175 lines
19 KiB
XML
175 lines
19 KiB
XML
<document id="0DC2F8804B39FECC78B21E6C7D1B9A66" ID-CLB-Dataset="299337" ID-DOI="10.5281/zenodo.11067222" ID-GBIF-Dataset="1cb742f7-205f-4cc5-9136-b613d5ab059d" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067222" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1714052073787" checkinUser="felipe" docAuthor="LaBonte, James R. & Maddison, David R." docDate="2023" docId="51533818FFE5477DFEE9B823FB3AFB45" docLanguage="en" docName="ProcCalifAcadSci.67.18.401-432.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (18)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Anillini" docType="key" docVersion="7" lastPageNumber="409" masterDocId="AD6A4060FFED4775FFFEBA21FFB5FFA3" masterDocTitle="Medusapyga LaBonte and Maddison, a New Genus of Anillini (Coleoptera: Carabidae: Trechinae) from the Pacific Northwest of the United States" masterLastPageNumber="432" masterPageNumber="401" pageNumber="409" updateTime="1719875109990" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="147AED7E051D2A06DD1045CF55620D41">
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<mods:title id="21483D14F46BA3A51789FCD527C645DE">Medusapyga LaBonte and Maddison, a New Genus of Anillini (Coleoptera: Carabidae: Trechinae) from the Pacific Northwest of the United States</mods:title>
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<mods:name id="34BE45EE48E9FFA376B215403FE56232" type="personal">
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<mods:roleTerm id="30C66E6E9C0E634DCB88917675B6D47B">Author</mods:roleTerm>
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<mods:namePart id="0557869231A652028976C112D2BB1CDC">LaBonte, James R.</mods:namePart>
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<mods:affiliation id="BEA334D18F05919BB07DA58DA0D4147C">1570 Kenard Street NW, Salem, OR 97304, USA.</mods:affiliation>
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<mods:name id="19A1B7BA7651A891A5BC6E1A873BD37E" type="personal">
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<mods:roleTerm id="20F74D20D5BC07E300431FA146C58451">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="05388743F7AD2A56BD0EFA7298D4A12F">Maddison, David R.</mods:namePart>
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<mods:affiliation id="CA62383CCA33DD87870A0876859AF220">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="BCE4047EB3790F0CC513F9B8F92B1A7B">text</mods:typeOfResource>
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<mods:title id="6207844DD0705116E6723C88C3408255">Proceedings of the California Academy of Sciences</mods:title>
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</mods:titleInfo>
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<mods:part id="97F8FE383870F58D4590281404827D89">
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<mods:date id="23192E870BA3A9D5E93021E328BA17D6">2023</mods:date>
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<mods:number id="43226A25C191E142FE44124F615D256F">2023-02-15</mods:number>
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<mods:detail id="6EDFE6118E6FF221E5ABE711566DD2C1" type="volume">
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<mods:number id="15902C6B53DB8EBEA16A15BB8D3F6C28">67</mods:number>
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<mods:number id="E65A3BBF45DBC2BAC55B8025B6A23FEE">18</mods:number>
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<mods:start id="7E16D27A0A72037E66917579BA027E58">401</mods:start>
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<mods:classification id="0AB8DA6227AD4D349A046C5D8D3BF8EE">journal article</mods:classification>
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<mods:identifier id="403E24F5C140F280CC6D8BD178BDCCBC" type="CLB-Dataset">299337</mods:identifier>
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<mods:identifier id="020E31BFCD416E39FCD530D9D2BB3BED" type="DOI">10.5281/zenodo.11067222</mods:identifier>
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<mods:identifier id="C4B4705DD3815D500C1BF0615DD05A1A" type="GBIF-Dataset">1cb742f7-205f-4cc5-9136-b613d5ab059d</mods:identifier>
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<mods:identifier id="D2EDAB1F3AD458B5A9ED27E9E5CDDA9C" type="ISSN">0068-547X</mods:identifier>
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<mods:identifier id="6B7DD51AF37029BA7AE0ECA72BA4B34D" type="Zenodo-Dep">11067222</mods:identifier>
