" collectorName="A. Bjornstad &amp; I. Lehmann" country="Tanzania" elevation="1" location="Tubira Forest" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Kigoma" typeStatus="holotype">
<typeStatus id="54CE63C2CC234C36F6C3FBD0FE6DFBEC" box="[395,499,1087,1113]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC234C36F50AFBD0FD34FBEC" box="[578,682,1087,1113]" name="Tanzania" pageId="18" pageNumber="19">Tanzania</collectingCountry>
,
<collectingRegion id="49B11382CC234C36F5FEFBD0FCF3FBEC" box="[694,877,1087,1113]" country="Tanzania" name="Kigoma" pageId="18" pageNumber="19">Kigoma Region</collectingRegion>
,
<collectingRegion id="49B11382CC234C36F432FBD0FBAAFBEC" box="[890,1076,1087,1113]" country="Tanzania" name="Kigoma" pageId="18" pageNumber="19">Kigoma District</collectingRegion>
,
<location id="8EAA8BBBCC234C36F308FBD0FB42FBEC" LSID="urn:lsid:plazi:treatment:03DC6C76CC234C30F7DFFC89FBF2FD1C:8EAA8BBBCC234C36F308FBD0FB42FBEC" box="[1088,1244,1087,1113]" country="Tanzania" name="Tubira Forest" pageId="18" pageNumber="19" stateProvince="Kigoma">Tubira Forest</location>
, 0501S, 3006E,
<quantity id="4C8D7085CC234C36F7DFFB8CFF67FBC8" box="[151,249,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="19" unit="m" value="1.1">
<elevation id="00583A53CC234C36F7DFFB8CFF67FBC8" box="[151,249,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="19" unit="m" value="1.1">1.100 m</elevation>
</quantity>
,
<date id="FFCBFBA0CC234C36F641FB8CFE3BFBC8" box="[265,421,1123,1149]" pageId="18" pageNumber="19" value="1989-07-28">
<collectingDate id="EF8F0248CC234C36F641FB8CFE3BFBC8" box="[265,421,1123,1149]" pageId="18" pageNumber="19" value="1989-07-28">28 July 1989</collectingDate>
</date>
, label number 15802,
<collectorName id="2680B8B6CC234C36F5E6FB8CFCDFFBC8" box="[686,833,1123,1149]" pageId="18" pageNumber="19">A. Bjørnstad</collectorName>
leg., genitalia slide number 16/012012
<collectorName id="2680B8B6CC234C36F25FFB8CFA03FBC8" box="[1303,1437,1123,1149]" pageId="18" pageNumber="19">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC234C36F7D7FB68FF65FB14" box="[159,251,1159,1185]" country="Norway" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="18" pageNumber="19" type="Museum">NHMO</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD73DCC234C36F646FB68FA06FB14" box="[270,1432,1158,1185]" collectingDate="1989-07-28" collectionCode="NMK" collectorName="I. Lehmann" country="Tanzania" elevation="1" location="Tubira Forest" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Kigoma" typeStatus="paratype">
<typeStatus id="54CE63C2CC234C36F646FB68FEE1FB14" box="[270,383,1159,1185]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7316E9CC234C36F6C1FB68FE6CFB14" box="[393,498,1159,1185]" pageId="18" pageNumber="19" type="male">one male</specimenCount>
, same data as holotype, label 15715, genitalia slide 21/022012
<collectorName id="2680B8B6CC234C36F3F0FB68FAA7FB14" box="[1208,1337,1159,1185]" pageId="18" pageNumber="19">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC234C36F200FB68FA11FB14" box="[1352,1423,1159,1185]" country="Kenya" httpUri="http://grbio.org/cool/h6fg-emd8" name="National Museums of Kenya" pageId="18" pageNumber="19" type="Museum">NMK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1DD73DCC234C36F7DFFB44FE68FB5C" collectingDate="1965-06" collectionCode="NMK" collectorName="I. Robertson &amp; I. Lehmann" country="Tanzania" location="Mbeya Region" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Mbeya" typeStatus="paratype">
<specimenCount id="9D7316E9CC234C36F7DFFB44FE9FFB70" box="[151,257,1195,1221]" pageId="18" pageNumber="19" type="male">one male</specimenCount>
,
<collectingCountry id="F3629DF0CC234C36F646FB44FEE8FB70" box="[270,374,1195,1221]" name="Tanzania" pageId="18" pageNumber="19">Tanzania</collectingCountry>
,
