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<mods:title id="DC2F9FA487AB7C7A6AE9A273884D5528">A remarkable new genus and species of Euryproctini (Hymenoptera: Ichneumonidae, Ctenopelmatinae) from Thailand</mods:title>
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Reshchikov &amp; Quicke
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<figureCitation id="130B898AFF89FF98FD38FBCEF779FB14" box="[743,845,1136,1163]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1960,1986]" captionStartId-1="7.[189,232,1924,1950]" captionTargetBox-0="[194,1395,496,1922]" captionTargetBox-1="[189,1398,501,1885]" captionTargetId-0="figure-81@6.[193,1395,480,1922]" captionTargetId-1="figure-89@7.[189,1398,500,1885]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." captionText-1="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi-0="http://doi.org/10.5281/zenodo.7010580" figureDoi-1="http://doi.org/10.5281/zenodo.7010593" httpUri-0="https://zenodo.org/record/7010580/files/figure.png" httpUri-1="https://zenodo.org/record/7010593/files/figure.png" pageId="5" pageNumber="107">Figs 12</figureCitation>
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<emphasis id="B944491DFF89FF98FF62FB0CF570FB53" bold="true" box="[189,324,1202,1228]" pageId="5" pageNumber="107">Etymology</emphasis>
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<paragraph id="8B8F950FFF89FF98FF62FB5EF72CFB65" blockId="5.[189,792,1248,1275]" box="[189,792,1248,1275]" pageId="5" pageNumber="107">The specific name refers to the species distribution.</paragraph>
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</subSubSection>
<subSubSection id="C32AC684FF89FF95FF62F93DF50DFB85" lastPageId="8" lastPageNumber="110" pageId="5" pageNumber="107" type="description">
<paragraph id="8B8F950FFF89FF98FF62F93DF57AF902" blockId="5.[189,334,1667,1693]" box="[189,334,1667,1693]" pageId="5" pageNumber="107">
<heading id="D0C72263FF89FF98FF62F93DF57AF902" bold="true" box="[189,334,1667,1693]" fontSize="11" level="3" pageId="5" pageNumber="107" reason="3">
<emphasis id="B944491DFF89FF98FF62F93DF57AF902" bold="true" box="[189,334,1667,1693]" pageId="5" pageNumber="107">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F950FFF89FF98FF35F90CF5F3F953" blockId="5.[189,1219,1714,1776]" box="[234,455,1714,1740]" pageId="5" pageNumber="107">
<emphasis id="B944491DFF89FF98FF35F90CF571F953" bold="true" box="[234,325,1714,1740]" pageId="5" pageNumber="107">Female</emphasis>
(
<typeStatus id="548B2BADFF89FF98FE8AF90CF58BF953" box="[341,447,1714,1740]" pageId="5" pageNumber="107" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8B8F950FFF89FF98FF62F968F0F7F96F" blockId="5.[189,1219,1714,1776]" box="[189,1219,1749,1776]" pageId="5" pageNumber="107">
MEASUREMENTS. Body length 6.0 mm, fore wing length 6.0 mm, antenna length
<quantity id="4CC838EAFF89FF98FBBDF96BF08BF96F" box="[1122,1215,1749,1776]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="107" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
<paragraph id="8B8F950FFF89FF98FF62F8A9F600F8AE" blockId="5.[189,564,1814,1841]" box="[189,564,1814,1841]" pageId="5" pageNumber="107">BODY. Matt and sparsely setose.</paragraph>
<paragraph id="8B8F950FFF89FF9BFF62F8E6F00AFE2F" blockId="5.[189,1399,1880,2012]" lastBlockId="6.[188,1400,265,432]" lastPageId="6" lastPageNumber="108" pageId="5" pageNumber="107">
HEAD. Antenna slender, as long as fore wing and with 37 flagellomeres. Scape 1.4 times as long as broad. First flagellomere with prominent basal annellus, 1.6 × as long as 2
<superScript id="7C453847FF89FF98FBE4F8C4F078F817" attach="left" box="[1083,1100,1914,1928]" fontSize="6" pageId="5" pageNumber="107">nd</superScript>
