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<document id="30FB480C8384C769EE01699496E00ECC" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC134C07F7DFFAF9FBDCFA84" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana hausmanni Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="C195A4C6-F91B-4BDD-A141-76C057E1C04F" docUuidSource="ZooBank" docVersion="3" lastPageNumber="36" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="35" updateTime="1699487052491" updateUser="plazi">
<mods:mods id="C1001CC839151DF67B0E16D720466525" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="84A9D9EFAE0495545E0CC98D4E572599">
<mods:title id="85EE7429488079460AB416CDB4F901D8">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
</mods:titleInfo>
<mods:name id="506ACF6E8412FB9E436DA55D6E4C2470" type="personal">
<mods:role id="1269D55C631CDC0C14F0CDC8F0BD44AB">
<mods:roleTerm id="0C473852102933B0EC91DF315F797D29">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B3684C85CB2E8425009CA998E06B1576">Lehmann, Ingo</mods:namePart>
<mods:nameIdentifier id="9432A952EF02B4EB0C528B88F7386E34" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
</mods:name>
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<mods:role id="B16F620C923A94A1BB8F4A09F04AE13D">
<mods:roleTerm id="E5DE3D3EB78B841C2B3AFCCD36E805C7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="051A2754C2C424904A28C3F35B9D7B86">Zahiri, Reza</mods:namePart>
<mods:nameIdentifier id="F576FF6BE5A3E0D76EBC33B8DAE0213E" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E4D58ED9DC1A54BF8058DD181214B4EF" type="personal">
<mods:role id="C36B7F51D97101AC17CAA3C7E2EC75E7">
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</mods:role>
<mods:namePart id="2BAC36FADA6C96C8F707F86DB2E15BE3">Husemann, Martin</mods:namePart>
<mods:nameIdentifier id="B89DB6AE8C3A792F41135150E5C5A303" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo id="B802AE39ECA8505980E3BD8FD56B8AAC">
<mods:title id="2C9855CB12B1E24BE323F152D6225135">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E1E6742D8D476D02DFC8577BF56A06E9">
<mods:date id="72033B45FEDF7B7B51D1F722905B4C69">2023</mods:date>
<mods:detail id="22B3386AB4182BF1EA484DE49622F7B9" type="pubDate">
<mods:number id="C38845DB7D4C5B529285AA26D2FA8080">2023-04-18</mods:number>
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<mods:detail id="19492EADA9EA97F0ECACA7DE8D25578C" type="volume">
<mods:number id="6F0AC089C6FC375F50F7A6CE98421A2B">5267</mods:number>
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<mods:number id="E1632C0DB49EC70B823300F10D5B1A6C">1</mods:number>
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<mods:start id="C353FF7CFDA2A592267AC4FAE2425AF9">1</mods:start>
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<mods:url id="E0826EF6BCA93B2742E5CFF5EAD9EBDA">http://dx.doi.org/10.11646/zootaxa.5267.1.1</mods:url>
</mods:location>
<mods:classification id="280E91F2F03EB65946369BA95F4E9B84">journal article</mods:classification>
<mods:identifier id="870D0A88B7BF12494455B50F72B98CE3" type="DOI">10.11646/zootaxa.5267.1.1</mods:identifier>
<mods:identifier id="5C3A10B8F7074E6902999B083537C9AB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EC56436D6AA5B6C8128717481D0A0D42" type="Zenodo-Dep">7840783</mods:identifier>
<mods:identifier id="3A9EA46236186813395D85327472C720" type="ZooBank">9CD59054-8D7D-413F-B9FD-29EAFE7E511D</mods:identifier>
</mods:mods>
<treatment id="03DC6C76CC134C07F7DFFAF9FBDCFA84" ID-DOI="http://doi.org/10.5281/zenodo.7840760" ID-Zenodo-Dep="7840760" LSID="urn:lsid:zoobank.org:act:C195A4C6-F91B-4BDD-A141-76C057E1C04F" httpUri="http://treatment.plazi.org/id/03DC6C76CC134C07F7DFFAF9FBDCFA84" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="C36F8EEBCC134C06F7DFFAF9FC03FA84" box="[151,925,1302,1329]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BCADD60CC134C06F7DFFAF9FC03FA84" blockId="34.[151,975,1302,1401]" box="[151,925,1302,1329]" pageId="34" pageNumber="35">
