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<document ID-DOI="10.5252/cryptogamie-mycologie2023v44a2" ID-ISSN="1776-100X" ID-Zenodo-Dep="7815402" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1681179038539" checkinUser="felipe" docAuthor="A, Lidiane, Santos, lves dos, Aptroot, André, F, Maria, Souza, ernanda de, Lücking, Robert, Guzmán-Guillermo, Jorge, E, Marcela, S, ugenia da &amp; Cáceres, ilva" docDate="2023" docId="03A54068CE39932AB676FA8AFBCBBBEC" docLanguage="en" docName="CryptogamieMycologie.44.2.11-25.pdf" docOrigin="Cryptogamie, Mycologie 20 (2)" docSource="http://dx.doi.org/10.5252/cryptogamie-mycologie2023v44a2" docStyle="DocumentStyle:81F49AB8619EB7E5D11A6328441E2653.1:CryptogamieMycologie.2019-.journal_article" docStyleId="81F49AB8619EB7E5D11A6328441E2653" docStyleName="CryptogamieMycologie.2019-.journal_article" docStyleVersion="1" docTitle="Tephromela obesimarginata L. A. Santos, Aptroot &amp; M. Caceres 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="18" masterDocId="FF9C3810CE309323B758FFB9FF8ABF42" masterDocTitle="Four new species of Tephromela M. Choisy (Ascomycota, Tephromelataceae), three containing lichexanthone, from Brazil and Mexico" masterLastPageNumber="25" masterPageNumber="11" pageNumber="18" updateTime="1681498483861" updateUser="juliana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Four new species of Tephromela M. Choisy (Ascomycota, Tephromelataceae), three containing lichexanthone, from Brazil and Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>A, Lidiane</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, lves dos</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aptroot, André</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>F, Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Souza, ernanda de</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lücking, Robert</mods:namePart>
<mods:affiliation>Botanischer Garten, Freie Universität Berlin, KÖnigin-Luise-Strasse 6 - 8, 14195 Berlin (Germany)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guzmán-Guillermo, Jorge</mods:namePart>
<mods:affiliation>Facultad de Ciencias Biológicas y Agropecuarías, Universidad Veracruzana, Carretera Peñuela-Amatlán Kilómetro 177, 94500 Córdoba, Veracruz (Mexico)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>E, Marcela</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>S, ugenia da</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cáceres, ilva</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Cryptogamie, Mycologie</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>20</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>11</mods:start>
<mods:end>25</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.5252/cryptogamie-mycologie2023v44a2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/cryptogamie-mycologie2023v44a2</mods:identifier>
<mods:identifier type="ISSN">1776-100X</mods:identifier>
<mods:identifier type="Zenodo-Dep">7815402</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7815438" ID-Zenodo-Dep="7815438" LSID="urn:lsid:plazi:treatment:03A54068CE39932AB676FA8AFBCBBBEC" httpUri="http://treatment.plazi.org/id/03A54068CE39932AB676FA8AFBCBBBEC" lastPageNumber="18" pageId="9" pageNumber="18">
<subSubSection pageId="9" pageNumber="18" type="nomenclature">
<paragraph blockId="9.[220,688,1331,1390]" pageId="9" pageNumber="18">
<heading centered="true" fontSize="11" level="2" pageId="9" pageNumber="18" reason="4">
<taxonomicName authority="L.A.Santos, Aptroot &amp; M.Caceres" authorityName="L. A. Santos, Aptroot &amp; M. Caceres" authorityYear="2023" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="9" pageNumber="18" phylum="Ascomycota" rank="species" species="obesimarginata" status="sp. nov.">