and 1.9 × as long as 3
<superScript id="7C453847FF89FF98FABEF8C4F15BF817" attach="none" box="[1377,1391,1914,1928]" fontSize="6" pageId="5" pageNumber="107">rd</superScript>
. Head narrowed behind eyes. Clypeus 0.4 times as high as wide, separated from face by distinct groove. Anterior tentorial pit rather large (
<figureCitation id="130B898AFF89FF98FD91F87FF69CF843" box="[590,680,1985,2012]" captionStart="Fig" captionStartId="6.[189,232,1960,1986]" captionTargetBox="[194,1395,496,1922]" captionTargetId="figure-81@6.[193,1395,480,1922]" captionTargetPageId="6" captionText="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.7010580" httpUri="https://zenodo.org/record/7010580/files/figure.png" pageId="5" pageNumber="107">Fig. 1B</figureCitation>
). Malar space 0.5 times basal mandible width. Inner margins of eyes (
<figureCitation id="130B898AFF8AFF9BFEC0FEB7F543FEBC" box="[287,375,265,291]" captionStart="Fig" captionStartId="6.[189,232,1960,1986]" captionTargetBox="[194,1395,496,1922]" captionTargetId="figure-81@6.[193,1395,480,1922]" captionTargetPageId="6" captionText="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.7010580" httpUri="https://zenodo.org/record/7010580/files/figure.png" pageId="6" pageNumber="108">Fig. 1B</figureCitation>
) almost parallel, scarcely concave near antennal sockets. Lateral ocellus separated from eye margin by 1.2 times its widest diameter. Face matt with dense setiferous punctures (
<figureCitation id="130B898AFF8AFF9BFB10FE92F11EFED9" box="[1231,1322,300,326]" captionStart="Fig" captionStartId="6.[189,232,1960,1986]" captionTargetBox="[194,1395,496,1922]" captionTargetId="figure-81@6.[193,1395,480,1922]" captionTargetPageId="6" captionText="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.7010580" httpUri="https://zenodo.org/record/7010580/files/figure.png" pageId="6" pageNumber="108">Fig. 1B</figureCitation>
), with short yellow setosity. Frons with weak curved groove behind antennal sockets; with deep, moderately dense setiferous punctures. Occipital carina broadly incomplete dorsally, joining hypostomal carina shortly before base of mandible. Gena matt, with sparse punctures (
<figureCitation id="130B898AFF8AFF9BFC0AFE28F005FE2F" box="[981,1073,406,432]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="6" pageNumber="108">Fig. 2E</figureCitation>
).
</paragraph>
<caption id="DF4FC587FF8AFF9BFF62F816F727F87A" ID-DOI="http://doi.org/10.5281/zenodo.7010580" ID-Zenodo-Dep="7010580" httpUri="https://zenodo.org/record/7010580/files/figure.png" pageId="6" pageNumber="108" startId="6.[189,232,1960,1986]" targetBox="[194,1395,496,1922]" targetPageId="6">
<paragraph id="8B8F950FFF8AFF9BFF62F816F727F87A" blockId="6.[189,1399,1960,2022]" pageId="6" pageNumber="108">
<emphasis id="B944491DFF8AFF9BFF62F816F527F85D" bold="true" box="[189,275,1960,1986]" pageId="6" pageNumber="108">Fig. 1.</emphasis>
<taxonomicName id="4C30EE8CFF8AFF9BFEFDF816F74EF85D" authority="Reshchikov &amp; Quicke" authorityName="Reshchikov &amp; Quicke" authorityYear="2022" box="[290,890,1960,1986]" class="Insecta" family="Ichneumonidae" genus="Rhytidaphora" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="108" phylum="Arthropoda" rank="species" species="thailandica" status="gen. et sp. nov.">
<emphasis id="B944491DFF8AFF9BFEFDF816F66CF85D" box="[290,600,1960,1986]" italics="true" pageId="6" pageNumber="108">Rhytidaphora thailandica</emphasis>
Reshchikov &amp; Quicke
</taxonomicName>
<taxonomicNameLabel id="A277F466FF8AFF9BFC56F817F066F85C" box="[905,1106,1961,1987]" pageId="6" pageNumber="108" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♀ (CUMZ).