<collectingRegion id="49B11382CC234C36F6CAFB44FDB2FB70" box="[386,556,1195,1221]" country="Tanzania" name="Mbeya" pageId="18" pageNumber="19">Mbeya Region</collectingRegion>
,
<collectingRegion id="49B11382CC234C36F570FB44FD78FB70" box="[568,742,1195,1221]" country="Tanzania" name="Mbeya" pageId="18" pageNumber="19">Mbeya District</collectingRegion>
,
<collectingRegion id="49B11382CC234C36F5BAFB44FCDDFB70" box="[754,835,1195,1221]" country="Tanzania" name="Mbeya" pageId="18" pageNumber="19">Mbeya</collectingRegion>
,
<date id="FFCBFBA0CC234C36F418FB44FC58FB70" box="[848,966,1194,1221]" pageId="18" pageNumber="19" value="1965-06">
<collectingDate id="EF8F0248CC234C36F418FB44FC58FB70" box="[848,966,1194,1221]" pageId="18" pageNumber="19" value="1965-06">June 1965</collectingDate>
</date>
,
<collectorName id="2680B8B6CC234C36F49BFB44FBC1FB70" box="[979,1119,1195,1221]" pageId="18" pageNumber="19">I. Robertson</collectorName>
leg., genitalia slide number 23/042012
<collectorName id="2680B8B6CC234C36F65EFB20FE09FB5C" box="[278,407,1231,1257]" pageId="18" pageNumber="19">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC234C36F6EEFB20FE70FB5C" box="[422,494,1231,1257]" country="Kenya" httpUri="http://grbio.org/cool/h6fg-emd8" name="National Museums of Kenya" pageId="18" pageNumber="19" type="Museum">NMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC234C37F78FFB1DFE3CFED0" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8BCADD60CC234C36F78FFB1DFEA6FACC" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F78FFB1DFEC7FAB9" bold="true" box="[199,345,1266,1292]" pageId="18" pageNumber="19">Description.</emphasis>
Male.
<emphasis id="B9010172CC234C36F6EFFB1CFE7AFAB8" box="[423,484,1267,1293]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
: Ochre and sepia, shiny; long, dense hair-like scales between eyes; eyes black with a vinaceous glint; antennae 0.50 length of forewing, bipectinate, with branches 6.0 width of shaft, branches and shaft covered with ochre scales dorsally; antennal tips not spatulate, scaled, bending towards apex; labial palpi ochre and sepia dorsally.
</paragraph>
<paragraph id="8BCADD60CC234C36F78FFA6CFE2DF908" blockId="18.[151,1437,1086,2013]" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F78FFA6CFE86FA28" box="[199,280,1411,1437]" italics="true" pageId="18" pageNumber="19">Thorax</emphasis>
: Patagia and tegulae with shiny and long hair-like scales of ochre mixed with sepia. A small crest of sepia on metathorax. Hind legs ochre with shiny, fine hair-like scales of sepia; a pair of narrow tibial spurs of unequal length, outer spur
<emphasis id="B9010172CC234C36F62CFA24FE18FA51" box="[356,390,1483,1508]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
1.0 mm, inner spur
<emphasis id="B9010172CC234C36F522FA24FD12FA51" box="[618,652,1483,1508]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
<quantity id="4C8D7085CC234C36F5DBFA24FD73FA50" box="[659,749,1483,1509]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="19" unit="mm" value="0.8">0.8 mm</quantity>
. Forewing length of
<typeStatus id="54CE63C2CC234C36F49FFA24FBA7FA50" box="[983,1081,1483,1509]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
15.0 mm (wingspan 33.0 mm), Forewing upperside with a dark appearance, largely light sepia mixed with ochre; costal margin with four striae of sepia near base of forewing, striae on whole upperside not distinctly coloured, only faded striae towards termen; narrow band of faded sepia along termen, without spots at end of veins; CuA
<subScript id="17F1DF25CC234C36F358F9A6FB86F9E2" attach="left" box="[1040,1048,1609,1623]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
broad, white, edged with a broad band of sepia above; remaining veins not distinctly coloured; cilia very long, 2.0 mm, ochre with sepia towards tips, shiny. Underside of forewing light ochre, glossy, costa ochre and sepia. Hindwing upperside ivory-yellow; cilia and underside as in forewing.
