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<document id="C0E5EDD015DFCA9490B9D8D33D4226F0" ID-CLB-Dataset="21948" ID-DOI="10.5852/ejt.2017.376" ID-GBIF-Dataset="4298f5cf-d85e-48fb-afe8-f12c99ad65b3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838544" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590088443336" checkinUser="carolina" docAuthor="Aharon, Shlomi, Huber, Bernhard A. &amp; Gavish-Regev, Efrat" docDate="2017" docId="516F87BB1315BC6AFDDEBAD1FC43EA17" docLanguage="en" docName="ejt-2017-376.pdf.imf" docOrigin="European Journal of Taxonomy 376 (376)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Artema kochi Kulczynski 1901" docType="treatment" docVersion="9" lastPageNumber="30" masterDocId="AD56FFC3130CBC77FFD0BC5EFF85E94D" masterDocTitle="Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="26" updateTime="1698840747163" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="35CA34FDD1856B9A618DD02039CD79BC">Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="ED91DF87EA68666765D7B428BE1C034F">Aharon, Shlomi</mods:namePart>
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<mods:affiliation id="5771EC26B3A71440EB498BEC849DB7EC">Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Sede Boqer Campus, Midreshet Ben-Gurion, Israel. Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Givat Ram, Jerusalem, Israel.</mods:affiliation>
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<mods:namePart id="B841329D1F15CC42C25C9880FB6B9024">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="3549DDB376F7DDE068A318AE74345CD5">Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. &amp; Email: b. huber @ zfmk. de &amp; urn: lsid: zoobank. org: author: 33607 F 65 - 19 BF- 4 DC 9 - 94 FD- 4 BB 88 CED 455 F</mods:affiliation>
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<mods:namePart id="71946FA30CD58FC924805452D9619B55">Gavish-Regev, Efrat</mods:namePart>
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<mods:affiliation id="FA648D3DD5047F02471D938858CE671F">The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Givat Ram, Jerusalem, Israel. &amp; urn: lsid: zoobank. org: author: FC 073 F 19 - 2202 - 4 C 89 - 8 B 43 - CEA 4 CC 5 E 2 D 50 &amp; Corresponding author: efrat. gavish-regev @ mail. huji. ac. il</mods:affiliation>
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<treatment id="516F87BB1315BC6AFDDEBAD1FC43EA17" ID-DOI="http://doi.org/10.5281/zenodo.3852330" ID-GBIF-Taxon="164246585" ID-Zenodo-Dep="3852330" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17" httpUri="http://treatment.plazi.org/id/516F87BB1315BC6AFDDEBAD1FC43EA17" lastPageId="29" lastPageNumber="30" pageId="25" pageNumber="26">
<subSubSection id="91DC65261315BC6EFDDEBAD1FBA0EFE7" box="[526,1061,1678,1706]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="D97936AD1315BC6EFDDEBAD1FBA0EFE7" blockId="25.[526,1061,1678,1741]" box="[526,1061,1678,1706]" pageId="25" pageNumber="26">
<heading id="823181C11315BC6EFDDEBAD1FBA0EFE7" box="[526,1061,1678,1706]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="1EC64D2E1315BC6EFDDEBAD1FC0DEFE4" ID-CoL="GVVG" authority="Kulczynski, 1901" authorityName="Kulczynski" authorityYear="1901" box="[526,904,1678,1705]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1315BC6EFDDEBAD1FD37EFE4" bold="true" box="[526,690,1678,1705]" italics="true" pageId="25" pageNumber="26">Artema kochi</emphasis>
<bibRefCitation id="BD574B5C1315BC6EFD69BAD1FC0DEFE4" author="Kulczynski" box="[697,904,1679,1705]" firstAuthor="Kulczynski" pageId="25" pageNumber="26" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński, 1901</bibRefCitation>
</taxonomicName>
(revalidated)
</heading>
</paragraph>
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<subSubSection id="91DC65261315BC6EFD5FBAECFC21EF80" box="[655,932,1714,1741]" pageId="25" pageNumber="26" type="description">
<paragraph id="D97936AD1315BC6EFD5FBAECFC21EF80" blockId="25.[526,1061,1678,1741]" box="[655,932,1714,1741]" pageId="25" pageNumber="26">
<figureCitation id="41FD2A281315BC6EFD5FBAECFD5DEF80" box="[655,728,1714,1741]" captionStart="Figs 12. 1" captionStartId="5.[189,241,1647,1673]" captionTargetBox="[190,1397,377,1570]" captionTargetId="figure@5.[187,1399,374,1571]" captionTargetPageId="5" captionText="Figs 12. 1. Known distributions of species of Artema except A. atlanta Walckenaer, 1837. The Japanese record of A. doriae Thorell, 1881 is not shown as it is based on a single introduced specimen. 2. Known distribution of Artema atlanta. Green dots: specimens examined; black dots: specimens not examined (data from: Banks 1898; Beatty et al. 2008; Blackwell 1958; Bösenberg &amp; Lenz 1895; Brignoli 1981; Bryant 1940, 1948; Caporiacco 1928, 1948, 1949; Chamberlin 1924; Chrysanthus 1967; Colmenares 2008; Denis 1953; Doleschall 1857; Dyal 1935; Franganillo 1926a; Gao &amp; Li 2010; Gertsch &amp; Davis 1942; Gonzalez-Sponga 2005; Karsch 1879, 1892; Leardi in Airaghi 1902; Lessert 1936; Majumder &amp; Biswas 1993; Marples 1955; Mello-Leitão 1942; Millot 1946; Moenkhaus 1898; F.O.Pickard-Cambridge 1902; Petrunkevitch 1925, 1929; Pocock 1900; Saaristo 2001; Simon 1885, 1894; Taczanowsky 1874; Thorell 1895, 1898; Vinson 1863; Yaginuma 1970). Records from Israel, Belgium, and Britain are not shown as they are based on single introduced specimens." figureDoi="http://doi.org/10.5281/zenodo.3838546" httpUri="https://zenodo.org/record/3838546/files/figure.png" pageId="25" pageNumber="26">Figs 1</figureCitation>
,
<figureCitation id="41FD2A281315BC6EFD37BAECFCB7EF81" box="[743,818,1714,1740]" captionStart-0="Figs 7682" captionStart-1="Figs 8389" captionStartId-0="26.[189,241,1928,1954]" captionStartId-1="28.[189,241,1858,1884]" captionTargetBox-0="[203,1390,1102,1875]" captionTargetBox-1="[195,1393,831,1810]" captionTargetId-0="figure@26.[189,1398,1094,1885]" captionTargetId-1="figure@28.[189,1399,816,1816]" captionTargetPageId-0="26" captionTargetPageId-1="28" captionText-0="Figs 7682. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen-Eritrea). 76. Male habitus. 7779. Male left pedipalp: prolateral, dorsal, and retrolateral views. 8082. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm." captionText-1="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3838566" figureDoi-1="http://doi.org/10.5281/zenodo.3838568" httpUri-0="https://zenodo.org/record/3838566/files/figure.png" httpUri-1="https://zenodo.org/record/3838568/files/figure.png" pageId="25" pageNumber="26">7689</figureCitation>
,
<figureCitation id="41FD2A281315BC6EFCEFBAEDFCEBEF80" box="[831,878,1715,1741]" captionStart="Figs 201207" captionStartId="49.[189,241,1824,1850]" captionTargetBox="[198,1391,724,1781]" captionTargetId="figure@49.[189,1398,716,1791]" captionTargetPageId="49" captionText="Figs 201207. Cleared female genitalia, dorsal views. 201. Artema atlanta Walckenaer, 1837 (AMNH, USA, Arizona, Tucson). 202. A. bunkpurugu Huber &amp; Kwapong, 2013 (ZFMK Ar 10516, Ghana, Bunkpurugu-Nakpanduri). 203. A. nephilit sp. nov. (HUJ Ara 16020, Israel, Samar). 204. A. kochi Kulczyński, 1901 (MNHN AR 10169 part). 205. A. doriae Thorell, 1881 (ZFMK Ar 15234, Iran, Tehran Prov., Tochal). 206. A. transcaspica Spassky, 1934 (ZIN type vials). 207. Artema sp. c. (ZMMU, Pakistan, Baluchistan, Muslim Bagh). Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838600" httpUri="https://zenodo.org/record/3838600/files/figure.png" pageId="25" pageNumber="26">204</figureCitation>
,
<figureCitation id="41FD2A281315BC6EFCA9BAEDFC21EF80" box="[889,932,1715,1741]" captionStart="Figs 208214" captionStartId="50.[189,241,1077,1103]" captionTargetBox="[196,1390,273,1028]" captionTargetId="figure@50.[188,1399,265,1037]" captionTargetPageId="50" captionText="Figs 208214. Cleared female genitalia, posterior dorsal views. 208. A. atlanta Walckenaer, 1837 (AMNH, USA, Arizona, Tucson). 209. A. bunkpurugu Huber &amp; Kwapong, 2013 (ZFMK Ar 10516, Ghana, Bunkpurugu-Nakpanduri). 210. A. nephilit sp. nov. (HUJAra 16020, Israel, Samar). 211. A. kochi Kulczyński, 1901 (MNHN AR 10169 part). 212. A. doriae Thorell, 1881 (ZFMK Ar 15234, Iran, Tehran Prov., Tochal). 213. A. transcaspica Spassky, 1934 (ZIN type vials). 214. Artema sp. c (ZMMU, Pakistan, Baluchistan, Muslim Bagh). Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838602" httpUri="https://zenodo.org/record/3838602/files/figure.png" pageId="25" pageNumber="26">211</figureCitation>
</paragraph>
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<paragraph id="D97936AD1315BC6EFF6DBAABFD4EEE7E" blockId="25.[189,1398,1780,1843]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="F9D611831315BC6EFF6DBAABFD4EEE7E" pageId="25" pageNumber="26">
<treatmentCitation id="586710BC1315BC6EFF6DBAABFD35EE42" author="Kulczynski W." box="[189,688,1780,1807]" page="20" pageId="25" pageNumber="26" year="1901">