<emphasis bold="true" box="[302,605,1331,1358]" italics="true" pageId="9" pageNumber="18">Tephromela obesimarginata</emphasis>
L.A.Santos, Aptroot &amp; M.Cáceres
</taxonomicName>
,
<taxonomicNameLabel box="[603,688,1364,1390]" pageId="9" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="description">
<paragraph blockId="9.[350,564,1395,1422]" box="[350,564,1395,1422]" pageId="9" pageNumber="18">
<heading box="[350,564,1395,1422]" centered="true" fontSize="11" level="3" pageId="9" pageNumber="18" reason="3">
(
<figureCitation box="[359,481,1395,1422]" captionStart="FIG" captionStartId="10.[132,143,1939,1957]" captionTargetBox="[238,1347,215,1878]" captionTargetId="figure-21@10.[236,793,769,1325]" captionTargetPageId="10" captionText="FIG. 3. — Habitus and UV reaction: A, B, Tephromela multireflexa Aptroot &amp; M.F.Souza, sp. nov. (holotype), apothecial margins UV+ yellow and UV+ bluish green; C, D, Tephromela obesimarginata L.A.Santos, Aptroot &amp; M.Cáceres, sp. nov. (holotype), thallus and apothecium margin UV+ yellow; E, F, Tephromela vinacea L.A.Santos, Aptroot, Lücking &amp; M.Cáceres, sp. nov.(holotype), medulla UV+ greenish white; G, H, Tephromela xanthonica Guzmán-Guillermo,sp.nov. (holotype), thallus and apothecium margin UV+ yellow. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7815406" httpUri="https://zenodo.org/record/7815406/files/figure.png" pageId="9" pageNumber="18">Figs 3C, D</figureCitation>
;
<figureCitation box="[493,556,1395,1422]" captionStart="FIG" captionStartId="12.[132,143,1738,1756]" captionTargetBox="[132,1455,219,1692]" captionTargetId="figure-89@12.[698,1267,769,1325]" captionTargetPageId="12" captionText="FIG. 4. — Section ascomata: A, B, Tephromela obesimarginata L.A.Santos, Aptroot &amp; M.Cáceres, sp. nov. (holotype), before and after KOH treatment; C, D, Tephromela vinacea L.A.Santos, Aptroot, Lücking &amp; M.Cáceres, sp. nov. (holotype), before and after KOH treatment; E, F, Tephromela xanthonica Guzmán- Guillermo, sp. nov. (holotype), before and after KOH treatment. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7815408" httpUri="https://zenodo.org/record/7815408/files/figure.png" pageId="9" pageNumber="18">4A, B</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="9.[132,777,1452,1528]" pageId="9" pageNumber="18">
Similar to
<taxonomicName box="[243,448,1452,1476]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="9" pageNumber="18" phylum="Ascomycota" rank="species" species="velloziae">
<emphasis box="[243,448,1452,1476]" italics="true" pageId="9" pageNumber="18">Tephromela velloziae</emphasis>
</taxonomicName>
but corticolous and differing in the unevenly thickened thalline apothecium margin and the larger ascospores.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="materials_examined">
<paragraph blockId="9.[130,776,1553,1630]" pageId="9" pageNumber="18">
<materialsCitation collectingDate="2021-05-17" collectingDateMax="2021-05-25" collectingDateMin="2021-05-17" collectionCode="ISE, CGMS" collectorName="L. A. Santos &amp; A. Aptroot" country="Brazil" elevation="1300" latitude="-20.1" location="Caraca" longLatPrecision="1269" longitude="-43.483334" municipality="Catas Altas" pageId="9" pageNumber="18" specimenCode="ISE52321" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus box="[132,238,1553,1576]" pageId="9" pageNumber="18">HOLOTYPE</typeStatus>
. —
<emphasis bold="true" box="[277,335,1554,1578]" pageId="9" pageNumber="18">
<collectingCountry box="[277,335,1554,1578]" name="Brazil" pageId="9" pageNumber="18">Brazil</collectingCountry>
</emphasis>
.