</paragraph>
<paragraph id="8BCADD60CC234C37F78FF928FD0EFEF4" blockId="18.[151,1437,1086,2013]" lastBlockId="19.[151,1436,151,357]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B9010172CC234C36F78FF928FEAFF954" box="[199,305,1735,1761]" italics="true" pageId="18" pageNumber="19">Abdomen</emphasis>
: Ochre mixed with ivory-yellow and sepia, glossy; abdominal tuft
<emphasis id="B9010172CC234C36F354F928FBA0F955" box="[1052,1086,1735,1760]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
30% length of abdomen. Genitalia (
<figureCitation id="134EC1E5CC234C36F79FF904FEAAF8B0" box="[215,308,1771,1797]" captionStart="FIGURE 13" captionStartId="64.[151,250,1283,1307]" captionTargetBox="[163,1424,181,1249]" captionTargetId="figure-28@64.[151,1436,166,1259]" captionTargetPageId="64" captionText="FIGURE 13. a. Lukeniana enaiposha, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha; b. L. obliqualinea, male, Kenya, County Nairobi, Nairobi; c. L. raymondrevellii, holotype, male, Namibia, Otjozondjupa Region, Ozombanda; d. L. rajaeii, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole; e. L. tubiraensis, holotype, male, Tanzania, Kigoma Region, Tubira Forest; f. L. lutztoepferi, holotype, male, Kenya, County Nairobi, Ololua Forest." figureDoi="http://doi.org/10.5281/zenodo.7840820" httpUri="https://zenodo.org/record/7840820/files/figure.png" pageId="18" pageNumber="19">Fig. 13e</figureCitation>
) with uncus subrectangular, outer edge slightly C-shaped, inner edge rounded at base, lobes densely covered with short setae ventrally; gnathos arms short, subequal o width of valva, not touching coastal margin of valva, bent towards uncus; valva narrow and long, rectangular, costa straight and without setae; sacculus with long setae near base of thorn-like process; weakly-sclerotized projection with short setae and a broad rectangular tip, longer than the two thorn-like processes (
<emphasis id="B9010172CC234C36F529F894FDE3F820" box="[609,637,1915,1941]" italics="true" pageId="18" pageNumber="19">cf.</emphasis>
<figureCitation id="134EC1E5CC234C36F5CAF894FD43F820" box="[642,733,1914,1941]" captionStart="FIGURE 15" captionStartId="66.[151,250,1656,1680]" captionTargetBox="[172,1419,191,1622]" captionTargetId="figure-28@66.[151,1436,181,1632]" captionTargetPageId="66" captionText="FIGURE 15. Examples of male genitalia in lateral view: a. Zambezia madambae, paratype, with extremely small thorn-like process and small weakly-sclerotized projection (red arrows); b. Lukeniana timdavenporti, paratype, with strongly bent ventral edge of valva and well developed thorn-like process (red arrows); c. L. bergsteni, paratype, without strongly bent ventral edge of valva and two thorn-like processes (red arrows); d. L. butleri, paratype, with many long setae on valva that appear to be black in colour and one thorn-like process (red arrows)." figureDoi="http://doi.org/10.5281/zenodo.7840824" httpUri="https://zenodo.org/record/7840824/files/figure.png" pageId="18" pageNumber="19">Fig. 15c</figureCitation>
for the two processes) below that extending beyond upper edge of weakly-sclerotized projection; thorn-like process strongly curved upward, hollow, acuminate, no dots with tiny setae, narrow at base without second thorn-like process (
<emphasis id="B9010172CC234C36F45BF82CFCB1F868" box="[787,815,1987,2013]" italics="true" pageId="18" pageNumber="19">cf.</emphasis>