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<treatment id="51533818FFE5477DFEE9B823FB3AFB45" ID-DOI="http://doi.org/10.5281/zenodo.12613591" ID-GBIF-Taxon="234396641" ID-Zenodo-Dep="12613591" LSID="urn:lsid:plazi:treatment:51533818FFE5477DFEE9B823FB3AFB45" httpUri="http://treatment.plazi.org/id/51533818FFE5477DFEE9B823FB3AFB45" lastPageNumber="409" pageId="8" pageNumber="409">
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<subSubSection id="91E0DA85FFE5477DFEE9B823FC4CFDBA" box="[279,1017,514,537]" pageId="8" pageNumber="409" type="nomenclature">
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<paragraph id="D945890EFFE5477DFEE9B823FC4CFDBA" blockId="8.[279,1017,514,537]" box="[279,1017,514,537]" pageId="8" pageNumber="409">
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<heading id="820D3E62FFE5477DFEE9B823FC4CFDBA" bold="true" box="[279,1017,514,537]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="409" reason="5">
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<emphasis id="EB8E551CFFE5477DFEE9B823FC4CFDBA" bold="true" box="[279,1017,514,537]" pageId="8" pageNumber="409">
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Key to the Genera of North American (North of
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<collectingRegion id="1B3E47ECFFE5477DFCBCB823FC22FDBA" box="[834,919,514,537]" country="Mexico" name="Mexico" pageId="8" pageNumber="409">Mexico</collectingRegion>
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)
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<taxonomicName id="1EFAF28DFFE5477DFC5DB823FC4CFDBA" box="[931,1017,514,537]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="403" phylum="Arthropoda" rank="tribe" tribe="Anillini">Anillini</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="91E0DA85FFE5477DFF85B80CFB3AFB45" pageId="8" pageNumber="409" type="key">
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<keyStep id="620E91ABFFE5477DFF85B80CFB3AFCAE" pageId="8" pageNumber="409">
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<paragraph id="D945890EFFE5477DFF85B80CFB3AFD3A" blockId="8.[123,1174,557,781]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85B80CFB3AFD3A" pageId="8" pageNumber="409">
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1. Mandibles extremely long and narrow, much longer than head behind the antennal insertions (
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<figureCitation id="41C1958BFFE5477DFBB2B80CFB3DFDE0" box="[1100,1160,557,579]" captionStart="FIGURE" captionStartId="9.[874,886,915,934]" captionTargetBox="[865,1147,221,892]" captionTargetId="figure-274@9.[838,1174,216,911]" captionText="FIGURE 3. Dorsal habitus of female Anillaspis explanata from USA: Cali- fornia: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12612687" httpUri="https://zenodo.org/record/12612687/files/figure.png" pageId="8" pageNumber="409">Fig. 3</figureCitation>
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); sides of head behind antennal insertions more-or-less parallel and straight; frontal furrows deep and long, extending to occiput (
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<figureCitation id="41C1958BFFE5477DFE7DB847FE0AFDDF" box="[387,447,614,636]" captionStart="FIGURE" captionStartId="9.[874,886,915,934]" captionTargetBox="[865,1147,221,892]" captionTargetId="figure-274@9.[838,1174,216,911]" captionText="FIGURE 3. Dorsal habitus of female Anillaspis explanata from USA: Cali- fornia: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12612687" httpUri="https://zenodo.org/record/12612687/files/figure.png" pageId="8" pageNumber="409">Fig. 3</figureCitation>
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); lateral explanation of elytron very broad in anterior half, abruptly narrowing thereafter (
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<figureCitation id="41C1958BFFE5477DFEA1B8A2FE2CFD3A" box="[351,409,643,665]" captionStart="FIGURE" captionStartId="9.[874,886,915,934]" captionTargetBox="[865,1147,221,892]" captionTargetId="figure-274@9.[838,1174,216,911]" captionText="FIGURE 3. Dorsal habitus of female Anillaspis explanata from USA: Cali- fornia: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12612687" httpUri="https://zenodo.org/record/12612687/files/figure.png" pageId="8" pageNumber="409">Fig. 3</figureCitation>
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); geographic range Sierra
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<collectingRegion id="1B3E47ECFFE5477DFD5FB8A2FCEDFD3A" box="[673,856,643,665]" isEnumeration="true" pageId="8" pageNumber="409">Nevada, California</collectingRegion>
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............