<emphasis id="B944491DFF8AFF9BFF62F875F4E6F87A" bold="true" box="[189,210,1995,2021]" pageId="6" pageNumber="108">A</emphasis>
. Habitus, in lateral view.
<emphasis id="B944491DFF8AFF9BFE20F875F627F87A" bold="true" box="[511,531,1995,2021]" pageId="6" pageNumber="108">B</emphasis>
. Face
<emphasis id="B944491DFF8AFF9BFDBFF875F641F87A" bold="true" box="[608,629,1995,2021]" pageId="6" pageNumber="108">C</emphasis>
. Propodeum.
</paragraph>
</caption>
<paragraph id="8B8F950FFF8BFF95FF62FEB7F5D8FD85" blockId="7.[188,1399,265,432]" lastBlockId="8.[188,1400,265,538]" lastPageId="8" lastPageNumber="110" pageId="7" pageNumber="109">
MESOSOMA. Mesoscutum shining, finely and densely punctate (
<figureCitation id="130B898AFF8BFF9AFC4EFEB7F7D9FEBC" box="[913,1005,265,291]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="7" pageNumber="109">Fig. 2C</figureCitation>
). Pronotum with crenulate groove only anteriorly, weakly shiny and with fine, moderately dense puncturation. Mesopleuron weakly shiny, setose and without punctures (
<figureCitation id="130B898AFF8BFF9AFDB1FEF1F6FEFEF6" box="[622,714,335,361]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="7" pageNumber="109">Fig. 2E</figureCitation>
). Scuto-scutellar groove deep with steep posterior margin, triangular and gradually merging with mesoscutum anteriorly. Scutellum strongly convex, matt, without punctures. Juxtacoxal carina absent. Area superomedia and area apicalis fused, hexagonal (
<figureCitation id="130B898AFF8BFF9AFAD7FE28F15CFE2F" box="[1288,1384,406,432]" captionStart="Fig" captionStartId="6.[189,232,1960,1986]" captionTargetBox="[194,1395,496,1922]" captionTargetId="figure-81@6.[193,1395,480,1922]" captionTargetPageId="6" captionText="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.7010580" httpUri="https://zenodo.org/record/7010580/files/figure.png" pageId="7" pageNumber="109">Fig. 1C</figureCitation>
). Spiracles almost round. Wings slightly hyaline (
<figureCitation id="130B898AFF84FF95FCD7FEB7F75FFEBC" box="[776,875,265,291]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2A</figureCitation>
). Fore wing with 1cu-a interstitial. Areolet absent, veins 2m-cu and 2rs-m interstitial. Pterostigma receiving vein Rs+2r at its basal 0.35. 2m-cu with a single, wide bulla anteriorly. Hind wing with cu-a receiving Cu1 well below middle (
<figureCitation id="130B898AFF84FF95FAD6FEF1F15DFEF6" box="[1289,1385,335,361]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2A</figureCitation>
). Propodeum smooth with strong lamelliform carination. Area basalis short and wide. Posterior transverse carina absent (
<figureCitation id="130B898AFF84FF95FEB7FE28F5F0FE2F" box="[360,452,406,432]" captionStart="Fig" captionStartId="6.[189,232,1960,1986]" captionTargetBox="[194,1395,496,1922]" captionTargetId="figure-81@6.[193,1395,480,1922]" captionTargetPageId="6" captionText="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." figureDoi="http://doi.org/10.5281/zenodo.7010580" httpUri="https://zenodo.org/record/7010580/files/figure.png" pageId="8" pageNumber="110">Fig. 1C</figureCitation>
). Claws pectinate (
<figureCitation id="130B898AFF84FF95FD77FE28F731FE2F" box="[680,773,406,432]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2B</figureCitation>
), fore claw with four teeth, mid- and hind claw with three teeth. Hind femur 4.2 times as long (including trochantellus) as maximally broad. Hind tibia 6.2 times as long as apically broad. First tarsomere of hind leg 8 times as long as broad and 2.5 times as long as second tarsomere.