<heading id="D0826A0CCC134C06F7DFFAF9FC03FA84" bold="true" box="[151,925,1302,1329]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
<emphasis id="B9010172CC134C06F7DFFAF9FC03FA84" bold="true" box="[151,925,1302,1329]" pageId="34" pageNumber="35">
<taxonomicName id="4C75A6E3CC134C06F7DFFAF9FCA6FA84" ID-CoL="B8R9J" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,824,1302,1329]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="hausmanni" status="sp. nov.">
<emphasis id="B9010172CC134C06F7DFFAF9FE32FA84" bold="true" box="[151,428,1302,1329]" italics="true" pageId="34" pageNumber="35">Lukeniana hausmanni</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC134C06F408FAF8FC03FA84" box="[832,925,1303,1329]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC134C06F7DFFAD4FBF7FA74" pageId="34" pageNumber="35" type="description">
<paragraph id="8BCADD60CC134C06F7DFFAD4FEBDFAE0" blockId="34.[151,975,1302,1401]" box="[151,291,1339,1365]" pageId="34" pageNumber="35">
<figureCitation id="134EC1E5CC134C06F7DFFAD4FF73FAE0" box="[151,237,1339,1365]" captionStart="FIGURE 6" captionStartId="37.[151,250,1791,1815]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-19@37.[151,1436,181,1767]" captionTargetPageId="37" captionText="FIGURE 6. a. Lukeniana kammeri, holotype, male, Kenya, County Kiambu, Thika; b. L. mbalaensis, paratype, female, Zambia, Northern Province, Mbala; c. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; g. L. utaheidenreichae, holotype, male, Mozambique, Manica Province, Barr De Chikamba Real; h. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840800" httpUri="https://zenodo.org/record/7840800/files/figure.png" pageId="34" pageNumber="35">Figs 6d</figureCitation>
,
<figureCitation id="134EC1E5CC134C06F7B0FAD4FEBDFAE0" box="[248,291,1339,1365]" captionStart="FIGURE 12" captionStartId="63.[151,250,1330,1354]" captionTargetBox="[160,1427,191,1293]" captionTargetId="figure-19@63.[151,1436,181,1306]" captionTargetPageId="63" captionText="FIGURE 12. a. Lukeniana mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; b. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; c. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840816" httpUri="https://zenodo.org/record/7840816/files/figure.png" pageId="34" pageNumber="35">12d</figureCitation>
</paragraph>
<paragraph id="8BCADD60CC134C06F7DFFAB1FC51FACC" blockId="34.[151,975,1302,1401]" box="[151,975,1374,1401]" pageId="34" pageNumber="35">
<uri id="FFE4D162CC134C06F7DFFAB1FC51FACC" box="[151,975,1374,1401]" pageId="34" pageNumber="35">
urn:lsid:zoobank.org:act:
<uuid id="FFD3E7B5CC134C06F6E5FAB1FC51FACC" box="[429,975,1374,1401]" pageId="34" pageNumber="35">C195A4C6-F91B-4BDD-A141-76C057E1C04F</uuid>
</uri>
</paragraph>
<paragraph id="8BCADD60CC134C06F7DFFA48FBF7FA74" blockId="34.[151,1129,1447,1473]" box="[151,1129,1447,1473]" pageId="34" pageNumber="35">
<materialsCitation id="3B1DD73DCC134C06F7DFFA48FBFBFA74" box="[151,1125,1447,1473]" collectionCode="ZSM" pageId="34" pageNumber="35" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC134C06F7DFFA48FF51FA74" box="[151,207,1447,1473]" pageId="34" pageNumber="35">Type</typeStatus>
locality and repository: Zamibia, the Zoological State Collection, Munich (
<collectionCode id="ED6445A5CC134C06F355FA48FBC2FA74" box="[1053,1116,1447,1473]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="34" pageNumber="35" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC134C06F7DFFA01FC61F92C" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BCADD60CC134C06F7DFFA01FC61F92C" blockId="34.[151,1437,1518,2013]" pageId="34" pageNumber="35">