<taxonomicName id="1EC64D2E1315BC6EFF6DBAABFD35EE42" authority="Kulczynski 1901: 19 - 20" authorityName="Kulczynski" authorityPageNumber="19 - 20" authorityYear="1901" box="[189,688,1780,1807]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kochii">
<emphasis id="EBB2EABF1315BC6EFF6DBAABFEEEEE43" box="[189,363,1780,1807]" italics="true" pageId="25" pageNumber="26">Artema kochii</emphasis>
<bibRefCitation id="BD574B5C1315BC6EFEABBAAAFD35EE42" author="Kulczynski" box="[379,688,1780,1807]" firstAuthor="Kulczynski" pageId="25" pageNumber="26" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński 1901: 1920</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; pl. 1, fig. 14. Erroneously synonymized with “
<taxonomicName id="1EC64D2E1315BC6EFAF0BAABFEF4EE7F" ID-CoL="GVVK" authorityName="Walckenaer" authorityYear="1837" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mauriciana">
<emphasis id="EBB2EABF1315BC6EFAF0BAABFEF4EE7F" italics="true" pageId="25" pageNumber="26">Artema mauriciana</emphasis>
</taxonomicName>
” by
<treatmentCitation id="586710BC1315BC6EFE7BBB46FD42EE7E" author="Simon E." box="[427,711,1816,1843]" page="426" pageId="25" pageNumber="26" year="1908">
<bibRefCitation id="BD574B5C1315BC6EFE7BBB46FD42EE7E" author="Simon" box="[427,711,1816,1843]" firstAuthor="Simon" pageId="25" pageNumber="26" pagination="419 - 438" refId="ref30437" refString="Simon E. 1908. Etude sur les arachnides recueillis par M. le Dr Klaptocz en Tripolitaine. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 26: 419 - 438." type="journal article" year="1908">Simon (1908: 426427)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D97936AD1315BC6EFF6DBB04FE19EE39" blockId="25.[189,412,1882,1908]" box="[189,412,1882,1908]" pageId="25" pageNumber="26">
<heading id="823181C11315BC6EFF6DBB04FE19EE39" bold="true" box="[189,412,1882,1908]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EBB2EABF1315BC6EFF6DBB04FE19EE39" bold="true" box="[189,412,1882,1908]" pageId="25" pageNumber="26">Misidentifications</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1315BC6EFF6DBBD7FDDEEE8A" blockId="25.[189,1398,1929,2027]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="F9D611831315BC6EFF6DBBD7FDDEEE8A" pageId="25" pageNumber="26">
<taxonomicName id="1EC64D2E1315BC6EFF1ABBD7FE32EEEE" authorityName="Vinson" authorityYear="1863" box="[202,439,1929,1955]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="borbonicus">
<emphasis id="EBB2EABF1315BC6EFF1ABBD7FE32EEEE" box="[202,439,1929,1955]" italics="true" pageId="25" pageNumber="26">Pholcus borbonicus</emphasis>
</taxonomicName>
L.
<treatmentCitation id="586710BC1315BC6EFDDABBD7FD68EEE9" author="Koch L." box="[522,749,1929,1956]" page="25" pageId="25" pageNumber="26" year="1875">
<bibRefCitation id="BD574B5C1315BC6EFDDABBD7FD68EEE9" author="Koch" box="[522,749,1929,1956]" firstAuthor="Koch" pageId="25" pageNumber="26" refId="ref29158" refString="Koch L. 1875. Aegyptische und abyssinische Arachniden gesammelt von Herrn C. Jickeli. Bauer and Raspe, Nurnberg." type="book" year="1875">Koch 1875: 2526</bibRefCitation>
</treatmentCitation>
; pl. 3, fig. 1, 1ac (
<specimenCount id="CFC0FD241315BC6EFC32BBD7FC73EEEE" box="[994,1014,1929,1955]" pageId="25" pageNumber="26" type="female"></specimenCount>
). —
<treatmentCitation id="586710BC1315BC6EFBE8BBD7FAC9EEEE" author="Thorell T." box="[1080,1356,1929,1956]" page="179" pageId="25" pageNumber="26" year="1881">
<bibRefCitation id="BD574B5C1315BC6EFBE8BBD7FAC9EEEE" author="Thorell" box="[1080,1356,1929,1956]" firstAuthor="Thorell" pageId="25" pageNumber="26" pagination="1 - 727" refId="ref30571" refString="Thorell T. 1881. Studi sui ragni Malesi e Papuani. III. Ragni dell'Austro Malesia e del Capo York, conservati nel Museo civico di storia naturale di Genova. Annali del Museo civico di storia naturale di Genova 17: 1 - 727." type="journal article" year="1881">Thorell 1881: 179180</bibRefCitation>
</treatmentCitation>
(in footnote). —
<treatmentCitation id="586710BC1315BC6EFE59BBF3FDD2EE8A" author="Simon E." box="[393,599,1965,1991]" page="234" pageId="25" pageNumber="26" year="1882">
<bibRefCitation id="BD574B5C1315BC6EFE59BBF3FDD2EE8A" author="Simon" box="[393,599,1965,1991]" firstAuthor="Simon" pageId="25" pageNumber="26" pagination="207 - 260" refId="ref30170" refString="Simon E. 1882. II. Etude sur les arachnides de l'Yemen meridional. In: Viaggio ad Assab nel Mar Rosso, dei signori G. Doria ed O. Beccari con il R. Aviso &quot; Esploratore &quot; dal 16 Novembre 1879 al 26 Febbraio 1880. Annali del Museo civico di storia naturale di Genova 18: 207 - 260." type="journal article" year="1882">Simon 1882: 234</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D97936AD1315BC6EFF6DBB8FFDFAEEA6" blockId="25.[189,1398,1929,2027]" box="[189,639,2001,2027]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="F9D611831315BC6EFF6DBB8FFDFAEEA6" box="[189,639,2001,2027]" pageId="25" pageNumber="26">
<taxonomicName id="1EC64D2E1315BC6EFF1ABB8FFE17EEA6" box="[202,402,2001,2027]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mauricia">
<emphasis id="EBB2EABF1315BC6EFF1ABB8FFE17EEA6" box="[202,402,2001,2027]" italics="true" pageId="25" pageNumber="26">Artema mauricia</emphasis>
</taxonomicName>
<treatmentCitation id="586710BC1315BC6EFE6CBB8FFDFEEEA6" author="Simon E." box="[444,635,2001,2027]" page="93" pageId="25" pageNumber="26" year="1890">
<bibRefCitation id="BD574B5C1315BC6EFE6CBB8FFDFEEEA6" author="Simon" box="[444,635,2001,2027]" firstAuthor="Simon" pageId="25" pageNumber="26" pagination="77 - 124" refId="ref30334" refString="Simon E. 1890. Etude arachnologiques. 22 e Memoire. XXXIV. Etude sur les arachnides de l'Yemen. Annales de la Societe Entomologique de France (6) 10: 77 - 124." type="journal article" year="1890">Simon 1890: 93</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="91DC65261316BC6DFF6DBD55FB7CEB20" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="D97936AD1316BC6DFF6DBD55FEB3E868" blockId="26.[189,310,267,293]" box="[189,310,267,293]" pageId="26" pageNumber="27">
<heading id="823181C11316BC6DFF6DBD55FEB3E868" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="EBB2EABF1316BC6DFF6DBD55FEB3E868" bold="true" box="[189,310,267,293]" pageId="26" pageNumber="27">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1316BC6DFF6DBD67FB7CEB20" blockId="26.[189,1399,313,621]" pageId="26" pageNumber="27">
Males are easily distinguished from known congeners by their bulbal process b (
<figureCitation id="41FD2A281316BC6DFBA2BD67FB4CE819" box="[1138,1225,313,340]" captionStart="Figs 7682" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[203,1390,1102,1875]" captionTargetId="figure@26.[189,1398,1094,1885]" captionTargetPageId="26" captionText="Figs 7682. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen-Eritrea). 76. Male habitus. 7779. Male left pedipalp: prolateral, dorsal, and retrolateral views. 8082. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838566" httpUri="https://zenodo.org/record/3838566/files/figure.png" pageId="26" pageNumber="27">Fig. 81</figureCitation>
), flattened and rectangular distally, rather than pointed or awl-shaped as in congeners; processes c, d, and e absent (process c present in all congeners, except
<taxonomicName id="1EC64D2E1316BC6DFD72BDDEFD7DE8D7" authorityName="Walckenaer" authorityYear="1837" box="[674,760,384,410]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="EBB2EABF1316BC6DFD72BDDEFD7DE8D7" box="[674,760,384,410]" italics="true" pageId="26" pageNumber="27">Artema</emphasis>
</taxonomicName>
sp. b from
<collectingCountry id="A1D1763D1316BC6DFCA3BDDEFC55E8D6" box="[883,976,384,411]" name="Algeria" pageId="26" pageNumber="27">Algeria</collectingCountry>
). Males differ also by their cheliceral processes that are strongly projecting proximally with frontal rows of modified hairs; with several cone-shaped, sclerotized processes situated frontally near median line (arrows in
<figureCitation id="41FD2A281316BC6DFBA8BD98FA85E8AC" box="[1144,1280,454,481]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="26" pageNumber="27">Figs 83, 85</figureCitation>
). Females differ from known congeners by entirely different shapes of anterior epigynal projections (AEP) and epigynal plate (
<figureCitation id="41FD2A281316BC6DFEA5BE52FD85EB6A" box="[373,512,524,551]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="26" pageNumber="27">Figs 86, 88</figureCitation>
): anterior epigynal projections sclerotized and elongated (in contrast to all congeners), not projecting (in contrast to
<taxonomicName id="1EC64D2E1316BC6DFD76BE6EFCD0EB07" authorityName="Huber &amp; Kwapong" authorityYear="2013" box="[678,853,560,586]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bunkpurugu">
<emphasis id="EBB2EABF1316BC6DFD76BE6EFCD0EB07" box="[678,853,560,586]" italics="true" pageId="26" pageNumber="27">A. bunkpurugu</emphasis>
</taxonomicName>
); epigynal plate sclerotized, wide rectangular, with triangular anterior projection between AEP, median posterior margin gently protruding.