<figureCitation id="134EC1E5CC234C36F47DF82CFC0FF868" box="[821,913,1986,2013]" captionStart="FIGURE 15" captionStartId="66.[151,250,1656,1680]" captionTargetBox="[172,1419,191,1622]" captionTargetId="figure-28@66.[151,1436,181,1632]" captionTargetPageId="66" captionText="FIGURE 15. Examples of male genitalia in lateral view: a. Zambezia madambae, paratype, with extremely small thorn-like process and small weakly-sclerotized projection (red arrows); b. Lukeniana timdavenporti, paratype, with strongly bent ventral edge of valva and well developed thorn-like process (red arrows); c. L. bergsteni, paratype, without strongly bent ventral edge of valva and two thorn-like processes (red arrows); d. L. butleri, paratype, with many long setae on valva that appear to be black in colour and one thorn-like process (red arrows)." figureDoi="http://doi.org/10.5281/zenodo.7840824" httpUri="https://zenodo.org/record/7840824/files/figure.png" pageId="18" pageNumber="19">Fig. 15c</figureCitation>
for the two processes); median sector of valva with rows comprising short setae on inner side; a short, broad, ovoid emargination extending between weakly-sclerotized projection and thorn-like process for
<emphasis id="B9010172CC224C37F535FF54FD01FF61" box="[637,671,187,212]" italics="true" pageId="19" pageNumber="20">ca.</emphasis>
30% length of valva; ventral side of valva not bent at middle. Saccus smaller than juxta, finger-like, juxta broader with two acuminate points, a flag-like process at each tip, emargination between points very deep (90% of length of juxta). Phallus long, 1.2 length of valva, not trumpet-like, strongly bent near middle, bilobed with a cleft at each end.
</paragraph>
<paragraph id="8BCADD60CC224C37F78FFEA4FE3CFED0" blockId="19.[151,1436,151,357]" box="[199,418,331,357]" pageId="19" pageNumber="20">
<emphasis id="B9010172CC224C37F78FFEA4FEBBFED0" bold="true" box="[199,293,331,357]" pageId="19" pageNumber="20">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF0A8DE8CC224C37F7DFF936FADCF8E9" ID-DOI="http://doi.org/10.5281/zenodo.7840792" ID-Zenodo-Dep="7840792" httpUri="https://zenodo.org/record/7840792/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1753,1777]" targetBox="[151,1436,381,1729]" targetPageId="19">
<paragraph id="8BCADD60CC224C37F7DFF936FADCF8E9" blockId="19.[151,1437,1752,1885]" pageId="19" pageNumber="20">
<emphasis id="B9010172CC224C37F7DFF936FD82F945" bold="true" box="[151,540,1752,1777]" pageId="19" pageNumber="20">
FIGURE 3. a.
<taxonomicName id="4C75A6E3CC224C37F67FF937FD82F945" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[311,540,1752,1776]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="enaiposha">
<emphasis id="B9010172CC224C37F67FF937FD82F945" bold="true" box="[311,540,1752,1776]" italics="true" pageId="19" pageNumber="20">Lukeniana enaiposha</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Kenya, Rift Valley Province, near Lake Naivasha;
<emphasis id="B9010172CC224C37F391F936FA08F945" bold="true" box="[1241,1430,1752,1777]" pageId="19" pageNumber="20">
b.
<taxonomicName id="4C75A6E3CC224C37F3BDF936FA08F945" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1909" box="[1269,1430,1752,1776]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC224C37F3BDF936FA08F945" bold="true" box="[1269,1430,1752,1776]" italics="true" pageId="19" pageNumber="20">L. obliqualinea</emphasis>
</taxonomicName>
</emphasis>
, male, Kenya, County Nairobi, Nairobi;
<emphasis id="B9010172CC224C37F563F912FD7CF8A1" bold="true" box="[555,738,1788,1813]" pageId="19" pageNumber="20">
c.