<collectingRegion box="[348,478,1553,1577]" country="Brazil" name="Minas Gerais" pageId="9" pageNumber="18">Minas Gerais</collectingRegion>
,
<collectingMunicipality box="[486,596,1553,1577]" pageId="9" pageNumber="18">Catas Altas</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03A54068CE39932AB676FA8AFBCBBBEC:8ED3A7A5CE39932AB504F9A8FD2BB96B" box="[604,673,1553,1577]" country="Brazil" latitude="-20.1" longLatPrecision="1269" longitude="-43.483334" municipality="Catas Altas" name="Caraca" pageId="9" pageNumber="18" stateProvince="Minas Gerais">Caraça</location>
, alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="9" pageNumber="18" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="9" pageNumber="18" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200- 1400 m</elevation>
</quantity>
,
<geoCoordinate box="[213,288,1579,1603]" degrees="20" direction="south" minutes="06" orientation="latitude" pageId="9" pageNumber="18" precision="925" value="-20.1">20°06S</geoCoordinate>
,
<geoCoordinate box="[296,380,1579,1603]" degrees="43" direction="west" minutes="29" orientation="longitude" pageId="9" pageNumber="18" precision="925" value="-43.483334">43°29W</geoCoordinate>
, on tree bark,
<date box="[510,642,1579,1603]" pageId="9" pageNumber="18" value="2021-05-17" valueMax="2021-05-25" valueMin="2021-05-17">
<collectingDate box="[510,642,1579,1603]" pageId="9" pageNumber="18" value="2021-05-17" valueMax="2021-05-25" valueMin="2021-05-17">17-25.V.2021</collectingDate>
</date>
,
<emphasis italics="true" pageId="9" pageNumber="18">
<collectorName box="[648,751,1579,1602]" pageId="9" pageNumber="18">L.A. Santos</collectorName>
&amp;
<collectorName box="[130,227,1606,1629]" pageId="9" pageNumber="18">A. Aptroot</collectorName>
s.n.
</emphasis>
(holo-,
<collectionCode box="[347,386,1605,1629]" country="Brazil" httpUri="http://grbio.org/cool/jqe8-8hrg" name="Universidade Federal de Sergipe, Campus Professor Alberto Carvalho" pageId="9" pageNumber="18" type="Herbarium">ISE</collectionCode>
[
<specimenCode box="[392,495,1605,1629]" collectionCode="ISE" country="Brazil" httpUri="http://grbio.org/cool/jqe8-8hrg" name="Universidade Federal de Sergipe, Campus Professor Alberto Carvalho" pageId="9" pageNumber="18" type="Herbarium">ISE52321</specimenCode>
];
</materialsCitation>
<materialsCitation box="[511,646,1605,1630]" collectingDate="2021-05-17" collectingDateMax="2021-05-25" collectingDateMin="2021-05-17" collectionCode="ISE, CGMS" collectorName="L. A. Santos &amp; A. Aptroot" country="Brazil" elevation="1300" latitude="-20.1" location="Caraca" longLatPrecision="1269" longitude="-43.483334" municipality="Catas Altas" pageId="9" pageNumber="18" specimenCode="ISE52321" specimenCount="1" stateProvince="Minas Gerais" typeStatus="isotype">
iso-,
<collectionCode box="[558,634,1606,1630]" collectionName="CGMS" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15107" lsid="urn:lsid:biocol.org:col:15107" name="Universidade Federal de Mato Grosso do Sul" pageId="9" pageNumber="18" type="Herbarium">CGMS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="9.[131,775,1653,1757]" pageId="9" pageNumber="18">
ADDITIONAL SPECIMENS EXAMINED. —
<materialsCitation collectingDate="2021-05-17" collectingDateMax="2021-05-25" collectingDateMin="2021-05-17" collectionCode="ISE, CGMS" collectorName="L. A. Santos &amp; A. Aptroot" country="Brazil" elevation="1300" latitude="-20.1" location="Caraca" longLatPrecision="1269" longitude="-43.483334" municipality="Catas Altas" pageId="9" pageNumber="18" specimenCount="1" stateProvince="Minas Gerais">
<emphasis bold="true" box="[510,568,1655,1679]" pageId="9" pageNumber="18">
<collectingCountry box="[510,568,1655,1679]" name="Brazil" pageId="9" pageNumber="18">Brazil</collectingCountry>
</emphasis>
.