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<taxonomicName id="1EFAF28DFFE5477DFC13B8A2FB3AFD3A" ID-CoL="8KV4S" authority="Casey" authorityName="Casey" authorityYear="1918" box="[1005,1167,643,665]" class="Insecta" family="Carabidae" genus="Anillaspis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
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<emphasis id="EB8E551CFFE5477DFC13B8A2FBF8FD3A" box="[1005,1101,643,665]" italics="true" pageId="8" pageNumber="409">Anillaspis</emphasis>
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Casey
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="D945890EFFE5477DFF85B880FB3AFCAE" blockId="8.[123,1174,557,781]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85B880FB3AFCAE" pageId="8" pageNumber="409">
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– Mandibles shorter and stouter, shorter than head behind the antennal insertions (
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<figureCitation id="41C1958BFFE5477DFC43B881FC4CFD15" box="[957,1017,672,694]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="8" pageNumber="409">Fig. 2</figureCitation>
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); sides of head behind antennal insertions rounded; frontal furrows shallow and short, extending no more than halfway to occiput (
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<figureCitation id="41C1958BFFE5477DFEE7B8FBFEE0FD53" box="[281,341,730,752]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="8" pageNumber="409">Fig. 2</figureCitation>
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); lateral explanation of elytra much narrower and broadest only near the humeri or in the anterior quarter, gradually narrowing until terminating before the apex (
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<figureCitation id="41C1958BFFE5477DFC7BB8D6FC0AFCAE" box="[901,959,759,781]" captionStart="FIGURE" captionStartId="2.[155,167,1150,1169]" captionTargetBox="[137,1130,244,1130]" captionTargetId="figure-257@2.[120,1176,216,1147]" captionTargetPageId="2" captionText="FIGURE 2. Dorsal habitus. (A) Medusapyga alsea, female. (B) M. chehalis, female. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11067226" httpUri="https://zenodo.org/record/11067226/files/figure.png" pageId="8" pageNumber="409">Fig. 2</figureCitation>
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)............... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="620E91ABFFE5477DFF85B901FB3AFC2E" pageId="8" pageNumber="409">
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<paragraph id="D945890EFFE5477DFF85B901FB3AFCF0" blockId="8.[123,1174,800,909]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85B901FB3AFCF0" pageId="8" pageNumber="409">
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2. Mentum with a pair of setae near apex of anteromedial tooth; geographic range
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<collectingRegion id="1B3E47ECFFE5477DFC4FB901FBDCFC95" box="[945,1129,800,822]" isEnumeration="true" pageId="8" pageNumber="409">California, Oregon</collectingRegion>
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, and
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<collectingRegion id="1B3E47ECFFE5477DFF55B91CFEA8FCF0" box="[171,285,829,851]" country="United States of America" name="Washington" pageId="8" pageNumber="409">Washington</collectingRegion>
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........................................................................ 3
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</keyLead>
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</paragraph>
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<paragraph id="D945890EFFE5477DFF85B97AFB3AFC2E" blockId="8.[123,1174,800,909]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85B97AFB3AFC2E" pageId="8" pageNumber="409">
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– If two setae are present on mentum tooth, then they are at base of tooth, well posterior of apex; geographic range eastern