</paragraph>
</subSubSection>
<subSubSection id="91DC65261316BC6CFF6DBECAFC5BEBD1" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="D97936AD1316BC6DFF6DBECAFE2FEBE3" blockId="26.[189,426,660,686]" box="[189,426,660,686]" pageId="26" pageNumber="27">
<heading id="823181C11316BC6DFF6DBECAFE2FEBE3" bold="true" box="[189,426,660,686]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="EBB2EABF1316BC6DFF6DBECAFE2FEBE3" bold="true" box="[189,426,660,686]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="69AE3CF01316BC6DFF3ABE9DFD97EA69" ID-GBIF-Occurrence="3763211429" collectingDate="1870-04" collectionCode="MSNG" collectorName="A. Yssel" country="Eritrea" latitude="15.61" location="Massaua" longLatPrecision="771" longitude="39.45" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<paragraph id="D97936AD1316BC6DFF3ABE9DFDBFEB93" blockId="26.[189,1398,707,804]" box="[234,570,707,734]" pageId="26" pageNumber="27">
<emphasis id="EBB2EABF1316BC6DFF3ABE9DFED4EB90" bold="true" box="[234,337,707,733]" pageId="26" pageNumber="27">
<typeStatus id="067D880F1316BC6DFF3ABE9DFED4EB90" box="[234,337,707,733]" pageId="26" pageNumber="27" type="neotype">Neotype</typeStatus>
</emphasis>
(designated herein)
</paragraph>
<paragraph id="D97936AD1316BC6DFF6DBEB8FD97EA69" blockId="26.[189,1398,707,804]" pageId="26" pageNumber="27">
<collectingCountry id="A1D1763D1316BC6DFF6DBEB8FEBBEA4D" box="[189,318,742,768]" name="Eritrea" pageId="26" pageNumber="27">ERITREA</collectingCountry>
:
<specimenCount id="CFC0FD241316BC6DFE9DBEB8FE05EA4C" box="[333,384,742,769]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
, “
<location id="DC1960761316BC6DFE4BBEB8FD82EA4D" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17:DC1960761316BC6DFE4BBEB8FD82EA4D" box="[411,519,742,768]" country="Eritrea" latitude="15.61" longLatPrecision="771" longitude="39.45" name="Massaua" pageId="26" pageNumber="27">Massaua</location>
” [=
<location id="DC1960761316BC6DFD94BEB9FD31EA4C" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17:DC1960761316BC6DFD94BEB9FD31EA4C" box="[580,692,743,769]" country="Eritrea" latitude="15.61" longLatPrecision="771" longitude="39.45" name="Massawa" pageId="26" pageNumber="27">Massawa</location>
,
<geoCoordinate id="BCF2506A1316BC6DFD15BEB8FCB2EA4C" box="[709,823,742,769]" degrees="15.61" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="555" value="15.61">15.61° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1316BC6DFC97BEB8FC32EA4C" box="[839,951,742,769]" degrees="39.45" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="39.45">39.45° E</geoCoordinate>
],
<date id="AD78106D1316BC6DFC1FBEB8FBCEEA4D" box="[975,1099,742,768]" pageId="26" pageNumber="27" value="1870-04">
<collectingDate id="BD3CE9851316BC6DFC1FBEB8FBCEEA4D" box="[975,1099,742,768]" pageId="26" pageNumber="27" value="1870-04">Apr. 1870</collectingDate>
</date>
,
<collectorName id="7433537B1316BC6DFB8BBEB8FB46EA4D" box="[1115,1219,742,768]" pageId="26" pageNumber="27">A. Yssel</collectorName>
leg. (
<collectionCode id="BFD7AE681316BC6DFADFBEB8FAEFEA4D" box="[1295,1386,742,768]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="26" pageNumber="27">MSNG</collectionCode>
), examined (see Notes below).
</paragraph>
</materialsCitation>
<paragraph id="D97936AD1316BC6DFF3ABF15FE23EA28" blockId="26.[189,867,843,905]" box="[234,422,843,869]" pageId="26" pageNumber="27">
<emphasis id="EBB2EABF1316BC6DFF3ABF15FE23EA28" bold="true" box="[234,422,843,869]" pageId="26" pageNumber="27">Other material</emphasis>
</paragraph>
<paragraph id="D97936AD1316BC6DFF6DBF30FCE6EAC5" blockId="26.[189,867,843,905]" box="[189,867,878,905]" pageId="26" pageNumber="27">
<materialsCitation id="69AE3CF01316BC6DFF6DBF30FCE5EAC5" ID-GBIF-Occurrence="3763211322" box="[189,864,878,905]" country="Eritrea" location="ERITREA" pageId="26" pageNumber="27" specimenCount="1" specimenCount-female="1" typeStatus="neotype">
<collectingCountry id="A1D1763D1316BC6DFF6DBF30FEBBEAC5" box="[189,318,878,904]" name="Eritrea" pageId="26" pageNumber="27">ERITREA</collectingCountry>
:
<specimenCount id="CFC0FD241316BC6DFE98BF30FEF1EAC4" box="[328,372,878,905]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
, 7 juvs, together with
<typeStatus id="067D880F1316BC6DFDABBF31FD5EEAC4" box="[635,731,879,905]" pageId="26" pageNumber="27" type="neotype">neotype</typeStatus>
, same data
</materialsCitation>
.
</paragraph>
<paragraph id="D97936AD1316BC6DFF6DBFEEFEC8EAA0" blockId="26.[189,1399,943,1005]" pageId="26" pageNumber="27">
<materialsCitation id="69AE3CF01316BC6DFF6DBFEEFEC8EAA0" ID-GBIF-Occurrence="3763211387" collectionCode="MNHN, AR" collectorName="E. Simon" location="Massawa" municipality="Massawa" pageId="26" pageNumber="27" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Aden">
YEMEN-ERITREA:
<specimenCount id="CFC0FD241316BC6DFE6CBFEEFD84EA87" box="[444,513,944,970]" pageId="26" pageNumber="27" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CFC0FD241316BC6DFDDFBFEEFDD6EA87" box="[527,595,944,970]" pageId="26" pageNumber="27" type="female">2 ♀♀</specimenCount>
, 4 juvs,
<collectingRegion id="1B02F84F1316BC6DFD68BFF1FD7DEA84" box="[696,760,943,969]" country="Yemen" name="'Adan" pageId="26" pageNumber="27">Aden</collectingRegion>
/
<collectingMunicipality id="391DACD71316BC6DFCD1BFF1FCE8EA84" box="[769,877,943,969]" pageId="26" pageNumber="27">Massawa</collectingMunicipality>
[1889] (
<collectorName id="7433537B1316BC6DFC06BFEEFBCDEA84" box="[982,1096,943,970]" pageId="26" pageNumber="27">E. Simon</collectorName>
, “393” part) (
<collectionCode id="BFD7AE681316BC6DFB3BBFEEFAC2EA87" box="[1259,1351,944,970]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
<collectionCode id="BFD7AE681316BC6DFA9EBFEEFAF2EA87" box="[1358,1399,944,970]" pageId="26" pageNumber="27">AR</collectionCode>
10169 part).
</materialsCitation>
</paragraph>
<caption id="8DB966251316BC6DFF6DBBD6FAC7EEA4" ID-DOI="http://doi.org/10.5281/zenodo.3838566" ID-Zenodo-Dep="3838566" httpUri="https://zenodo.org/record/3838566/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,241,1928,1954]" targetBox="[203,1390,1102,1875]" targetPageId="26">
<paragraph id="D97936AD1316BC6DFF6DBBD6FAC7EEA4" blockId="26.[189,1399,1928,2025]" pageId="26" pageNumber="27">
<emphasis id="EBB2EABF1316BC6DFF6DBBD6FECDEEEF" bold="true" box="[189,328,1928,1954]" pageId="26" pageNumber="27">Figs 7682.</emphasis>
<taxonomicName id="1EC64D2E1316BC6DFE9FBBD7FD44EEEF" authority="Kulczynski, 1901" authorityName="Kulczynski" authorityYear="1901" box="[335,705,1928,1955]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1316BC6DFE9FBBD7FE69EEEF" box="[335,492,1928,1955]" italics="true" pageId="26" pageNumber="27">Artema kochi</emphasis>
<bibRefCitation id="BD574B5C1316BC6DFE22BBD6FD44EEEF" author="Kulczynski" box="[498,705,1928,1954]" firstAuthor="Kulczynski" pageId="26" pageNumber="27" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński, 1901</bibRefCitation>
</taxonomicName>
(MNHN AR 10169 part, Yemen-Eritrea).
<emphasis id="EBB2EABF1316BC6DFB7CBBD6FB4CEEEF" bold="true" box="[1196,1225,1928,1954]" pageId="26" pageNumber="27">76</emphasis>
. Male habitus.
<emphasis id="EBB2EABF1316BC6DFF6DBBF5FE83EE88" bold="true" box="[189,262,1963,1989]" pageId="26" pageNumber="27">7779</emphasis>
. Male left pedipalp: prolateral, dorsal, and retrolateral views.
<emphasis id="EBB2EABF1316BC6DFC1EBBF5FB92EE88" bold="true" box="[974,1047,1963,1989]" pageId="26" pageNumber="27">8082</emphasis>
. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="D97936AD1317BC6CFF6DBD52FD70E8DD" blockId="27.[189,1399,267,400]" pageId="27" pageNumber="28">
<materialsCitation id="69AE3CF01317BC6CFF6DBD52FB71E804" ID-GBIF-Occurrence="3763211326" collectingDate="1880-01" collectionCode="MSNG" country="Yemen" latitude="12.86" location="Simon" longLatPrecision="775" longitude="44.98" municipality="Doria and Beccari" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Aden">
<collectingCountry id="A1D1763D1317BC6CFF6DBD52FEAFE86B" box="[189,298,268,294]" name="Yemen" pageId="27" pageNumber="28">YEMEN</collectingCountry>
:
<specimenCount id="CFC0FD241317BC6CFEE5BD55FEE4E86B" box="[309,353,267,294]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
, 3 juvs, “Sceik Osman” [= Shaykh Uthman,
<collectingRegion id="1B02F84F1317BC6CFCBEBD55FC34E868" box="[878,945,267,293]" country="Yemen" name="'Adan" pageId="27" pageNumber="28">Aden</collectingRegion>
,
<geoCoordinate id="BCF2506A1317BC6CFC6CBD55FBACE86B" box="[956,1065,267,294]" degrees="12.86" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="12.86">12.86° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1317BC6CFBE5BD55FB25E86B" box="[1077,1184,267,294]" degrees="44.98" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="555" value="44.98">44.98° E</geoCoordinate>
],
<date id="AD78106D1317BC6CFB65BD52FAA2E868" box="[1205,1319,267,294]" pageId="27" pageNumber="28" value="1880-01">
<collectingDate id="BD3CE9851317BC6CFB65BD52FAA2E868" box="[1205,1319,267,294]" pageId="27" pageNumber="28" value="1880-01">Jan. 1880</collectingDate>
</date>
,
<collectingMunicipality id="391DACD71317BC6CFAE4BD55FECCE804" pageId="27" pageNumber="28">Doria and Beccari</collectingMunicipality>
leg. (this is the material cited in
<bibRefCitation id="BD574B5C1317BC6CFD00BD71FCDAE804" author="Simon" box="[720,863,303,329]" firstAuthor="Simon" pageId="27" pageNumber="28" pagination="207 - 260" refId="ref30170" refString="Simon E. 1882. II. Etude sur les arachnides de l'Yemen meridional. In: Viaggio ad Assab nel Mar Rosso, dei signori G. Doria ed O. Beccari con il R. Aviso &quot; Esploratore &quot; dal 16 Novembre 1879 al 26 Febbraio 1880. Annali del Museo civico di storia naturale di Genova 18: 207 - 260." type="journal article" year="1882">
<location id="DC1960761317BC6CFD00BD71FC98E804" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17:DC1960761317BC6CFD00BD71FC98E804" box="[720,797,303,329]" country="Yemen" latitude="12.86" longLatPrecision="775" longitude="44.98" municipality="Doria and Beccari" name="Simon" pageId="27" pageNumber="28" stateProvince="Aden">Simon</location>
1882
</bibRefCitation>
as
<taxonomicName id="1EC64D2E1317BC6CFC58BD71FBF7E804" authorityName="Vinson" authorityYear="1863" box="[904,1138,303,329]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="borbonicus">
<emphasis id="EBB2EABF1317BC6CFC58BD71FBF7E804" box="[904,1138,303,329]" italics="true" pageId="27" pageNumber="28">
<location id="DC1960761317BC6CFC58BD71FC62E804" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17:DC1960761317BC6CFC58BD71FC62E804" box="[904,999,303,329]" country="Yemen" latitude="12.86" longLatPrecision="775" longitude="44.98" municipality="Doria and Beccari" name="Pholcus" pageId="27" pageNumber="28" stateProvince="Aden">Pholcus</location>
borbonicus
</emphasis>
</taxonomicName>
) (
<collectionCode id="BFD7AE681317BC6CFB5CBD71FB62E804" box="[1164,1255,303,329]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="27" pageNumber="28">MSNG</collectionCode>
);
</materialsCitation>
<materialsCitation id="69AE3CF01317BC6CFB2CBD71FBAEE821" ID-GBIF-Occurrence="3763211465" collectorName="E. A. Emerton" latitude="12.81" location="Aden" longLatPrecision="775" longitude="45.03" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Aden">
<specimenCount id="CFC0FD241317BC6CFB2CBD71FAACE804" box="[1276,1321,303,329]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
,
<collectingRegion id="1B02F84F1317BC6CFAE7BD71FAF2E804" box="[1335,1399,303,329]" country="Yemen" name="'Adan" pageId="27" pageNumber="28">Aden</collectingRegion>
[
<geoCoordinate id="BCF2506A1317BC6CFF14BD0CFEBDE820" box="[196,312,338,365]" degrees="12.81" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="12.81">12.81° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1317BC6CFE98BD0CFE32E820" box="[328,439,338,365]" degrees="45.03" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="555" value="45.03">45.03° E</geoCoordinate>
], date not given,
<collectorName id="7433537B1317BC6CFD42BD0CFCBDE820" box="[658,824,338,365]" pageId="27" pageNumber="28">E.A. Emerton</collectorName>
leg. (MCZ 34048);
</materialsCitation>
<materialsCitation id="69AE3CF01317BC6CFBE6BD0CFD70E8DD" ID-GBIF-Occurrence="3763211437" collectingDate="1938-10-15" collectionCode="NHMW" pageId="27" pageNumber="28" specimenCode="NHMW 353" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<specimenCount id="CFC0FD241317BC6CFBE6BD0CFBEDE820" box="[1078,1128,338,365]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="CFC0FD241317BC6CFBA9BD0CFB44E821" box="[1145,1217,338,364]" pageId="27" pageNumber="28" type="juvenile">1 juv.</specimenCount>
, same locality,
<date id="AD78106D1317BC6CFF6DBD2BFEDDE8DD" box="[189,344,373,400]" pageId="27" pageNumber="28" value="1938-10-15">
<collectingDate id="BD3CE9851317BC6CFF6DBD2BFEDDE8DD" box="[189,344,373,400]" pageId="27" pageNumber="28" value="1938-10-15">15 Oct. 1938</collectingDate>
</date>
, collector not given (
<specimenCode id="89609ED61317BC6CFD9FBD28FD6DE8DD" box="[591,744,373,400]" pageId="27" pageNumber="28">NHMW 353</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBDE9FA90E89C" blockId="27.[189,1301,439,466]" box="[189,1301,439,466]" pageId="27" pageNumber="28">
<materialsCitation id="69AE3CF01317BC6CFF6DBDE9FA90E89C" ID-GBIF-Occurrence="3763211376" box="[189,1301,439,466]" collectingDate="2013-06" collectionCode="ZMB" collectorName="M. Siyam" country="Sudan" elevation="450" latitude="15.32" location="New Halfa" longLatPrecision="771" longitude="35.6" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A1D1763D1317BC6CFF6DBDE9FEA2E89C" box="[189,295,439,465]" name="Sudan" pageId="27" pageNumber="28">SUDAN</collectingCountry>