<emphasis id="B9010172CC134C06F7DFFA01FE1AF9BD" bold="true" box="[151,388,1518,1544]" pageId="34" pageNumber="35">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC134C06F6C5FA00FF79F9C0" collectingDate="2009-10-13" collectionCode="ZSM" collectorName="J. Lenz &amp; I. Lehmann" country="Zambia" county="Mbala District" location="Mbala" municipality="Mbala" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Northern" typeStatus="holotype">
<typeStatus id="54CE63C2CC134C06F6C5FA00FE6BF9BC" box="[397,501,1519,1545]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC134C06F50FFA00FD01F9BC" box="[583,671,1519,1545]" name="Zambia" pageId="34" pageNumber="35">Zambia</collectingCountry>
,
<collectingRegion id="49B11382CC134C06F5E5FA00FC1CF9BC" box="[685,898,1519,1545]" country="Zambia" name="Northern" pageId="34" pageNumber="35">Northern Province</collectingRegion>
,
<collectingCounty id="62ABA5ECCC134C06F4D9FA00FBA7F9BC" box="[913,1081,1519,1545]" pageId="34" pageNumber="35">Mbala District</collectingCounty>
,
<quantity id="4C8D7085CC134C06F30FFA00FB0FF9BC" box="[1095,1169,1519,1545]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="35" unit="km" value="30.0">30 km</quantity>
southeast from
<collectingMunicipality id="6BAE471ACC134C06F206FA00FA06F9BC" box="[1358,1432,1519,1545]" pageId="34" pageNumber="35">Mbala</collectingMunicipality>
, 905′21 S, 3127′65 E (=
<emphasis id="B9010172CC134C06F6D6F9FCFE21F999" box="[414,447,1555,1580]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
<quantity id="4C8D7085CC134C06F68DF9FCFD92F998" box="[453,524,1555,1581]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="34" pageNumber="35" unit="km" value="23.0">23 km</quantity>
southeast
<location id="8EAA8BBBCC134C06F5C8F9FCFD56F998" LSID="urn:lsid:plazi:treatment:03DC6C76CC134C07F7DFFAF9FBDCFA84:8EAA8BBBCC134C06F5C8F9FCFD56F998" box="[640,712,1555,1581]" country="Zambia" county="Mbala District" municipality="Mbala" name="Mbala" pageId="34" pageNumber="35" stateProvince="Northern">Mbala</location>
and
<emphasis id="B9010172CC134C06F5B5F9FCFC80F999" box="[765,798,1555,1580]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
<quantity id="4C8D7085CC134C06F46CF9FCFCECF998" box="[804,882,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="34" pageNumber="35" unit="km" value="1.4">1.4 km</quantity>
southeast
<location id="8EAA8BBBCC134C06F4AEF9FCFB2EF998" LSID="urn:lsid:plazi:treatment:03DC6C76CC134C07F7DFFAF9FBDCFA84:8EAA8BBBCC134C06F4AEF9FCFB2EF998" box="[998,1200,1555,1581]" country="Zambia" county="Mbala District" municipality="Mbala" name="Chalumba village" pageId="34" pageNumber="35" stateProvince="Northern">Chalumba village</location>
),
<date id="FFCBFBA0CC134C06F38AF9FCFA1EF998" box="[1218,1408,1555,1581]" pageId="34" pageNumber="35" value="2009-10-13">
<collectingDate id="EF8F0248CC134C06F38AF9FCFA1EF998" box="[1218,1408,1555,1581]" pageId="34" pageNumber="35" value="2009-10-13">13 October 2009</collectingDate>
</date>
,
<collectorName id="2680B8B6CC134C06F2C2F9FCFF51F9E4" pageId="34" pageNumber="35">J. Lenz</collectorName>
leg.; second label (barcode): “BC
<collectionCode id="ED6445A5CC134C06F519F9D8FD15F9E4" box="[593,651,1591,1617]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="34" pageNumber="35" type="Museum">ZSM</collectionCode>
Lep33737”; third label: genitalia slide number 17/052015
<collectorName id="2680B8B6CC134C06F253F9D8FA02F9E4" box="[1307,1436,1591,1617]" pageId="34" pageNumber="35">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC134C06F7D7F9B4FF40F9C0" box="[159,222,1627,1653]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="34" pageNumber="35" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1DD73DCC134C06F7B9F9B4FC65F92C" collectingDate="2009-10-13" collectionCode="ZSM" collectorName="I. Lehmann" country="Zambia" county="Mbala District" location="Mbala" municipality="Mbala" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Northern" typeStatus="paratype">