<taxonomicName id="4C75A6E3CC224C37F50BF912FD7CF8A1" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1909" box="[579,738,1788,1812]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC224C37F50BF912FD7CF8A1" bold="true" box="[579,738,1788,1812]" italics="true" pageId="19" pageNumber="20">L. obliqualinea</emphasis>
</taxonomicName>
</emphasis>
, female, Kenya, County Nairobi, Loresho Ridge;
<emphasis id="B9010172CC224C37F397F912FF20F88D" bold="true" pageId="19" pageNumber="20">
d.
<taxonomicName id="4C75A6E3CC224C37F3B2F912FF20F88D" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="raymondrevellii">
<emphasis id="B9010172CC224C37F3B2F912FF20F88D" bold="true" italics="true" pageId="19" pageNumber="20">L. raymondrevellii</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Namibia, Otjozondjupa Region, Ozombanda;
<emphasis id="B9010172CC224C37F406F8CEFC55F88C" bold="true" box="[846,971,1825,1849]" pageId="19" pageNumber="20">
e.
<taxonomicName id="4C75A6E3CC224C37F420F8CEFC55F88C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[872,971,1825,1849]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="rajaeii">
<emphasis id="B9010172CC224C37F420F8CEFC55F88C" bold="true" box="[872,971,1825,1849]" italics="true" pageId="19" pageNumber="20">L. rajaeii</emphasis>
</taxonomicName>
</emphasis>
, paratype, male, Ethiopia, Southern Nations, Nationalities and PeoplesRegion, Gidole;
<emphasis id="B9010172CC224C37F501F8AAFD70F8E9" bold="true" box="[585,750,1860,1885]" pageId="19" pageNumber="20">
f.
<taxonomicName id="4C75A6E3CC224C37F516F8AAFD70F8E9" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[606,750,1860,1884]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC224C37F516F8AAFD70F8E9" bold="true" box="[606,750,1860,1884]" italics="true" pageId="19" pageNumber="20">L. tubiraensis</emphasis>
</taxonomicName>
</emphasis>
, holotype, male, Tanzania, Kigoma Region, Tubira Forest.
</paragraph>
</caption>
<subSubSection id="C36F8EEBCC224C30F78FF864FAA6FEF4" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BCADD60CC224C30F78FF864FAA6FEF4" blockId="19.[151,1436,1931,2030]" lastBlockId="20.[151,1437,151,681]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B9010172CC224C37F78FF864FEDCF810" bold="true" box="[199,322,1931,1957]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC224C37F601F864FDDAF810" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[329,580,1931,1957]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC224C37F601F864FDDAF810" box="[329,580,1931,1957]" italics="true" pageId="19" pageNumber="20">Lukeniana tubiraensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C75A6E3CC224C37F451F863FC5BF810" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1909" box="[793,965,1931,1957]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC224C37F451F863FC5BF810" box="[793,965,1931,1957]" italics="true" pageId="19" pageNumber="20">L. obliqualinea</emphasis>
</taxonomicName>
on the basis of the rectangular uncus and the well-developed thorn-like process. Four characters separate
<taxonomicName id="4C75A6E3CC224C37F42CF85FFC60F87C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[868,1022,1967,1993]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC224C37F42CF85FFC60F87C" box="[868,1022,1967,1993]" italics="true" pageId="19" pageNumber="20">L. tubiraensis</emphasis>
</taxonomicName>
from the latter species: the forewing upperside has a dark appearance and ground-colour of dark ochre intermixed with sepia (
<typeStatus id="54CE63C2CC224C37F332F83CFB79F858" box="[1146,1255,2003,2029]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
are slightly less sepia coloured); the thorn-like process is acuminate, strongly curved upward, extending well beyond the weaklysclerotized projection, hence,
<taxonomicName id="4C75A6E3CC254C30F6B9FF54FD10FF60" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[497,654,187,213]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC254C30F6B9FF54FD10FF60" box="[497,654,187,213]" italics="true" pageId="20" pageNumber="21">L. tubiraensis</emphasis>
</taxonomicName>
has the longest thorn-like process among species of
<taxonomicName id="4C75A6E3CC254C30F3B4FF54FAECFF60" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1276,1394,187,213]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B9010172CC254C30F3B4FF54FAECFF60" box="[1276,1394,187,213]" italics="true" pageId="20" pageNumber="21">Lukeniana</emphasis>