:
<specimenCount id="CFC0FD241317BC6CFEE1BDE9FEDAE89F" box="[305,351,439,466]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<location id="DC1960761317BC6CFEBCBDE6FE69E89C" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17:DC1960761317BC6CFEBCBDE6FE69E89C" box="[364,492,439,466]" country="Sudan" latitude="15.32" longLatPrecision="771" longitude="35.6" name="New Halfa" pageId="27" pageNumber="28">New Halfa</location>
[
<geoCoordinate id="BCF2506A1317BC6CFE2ABDE9FDEFE89F" box="[506,618,439,466]" degrees="15.32" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="555" value="15.32">15.32° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1317BC6CFDA6BDE9FD64E89F" box="[630,737,439,466]" degrees="35.60" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="555" value="35.6">35.60° E</geoCoordinate>
],
<quantity id="1E3E9B481317BC6CFD25BDE9FCC5E89F" box="[757,832,439,466]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="27" pageNumber="28" unit="m" value="450.0">
<elevation id="52EBD19E1317BC6CFD25BDE9FCC5E89F" box="[757,832,439,466]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="27" pageNumber="28" unit="m" value="450.0">450 m</elevation>
</quantity>
a.s.l.,
<date id="AD78106D1317BC6CFC5CBDE6FB84E89C" box="[908,1025,439,466]" pageId="27" pageNumber="28" value="2013-06">
<collectingDate id="BD3CE9851317BC6CFC5CBDE6FB84E89C" box="[908,1025,439,466]" pageId="27" pageNumber="28" value="2013-06">Jun. 2013</collectingDate>
</date>
,
<collectorName id="7433537B1317BC6CFBDDBDE6FB04E89C" box="[1037,1153,439,466]" pageId="27" pageNumber="28">M. Siyam</collectorName>
leg. (
<collectionCode id="BFD7AE681317BC6CFB13BDE9FA8CE89C" box="[1219,1289,439,465]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="27" pageNumber="28" type="Museum">ZMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBDA7FEB7EB7B" blockId="27.[189,1399,505,566]" pageId="27" pageNumber="28">
<materialsCitation id="69AE3CF01317BC6CFF6DBDA7FEB7EB7B" ID-GBIF-Occurrence="3763211341" collectingDate="1920-06-20" collectingDateMax="1920-06-26" collectingDateMin="1920-06-20" collectionCode="AMNH" collectorName="B. Brown" country="Ethiopia" location="Didi Davvs" municipality="Abyssinia" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1">
<collectingCountry id="A1D1763D1317BC6CFF6DBDA7FECCEB5E" box="[189,329,505,531]" name="Ethiopia" pageId="27" pageNumber="28">ETHIOPIA</collectingCountry>
:
<specimenCount id="CFC0FD241317BC6CFE8ABDA7FE09EB5E" box="[346,396,505,531]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
, “
<collectingMunicipality id="391DACD71317BC6CFE7CBDA7FDA2EB5E" box="[428,551,505,531]" pageId="27" pageNumber="28">Abyssinia</collectingMunicipality>
,
<location id="DC1960761317BC6CFDE9BDA7FD4CEB5E" LSID="urn:lsid:plazi:treatment:516F87BB1315BC6AFDDEBAD1FC43EA17:DC1960761317BC6CFDE9BDA7FD4CEB5E" box="[569,713,505,531]" country="Ethiopia" municipality="Abyssinia" name="Didi Davvs" pageId="27" pageNumber="28">Didi Davvs</location>
” (uncertain spelling),
<date id="AD78106D1317BC6CFC31BDA7FB37EB5E" box="[993,1202,505,531]" pageId="27" pageNumber="28" value="1920-06-20" valueMax="1920-06-26" valueMin="1920-06-20">
<collectingDate id="BD3CE9851317BC6CFC31BDA7FB37EB5E" box="[993,1202,505,531]" pageId="27" pageNumber="28" value="1920-06-20" valueMax="1920-06-26" valueMin="1920-06-20">2026 Jun. 1920</collectingDate>
</date>
,
<collectorName id="7433537B1317BC6CFB15BDA7FABBEB5E" box="[1221,1342,505,531]" pageId="27" pageNumber="28">B. Brown</collectorName>
leg. (
<collectionCode id="BFD7AE681317BC6CFF15BE42FEA3EB7B" box="[197,294,540,566]" collectionName="USA, New York, New York, American Museum of Natural History" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="27" pageNumber="28">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBE00FC5BEBD1" blockId="27.[189,1398,606,668]" pageId="27" pageNumber="28">
Localities not clear:
<specimenCount id="CFC0FD241317BC6CFE60BE00FE5AEB34" box="[432,479,606,633]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
, “Urso 1283”, date not given, Kovacs leg. (
<collectionCode id="BFD7AE681317BC6CFC37BE00FBBAEB35" box="[999,1087,606,632]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="27" pageNumber="28">MZUF</collectionCode>
);
<specimenCount id="CFC0FD241317BC6CFB85BE00FB01EB34" box="[1109,1156,606,633]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
, “Afrique Orientale, Gibdo” [=
<collectingCountry id="A1D1763D1317BC6CFEECBEDFFE27EBD1" box="[316,418,641,668]" name="Djibouti" pageId="27" pageNumber="28">Djibouti</collectingCountry>
/
<collectingCountry id="A1D1763D1317BC6CFE7CBEDFFE7AEBD1" box="[428,511,641,668]" name="Eritrea" pageId="27" pageNumber="28">Eritrea</collectingCountry>
?],
<date id="AD78106D1317BC6CFDCABEDCFD53EBD1" box="[538,726,642,668]" pageId="27" pageNumber="28" value="1907-04" valueMax="1907-05-31" valueMin="1907-04-00">Apr.May 1907</date>
, Katona leg. (
<collectionCode id="BFD7AE681317BC6CFCA9BEDCFC54EBD1" box="[889,977,642,668]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="27" pageNumber="28">MZUF</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="91DC65261317BC6BFF6DBE9DFB73E8FF" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="D97936AD1317BC6CFF6DBE9DFECBEB90" blockId="27.[189,334,707,733]" box="[189,334,707,733]" pageId="27" pageNumber="28">
<heading id="823181C11317BC6CFF6DBE9DFECBEB90" bold="true" box="[189,334,707,733]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="EBB2EABF1317BC6CFF6DBE9DFECBEB90" bold="true" box="[189,334,707,733]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1317BC6CFF3ABEACFD91EA40" blockId="27.[189,1399,754,887]" box="[234,532,754,781]" pageId="27" pageNumber="28">
<emphasis id="EBB2EABF1317BC6CFF3ABEACFEAFEA41" bold="true" box="[234,298,754,780]" pageId="27" pageNumber="28">Male</emphasis>
(MNHN Ar 10169)
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBF48FB65EA3A" blockId="27.[189,1399,754,887]" pageId="27" pageNumber="28">MEASUREMENTS. Total body length 6.8, carapace width 3.5. Leg 1: 46.2 (12.1 + 1.6 + 13.4 + 16.2 + 2.9), tibia 2: 10.0, tibia 3: 7.6, tibia 4: 10.5; tibia 1 L/d: 34. Distance PMEPME 190 μm, diameter PME 190 μm, distance PMEALE 90 μm, distance AMEAME 40 μm, diameter AME 180 μm.</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBFC8FE68EDE5" blockId="27.[189,1399,918,1192]" pageId="27" pageNumber="28">
COLOR. Carapace ochre to light brown, with brown median band and narrow light brown margins; brown to light brown stripes radiate from median pit to margins; ocular area ochre to brown, clypeus with light brown band below AME, and dark brown rim (
<figureCitation id="41FD2A281317BC6CFD3BBF83FCC6EAB5" box="[747,835,989,1016]" captionStart="Figs 7682" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[203,1390,1102,1875]" captionTargetId="figure@26.[189,1398,1094,1885]" captionTargetPageId="26" captionText="Figs 7682. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen-Eritrea). 76. Male habitus. 7779. Male left pedipalp: prolateral, dorsal, and retrolateral views. 8082. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838566" httpUri="https://zenodo.org/record/3838566/files/figure.png" pageId="27" pageNumber="28">Fig. 76</figureCitation>
); legs ochre to light brown, with reddish brown to ochre prolateral marks on femora proximally, with brown rings on femora subdistally, patellae + tibiae proximally, and tibiae subdistally, tips of femora and tibiae not whitish (probably an artifact, see variation below); sternum ochre to light brown with narrow brown margins; abdomen beige to light brown with pale yellow and dark dots forming indistinct stripes from dorsal to lateral side of abdomen with large marks dorsally.