<typeStatus id="54CE63C2CC134C06F7B9F9B4FECDF9C0" box="[241,339,1627,1653]" pageId="34" pageNumber="35" type="paratype">Paratype</typeStatus>
male,
<collectingCountry id="F3629DF0CC134C06F6D5F9B4FE6BF9C0" box="[413,501,1627,1653]" name="Zambia" pageId="34" pageNumber="35">Zambia</collectingCountry>
,
<collectingRegion id="49B11382CC134C06F548F9B4FD4CF9C0" box="[512,722,1627,1653]" country="Zambia" name="Northern" pageId="34" pageNumber="35">Northern Province</collectingRegion>
, same locality, same date and collector; second label: “BC
<collectionCode id="ED6445A5CC134C06F22BF9B4FA03F9C0" box="[1379,1437,1627,1653]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="34" pageNumber="35" type="Museum">ZSM</collectionCode>
Lep33735”; third label: genitalia slide number 08/122015
<collectorName id="2680B8B6CC134C06F46CF990FC3BF92C" box="[804,933,1663,1689]" pageId="34" pageNumber="35">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC134C06F4FCF990FC6DF92C" box="[948,1011,1663,1689]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="34" pageNumber="35" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC134C07F78FF94DFE3CFC8C" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="8BCADD60CC134C06F78FF94DFC96F8F8" blockId="34.[151,1437,1518,2013]" pageId="34" pageNumber="35">
<emphasis id="B9010172CC134C06F78FF94DFEC7F909" bold="true" box="[199,345,1698,1724]" pageId="34" pageNumber="35">Description.</emphasis>
Male.
<emphasis id="B9010172CC134C06F6E4F94CFE77F908" box="[428,489,1699,1725]" italics="true" pageId="34" pageNumber="35">Head</emphasis>
: Yellow-ochre mixed with some shiny sepia-coloured hair-like scales; long, dense hair-like scales between eyes; eyes vinaceous brown with small black patches; antenna 0.56‒0.58 length of forewing, bipectinate, with branches 6.5 width of shaft, covered with cream-coloured scales laterally, shaft covered with cream-coloured scales dorsally; antennal tips not spatulate, and with two long scales bending towards apex; labial palpi cream coloured, slightly longer than eye diameter.
</paragraph>
<paragraph id="8BCADD60CC134C07F78FF8B8FB42FE3C" blockId="34.[151,1437,1518,2013]" lastBlockId="35.[151,1437,151,1329]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B9010172CC134C06F78FF8B8FE86F8C4" box="[199,280,1879,1905]" italics="true" pageId="34" pageNumber="35">Thorax</emphasis>
: Patagia and tegulae with and long, shiny cream-coloured hair-like scales; a small cream-coloured metathoracic crest with ivory-yellow scales intermixed. Hindlegs yellow-ochre with fine, shiny hair-like scales; in
<typeStatus id="54CE63C2CC134C06F7DFF870FF67F80C" box="[151,249,1951,1977]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
and
<typeStatus id="54CE63C2CC134C06F678F870FE0EF80C" box="[304,400,1951,1977]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
two pairs of narrow tibial spurs present on hindlegs, outer spur of lower pair of
<typeStatus id="54CE63C2CC134C06F25AF870FAEAF80C" box="[1298,1396,1951,1977]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
<emphasis id="B9010172CC134C06F233F870FA02F80D" box="[1403,1436,1951,1976]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
1.0 mm, inner spur
<emphasis id="B9010172CC134C06F63CF82CFE08F869" box="[372,406,1987,2012]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
<quantity id="4C8D7085CC134C06F6D5F82CFE6BF868" box="[413,501,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="34" pageNumber="35" unit="mm" value="0.7">0.7 mm</quantity>
, outer spur of