</paragraph>
<caption id="DF4FC587FF8BFF9AFF62F83AF7E4F87A" ID-DOI="http://doi.org/10.5281/zenodo.7010593" ID-Zenodo-Dep="7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="7" pageNumber="109" startId="7.[189,232,1924,1950]" targetBox="[189,1398,501,1885]" targetPageId="7">
<paragraph id="8B8F950FFF8BFF9AFF62F83AF7E4F87A" blockId="7.[187,1399,1924,2021]" pageId="7" pageNumber="109">
<emphasis id="B944491DFF8BFF9AFF62F83AF53DF801" bold="true" box="[189,265,1924,1950]" pageId="7" pageNumber="109">Fig. 2.</emphasis>
<taxonomicName id="4C30EE8CFF8BFF9AFED0F83AF776F801" authority="Reshchikov &amp; Quicke" authorityName="Reshchikov &amp; Quicke" authorityYear="2022" box="[271,834,1924,1950]" class="Insecta" family="Ichneumonidae" genus="Rhytidaphora" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="109" phylum="Arthropoda" rank="species" species="thailandica" status="gen. et sp. nov.">
<emphasis id="B944491DFF8BFF9AFED0F83AF608F801" box="[271,572,1924,1950]" italics="true" pageId="7" pageNumber="109">Rhytidaphora thailandica</emphasis>
Reshchikov &amp; Quicke
</taxonomicName>
<taxonomicNameLabel id="A277F466FF8BFF9AFC98F83BF7C0F800" box="[839,1012,1925,1951]" pageId="7" pageNumber="109" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♀ (CUMZ).
<emphasis id="B944491DFF8BFF9AFADDF83AF123F801" bold="true" box="[1282,1303,1924,1950]" pageId="7" pageNumber="109">A</emphasis>
. Wings.
<emphasis id="B944491DFF8BFF9AFF64F816F4FBF85D" bold="true" box="[187,207,1960,1986]" pageId="7" pageNumber="109">B</emphasis>
. Tarsal claw.
<emphasis id="B944491DFF8BFF9AFEB5F819F54BF85E" bold="true" box="[362,383,1959,1985]" pageId="7" pageNumber="109">C</emphasis>
. Head and mesosoma dorsally.
<emphasis id="B944491DFF8BFF9AFD33F816F735F85D" bold="true" box="[748,769,1960,1986]" pageId="7" pageNumber="109">D</emphasis>
. Metasoma laterally.
<emphasis id="B944491DFF8BFF9AFC26F816F039F85D" bold="true" box="[1017,1037,1960,1986]" pageId="7" pageNumber="109">E</emphasis>
. Head and mesosoma laterally.
<emphasis id="B944491DFF8BFF9AFF64F875F4F9F87A" bold="true" box="[187,205,1995,2021]" pageId="7" pageNumber="109">F</emphasis>
. Metasoma dorsally.
<emphasis id="B944491DFF8BFF9AFE15F874F5D5F87B" bold="true" box="[458,481,1994,2020]" pageId="7" pageNumber="109">G</emphasis>
. Fore tibia laterally.
<emphasis id="B944491DFF8BFF9AFD0AF875F6D8F87A" bold="true" box="[725,748,1995,2021]" pageId="7" pageNumber="109">H</emphasis>
. Fore tibia distally.