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DB897FE5DECDE" blockId="27.[189,1399,1224,1427]" pageId="27" pageNumber="28">
BODY. Ocular area slightly elevated; carapace with distinct median pit and distinctive posterior furrow; clypeus unmodified; sternum wider than long (2.4/1.7); chelicerae as in
<figureCitation id="41FD2A281317BC6CFB98B8B2FB51EC4B" box="[1096,1236,1259,1286]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="27" pageNumber="28">Figs 8385</figureCitation>
, with frontal row of 1619 modified (cone-shaped) hairs on each side situated on proximally strongly projecting processes; with several cone-shaped, sclerotized processes situated frontally near median line (arrows in
<figureCitation id="41FD2A281317BC6CFF0CB908FEE0EC3C" box="[220,357,1366,1393]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="27" pageNumber="28">Figs 83, 85</figureCitation>
); without stridulatory ridges (as in
<figureCitation id="41FD2A281317BC6CFCD7B908FCDAEC3D" box="[775,863,1366,1392]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="27" pageNumber="28">Fig. 83</figureCitation>
); Abdomen globose and high; gonopore with four epiandrous spigots.
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DB9EAFBE2EFEF" blockId="27.[189,1398,1459,1698]" pageId="27" pageNumber="28">
PALPS. As in
<figureCitation id="41FD2A281317BC6CFE84B9EDFE5EEC83" box="[340,475,1459,1486]" captionStart="Figs 7682" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[203,1390,1102,1875]" captionTargetId="figure@26.[189,1398,1094,1885]" captionTargetPageId="26" captionText="Figs 7682. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen-Eritrea). 76. Male habitus. 7779. Male left pedipalp: prolateral, dorsal, and retrolateral views. 8082. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838566" httpUri="https://zenodo.org/record/3838566/files/figure.png" pageId="27" pageNumber="28">Figs 7779</figureCitation>
; coxa unmodified, trochanter with short ventral projection, femur with distinct retrolateral process proximally, ventral membranous area proximally bordered on both sides by heavily sclerotized ridges, and small dorsal projection proximally; femur-patella hinges close together dorsally; patella very short; procursus with proximal dorsal process and weakly developed ventral pocket (
<figureCitation id="41FD2A281317BC6CFF16BA1FFE9AEF16" box="[198,287,1601,1627]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="27" pageNumber="28">Fig. 89</figureCitation>
), and distal dorsal notch on retrolateral margin; bulb with membranous embolus rising from base of process a; process a elongated, projecting towards retrolaterally with large subdistal hump; process b flattened, band-like, rectangular (
<figureCitation id="41FD2A281317BC6CFD67BAD6FC8BEFEF" box="[695,782,1672,1698]" captionStart="Figs 7682" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[203,1390,1102,1875]" captionTargetId="figure@26.[189,1398,1094,1885]" captionTargetPageId="26" captionText="Figs 7682. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen-Eritrea). 76. Male habitus. 7779. Male left pedipalp: prolateral, dorsal, and retrolateral views. 8082. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838566" httpUri="https://zenodo.org/record/3838566/files/figure.png" pageId="27" pageNumber="28">Fig. 81</figureCitation>
); processes c, d and e absent.
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBA9DFAA0EFB2" blockId="27.[189,1398,1730,1792]" pageId="27" pageNumber="28">LEGS.Without spines;with long curved hairs, especially on tibiae and metatarsi; retrolateral trichobothrium on tibia 1 at 8%; prolateral trichobothrium present on all tibiae; pseudosegmentation not visible.</paragraph>
<paragraph id="D97936AD1317BC6CFF3ABB7EFE28EE77" blockId="27.[189,1399,1823,2027]" box="[234,429,1823,1850]" pageId="27" pageNumber="28">
<emphasis id="EBB2EABF1317BC6CFF3ABB7EFEAFEE77" bold="true" box="[234,298,1824,1850]" pageId="27" pageNumber="28">Male</emphasis>
(variation)
</paragraph>
<paragraph id="D97936AD1317BC6CFF6DBB1DFC5DEEA6" blockId="27.[189,1399,1823,2027]" pageId="27" pageNumber="28">
Tibia
<quantity id="1E3E9B481317BC6CFED1BB1DFEABEE10" box="[257,302,1859,1885]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="28" unit="in" value="1.0">1 in</quantity>
3 other males: 11.4, 13.7, 14.2; color pattern on abdomen varies from pale without any marks to light brown with dark and pale dots forming large dorsal marks and fine lateral stripes; leg color varies from light brown to ochre, in fresh specimen from
<collectingCountry id="A1D1763D1317BC6CFCF7BBD4FCF5EEE9" box="[807,880,1930,1956]" name="Sudan" pageId="27" pageNumber="28">Sudan</collectingCountry>
tips of femora and tibiae whitish; elevated ocular area usually light brown; carapace pattern varies from narrow brown stripes to wide brown marks radiating from median pit to margin (
<figureCitation id="41FD2A281317BC6CFDA2BB8FFD4DEEA6" box="[626,712,2001,2027]" captionStart="Figs 7682" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[203,1390,1102,1875]" captionTargetId="figure@26.[189,1398,1094,1885]" captionTargetPageId="26" captionText="Figs 7682. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen-Eritrea). 76. Male habitus. 7779. Male left pedipalp: prolateral, dorsal, and retrolateral views. 8082. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. Scale lines: 76 = 2 mm; 7779 = 1 mm; 8082 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838566" httpUri="https://zenodo.org/record/3838566/files/figure.png" pageId="27" pageNumber="28">Fig. 76</figureCitation>
, cf.
<bibRefCitation id="BD574B5C1317BC6CFD2ABB8FFCF9EEA6" author="Koch" box="[762,892,2000,2027]" firstAuthor="Koch" pageId="27" pageNumber="28" refId="ref29158" refString="Koch L. 1875. Aegyptische und abyssinische Arachniden gesammelt von Herrn C. Jickeli. Bauer and Raspe, Nurnberg." type="book" year="1875">Koch 1875</bibRefCitation>
: fig. 1).
</paragraph>
<paragraph id="D97936AD1310BC6BFF3ABD55FEC0E868" blockId="28.[189,1399,267,434]" box="[234,325,267,293]" pageId="28" pageNumber="29">
<emphasis id="EBB2EABF1310BC6BFF3ABD55FEC0E868" bold="true" box="[234,325,267,293]" pageId="28" pageNumber="29">Female</emphasis>
</paragraph>
<paragraph id="D97936AD1310BC6BFF6DBD71FB73E8FF" blockId="28.[189,1399,267,434]" pageId="28" pageNumber="29">
In general similar to male; tibia
<quantity id="1E3E9B481310BC6BFDE2BD70FDDAE805" box="[562,607,302,328]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="28" pageNumber="29" unit="in" value="1.0">1 in</quantity>
<specimenCount id="CFC0FD241310BC6BFDB5BD70FD5EE805" box="[613,731,302,328]" pageId="28" pageNumber="29" type="female">2 females</specimenCount>
: 5.6, 9.8; stridulatory files laterally on chelicerae present; anterior epigynal projections (AEP) sclerotized, elongated (
<figureCitation id="41FD2A281310BC6BFC58BD0FFC67E821" box="[904,994,337,364]" captionStart="Figs 8389" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[195,1393,831,1810]" captionTargetId="figure@28.[189,1399,816,1816]" captionTargetPageId="28" captionText="Figs 8389. Artema kochi Kulczyński, 1901 (MNHN AR 10169 part, Yemen/Eritrea). 8385. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line). 86. Epigynum and anterior epigynal projections (AEP). 8788. Female abdomen, lateral and ventral views. 89. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="28" pageNumber="29">Fig. 86</figureCitation>
); epigynal plate sclerotized, wide rectangular (~3 × wider than long) projecting anteriorly between AEP, with median posterior margin gently protruding; some females with two dark dots at base of triangular anterior projection.