<typeStatus id="54CE63C2CC134C06F5D0F82CFD66F868" box="[664,760,1987,2013]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
<emphasis id="B9010172CC134C06F448F82CFCBCF869" box="[768,802,1987,2012]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
<quantity id="4C8D7085CC134C06F461F82CFC1FF868" box="[809,897,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="35" unit="mm" value="0.8">0.8 mm</quantity>
, inner spur
<emphasis id="B9010172CC134C06F34EF82CFBB6F869" box="[1030,1064,1987,2012]" italics="true" pageId="34" pageNumber="35">ca.</emphasis>
<quantity id="4C8D7085CC134C06F367F82DFB16F868" box="[1071,1160,1986,2013]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="35" unit="mm" value="0.5">0.5 mm</quantity>
; outer spur in upper pair of
<typeStatus id="54CE63C2CC124C07F7FDFF78FE89FF04" box="[181,279,151,177]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
<emphasis id="B9010172CC124C07F656FF78FEA1FF05" box="[286,319,151,176]" italics="true" pageId="35" pageNumber="36">ca.</emphasis>
<quantity id="4C8D7085CC124C07F60DFF78FE03FF04" box="[325,413,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="35" pageNumber="36" unit="mm" value="0.9">0.9 mm</quantity>
, inner spur
<emphasis id="B9010172CC124C07F557FF78FDDEFF05" box="[543,576,151,176]" italics="true" pageId="35" pageNumber="36">ca.</emphasis>
<quantity id="4C8D7085CC124C07F50FFF78FD00FF04" box="[583,670,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="mm" value="0.3">0.3 mm</quantity>
, outer spur of
<typeStatus id="54CE63C2CC124C07F476FF78FC00FF04" box="[830,926,151,177]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
<emphasis id="B9010172CC124C07F4EDFF78FC59FF05" box="[933,967,151,176]" italics="true" pageId="35" pageNumber="36">ca.</emphasis>
<quantity id="4C8D7085CC124C07F485FF78FBBAFF04" box="[973,1060,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="35" pageNumber="36" unit="mm" value="0.6">0.6 mm</quantity>
, inner spur
<emphasis id="B9010172CC124C07F3EEFF78FB56FF05" box="[1190,1224,151,176]" italics="true" pageId="35" pageNumber="36">ca.</emphasis>
<quantity id="4C8D7085CC124C07F386FF78FAB9FF04" box="[1230,1319,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="mm" value="0.2">0.2 mm</quantity>
. Forewing length in
<typeStatus id="54CE63C2CC124C07F64BFF54FEFBFF60" box="[259,357,187,213]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
12.0 mm (wingspan
<quantity id="4C8D7085CC124C07F51DFF55FD20FF60" box="[597,702,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="35" pageNumber="36" unit="mm" value="25.5">25.5 mm</quantity>
), in
<typeStatus id="54CE63C2CC124C07F5B8FF54FCCEFF60" box="[752,848,187,213]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
<quantity id="4C8D7085CC124C07F410FF55FC22FF60" box="[856,956,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="35" pageNumber="36" unit="mm" value="11.5">11.5 mm</quantity>
(wingspan 25.0 mm). Forewing upperside cream-coloured mixed with ochre; costal margin without striae; termen without lunules or faded lunules; terminal and subterminal lines absent in both males; CuA
<subScript id="17F1DF25CC124C07F5F2FEFAFD5CFE96" attach="left" box="[698,706,277,291]" fontSize="6" pageId="35" pageNumber="36">2</subScript>
light ochre, edged broadly with sepia above; all remaining veins cream-coloured hence indistinct; cilia long in forewing and hindwing,
<quantity id="4C8D7085CC124C07F498FEC8FBB7FEF4" box="[976,1065,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="mm" value="1.2">1.2 mm</quantity>
in both males, cream coloured, glossy. Underside of forewing roughly scaled with many fine hair-like cream-coloured glossy scales; costal margin dark ochre, without striae. Hindwing upperside cream coloured, glossy; underside as in forewing.
</paragraph>
<paragraph id="8BCADD60CC124C07F78FFE7CFF59FCA0" blockId="35.[151,1437,151,1329]" pageId="35" pageNumber="36">
<emphasis id="B9010172CC124C07F78FFE7CFEAFFE18" box="[199,305,403,429]" italics="true" pageId="35" pageNumber="36">Abdomen</emphasis>
: Cream-coloured mixed with ochre, shiny; abdominal tuft short, one-fifth abdomen length. Genitalia (
<figureCitation id="134EC1E5CC124C07F7D7FE58FE9EFE64" box="[159,256,439,465]" captionStart="FIGURE 12" captionStartId="63.[151,250,1330,1354]" captionTargetBox="[160,1427,191,1293]" captionTargetId="figure-19@63.[151,1436,181,1306]" captionTargetPageId="63" captionText="FIGURE 12. a. Lukeniana mzuzuensis, holotype, male, Malawi, Northern Region, Mzuzu, Nkhorongo; b. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; c. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840816" httpUri="https://zenodo.org/record/7840816/files/figure.png" pageId="35" pageNumber="36">Fig. 12d</figureCitation>
) with uncus lobes subtriangular, with short and long setae ventrally, basal edge of uncus almost straight (slightly bent at middle, ventral view); gnathos arms 1.4 longer than basal width of valva; valva rectangular, costa with few setae; sacculus with few setae; inner side of valva with short setae except in centre, weakly-sclerotized projection setose, with a subrectangular tip longer than single thorn-like process below; the latter well developed, hollow, thick, bent and with few scattered tiny setae; long, rounded emargination extending 45% of length of valva between weakly-sclerotized projection and thorn-like process; ventral side of valva not bent at middle. Saccus long, narrowly acuminate, attached to a broader ventral part of vinculum that forms no plate. Juxta not longer than saccus, with two acuminate tips, each bearing a tiny process, between tips a deep emargination extending 90% of length of juxta. Phallus short, 1.2 as long as width of valva, not trumpet-like, not bent at middle, bilobate with cleft on each end.
</paragraph>
<paragraph id="8BCADD60CC124C07F78FFCF0FE3CFC8C" blockId="35.[151,1437,151,1329]" box="[199,418,799,825]" pageId="35" pageNumber="36">
<emphasis id="B9010172CC124C07F78FFCF0FEBBFC8C" bold="true" box="[199,293,799,825]" pageId="35" pageNumber="36">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC124C07F78FFCADFBEDFC58" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BCADD60CC124C07F78FFCADFBEDFC58" blockId="35.[151,1437,151,1329]" pageId="35" pageNumber="36">
<emphasis id="B9010172CC124C07F78FFCADFEDCFCE9" bold="true" box="[199,322,834,860]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC124C07F601FCACFDDCFCE8" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[329,578,835,861]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="hausmanni">
<emphasis id="B9010172CC124C07F601FCACFDDCFCE8" box="[329,578,835,861]" italics="true" pageId="35" pageNumber="36">Lukeniana hausmanni</emphasis>
</taxonomicName>
has two characters that separate it from all remaining species of
<taxonomicName id="4C75A6E3CC124C07F256FCACFA0AFCE8" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1310,1428,835,861]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B9010172CC124C07F256FCACFA0AFCE8" box="[1310,1428,835,861]" italics="true" pageId="35" pageNumber="36">Lukeniana</emphasis>
</taxonomicName>
: the hindlegs have two pairs of tibial spurs, of which the inner spur of upper pair is strongly reduced; and the lobes of the uncus are nearly triangular with rounded tips. Two characters in the shape of the uncus lobes and saccus are similar only to
<taxonomicName id="4C75A6E3CC124C07F60CFC40FE4BFC7C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[324,469,943,969]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="enaiposha">
<emphasis id="B9010172CC124C07F60CFC40FE4BFC7C" box="[324,469,943,969]" italics="true" pageId="35" pageNumber="36">L. enaiposha</emphasis>
</taxonomicName>
(see diagnosis). Both species have a well-developed, thorn-like process and no platelike structure associated with the vinculum, from which the thorn-like saccus originates.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC124C07F78FFC19FE77FB70" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BCADD60CC124C07F78FFC19FE10FB14" blockId="35.[151,1437,151,1329]" pageId="35" pageNumber="36">