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<collectingCountry id="A1EDC99EFFE5477DFE7EB956FDB4FC2E" box="[384,513,887,909]" name="United States of America" pageId="8" pageNumber="409">United States</collectingCountry>
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west to
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<collectingRegion id="1B3E47ECFFE5477DFDADB956FD1DFC2E" box="[595,680,887,909]" country="United States of America" name="Missouri" pageId="8" pageNumber="409">Missouri</collectingRegion>
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and
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<collectingRegion id="1B3E47ECFFE5477DFD26B959FCA5FC2E" box="[728,784,888,909]" country="United States of America" name="Texas" pageId="8" pageNumber="409">Texas</collectingRegion>
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.............................. 4
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="620E91ABFFE5477DFF85B981FB3BFBC7" pageId="8" pageNumber="409">
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<paragraph id="D945890EFFE5477DFF85B981FB3AFBAE" blockId="8.[123,1174,928,1124]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85B981FB3AFBAE" pageId="8" pageNumber="409">
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3. Male profemur with a spine on ventrobasal face (
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<figureCitation id="41C1958BFFE5477DFD70B981FD4FFC15" box="[654,762,928,950]" captionStart="FIGURE" captionStartId="9.[157,169,1658,1677]" captionTargetBox="[129,1168,1135,1647]" captionTargetId="figure-299@9.[120,1176,1127,1655]" captionTargetPageId="9" captionText="FIGURE 4. Left profemora, anterior view. (A) Medusapyga alsea, male. (B) M. alsea, female. (C) M. chehalis, male. (D) M. chehalis, female. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.11067228" httpUri="https://zenodo.org/record/11067228/files/figure.png" pageId="8" pageNumber="409">Fig. 4A, C</figureCitation>
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); male protarsus with tarsomeres 1 and 2 strongly asymmetrically dilated (
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<figureCitation id="41C1958BFFE5477DFE12B99CFD9DFC70" box="[492,552,957,979]" captionStart="Figure 5" captionStartId="10.[879,935,563,582]" captionTargetBox="[849,1170,220,542]" captionTargetId="figure-335@10.[844,1176,216,547]" captionText="Figure 5 (above). Male right pro- tarsi, dorsal view. (A) Medusapyga alsea. (B) M. chehalis. The view of the two specimens is not exactly equivalent, which accounts for some differences between the two images. Scale bars 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12612690" httpUri="https://zenodo.org/record/12612690/files/figure.png" pageId="8" pageNumber="409">Fig. 5</figureCitation>
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); female genitalia with a field of long bifurcate setae scattered throughout the apical region of laterotergite IX (
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<figureCitation id="41C1958BFFE5477DFD83B9FBFD0CFC53" box="[637,697,986,1008]" captionStart="Figure 6" captionStartId="10.[155,211,1666,1685]" captionTargetBox="[143,1176,1168,1651]" captionTargetId="figure-375@10.[120,1176,1168,1651]" captionTargetPageId="10" captionText="Figure 6. Female genitalia of Medusapyga alsea from the type locality, ventral view. (A) Overview of voucher V101505. Scale bar 100 µm. (B) Closeup of voucher V101504, showing bifurcate setae on laterotergites. Scale bar 50 µm." figureDoi="http://doi.org/10.5281/zenodo.11067230" httpUri="https://zenodo.org/record/11067230/files/figure.png" pageId="8" pageNumber="409">Fig. 6</figureCitation>
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); geographical range Pacific Northwestern United States (
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<collectingRegion id="1B3E47ECFFE5477DFEF0B9D6FE4EFBAE" box="[270,507,1015,1037]" isEnumeration="true" pageId="8" pageNumber="409">Oregon and Washington</collectingRegion>
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)........................