</paragraph>
</subSubSection>
<subSubSection id="91DC65261310BC6BFF6DBD87FB34EB0B" pageId="28" pageNumber="29" type="distribution">
<paragraph id="D97936AD1310BC6BFF6DBD87FED3E8BE" blockId="28.[189,342,473,499]" box="[189,342,473,499]" pageId="28" pageNumber="29">
<heading id="823181C11310BC6BFF6DBD87FED3E8BE" bold="true" box="[189,342,473,499]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="EBB2EABF1310BC6BFF6DBD87FED3E8BE" bold="true" box="[189,342,473,499]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1310BC6BFF6DBE56FB34EB0B" blockId="28.[189,1398,520,582]" pageId="28" pageNumber="29">
We examined specimens from
<collectingCountry id="A1D1763D1310BC6BFDEFBE56FD20EB6F" box="[575,677,520,546]" name="Ethiopia" pageId="28" pageNumber="29">Ethiopia</collectingCountry>
,
<collectingCountry id="A1D1763D1310BC6BFD67BE56FC8FEB6F" box="[695,778,520,546]" name="Eritrea" pageId="28" pageNumber="29">Eritrea</collectingCountry>
, and
<collectingCountry id="A1D1763D1310BC6BFC83BE56FC20EB6F" box="[851,933,520,546]" name="Yemen" pageId="28" pageNumber="29">Yemen</collectingCountry>
(
<figureCitation id="41FD2A281310BC6BFC6BBE56FB83EB6F" box="[955,1030,520,546]" captionStart="Figs 12. 1" captionStartId="5.[189,241,1647,1673]" captionTargetBox="[190,1397,377,1570]" captionTargetId="figure@5.[187,1399,374,1571]" captionTargetPageId="5" captionText="Figs 12. 1. Known distributions of species of Artema except A. atlanta Walckenaer, 1837. The Japanese record of A. doriae Thorell, 1881 is not shown as it is based on a single introduced specimen. 2. Known distribution of Artema atlanta. Green dots: specimens examined; black dots: specimens not examined (data from: Banks 1898; Beatty et al. 2008; Blackwell 1958; Bösenberg &amp; Lenz 1895; Brignoli 1981; Bryant 1940, 1948; Caporiacco 1928, 1948, 1949; Chamberlin 1924; Chrysanthus 1967; Colmenares 2008; Denis 1953; Doleschall 1857; Dyal 1935; Franganillo 1926a; Gao &amp; Li 2010; Gertsch &amp; Davis 1942; Gonzalez-Sponga 2005; Karsch 1879, 1892; Leardi in Airaghi 1902; Lessert 1936; Majumder &amp; Biswas 1993; Marples 1955; Mello-Leitão 1942; Millot 1946; Moenkhaus 1898; F.O.Pickard-Cambridge 1902; Petrunkevitch 1925, 1929; Pocock 1900; Saaristo 2001; Simon 1885, 1894; Taczanowsky 1874; Thorell 1895, 1898; Vinson 1863; Yaginuma 1970). Records from Israel, Belgium, and Britain are not shown as they are based on single introduced specimens." figureDoi="http://doi.org/10.5281/zenodo.3838546" httpUri="https://zenodo.org/record/3838546/files/figure.png" pageId="28" pageNumber="29">Fig. 1</figureCitation>
). The original description by
<bibRefCitation id="BD574B5C1310BC6BFF6DBE75FE1DEB08" author="Kulczynski" box="[189,408,555,581]" firstAuthor="Kulczynski" pageId="28" pageNumber="29" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński (1901)</bibRefCitation>
was apparently based on specimens from
<collectingCountry id="A1D1763D1310BC6BFC58BE75FC4BEB08" box="[904,974,555,581]" name="Egypt" pageId="28" pageNumber="29">Egypt</collectingCountry>
(see Notes below).
</paragraph>
</subSubSection>
<subSubSection id="91DC65261310BC6AFF6DBE32FC43EA17" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
<paragraph id="D97936AD1310BC6BFF6DBE32FE86EBCB" blockId="28.[189,259,620,646]" box="[189,259,620,646]" pageId="28" pageNumber="29">
<heading id="823181C11310BC6BFF6DBE32FE86EBCB" bold="true" box="[189,259,620,646]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="EBB2EABF1310BC6BFF6DBE32FE86EBCB" bold="true" box="[189,259,620,646]" pageId="28" pageNumber="29">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1310BC6AFF6DBEC5FB67EB52" blockId="28.[189,1399,667,764]" lastBlockId="29.[189,1399,267,543]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
The original description by
<bibRefCitation id="BD574B5C1310BC6BFDC7BEC5FD7DEBF8" author="Kulczynski" box="[535,760,667,693]" firstAuthor="Kulczynski" pageId="28" pageNumber="29" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński (1901)</bibRefCitation>
is based on two female specimens, but it does not specify where these females were collected. We assume that Kulczyński examined and illustrated the specimens from
<collectingCountry id="A1D1763D1310BC6BFEACBEBCFE40EBB1" box="[380,453,738,764]" name="Egypt" pageId="28" pageNumber="29">Egypt</collectingCountry>
, near
<collectingRegion id="1B02F84F1310BC6BFDD8BEBFFDC9EBB6" box="[520,588,737,763]" country="Egypt" name="Al Qahirah" pageId="28" pageNumber="29">Cairo</collectingRegion>
, that L.
<bibRefCitation id="BD574B5C1310BC6BFD77BEBFFCBEEBB1" author="Koch" box="[679,827,737,764]" firstAuthor="Koch" pageId="28" pageNumber="29" refId="ref29158" refString="Koch L. 1875. Aegyptische und abyssinische Arachniden gesammelt von Herrn C. Jickeli. Bauer and Raspe, Nurnberg." type="book" year="1875">Koch (1875)</bibRefCitation>
had misidentified as
<taxonomicName id="1EC64D2E1310BC6AFBFEBEBFFF7DE868" authority="Vinson, 1863" authorityName="Vinson" authorityYear="1863" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="29" lastPageNumber="30" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="borbonicus">
<emphasis id="EBB2EABF1310BC6BFBFEBEBFFA92EBB6" box="[1070,1303,737,763]" italics="true" pageId="28" pageNumber="29">Pholcus borbonicus</emphasis>
<bibRefCitation id="BD574B5C1310BC6AFACCBEBFFF7DE868" author="Vinson" firstAuthor="Vinson" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" refId="ref30850" refString="Vinson A. 1863. Araneides des iles de la Reunion, Maurice et Madagascar. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 125517" type="book" year="1863">Vinson, 1863</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="1EC64D2E1311BC6AFEF7BD52FE1BE868" authorityName="Walckenaer" authorityYear="1837" box="[295,414,267,294]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="atlanta">
<emphasis id="EBB2EABF1311BC6AFEF7BD52FE1BE868" box="[295,414,267,294]" italics="true" pageId="29" pageNumber="30">A. atlanta</emphasis>
</taxonomicName>
). Kulczyński also mentioned specimens from “Massaua” (
<collectingCountry id="A1D1763D1311BC6AFBB6BD55FB3EE868" box="[1126,1211,267,293]" name="Eritrea" pageId="29" pageNumber="30">Eritrea</collectingCountry>
, Massawa) that
<bibRefCitation id="BD574B5C1311BC6AFF6DBD71FEEDE804" author="Thorell" box="[189,360,303,329]" firstAuthor="Thorell" pageId="29" pageNumber="30" pagination="1 - 727" refId="ref30571" refString="Thorell T. 1881. Studi sui ragni Malesi e Papuani. III. Ragni dell'Austro Malesia e del Capo York, conservati nel Museo civico di storia naturale di Genova. Annali del Museo civico di storia naturale di Genova 17: 1 - 727." type="journal article" year="1881">Thorell (1881)</bibRefCitation>
had described in some detail but that he had equally misidentified as
<taxonomicName id="1EC64D2E1311BC6AFB57BD71FAEAE804" authorityName="Vinson" authorityYear="1863" box="[1159,1391,303,329]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="borbonicus">
<emphasis id="EBB2EABF1311BC6AFB57BD71FAEAE804" box="[1159,1391,303,329]" italics="true" pageId="29" pageNumber="30">Pholcus borbonicus</emphasis>
</taxonomicName>
. Since this material from Massawa contains males and females, we conclude that it was not available to Kulczyński. However, we chose to designate a
<typeStatus id="067D880F1311BC6AFCD8BD29FCE0E8DC" box="[776,869,375,401]" pageId="29" pageNumber="30" type="neotype">neotype</typeStatus>
from the material originating from Massawa for three reasons: (1) Kulczyński neither designated type specimens nor specified a type locality; (2) we were not able to locate Kochs specimens from
<collectingCountry id="A1D1763D1311BC6AFCF4BDE0FCEFE895" box="[804,874,446,472]" name="Egypt" pageId="29" pageNumber="30">Egypt</collectingCountry>
(they are not in ZMH or ZMB); (3) a good illustration by
<bibRefCitation id="BD574B5C1311BC6AFEB9BDBFFDC0E8B6" author="Kulczynski" box="[361,581,481,507]" firstAuthor="Kulczynski" pageId="29" pageNumber="30" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński (1901)</bibRefCitation>
of the highly distinctive epigynum of
<taxonomicName id="1EC64D2E1311BC6AFBD9BDBFFBEFE8B6" authorityName="Kulczynski" authorityYear="1901" box="[1033,1130,481,507]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1311BC6AFBD9BDBFFBEFE8B6" box="[1033,1130,481,507]" italics="true" pageId="29" pageNumber="30">A. kochi</emphasis>
</taxonomicName>
leaves little doubt that the specimens available to him are, in fact, conspecific with the specimens from Massawa.