</paragraph>
</caption>
<paragraph id="8B8F950FFF84FF95FF62FDFFF135FC94" blockId="8.[189,1399,577,779]" pageId="8" pageNumber="110">
METASOMA. Without obvious punctures but densely finely setose (
<figureCitation id="130B898AFF84FF95FC63FDFFF00DFDC4" box="[956,1081,577,603]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2D, F</figureCitation>
). T1 as long as broad, with strong dorsal longitudinal carina continuing in dorsal carinae of T2 (
<figureCitation id="130B898AFF84FF95FBDEFDDAF06AFDE1" box="[1025,1118,612,638]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2F</figureCitation>
). Ventro-lateral carinae complete (
<figureCitation id="130B898AFF84FF95FEE3FD39F5AAFD3E" box="[316,414,647,673]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2D</figureCitation>
). Glymma absent (
<figureCitation id="130B898AFF84FF95FD59FD39F6DCFD3E" box="[646,744,647,673]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2D</figureCitation>
). S1 rather short, 0.3 times as long as T1. T2 and T3 immovably fused, second suture deep and crenulate (
<figureCitation id="130B898AFF84FF95FCF8FD14F790FD5A" box="[807,932,682,709]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2D, F</figureCitation>
). T15 with distinct basal and subapical transverse impressions distinctly striate (
<figureCitation id="130B898AFF84FF95FD7CFD70F6CBFD77" box="[675,767,718,744]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2F</figureCitation>
). Ovipositor sheath very short, sheaths 0.13 times as long as hind tibia. Ovipositor straight, with a long, shallow, subapical dorsal notch (
<figureCitation id="130B898AFF84FF95FB4AFD4FF0C0FC94" box="[1173,1268,753,779]" captionStart="Fig" captionStartId="7.[189,232,1924,1950]" captionTargetBox="[189,1398,501,1885]" captionTargetId="figure-89@7.[189,1398,500,1885]" captionTargetPageId="7" captionText="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi="http://doi.org/10.5281/zenodo.7010593" httpUri="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8B8F950FFF84FF95FF62FC8CF64FFC29" blockId="8.[189,1398,818,950]" pageId="8" pageNumber="110">
COLORATION. Yellow, except for the following which are marked with black: antenna (dorsally), teeth of mandible, dorsal part of head, pterostigma, hind coxa dorsoapically, hind femur and tibia apically, hind tarsus (except first tarsomere basally), lateral parts of mesoscutum entirely and its anterior medial part, pair of spots on T3 and T4 (
<figureCitation id="130B898AFF84FF95FDDCFC22F659FC29" box="[515,621,924,950]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1960,1986]" captionStartId-1="7.[189,232,1924,1950]" captionTargetBox-0="[194,1395,496,1922]" captionTargetBox-1="[189,1398,501,1885]" captionTargetId-0="figure-81@6.[193,1395,480,1922]" captionTargetId-1="figure-89@7.[189,1398,500,1885]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 1. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Habitus, in lateral view. B. Face C. Propodeum." captionText-1="Fig. 2. Rhytidaphora thailandica Reshchikov &amp; Quicke gen. et sp. nov., holotype, ♀ (CUMZ). A. Wings. B. Tarsal claw.C. Head and mesosoma dorsally.D. Metasoma laterally.E. Head and mesosoma laterally. F. Metasoma dorsally. G. Fore tibia laterally. H. Fore tibia distally." figureDoi-0="http://doi.org/10.5281/zenodo.7010580" figureDoi-1="http://doi.org/10.5281/zenodo.7010593" httpUri-0="https://zenodo.org/record/7010580/files/figure.png" httpUri-1="https://zenodo.org/record/7010593/files/figure.png" pageId="8" pageNumber="110">Figs 12</figureCitation>
).