<collectingRegion box="[581,712,1653,1677]" country="Brazil" name="Minas Gerais" pageId="9" pageNumber="18">Minas Gerais</collectingRegion>
,
<collectingMunicipality pageId="9" pageNumber="18">Catas Altas</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03A54068CE39932AB676FA8AFBCBBBEC:8ED3A7A5CE39932AB799F929FE8DB9EA" box="[193,263,1680,1704]" country="Brazil" latitude="-20.1" longLatPrecision="1269" longitude="-43.483334" municipality="Catas Altas" name="Caraca" pageId="9" pageNumber="18" stateProvince="Minas Gerais">Caraça</location>
, alt.
<quantity box="[310,449,1680,1704]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="9" pageNumber="18" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">
<elevation box="[310,449,1680,1704]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="9" pageNumber="18" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200-1400 m</elevation>
</quantity>
,
<geoCoordinate box="[462,541,1680,1704]" degrees="20" direction="south" minutes="06" orientation="latitude" pageId="9" pageNumber="18" precision="925" value="-20.1">20°06S</geoCoordinate>
,
<geoCoordinate box="[551,637,1680,1704]" degrees="43" direction="west" minutes="29" orientation="longitude" pageId="9" pageNumber="18" precision="925" value="-43.483334">43°29W</geoCoordinate>
, on tree bark,
<date box="[133,276,1706,1730]" pageId="9" pageNumber="18" value="2021-05-17" valueMax="2021-05-25" valueMin="2021-05-17">
<collectingDate box="[133,276,1706,1730]" pageId="9" pageNumber="18" value="2021-05-17" valueMax="2021-05-25" valueMin="2021-05-17">17-25.V.2021</collectingDate>
</date>
,
<emphasis box="[288,614,1706,1730]" italics="true" pageId="9" pageNumber="18">
<collectorName box="[288,541,1706,1729]" pageId="9" pageNumber="18">L.A. Santos &amp; A. Aptroot</collectorName>
52143
</emphasis>
,
<emphasis box="[629,694,1706,1730]" italics="true" pageId="9" pageNumber="18">52312</emphasis>
,
<emphasis box="[710,775,1706,1730]" italics="true" pageId="9" pageNumber="18">52309</emphasis>
(
<collectionCode box="[139,178,1733,1757]" country="Brazil" httpUri="http://grbio.org/cool/jqe8-8hrg" name="Universidade Federal de Sergipe, Campus Professor Alberto Carvalho" pageId="9" pageNumber="18" type="Herbarium">ISE</collectionCode>
,
<collectionCode box="[189,265,1733,1757]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15107" lsid="urn:lsid:biocol.org:col:15107" name="Universidade Federal de Mato Grosso do Sul" pageId="9" pageNumber="18" type="Herbarium">CGMS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="etymology">
<paragraph blockId="9.[131,775,1780,1831]" pageId="9" pageNumber="18">ETYMOLOGY. — The species is named after the irregularly thickened, plump, thalline apothecium margin.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="conservation">
<paragraph blockId="9.[131,775,1854,1905]" pageId="9" pageNumber="18">CHEMISTRY. — Thallus UV+ yellow, C-, P-, K-, apothecium margin UV+ yellow. With lichexanthone.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="biology_ecology">
<paragraph blockId="9.[132,775,1928,1979]" pageId="9" pageNumber="18">
ECOLOGY AND DISTRIBUTION. — On exposed trees in Campo rupestre vegetation in SE
<collectingCountry box="[349,408,1955,1979]" name="Brazil" pageId="9" pageNumber="18">Brazil</collectingCountry>
; known only from
<collectingCountry box="[595,654,1955,1979]" name="Brazil" pageId="9" pageNumber="18">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="description">
<paragraph blockId="9.[132,422,2002,2026]" box="[132,422,2002,2026]" pageId="9" pageNumber="18">MYCOBANK. — MB 846315.</paragraph>
<paragraph blockId="9.[813,1073,216,240]" box="[813,1073,216,240]" pageId="9" pageNumber="18">GENBANK. — OP881898.</paragraph>
<paragraph blockId="9.[813,957,280,306]" box="[813,957,280,306]" pageId="9" pageNumber="18">
<heading box="[813,957,280,306]" fontSize="9" level="3" pageId="9" pageNumber="18" reason="5">DESCRIPTION</heading>
</paragraph>
<paragraph blockId="9.[810,1457,311,1006]" pageId="9" pageNumber="18">
Thallus crustose, whitish to pale ochraceous-grey, rimose to areolate, verrucose,somewhat glossy,
<emphasis box="[1182,1191,344,369]" italics="true" pageId="9" pageNumber="18">c</emphasis>
.