</taxonomicName>
(
<taxonomicName id="4C75A6E3CC254C30F2CEFF54FEBBFF4C" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1909" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC254C30F2CEFF54FEBBFF4C" italics="true" pageId="20" pageNumber="21">L. obliqualinea</emphasis>
</taxonomicName>
has a shorter thorn-like process which does not extend beyond the weakly-sclerotized projection); the gnathos arms are broad and short, almost equal in length to the width of valva, but are longer than in
<taxonomicName id="4C75A6E3CC254C30F3A3FEECFA0BFEA8" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1909" box="[1259,1429,259,285]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC254C30F3A3FEECFA0BFEA8" box="[1259,1429,259,285]" italics="true" pageId="20" pageNumber="21">L. obliqualinea</emphasis>
</taxonomicName>
; and the saccus is narrowly finger-shaped in
<taxonomicName id="4C75A6E3CC254C30F5CBFEC8FC82FEF4" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[643,796,295,321]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC254C30F5CBFEC8FC82FEF4" box="[643,796,295,321]" italics="true" pageId="20" pageNumber="21">L. tubiraensis</emphasis>
</taxonomicName>
but broad with a rounded tip in
<taxonomicName id="4C75A6E3CC254C30F3CEFEC8FAAFFEF4" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1909" box="[1158,1329,295,321]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC254C30F3CEFEC8FAAFFEF4" box="[1158,1329,295,321]" italics="true" pageId="20" pageNumber="21">L. obliqualinea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC254C30F78FFEA5FE77FD30" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BCADD60CC254C30F78FFEA5FD92FDD4" blockId="20.[151,1437,151,681]" pageId="20" pageNumber="21">
<emphasis id="B9010172CC254C30F78FFEA5FEC7FED1" bold="true" box="[199,345,330,356]" pageId="20" pageNumber="21">Distribution</emphasis>
.
<taxonomicName id="4C75A6E3CC254C30F62FFEA4FDC1FED0" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[359,607,331,357]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC254C30F62FFEA4FDC1FED0" box="[359,607,331,357]" italics="true" pageId="20" pageNumber="21">Lukeniana tubiraensis</emphasis>
</taxonomicName>
has a disjunct distribution, occurring in northwest and southwest
<collectingCountry id="F3629DF0CC254C30F270FEA4FA03FED0" box="[1336,1437,331,357]" name="Tanzania" pageId="20" pageNumber="21">Tanzania</collectingCountry>
in areas linked to the EARS. It is known from Tubira Forest patches ca.
<quantity id="4C8D7085CC254C30F490FE81FBBEFE3C" box="[984,1056,366,393]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="km" value="25.0">25 km</quantity>
east of Lake Tanganyika and the Tanganyika Rift.
<collectingRegion id="49B11382CC254C30F613FE7CFE32FE18" box="[347,428,403,429]" country="Tanzania" name="Mbeya" pageId="20" pageNumber="21">Mbeya</collectingRegion>
, where
<specimenCount id="9D7316E9CC254C30F54BFE7CFD0DFE18" box="[515,659,403,429]" pageId="20" pageNumber="21" type="generic" typeStatus="paratype">one paratype</specimenCount>
was collected, is located within the
<collectingRegion id="49B11382CC254C30F366FE7CFBE3FE18" box="[1070,1149,403,429]" country="Tanzania" name="Rukwa" pageId="20" pageNumber="21">Rukwa</collectingRegion>
Rift Basin
<emphasis id="B9010172CC254C30F3B7FE7CFAA3FE19" box="[1279,1341,403,428]" italics="true" pageId="20" pageNumber="21">sensu</emphasis>
<bibRefCitation id="EFE4A091CC254C30F20DFE7CFEBDFE64" author="Roberts, E. M. &amp; Stevens, N. J. &amp; O'Connor, P. M. &amp; Dirks, P. H. G. M. &amp; Gottfried, M. D. &amp; Clyde, W. C. &amp; Armstrong, R. A. &amp; Kemp, A. I. S. &amp; Hemming, S." pageId="20" pageNumber="21" pagination="289 - 294" refId="ref82238" refString="Roberts, E. M., Stevens, N. J., O'Connor, P. M., Dirks, P. H. G. M., Gottfried, M. D., Clyde, W. C., Armstrong, R. A., Kemp, A. I. S. &amp; Hemming, S. (2012) Initiation of the western branch of the East African Rift coeval with the eastern branch. Nature Geoscience, 5, 289 - 294. https: // doi. org / 10.1038 / NGEO 1432" type="journal article" year="2012">
Roberts
<emphasis id="B9010172CC254C30F7DFFE58FF51FE64" box="[151,207,439,465]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
(2012)
</bibRefCitation>
, ca.