</paragraph>
<paragraph id="8B8F950FFF84FF95FF35FC63F51EFC68" blockId="8.[189,313,989,1050]" box="[234,298,989,1015]" pageId="8" pageNumber="110">
<emphasis id="B944491DFF84FF95FF35FC63F51EFC68" bold="true" box="[234,298,989,1015]" pageId="8" pageNumber="110">Male</emphasis>
</paragraph>
<paragraph id="8B8F950FFF84FF95FF62FBBEF50DFB85" blockId="8.[189,313,989,1050]" box="[189,313,1024,1050]" pageId="8" pageNumber="110">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C32AC684FF84FF95FF62FBFCF519FB15" box="[189,342,1090,1162]" pageId="8" pageNumber="110" type="distribution">
<paragraph id="8B8F950FFF84FF95FF62FBFCF562FBC3" blockId="8.[189,342,1090,1116]" box="[189,342,1090,1116]" pageId="8" pageNumber="110">
<heading id="D0C72263FF84FF95FF62FBFCF562FBC3" bold="true" box="[189,342,1090,1116]" fontSize="11" level="3" pageId="8" pageNumber="110" reason="3">
<emphasis id="B944491DFF84FF95FF62FBFCF562FBC3" bold="true" box="[189,342,1090,1116]" pageId="8" pageNumber="110">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F950FFF84FF95FF62FBCEF519FB15" blockId="8.[189,301,1136,1162]" box="[189,301,1136,1162]" pageId="8" pageNumber="110">
<collectingCountry id="F327D59FFF84FF95FF62FBCEF51DFB15" box="[189,297,1136,1162]" name="Thailand" pageId="8" pageNumber="110">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32AC684FF84FF95FF62FB0DF14DFA67" pageId="8" pageNumber="110" type="description">
<paragraph id="8B8F950FFF84FF95FF62FB0DF58DFB4F" blockId="8.[189,441,1203,1232]" box="[189,441,1203,1232]" pageId="8" pageNumber="110">
<emphasis id="B944491DFF84FF95FF62FB0DF58DFB4F" bold="true" box="[189,441,1203,1232]" italics="true" pageId="8" pageNumber="110">Molecular analysis</emphasis>
</paragraph>
<paragraph id="8B8F950FFF84FF95FF62FB56F14DFA67" blockId="8.[187,1401,1255,1528]" pageId="8" pageNumber="110">
As in the only previous study in which
<taxonomicName id="4C30EE8CFF84FF95FD55FB59F765FA9E" box="[650,849,1255,1281]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="112" phylum="Arthropoda" rank="subFamily" subFamily="Ctenopelmatinae">Ctenopelmatinae</taxonomicName>
were relatively well represented (
<bibRefCitation id="EFA1E8FEFF84FF95FB01FB59F4C8FABA" author="Quicke D. &amp; Laurenne N. M. &amp; Fitton M. G. &amp; Broad G. R." pageId="8" pageNumber="110" pagination="1305 - 1421" refId="ref6454" refString="Quicke D., Laurenne N. M., Fitton M. G. &amp; Broad G. R. 2009. A thousand and one wasps: a 28 S rDNA and morphological phylogeny of the Ichneumonidae (Insecta: Hymenoptera) with an investigation into alignment parameter space and elision. Journal of Natural History 43 (23 - 24): 1305 - 1421. https: // doi. org / 10.1080 / 00222930902807783" type="journal article" year="2009">
Quicke
<emphasis id="B944491DFF84FF95FAE3FB56F15BFA9E" box="[1340,1391,1255,1281]" italics="true" pageId="8" pageNumber="110">et al</emphasis>
. 2009
</bibRefCitation>
), the molecular analysis did not recover any large ctenopelmatine tribes as monophyletic (
<figureCitation id="130B898AFF84FF95FAFFFAB5F153FABA" box="[1312,1383,1291,1317]" captionStart="Fig" captionStartId="9.[190,233,1922,1948]" captionTargetBox="[221,1327,766,1813]" captionTargetId="figure-202@9.[189,1398,729,1867]" captionTargetPageId="9" captionText="Fig. 3. Maximum likelihood tree based on a 658 base pair segment near the 5, terminus of the CO1 gene. This tree includes the new genus plus at least one representative of each genus of Ctenopelmatinae Förster, 1869 with a DNA barcode, plus two tryphonines as outgroups." figureDoi="http://doi.org/10.5281/zenodo.7010630" httpUri="https://zenodo.org/record/7010630/files/figure.png" pageId="8" pageNumber="110">Fig. 3</figureCitation>
). The placement of
<taxonomicName id="4C30EE8CFF84FF95FE45FA90F63EFAD7" authorityName="Townes &amp; Townes" authorityYear="1950" box="[410,522,1326,1352]" class="Insecta" family="Ichneumonidae" genus="Scolomus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="B944491DFF84FF95FE45FA90F63EFAD7" box="[410,522,1326,1352]" italics="true" pageId="8" pageNumber="110">Scolomus</emphasis>