<quantity box="[1204,1285,343,369]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="18" unit="mm" value="0.1">0.1 mm</quantity>
thick.Prothallus absent. Photobiont an unicellular green alga, arranged in one layer below the cortex, algal layer
<emphasis box="[1180,1189,407,432]" italics="true" pageId="9" pageNumber="18">c</emphasis>
. 50 µm high. Apothecia always present, sessile, roundish, homogeneously distributed on the thallus, with unevenly crenate, irregularly thickened margin. Disk black,
<quantity box="[1041,1153,502,529]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="9" pageNumber="18" unit="mm" value="1.35" valueMax="2.0" valueMin="0.7">0.7-2 mm</quantity>
in diam., flat to somewhat concave or saddle-shaped, epruinose. Thalline margin concolorous with the thallus,
<emphasis box="[1096,1105,567,592]" italics="true" pageId="9" pageNumber="18">c</emphasis>
.
<quantity box="[1120,1250,566,593]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="9" pageNumber="18" unit="mm" value="0.4" valueMax="0.6" valueMin="0.2">0.2-0.6 mm</quantity>
wide, raised above the level of the disk (100-150 µm). Proper exciple thickest at the base, becoming thinner at the edges, pale to intense yellow-brown, intensifying yellow in K. Exciple with crystals that after treatment with K partly or completely dissolve. Hymenium
<emphasis box="[947,956,726,751]" italics="true" pageId="9" pageNumber="18">c</emphasis>
. 100-150 µm thick, violet, K+ greenish-grey. Epihymenium violet, K+ greenish-grey. Paraphyses simple, septate, 2-3 µm wide, with 3-5 µm wide hyaline apical cells, surrounded by violet pigmented gel.Hypothecium 100-150 µm thick, violet, K+ greenish-grey, with a narrow golden-brown band below the hymenium (25-50 µm thick). Asci slightly clavate, 50-75 µm tall, 8-spored. Ascospores simple, hyaline, ellipsoid, 14-16 × 6.5-7.5 µm. Pycnidia black. Conidia not observed.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="18" type="discussion">
<paragraph blockId="9.[813,942,1045,1071]" box="[813,942,1045,1071]" pageId="9" pageNumber="18">
<heading allCaps="true" box="[813,942,1045,1071]" fontSize="9" level="2" pageId="9" pageNumber="18" reason="2">DISCUSSION</heading>
</paragraph>
<paragraph blockId="9.[812,1455,1076,1198]" pageId="9" pageNumber="18">
The new species somewhat resembles
<taxonomicName box="[1232,1448,1076,1102]" class="Lecanoromycetes" family="Tephromelataceae" genus="Tephromela" kingdom="Fungi" order="Lecanorales" pageId="9" pageNumber="18" phylum="Ascomycota" rank="species" species="velloziae">
<emphasis box="[1232,1448,1076,1102]" italics="true" pageId="9" pageNumber="18">Tephromela velloziae</emphasis>
</taxonomicName>
, but differs in the irregularly thickened apothecium margin, the larger ascospores (14-16×6.5-7.5 µm vs 9-12× 5-7 µm), and the corticolous habit.
</paragraph>
</subSubSection>
</treatment>
</document>