<quantity id="4C8D7085CC254C30F61DFE59FE05FE64" box="[341,411,438,465]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="21" unit="km" value="85.0">85 km</quantity>
southeast of Lake
<collectingRegion id="49B11382CC254C30F525FE58FD22FE64" box="[621,700,439,465]" country="Tanzania" name="Rukwa" pageId="20" pageNumber="21">Rukwa</collectingRegion>
and ca.
<quantity id="4C8D7085CC254C30F451FE59FCF0FE64" box="[793,878,438,465]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="km" value="250.0">250 km</quantity>
east of Lake Tanganyika. Elevations of collecting localities range from
<quantity id="4C8D7085CC254C30F6C1FE35FDA9FE40" box="[393,567,474,501]" metricMagnitude="3" metricUnit="m" metricValue="1.4625" metricValueMax="1.835" metricValueMin="1.09" pageId="20" pageNumber="21" unit="m" value="1462.5" valueMax="1835.0" valueMin="1090.0">
<geoCoordinate id="EE41BBA7CC254C30F6C1FE35FE56FE40" box="[393,456,474,501]" degrees="1,090" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="55" value="1.09">1,090</geoCoordinate>
<geoCoordinate id="EE41BBA7CC254C30F681FE35FD88FE40" box="[457,534,474,501]" degrees="1,835" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="55" value="-1.835">1,835</geoCoordinate>
m
</quantity>
. The area between Tubira Forest and
<collectingRegion id="49B11382CC254C30F4AAFE34FBAEFE40" box="[994,1072,475,501]" country="Tanzania" name="Mbeya" pageId="20" pageNumber="21">Mbeya</collectingRegion>
represents a distance of ca.
<quantity id="4C8D7085CC254C30F23AFE35FF20FDAC" metricMagnitude="5" metricUnit="m" metricValue="5.8" pageId="20" pageNumber="21" unit="km" value="580.0">580 km</quantity>
. Because of its disjunct distribution it is possible that
<taxonomicName id="4C75A6E3CC254C30F46EFE10FC5FFDAC" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[806,961,511,537]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tubiraensis">
<emphasis id="B9010172CC254C30F46EFE10FC5FFDAC" box="[806,961,511,537]" italics="true" pageId="20" pageNumber="21">L. tubiraensis</emphasis>
</taxonomicName>
occurs in various scattered forest patches located between both localities. Based on its geographic distribution, this species is considered an Afromontane linking species as defined herein.
</paragraph>
<paragraph id="8BCADD60CC254C30F78FFD85FE77FD30" blockId="20.[151,1437,151,681]" box="[199,489,618,645]" pageId="20" pageNumber="21">
<emphasis id="B9010172CC254C30F78FFD85FEB4FD31" bold="true" box="[199,298,618,644]" pageId="20" pageNumber="21">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC254C30F78FFD60FBF2FD1C" box="[199,1132,655,681]" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BCADD60CC254C30F78FFD60FBF2FD1C" blockId="20.[151,1437,151,681]" box="[199,1132,655,681]" pageId="20" pageNumber="21">
<emphasis id="B9010172CC254C30F78FFD60FED0FD1C" bold="true" box="[199,334,655,681]" pageId="20" pageNumber="21">Etymology.</emphasis>
The new species is named for the Tubira Forest in northwest
<collectingCountry id="F3629DF0CC254C30F348FD60FBF6FD1C" box="[1024,1128,655,681]" name="Tanzania" pageId="20" pageNumber="21">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>