<emphasis id="B9010172CC124C07F78FFC19FEFFFBA5" bold="true" box="[199,353,1014,1040]" pageId="35" pageNumber="36">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC124C07F62FFC18FDC1FBA4" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[359,607,1015,1041]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="hausmanni">
<emphasis id="B9010172CC124C07F62FFC18FDC1FBA4" box="[359,607,1015,1041]" italics="true" pageId="35" pageNumber="36">Lukeniana hausmanni</emphasis>
</taxonomicName>
is known only from Chalumba village (elevation
<quantity id="4C8D7085CC124C07F3C3FC19FB76FBA4" box="[1163,1256,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.565" pageId="35" pageNumber="36" unit="m" value="1565.0">1,565 m</quantity>
, average annual rainfall
<emphasis id="B9010172CC124C07F7A7FBF4FE8FFB81" box="[239,273,1051,1076]" italics="true" pageId="35" pageNumber="36">ca.</emphasis>
<quantity id="4C8D7085CC124C07F650FBF4FE19FB80" box="[280,391,1051,1077]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="mm" value="1200.0">1200 mm</quantity>
), located near, but still on, the northeastern edge of the Southern African
<collectingRegion id="49B11382CC124C07F3F6FBF4FA8EFB80" box="[1214,1296,1051,1077]" country="Nigeria" name="Plateau" pageId="35" pageNumber="36">Plateau</collectingRegion>
south of the Tanganyika Rift and
<emphasis id="B9010172CC124C07F6CEFBD0FE36FBED" box="[390,424,1087,1112]" italics="true" pageId="35" pageNumber="36">ca.</emphasis>
<quantity id="4C8D7085CC124C07F6F8FBD0FE66FBEC" box="[432,504,1087,1113]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="km" value="20.0">20 km</quantity>
south of the Albertine Rift Region (
<emphasis id="B9010172CC124C07F4DEFBD0FC4AFBED" box="[918,980,1087,1112]" italics="true" pageId="35" pageNumber="36">sensu</emphasis>
<bibRefCitation id="EFE4A091CC124C07F495FBD0FB56FBEC" author="Plumptre, A. J. &amp; Davenport, T. R. B. &amp; Behangana, M. &amp; Kityo, R. &amp; Eilu, G. &amp; Ssegawa, P. &amp; Ewango, C. &amp; Meirte, D. &amp; Kahindo, C. &amp; Herremans, M. &amp; Kerbis Peterhans, J. &amp; Pilgrim, J. D. &amp; Wilson, M. &amp; Languy, M. &amp; Moyer, D." box="[989,1224,1087,1113]" pageId="35" pageNumber="36" pagination="178 - 194" refId="ref81834" refString="Plumptre, A. J., Davenport, T. R. B., Behangana, M., Kityo, R., Eilu, G., Ssegawa, P., Ewango, C., Meirte, D., Kahindo, C., Herremans, M., Kerbis Peterhans, J., Pilgrim, J. D., Wilson, M., Languy, M. &amp; Moyer, D. (2007) The biodiversity of the Albertine Rift. Biological Conservation, 134, 178 - 194. https: // doi. org / 10.1016 / j. biocon. 2006.08.021" type="journal article" year="2007">
Plumptre
<emphasis id="B9010172CC124C07F304FBD0FB18FBEC" box="[1100,1158,1087,1113]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2007
</bibRefCitation>
) in the Zambezian regional centre of endemism (
<emphasis id="B9010172CC124C07F6A4FB8CFDB4FBC9" box="[492,554,1123,1148]" italics="true" pageId="35" pageNumber="36">sensu</emphasis>
<bibRefCitation id="EFE4A091CC124C07F57AFB8CFD29FBC8" author="White, F." box="[562,695,1123,1149]" pageId="35" pageNumber="36" refId="ref83858" refString="White, F. (1983) The Vegetation of Africa: a Descriptive Memoir to Accompany the Unesco / AETFAT / UNSO Vegetation Map of Africa. Natural Resources Research No. XX. Unesco, Paris, 356 pp." type="book" year="1983">White 1983</bibRefCitation>
) (
<figureCitation id="134EC1E5CC124C07F598FB8CFCAAFBC8" box="[720,820,1123,1149]" captionStart="FIGURE 20" captionStartId="72.[151,250,1115,1139]" captionTargetBox="[151,1436,181,1091]" captionTargetId="figure-28@72.