</paragraph>
<caption id="8DB966251310BC6BFF6DBB1CFD4EEEA7" ID-DOI="http://doi.org/10.5281/zenodo.3838568" ID-Zenodo-Dep="3838568" httpUri="https://zenodo.org/record/3838568/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,241,1858,1884]" targetBox="[195,1393,831,1810]" targetPageId="28">
<paragraph id="D97936AD1310BC6BFF6DBB1CFD4EEEA7" blockId="28.[189,1399,1858,2026]" pageId="28" pageNumber="29">
<emphasis id="EBB2EABF1310BC6BFF6DBB1CFEC8EE11" bold="true" box="[189,333,1858,1884]" pageId="28" pageNumber="29">Figs 8389.</emphasis>
<taxonomicName id="1EC64D2E1310BC6BFE88BB1DFD5DEE10" authority="Kulczynski, 1901" authorityName="Kulczynski" authorityYear="1901" box="[344,728,1858,1885]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1310BC6BFE88BB1DFE7FEE10" box="[344,506,1859,1885]" italics="true" pageId="28" pageNumber="29">Artema kochi</emphasis>
<bibRefCitation id="BD574B5C1310BC6BFDD5BB1CFD5DEE10" author="Kulczynski" box="[517,728,1858,1885]" firstAuthor="Kulczynski" pageId="28" pageNumber="29" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński, 1901</bibRefCitation>
</taxonomicName>
(MNHN AR 10169 part, Yemen/Eritrea).
<emphasis id="EBB2EABF1310BC6BFB0EBB1CFAA2EE11" bold="true" box="[1246,1319,1858,1884]" pageId="28" pageNumber="29">8385</emphasis>
. Male chelicerae, lateral and frontal views (arrows: distinctive sclerotized processes frontally near median line).
<emphasis id="EBB2EABF1310BC6BFED6BBD7FEA6EEEE" bold="true" box="[262,291,1929,1955]" pageId="28" pageNumber="29">86</emphasis>
. Epigynum and anterior epigynal projections (AEP).
<emphasis id="EBB2EABF1310BC6BFC69BBD7FB87EEEE" bold="true" box="[953,1026,1929,1955]" pageId="28" pageNumber="29">8788</emphasis>
. Female abdomen, lateral and ventral views.
<emphasis id="EBB2EABF1310BC6BFEBBBBF2FE0DEE8B" bold="true" box="[363,392,1964,1990]" pageId="28" pageNumber="29">89</emphasis>
. Procursus, showing ventral pocket (vp) and dorsal process (dp). Scale lines: 8385, 89 = 0.25 mm; 86 = 0.5 mm; 8788 = 1 mm.
</paragraph>
</caption>
<paragraph id="D97936AD1311BC6AFF6DBE18FC43EA17" blockId="29.[189,1399,582,858]" pageId="29" pageNumber="30">
Much less obvious is how
<bibRefCitation id="BD574B5C1311BC6AFE38BE18FD0CEB2C" author="Simon" box="[488,649,582,609]" firstAuthor="Simon" pageId="29" pageNumber="30" pagination="419 - 438" refId="ref30437" refString="Simon E. 1908. Etude sur les arachnides recueillis par M. le Dr Klaptocz en Tripolitaine. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 26: 419 - 438." type="journal article" year="1908">Simon (1908)</bibRefCitation>
, after having studied specimens of “
<taxonomicName id="1EC64D2E1311BC6AFBF5BE19FB4FEB2D" authorityName="Walckenaer" authorityYear="1837" box="[1061,1226,583,609]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mauriciana">
<emphasis id="EBB2EABF1311BC6AFBF5BE19FB4FEB2D" box="[1061,1226,583,609]" italics="true" pageId="29" pageNumber="30">A. mauriciana</emphasis>
</taxonomicName>
” (=
<taxonomicName id="1EC64D2E1311BC6AFB2CBE19FAE8EB2D" authorityName="Walckenaer" authorityYear="1837" box="[1276,1389,582,609]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="atlanta">
<emphasis id="EBB2EABF1311BC6AFB2CBE19FAE8EB2D" box="[1276,1389,582,609]" italics="true" pageId="29" pageNumber="30">A. atlanta</emphasis>
</taxonomicName>
) from many localities, arrived at the conclusion that the drawings of
<bibRefCitation id="BD574B5C1311BC6AFBD5BE34FB61EBC8" author="Kulczynski" box="[1029,1252,618,645]" firstAuthor="Kulczynski" pageId="29" pageNumber="30" pagination="1 - 64" refId="ref29180" refString="Kulczynski W. 1901. Arachnoidea in Colonia Erythraea a Dre K. M. Levander collecta. Rozprawy i Sprawozdania z Posiedzen Wydzialu Matematyczno Przyrodniczego Akademji Umiejetnosci, Krakow 41: 1 - 64." type="journal article" year="1901">Kulczyński (1901)</bibRefCitation>
represented merely slight individual variation and that
<taxonomicName id="1EC64D2E1311BC6AFD68BED0FC9FEBE5" authorityName="Kulczynski" authorityYear="1901" box="[696,794,654,680]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1311BC6AFD68BED0FC9FEBE5" box="[696,794,654,680]" italics="true" pageId="29" pageNumber="30">A. kochi</emphasis>
</taxonomicName>
was a synonym of “
<taxonomicName id="1EC64D2E1311BC6AFBDFBED0FB3DEBEA" authorityName="Walckenaer" authorityYear="1837" box="[1039,1208,654,680]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mauriciana">
<emphasis id="EBB2EABF1311BC6AFBDFBED0FB3DEBEA" box="[1039,1208,654,680]" italics="true" pageId="29" pageNumber="30">A. mauriciana</emphasis>
</taxonomicName>
”. This is all the more mysterious because Simon himself had collected
<taxonomicName id="1EC64D2E1311BC6AFC91BEECFBB7EB81" authority="in Massawa" authorityName="in Massawa" box="[833,1074,689,716]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1311BC6AFC91BEECFC24EB86" box="[833,929,689,716]" italics="true" pageId="29" pageNumber="30">A. kochi</emphasis>
in Massawa
</taxonomicName>
and
<collectingRegion id="1B02F84F1311BC6AFBB9BEEFFB2CEB86" box="[1129,1193,689,715]" country="Yemen" name="'Adan" pageId="29" pageNumber="30">Aden</collectingRegion>
(in 1889) and his own material (in MNHN, examined) includes not only females, but also the equally distinctive males. We found both species that Simon collected from Massawa and
<collectingRegion id="1B02F84F1311BC6AFC72BEA6FC60EA5F" box="[930,997,760,786]" country="Yemen" name="'Adan" pageId="29" pageNumber="30">Aden</collectingRegion>
,
<taxonomicName id="1EC64D2E1311BC6AFC3FBEA7FBE7EA5F" authorityName="Walckenaer" authorityYear="1837" box="[1007,1122,760,787]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="atlanta">
<emphasis id="EBB2EABF1311BC6AFC3FBEA7FBE7EA5F" box="[1007,1122,760,787]" italics="true" pageId="29" pageNumber="30">A. atlanta</emphasis>
</taxonomicName>
and
<taxonomicName id="1EC64D2E1311BC6AFB49BEA7FB7CEA5F" box="[1177,1273,760,787]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kochi">
<emphasis id="EBB2EABF1311BC6AFB49BEA7FB7CEA5F" box="[1177,1273,760,787]" italics="true" pageId="29" pageNumber="30">A. kochi</emphasis>
</taxonomicName>
, deposited in the same vial (Simon collection number 393), so we assume that Simon did not examine all the material in detail but only part of it that happened to be
<taxonomicName id="1EC64D2E1311BC6AFC9BBF1EFC3AEA17" authorityName="Walckenaer" authorityYear="1837" box="[843,959,832,858]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="atlanta">
<emphasis id="EBB2EABF1311BC6AFC9BBF1EFC3AEA17" box="[843,959,832,858]" italics="true" pageId="29" pageNumber="30">A. atlanta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>