</taxonomicName>
within
<taxonomicName id="4C30EE8CFF84FF95FDB5FA90F700FAD7" box="[618,820,1326,1352]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="112" phylum="Arthropoda" rank="subFamily" subFamily="Ctenopelmatinae">Ctenopelmatinae</taxonomicName>
, even though this genus has been transferred to Metopiinae (
<bibRefCitation id="EFA1E8FEFF84FF95FE8FFAEFF608FAF4" author="Gauld I. D. &amp; Wahl D. B." box="[336,572,1361,1387]" pageId="8" pageNumber="110" pagination="35 - 41" refId="ref5266" refString="Gauld I. D. &amp; Wahl D. B. 2006. The relationship and taxonomic position of the genera Apolophus and Scolomus (Hymenoptera: Ichneumonidae). Zootaxa 1130 (1): 35 - 41. https: // doi. org / 10.11646 / zootaxa. 1130.1.2" type="journal article" year="2006">Gauld &amp; Wahl 2006</bibRefCitation>
), is consistent with the idea that Metopiinae may be derived from within the Ctenopelmatinae (
<bibRefCitation id="EFA1E8FEFF84FF95FE1CFACAF69BFA11" author="Quicke D. &amp; Laurenne N. M. &amp; Fitton M. G. &amp; Broad G. R." box="[451,687,1396,1423]" pageId="8" pageNumber="110" pagination="1305 - 1421" refId="ref6454" refString="Quicke D., Laurenne N. M., Fitton M. G. &amp; Broad G. R. 2009. A thousand and one wasps: a 28 S rDNA and morphological phylogeny of the Ichneumonidae (Insecta: Hymenoptera) with an investigation into alignment parameter space and elision. Journal of Natural History 43 (23 - 24): 1305 - 1421. https: // doi. org / 10.1080 / 00222930902807783" type="journal article" year="2009">
Quicke
<emphasis id="B944491DFF84FF95FDF9FACBF669FA11" box="[550,605,1396,1422]" italics="true" pageId="8" pageNumber="110">et al</emphasis>
. 2009
</bibRefCitation>
). The
<taxonomicName id="4C30EE8CFF84FF95FD22FACAF7A5FA11" authorityName="Thomson" authorityYear="1883" box="[765,913,1396,1422]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="106" phylum="Arthropoda" rank="tribe" tribe="Euryproctini">Euryproctini</taxonomicName>
as currently constituted were recovered spread across the tree in five separate clades but with low support. Nevertheless, nine of the included euryproctine genera formed a monophyletic grouping (
<figureCitation id="130B898AFF84FF95FC91FA05F7A3FA4A" box="[846,919,1467,1493]" captionStart="Fig" captionStartId="9.[190,233,1922,1948]" captionTargetBox="[221,1327,766,1813]" captionTargetId="figure-202@9.[189,1398,729,1867]" captionTargetPageId="9" captionText="Fig. 3. Maximum likelihood tree based on a 658 base pair segment near the 5, terminus of the CO1 gene. This tree includes the new genus plus at least one representative of each genus of Ctenopelmatinae Förster, 1869 with a DNA barcode, plus two tryphonines as outgroups." figureDoi="http://doi.org/10.5281/zenodo.7010630" httpUri="https://zenodo.org/record/7010630/files/figure.png" pageId="8" pageNumber="110">Fig. 3</figureCitation>
, top) including
<taxonomicName id="4C30EE8CFF84FF95FB85FA05F0CFFA4A" authority="Reshchikov &amp; Quicke &amp; Butcher, 2022" authorityName="Reshchikov &amp; Quicke &amp; Butcher" authorityYear="2022" box="[1114,1275,1467,1493]" class="Insecta" family="Ichneumonidae" genus="Rhytidaphora" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="110" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944491DFF84FF95FB85FA05F0CFFA4A" box="[1114,1275,1467,1493]" italics="true" pageId="8" pageNumber="110">Rhytidaphora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277F466FF84FF95FADDFA05F146FA4A" box="[1282,1394,1467,1493]" pageId="8" pageNumber="110" rank="genus">gen. nov.</taxonomicNameLabel>
, and is therefore consistent with the placement of the new genus based on our morphological assessment.
</paragraph>
</subSubSection>
</treatment>
</document>