[151,1436,181,1091]" captionTargetPageId="72" captionText="FIGURE 20. a. Habitat of Lukeniana enaiposha with Rothschild’s Giraffes, Giraffa camelopardalis rothschildi Lydekker (1903) graze in a woodland site at 1.786 m elevation within Lake Nakuru National Park (Kenya) ca. 1 km south of the lake dominated by tall (&gt; 15 m) and young trees of Acacia xanthophloea Benth. (Fabaceae: Mimosoideae) mixed with shrubs of Rhus natalensis Bernh. (Anacardiaceae) (right) and Solanum incanum L. (Solanaceae) (foreground). Picture taken by Shimoni Lehmann in March 2016. b. Between 1600 and 2000 m Lower montane Cassipourea forest sensu Hemp (2006b) covers the slopes of Mount Meru (4.566 m). The picture shows an undisturbed forest of this type comprising trees of Ficus thonningii Blume (Moraceae), Olea capensis L. (Oleaceae) and Ekebergia capensis Sparrm. (Meliaceae) with smaller trees of Vepris simplicifolia (Verd.) Mziray (Rutaceae) and Casearia battiscombei R.E. Fries (Flacourtiaceae). Ferns are present on the ground indicating a wetter site with species of Pteris dentata Forssk. (Pteridaceae) and Asplenium aethiopicum (Burm. f.) Bech. (Aspleniaceae). Note that Lukeniana timdavenporti occurs in a similar habitat on Mount Elgon (Kenya/Uganda), but is not closely related to L. andreashempi and L. michaelgrzimeki, that both occur on Mount Meru. Picture by Dr. Andreas Hemp. c. Habitat of Lukeniana mzuzuensis: wetter, ca. 50 years old Zambezian miombo woodland sensu White (1983) at Nkhorongo (Malawi) with diverse structure, high species diversity, e.g. leguminous trees. The treelet with broad leaves in the bottom right corner is Uapaca nitida Müll. Arg. (Euphorbiaceae); in the centre left is U. kirkiana Müll. Arg.; background trees are Brachystegia utilis Burtt Davy &amp; Hutch., B. floribunda Benth., B. spiciformis Benth. (Fabaceae: Caesalpiniaceae), Uapaca sansibarica Pax., Julbernardia paniculata (Benth.) Troupin (Caesalpiniaceae), Dalbergia melanoxylon Guill. &amp; Perr. (Papilionaceae). In the top right centre is Cussonia arborea Hochst. (Araliaceae) with light green leaves. Picture by Raymond Murphy (Nkhorongo), May 2016. d. Road ca. 23 km southeast Mbala (Zambia), close to the habitat of Lukeniana hausmanni, on the Southern African Plateau with northern wet miombo sensu Chidumayo (1997); Brachystegia-Isoberlinia-Julbernardia association mixed with Albizia antunesiana Harms (Fabaceae: Mimosoideae). Picture by Jürgen Lenz (Leipzig), October 2009." figureDoi="http://doi.org/10.5281/zenodo.7840837" httpUri="https://zenodo.org/record/7840837/files/figure.png" pageId="35" pageNumber="36">Fig. 20d</figureCitation>
).
<taxonomicName id="4C75A6E3CC124C07F400FB8CFBDFFBC8" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[840,1089,1123,1149]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="hausmanni">
<emphasis id="B9010172CC124C07F400FB8CFBDFFBC8" box="[840,1089,1123,1149]" italics="true" pageId="35" pageNumber="36">Lukeniana hausmanni</emphasis>
</taxonomicName>
is considered an Afromontane near-endemic species.
</paragraph>
<paragraph id="8BCADD60CC124C07F78FFB45FE77FB70" blockId="35.[151,1437,151,1329]" box="[199,489,1194,1221]" pageId="35" pageNumber="36">
<emphasis id="B9010172CC124C07F78FFB45FEB4FB71" bold="true" box="[199,298,1194,1220]" pageId="35" pageNumber="36">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC124C07F78FFB20FBDCFA84" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8BCADD60CC124C07F78FFB20FBDCFA84" blockId="35.[151,1437,151,1329]" pageId="35" pageNumber="36">
<emphasis id="B9010172CC124C07F78FFB20FED0FB5C" bold="true" box="[199,334,1231,1257]" pageId="35" pageNumber="36">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC124C07F61DFB20FDD0FB5C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[341,590,1231,1257]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="hausmanni">
<emphasis id="B9010172CC124C07F61DFB20FDD0FB5C" box="[341,590,1231,1257]" italics="true" pageId="35" pageNumber="36">Lukeniana hausmanni</emphasis>
</taxonomicName>
is named after Dr. Axel Hausmann, head of the
<taxonomicName id="4C75A6E3CC124C07F338FB20FB69FB5C" box="[1136,1271,1231,1257]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
Section of the ZSM. The first author is grateful to Axel for his support during the studies on the
<taxonomicName id="4C75A6E3CC124C07F357FB1CFB28FAB8" baseAuthorityName="Lehmann" baseAuthorityYear="2019" box="[1055,1206,1267,1293]" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
, and for the comprehensive loans of more than 50 undescribed species from the collections at the ZSM.
</paragraph>
</subSubSection>
</treatment>
</document>
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