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<taxonomicName id="1EFAF28DFFE5477DFCD6B9D9FB3AFBAE" authority="LaBonte and Maddison" authorityName="LaBonte and Maddison" box="[808,1167,1015,1037]" class="Insecta" family="Carabidae" genus="Medusapyga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
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<emphasis id="EB8E551CFFE5477DFCD6B9D9FC10FBAE" bold="true" box="[808,933,1016,1037]" italics="true" pageId="8" pageNumber="409">Medusapyga</emphasis>
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LaBonte and Maddison
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="D945890EFFE5477DFF85BE34FB3BFBC7" blockId="8.[123,1174,928,1124]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85BE34FB3BFBC7" pageId="8" pageNumber="409">
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– Male profemur without a spine on ventrobasal face; male protarsus with only tarsomere 1 dilated; each lateral tergite of female genitalia with a field of simple setae on the venter along medial or apical edges; geographical range
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<collectingRegion id="1B3E47ECFFE5477DFE97BE6FFE7FFBC7" box="[361,458,1102,1124]" country="United States of America" name="California" pageId="8" pageNumber="409">California</collectingRegion>
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............................................
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<taxonomicName id="1EFAF28DFFE5477DFC1BBE6FFB3BFBC7" ID-CoL="8KV4V" authority="Jeannel" authorityName="Jeannel" box="[997,1166,1102,1124]" class="Insecta" family="Carabidae" genus="Anillodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
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<emphasis id="EB8E551CFFE5477DFC1BBE6FFBF5FBC7" box="[997,1088,1102,1124]" italics="true" pageId="8" pageNumber="409">Anillodes</emphasis>
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Jeannel
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="620E91ABFFE5477DFF85BE56FB3AFB45" pageId="8" pageNumber="409">
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<paragraph id="D945890EFFE5477DFF85BE56FB3BFB09" blockId="8.[123,1174,1143,1254]" pageId="8" pageNumber="409">
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<keyLead id="620B2A3BFFE5477DFF85BE56FB3BFB09" pageId="8" pageNumber="409">
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4. Both sexes with apical margin of last visible abdominal ventrite evenly arcuate; right paramere present; geographic range eastern
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<collectingCountry id="A1EDC99EFFE5477DFE5CBEB5FD96FB09" box="[418,547,1172,1194]" name="United States of America" pageId="8" pageNumber="409">United States</collectingCountry>
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west to
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<collectingRegion id="1B3E47ECFFE5477DFD88BEB5FD7EFB09" box="[630,715,1172,1194]" country="United States of America" name="Missouri" pageId="8" pageNumber="409">Missouri</collectingRegion>
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and
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<collectingRegion id="1B3E47ECFFE5477DFD05BEB4FC86FB09" box="[763,819,1173,1194]" country="United States of America" name="Texas" pageId="8" pageNumber="409">Texas</collectingRegion>
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...............
|
||
<taxonomicName id="1EFAF28DFFE5477DFC08BEB5FB3BFB09" ID-CoL="8KV4T" authority="Casey" authorityName="Casey" authorityYear="1918" box="[1014,1166,1172,1194]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EB8E551CFFE5477DFC08BEB5FBF8FB09" box="[1014,1101,1172,1194]" italics="true" pageId="8" pageNumber="409">Anillinus</emphasis>
|
||
Casey
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="D945890EFFE5477DFF85BE93FB3AFB45" blockId="8.[123,1174,1143,1254]" pageId="8" pageNumber="409">
|
||
<keyLead id="620B2A3BFFE5477DFF85BE93FB3AFB45" pageId="8" pageNumber="409">
|
||
– Male with apical margin of last visible abdominal ventrite toothed; right paramere apparently absent; geographic range southeastern
|
||
<collectingCountry id="A1EDC99EFFE5477DFE28BEF1FDE2FB45" box="[470,599,1232,1254]" name="United States of America" pageId="8" pageNumber="409">United States</collectingCountry>
|
||
.................................
|
||
<taxonomicName id="1EFAF28DFFE5477DFC0DBEF1FB3AFB45" ID-CoL="8KYVQ" authority="Barr" authorityName="Barr" authorityYear="1995" box="[1011,1167,1232,1254]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EB8E551CFFE5477DFC0DBEF1FBEBFB45" box="[1011,1118,1232,1254]" italics="true" pageId="8" pageNumber="409">Serranillus</emphasis>
|
||
Barr
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |