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<document id="CC0146AB8BDC4C77DF9BFB477EBEF7BD" ID-CLB-Dataset="8014" ID-DOI="10.11646/zootaxa.4877.3.4" ID-GBIF-Dataset="05b76ae0-dcec-434c-8897-bd71572268c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425545" ID-ZooBank="5DAFB47B-1A91-45C6-891C-A8D2A4F6BB74" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610031104548" checkinUser="plazi" docAuthor="Grogan, William L." docDate="2020" docId="514787962813FFBDFF1CFE20FA59FA2C" docLanguage="en" docName="zootaxa.4877.3.4.pdf" docOrigin="Zootaxa 4877 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bezzia (Bezzia) pulverea" docType="key" docVersion="8" lastPageNumber="433" masterDocId="AD7EFFEE2810FFB9FF8BFFDFFFC5FF9B" masterDocTitle="A Revision of the Nearctic Predaceous Midges in the Bezzia (Bezzia) pulverea complex (Diptera: Ceratopogonidae)" masterLastPageNumber="467" masterPageNumber="429" pageNumber="432" updateTime="1698923500066" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="66345A3EF8B413F7AC15B6290A6FFC24">A Revision of the Nearctic Predaceous Midges in the Bezzia (Bezzia) pulverea complex (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="11AE2E4B15DBF0FABE2563E5528DD961">Grogan, William L.</mods:namePart>
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<mods:title id="D5A8A3C900BF6D9E7D806EB12259172A">Zootaxa</mods:title>
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<mods:date id="AB95B2C6CE5F0F8C0551C0343B3550B1">2020</mods:date>
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<mods:identifier id="88C50D58875A5796889536A9DACEBA04" type="DOI">10.11646/zootaxa.4877.3.4</mods:identifier>
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<treatment id="514787962813FFBDFF1CFE20FA59FA2C" ID-DOI="http://doi.org/10.5281/zenodo.4562921" ID-GBIF-Taxon="179911450" ID-Zenodo-Dep="4562921" LSID="urn:lsid:plazi:treatment:514787962813FFBDFF1CFE20FA59FA2C" httpUri="http://treatment.plazi.org/id/514787962813FFBDFF1CFE20FA59FA2C" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432">
<subSubSection id="91F4650B2813FFBAFF1CFE20FCECFD82" box="[151,809,510,537]" pageId="3" pageNumber="432" type="nomenclature">
<paragraph id="D95136802813FFBAFF1CFE20FCECFD82" blockId="3.[151,809,510,574]" box="[151,809,510,537]" pageId="3" pageNumber="432">
<heading id="821981EC2813FFBAFF1CFE20FCECFD82" bold="true" box="[151,809,510,537]" fontSize="11" level="1" pageId="3" pageNumber="432" reason="1">
<emphasis id="EB9AEA922813FFBAFF1CFE20FCECFD82" bold="true" box="[151,809,510,537]" pageId="3" pageNumber="432">
Key to adults in the
<taxonomicName id="1EEE4D032813FFBAFE18FE20FD7FFD82" ID-CoL="8B6RF" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[403,698,510,537]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="pulverea" subGenus="Bezzia">
<emphasis id="EB9AEA922813FFBAFE18FE20FE24FD82" bold="true" box="[403,481,511,537]" italics="true" pageId="3" pageNumber="432">Bezzia</emphasis>
(
<emphasis id="EB9AEA922813FFBAFE79FE20FD85FD82" bold="true" box="[498,576,511,537]" italics="true" pageId="3" pageNumber="432">Bezzia</emphasis>
)
<emphasis id="EB9AEA922813FFBAFDD9FE21FD7FFD82" bold="true" box="[594,698,510,537]" italics="true" pageId="3" pageNumber="432">pulverea</emphasis>
</taxonomicName>
complex
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91F4650B2813FFBAFF1CFDFCFD5AFDA5" box="[151,671,546,574]" pageId="3" pageNumber="432" type="discussion">
<paragraph id="D95136802813FFBAFF1CFDFCFD5AFDA5" blockId="3.[151,809,510,574]" box="[151,671,546,574]" pageId="3" pageNumber="432">
(Females of
<emphasis id="EB9AEA922813FFBAFEA8FDFCFE38FDA6" bold="true" box="[291,509,546,573]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFEA8FDFCFE74FDA6" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[291,433,546,573]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="leptostyla" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFEA8FDFCFEF3FDA6" bold="true" box="[291,310,547,573]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFECFFDFDFE74FDA6" bold="true" box="[324,433,546,573]" italics="true" pageId="3" pageNumber="432">leptostyla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92813FFBAFE34FDFCFE38FDA6" box="[447,509,547,573]" pageId="3" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are unknown)
</paragraph>
</subSubSection>
<subSubSection id="91F4650B2813FFBDFF1CFDB7FA59FA2C" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432" type="key">
<keyStep id="621A2E252813FFBAFF1CFDB7FA59FD01" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFDB7FA59FDE6" blockId="3.[151,1437,616,2016]" box="[151,1436,616,638]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFDB7FA59FDE6" box="[151,1436,616,638]" pageId="3" pageNumber="432">
<specimenCount id="CFE8FD092813FFBAFF1CFDB7FED8FDE5" box="[151,285,616,638]" pageId="3" pageNumber="432" type="male">1 Males</specimenCount>
.............................................................................................. 2
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFD5AFA59FD01" blockId="3.[151,1437,616,2016]" box="[151,1436,645,667]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFD5AFA59FD01" box="[151,1436,645,667]" pageId="3" pageNumber="432">- Females............................................................................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CFD7DFA59FD69" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFD7DFA59FD2C" blockId="3.[151,1437,616,2016]" box="[151,1436,674,696]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFD7DFA59FD2C" box="[151,1436,674,696]" pageId="3" pageNumber="432">
2. Apex of gonostylus rounded or blunt, without apical inner or outer prongs (
<figureCitation id="41D52A052813FFBAFC3DFD7DFBC3FD2C" box="[950,1030,674,696]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="3" pageNumber="432">Figs. 33</figureCitation>
,
<figureCitation id="41D52A052813FFBAFB9BFD7DFBECFD23" box="[1040,1065,674,696]" captionStart="FIGURES 43–48" captionStartId="25.[151,264,1894,1918]" captionTargetBox="[161,1420,256,1859]" captionTargetId="figure-16@25.[151,1436,181,1871]" captionTargetPageId="25" captionText="FIGURES 43–48. Bezzia folkertsi n. sp. Male, 43–45, Female, 46–48. 43, 47 Thorax and legs. 44 Genitalia at focal level of parameres, gonocoxites and gonostyli. 45 Genitalia at focal level of aedeagus. 46 Head. 48 Apical hind tibial spines." figureDoi="http://doi.org/10.5281/zenodo.4425567" httpUri="https://zenodo.org/record/4425567/files/figure.png" pageId="3" pageNumber="432">44</figureCitation>
)............................. 3
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFD1FFA59FD69" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFD1FFA59FD69" pageId="3" pageNumber="432">
- Apex of gonostylus with apical inner and outer prongs (
<figureCitation id="41D52A052813FFBAFD7AFD1FFCF1FD4E" box="[753,820,704,726]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="3" pageNumber="432">Figs. 3</figureCitation>
,
<figureCitation id="41D52A052813FFBAFCB5FD1FFCB9FD4E" box="[830,892,704,725]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">10–14</figureCitation>
,
<figureCitation id="41D52A052813FFBAFC0EFD60FC06FD4E" box="[901,963,703,725]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="3" pageNumber="432">25–26</figureCitation>
,
<figureCitation id="41D52A052813FFBAFC47FD60FC23FD4E" box="[972,998,703,725]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="3" pageNumber="432">52</figureCitation>
,
<figureCitation id="41D52A052813FFBAFC64FD1FFBCCFD4D" box="[1007,1033,704,726]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">63</figureCitation>
,
<figureCitation id="41D52A052813FFBAFB99FD1FFB95FD4D" box="[1042,1104,704,726]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">69–70</figureCitation>
) or with only an apical outer prong (
<figureCitation id="41D52A052813FFBAFF62FD02FEFCFD69" box="[233,313,733,755]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">Figs. 20</figureCitation>
,
<figureCitation id="41D52A052813FFBAFEC8FD02FE98FD69" box="[323,349,733,754]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="3" pageNumber="432">38</figureCitation>
,
<figureCitation id="41D52A052813FFBAFEECFD02FE61FD68" box="[359,420,733,755]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">59–60</figureCitation>
)................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CFD25FA58FC39" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFD25FA59FCD1" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFD25FA59FCD1" pageId="3" pageNumber="432">
3. Gonocoxite and gonostylus short, apex of gonostylus blunt or slightly rounded (
<figureCitation id="41D52A052813FFBAFC50FD25FBE6FC8B" box="[987,1059,762,784]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="3" pageNumber="432">Fig. 33</figureCitation>
); aedeagus with moderately long apical section with sharply pointed tip (
<figureCitation id="41D52A052813FFBAFD97FCC7FDA6FCB5" box="[540,611,792,814]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="3" pageNumber="432">Fig. 33</figureCitation>
); hind tibia brown with narrow pale yellow to light brown basal and broader subapical bands (
<figureCitation id="41D52A052813FFBAFEACFCEAFEABFCD0" box="[295,366,821,843]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="3" pageNumber="432">Fig. 31</figureCitation>
); hind tarsomere 1 yellow (
<figureCitation id="41D52A052813FFBAFDFDFCEAFD78FCD0" box="[630,701,821,843]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="3" pageNumber="432">Fig. 31</figureCitation>
); mid, hind femur with 0–1 ventral spines (
<figureCitation id="41D52A052813FFBAFBD5FCEAFB66FCD0" box="[1118,1187,821,843]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="3" pageNumber="432">Fig. 31</figureCitation>
)...
<emphasis id="EB9AEA922813FFBAFB5EFCEAFAA5FCD1" bold="true" box="[1237,1376,820,842]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFB5EFCEAFA9FFCD1" ID-CoL="9HSM5" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1237,1370,820,842]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="amblystyla" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFB5EFCEAFB20FCD1" bold="true" box="[1237,1253,821,842]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB7AFCEBFA9FFCD1" bold="true" box="[1265,1370,820,842]" italics="true" pageId="3" pageNumber="432">amblystyla</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="F0A957E92813FFBAFAEDFCEAFA59FCD1" box="[1382,1436,821,842]" pageId="3" pageNumber="432" rank="species">
<emphasis id="EB9AEA922813FFBAFAEDFCEAFA52FCD1" bold="true" box="[1382,1431,821,842]" pageId="3" pageNumber="432">n. sp</emphasis>
.
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFC8CFA58FC39" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFC8CFA58FC39" pageId="3" pageNumber="432">
- Gonocoxite and gonostylus moderately elongate, apex of gonostylus rounded (
<figureCitation id="41D52A052813FFBAFC6EFC8DFBEAFCF3" box="[997,1071,850,872]" captionStart="FIGURES 43–48" captionStartId="25.[151,264,1894,1918]" captionTargetBox="[161,1420,256,1859]" captionTargetId="figure-16@25.[151,1436,181,1871]" captionTargetPageId="25" captionText="FIGURES 43–48. Bezzia folkertsi n. sp. Male, 43–45, Female, 46–48. 43, 47 Thorax and legs. 44 Genitalia at focal level of parameres, gonocoxites and gonostyli. 45 Genitalia at focal level of aedeagus. 46 Head. 48 Apical hind tibial spines." figureDoi="http://doi.org/10.5281/zenodo.4425567" httpUri="https://zenodo.org/record/4425567/files/figure.png" pageId="3" pageNumber="432">Fig. 44</figureCitation>
); aedeagus with very slender, greatly elongate apical section with bifid tip (
<figureCitation id="41D52A052813FFBAFDC5FCAFFD53FC1E" box="[590,662,879,902]" captionStart="FIGURES 43–48" captionStartId="25.[151,264,1894,1918]" captionTargetBox="[161,1420,256,1859]" captionTargetId="figure-16@25.[151,1436,181,1871]" captionTargetPageId="25" captionText="FIGURES 43–48. Bezzia folkertsi n. sp. Male, 43–45, Female, 46–48. 43, 47 Thorax and legs. 44 Genitalia at focal level of parameres, gonocoxites and gonostyli. 45 Genitalia at focal level of aedeagus. 46 Head. 48 Apical hind tibial spines." figureDoi="http://doi.org/10.5281/zenodo.4425567" httpUri="https://zenodo.org/record/4425567/files/figure.png" pageId="3" pageNumber="432">Fig. 45</figureCitation>
); hind tibia dark brown without pale yellow or light brown bands (
<figureCitation id="41D52A052813FFBAFA93FCAFFAA5FC1D" box="[1304,1376,880,902]" captionStart="FIGURES 43–48" captionStartId="25.[151,264,1894,1918]" captionTargetBox="[161,1420,256,1859]" captionTargetId="figure-16@25.[151,1436,181,1871]" captionTargetPageId="25" captionText="FIGURES 43–48. Bezzia folkertsi n. sp. Male, 43–45, Female, 46–48. 43, 47 Thorax and legs. 44 Genitalia at focal level of parameres, gonocoxites and gonostyli. 45 Genitalia at focal level of aedeagus. 46 Head. 48 Apical hind tibial spines." figureDoi="http://doi.org/10.5281/zenodo.4425567" httpUri="https://zenodo.org/record/4425567/files/figure.png" pageId="3" pageNumber="432">Fig. 43</figureCitation>
); hind tarsomere 1 dark brown (
<figureCitation id="41D52A052813FFBAFE5FFC52FDDEFC38" box="[468,539,909,931]" captionStart="FIGURES 43–48" captionStartId="25.[151,264,1894,1918]" captionTargetBox="[161,1420,256,1859]" captionTargetId="figure-16@25.[151,1436,181,1871]" captionTargetPageId="25" captionText="FIGURES 43–48. Bezzia folkertsi n. sp. Male, 43–45, Female, 46–48. 43, 47 Thorax and legs. 44 Genitalia at focal level of parameres, gonocoxites and gonostyli. 45 Genitalia at focal level of aedeagus. 46 Head. 48 Apical hind tibial spines." figureDoi="http://doi.org/10.5281/zenodo.4425567" httpUri="https://zenodo.org/record/4425567/files/figure.png" pageId="3" pageNumber="432">Fig. 43</figureCitation>
); mid, hind femora without ventral spines (
<figureCitation id="41D52A052813FFBAFC31FC52FBC5FC38" box="[954,1024,909,931]" captionStart="FIGURES 43–48" captionStartId="25.[151,264,1894,1918]" captionTargetBox="[161,1420,256,1859]" captionTargetId="figure-16@25.[151,1436,181,1871]" captionTargetPageId="25" captionText="FIGURES 43–48. Bezzia folkertsi n. sp. Male, 43–45, Female, 46–48. 43, 47 Thorax and legs. 44 Genitalia at focal level of parameres, gonocoxites and gonostyli. 45 Genitalia at focal level of aedeagus. 46 Head. 48 Apical hind tibial spines." figureDoi="http://doi.org/10.5281/zenodo.4425567" httpUri="https://zenodo.org/record/4425567/files/figure.png" pageId="3" pageNumber="432">Fig. 43</figureCitation>
)..................
<emphasis id="EB9AEA922813FFBAFB66FC52FA58FC39" bold="true" box="[1261,1437,908,930]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFB66FC52FA9FFC39" ID-CoL="9HWGK" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1261,1370,908,930]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="folkertsi" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFB66FC52FB38FC39" bold="true" box="[1261,1277,909,930]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFA82FC53FA9FFC39" bold="true" box="[1289,1370,908,930]" italics="true" pageId="3" pageNumber="432">folkertsi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92813FFBAFAEDFC52FA58FC39" box="[1382,1437,909,930]" pageId="3" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CFC75FA59FBAE" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFC75FA58FC61" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFC75FA58FC61" pageId="3" pageNumber="432">
4. Hind tibia brown with moderately broad (0.25 length of tibia) pale yellowish subapical band (
<figureCitation id="41D52A052813FFBAFBFAFC75FB7CFC5B" box="[1137,1209,938,960]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="3" pageNumber="432">Fig. 49</figureCitation>
); gonostylus very short without narrow neck-like base, apex broad with long slender pointed outer prong and shorter broader blade-like inner prong (
<figureCitation id="41D52A052813FFBAFF62FC3AFEEAFC60" box="[233,303,997,1019]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="3" pageNumber="432">Fig. 52</figureCitation>
)...............................................................................
<emphasis id="EB9AEA922813FFBAFB7DFC3AFA58FC61" bold="true" box="[1270,1437,996,1018]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFB7DFC3AFA9FFC61" ID-CoL="9HTJR" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1270,1370,996,1018]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="huberti" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFB7DFC3AFAC3FC61" bold="true" box="[1270,1286,997,1018]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFA99FC3BFA9FFC61" bold="true" box="[1298,1370,996,1018]" italics="true" pageId="3" pageNumber="432">huberti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92813FFBAFAEDFC3AFA58FC61" box="[1382,1437,997,1018]" pageId="3" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFBDCFA59FBAE" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFBDCFA59FBAE" pageId="3" pageNumber="432">- Hind tibia brown with or without narrow pale yellowish subapical band; gonostylus moderately long to elongate with narrow neck-like base, apex with inner and outer prongs or only outer prong............................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CFBE2FA59FBF4" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFBE2FA59FBC8" blockId="3.[151,1437,616,2016]" box="[151,1436,1085,1107]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFBE2FA59FBC8" box="[151,1436,1085,1107]" pageId="3" pageNumber="432">
5. Hind femur and tibia brown to dark brown without yellowish or light brown bands (
<figureCitation id="41D52A052813FFBAFB83FBE2FB8EFBC9" box="[1032,1099,1085,1107]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">Figs. 8</figureCitation>
,
<figureCitation id="41D52A052813FFBAFBDDFBE2FBB5FBC8" box="[1110,1136,1085,1107]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="3" pageNumber="432">16</figureCitation>
,
<figureCitation id="41D52A052813FFBAFBF1FBE2FB51FBC9" box="[1146,1172,1085,1106]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="3" pageNumber="432">28</figureCitation>
,
<figureCitation id="41D52A052813FFBAFB15FBE2FB7DFBC9" box="[1182,1208,1085,1106]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="3" pageNumber="432">37</figureCitation>
,
<figureCitation id="41D52A052813FFBAFB49FBE2FB19FBC8" box="[1218,1244,1085,1107]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">56</figureCitation>
,
<figureCitation id="41D52A052813FFBAFB6DFBE2FB3AFBC8" box="[1254,1279,1085,1107]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">62</figureCitation>
)........... 6
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFB84FA59FBF4" blockId="3.[151,1437,616,2016]" box="[151,1436,1114,1136]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFB84FA59FBF4" box="[151,1436,1114,1136]" pageId="3" pageNumber="432">
- Hind femur and tibia brown with yellowish or light brown bands on both or only one of these segments(
<figureCitation id="41D52A052813FFBAFB47FB85FAC8FBF4" box="[1228,1293,1114,1136]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="3" pageNumber="432">Figs. 4</figureCitation>
,
<figureCitation id="41D52A052813FFBAFA9CFB85FAF4FBF4" box="[1303,1329,1114,1135]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">21</figureCitation>
,
<figureCitation id="41D52A052813FFBAFAB0FB85FA91FBEB" box="[1339,1364,1114,1136]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">67</figureCitation>
).... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CFBA7FA59FA99" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFBA7FA58FB31" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFBA7FA58FB31" pageId="3" pageNumber="432">
6. Hind tarsomere 1 dark brown (
<figureCitation id="41D52A052813FFBAFD81FBA7FD94FB15" box="[522,593,1144,1166]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="3" pageNumber="432">Fig. 37</figureCitation>
); gonostylus short, very broad with slender sharply pointed outer prong (
<figureCitation id="41D52A052813FFBAFA85FBA7FA96FB15" box="[1294,1363,1144,1166]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="3" pageNumber="432">Fig. 38</figureCitation>
)......................................................................................
<emphasis id="EB9AEA922813FFBAFB31FB4AFA58FB31" bold="true" box="[1210,1437,1172,1194]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFB31FB4AFA9FFB31" ID-CoL="9HV5F" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1210,1370,1172,1194]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="brunneipedia" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFB31FB4AFB0FFB31" bold="true" box="[1210,1226,1173,1194]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB5DFB4BFA9FFB31" bold="true" box="[1238,1370,1172,1194]" italics="true" pageId="3" pageNumber="432">brunneipedia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92813FFBAFAEDFB4AFA58FB31" box="[1382,1437,1173,1194]" pageId="3" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFB6CFA59FA99" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFB6CFA59FA99" pageId="3" pageNumber="432">
- Hind tarsomere 1 yellowish to light brown (
<figureCitation id="41D52A052813FFBAFD02FB6DFD08FB5C" box="[649,717,1202,1224]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">Figs. 8</figureCitation>
,
<figureCitation id="41D52A052813FFBAFD53FB6DFD37FB53" box="[728,754,1202,1224]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">56</figureCitation>
,
<figureCitation id="41D52A052813FFBAFD76FB6DFCD2FB53" box="[765,791,1202,1224]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">62</figureCitation>
,
<figureCitation id="41D52A052813FFBAFCAAFB6DFCF9FB53" box="[801,828,1202,1224]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">67</figureCitation>
); gonostylus elongate with apical inner and outer prongs (
<figureCitation id="41D52A052813FFBAFAE7FB6DFF2AFB7E" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="3" pageNumber="432">Figs. 3</figureCitation>
,
<figureCitation id="41D52A052813FFBAFF70FB0FFEFCFB7E" box="[251,313,1232,1253]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">10–14</figureCitation>
,
<figureCitation id="41D52A052813FFBAFEC8FB10FE47FB7E" box="[323,386,1231,1253]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="3" pageNumber="432">25–26</figureCitation>
,
<figureCitation id="41D52A052813FFBAFE07FB0FFE63FB7D" box="[396,422,1232,1254]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">63</figureCitation>
), elongate with only a long outer prong (
<figureCitation id="41D52A052813FFBAFCBFFB0FFC6FFB7E" box="[820,938,1231,1254]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">Figs. 59–60</figureCitation>
), or short, broad with apical inner and outer prongs (
<figureCitation id="41D52A052813FFBAFF62FB32FE9BFA98" box="[233,350,1261,1283]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">Figs. 69–70</figureCitation>
), or with only an outer prong (
<figureCitation id="41D52A052813FFBAFD0FFB32FD0CFA98" box="[644,713,1261,1283]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">Fig. 20</figureCitation>
).......................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CFAD4FA59FAEC" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFAD4FA58FAA6" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFAD4FA58FAA6" pageId="3" pageNumber="432">
7. Inner prong of gonostylus longer than outer prong (
<figureCitation id="41D52A052813FFBAFD5EFAD5FCDBFABB" box="[725,798,1290,1312]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">Fig. 63</figureCitation>
); fore femur with 8–9 ventral spines (
<figureCitation id="41D52A052813FFBAFB06FAD5FB13FABB" box="[1165,1238,1290,1312]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">Fig. 62</figureCitation>
); antennal flagellum very long, length
<quantity id="1E169B652813FFBAFE00FAF7FDD3FAA6" box="[395,534,1320,1341]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" metricValueMax="1.83" metricValueMin="1.79" pageId="3" pageNumber="432" unit="mm" value="1.81" valueMax="1.83" valueMin="1.79">1.79–1.83 mm</quantity>
(
<figureCitation id="41D52A052813FFBAFDAFFAF7FDAFFAA5" box="[548,618,1320,1342]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="3" pageNumber="432">Fig. 61</figureCitation>
)...............................................
<emphasis id="EB9AEA922813FFBAFB25FAF7FA58FAA6" bold="true" box="[1198,1437,1319,1341]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFB25FAF7FA9FFAA6" ID-CoL="9HTYF" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1198,1370,1319,1341]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="marylandensis" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFB25FAF7FB7BFAA6" bold="true" box="[1198,1214,1320,1341]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB41FAF8FA9FFAA6" bold="true" box="[1226,1370,1319,1341]" italics="true" pageId="3" pageNumber="432">marylandensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92813FFBAFAEDFAF7FA58FAA6" box="[1382,1437,1320,1341]" pageId="3" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="D95136802813FFBAFF1CFA9AFA59FAEC" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFA9AFA59FAEC" pageId="3" pageNumber="432">
- Inner prong of gonostylus much shorter than outer prong (
<figureCitation id="41D52A052813FFBAFC9DFA9AFCA3FAC1" box="[790,870,1349,1371]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">Figs. 20</figureCitation>
,
<figureCitation id="41D52A052813FFBAFCFAFA9AFC6AFAC0" box="[881,943,1349,1371]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="3" pageNumber="432">25–26</figureCitation>
,
<figureCitation id="41D52A052813FFBAFC31FA9AFC11FAC0" box="[954,980,1349,1371]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">60</figureCitation>
,
<figureCitation id="41D52A052813FFBAFC55FA9AFBDAFAC0" box="[990,1055,1349,1371]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">69–70</figureCitation>
); fore femur with 2–7 (rarely 8) ventral spines; antennal flagellum shorter, length
<quantity id="1E169B652813FFBAFDF8FABDFD3BFAE3" box="[627,766,1378,1400]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.69" metricValueMin="1.06" pageId="3" pageNumber="432" unit="mm" value="1.375" valueMax="1.69" valueMin="1.06">1.06–1.69 mm</quantity>
(
<figureCitation id="41D52A052813FFBAFC87FABDFC83FAE3" box="[780,838,1378,1400]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">Fig. 7</figureCitation>
)................................................ 8
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<keyStep id="621A2E252813FFBAFF1CFA5FFA59F9DE" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CFA5FFA59FA54" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<materialsCitation id="69863CDD2813FFBAFF1CFA5FFA59FA54" ID-GBIF-Occurrence="3042855419" country="United States of America" location="Maryland" pageId="3" pageNumber="432" specimenCount="1" stateProvince="Maryland">
<keyLead id="621F95B52813FFBAFF1CFA5FFA59FA54" pageId="3" pageNumber="432">
8. Gonostylus usually slender distally with blunt, folded, flap-like inner prong and very short outer prong (
<figureCitation id="41D52A052813FFBAFB34FA5FFAF1FA0D" box="[1215,1332,1408,1430]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">Figs. 10–11</figureCitation>
); aedeagus very broad with relatively short apical section (
<figureCitation id="41D52A052813FFBAFD2EFA42FD2FFA28" box="[677,746,1437,1459]" captionStart="FIGURES 7–14" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[221,1385,191,1900]" captionTargetId="figure-16@9.[189,1398,181,1919]" captionTargetPageId="9" captionText="FIGURES 7–14. Bezzia laciniastyla. Male. 7 Head. 8 Thorax and legs. 9 wing. 10 Genitalia at focal level of sternite 9, parameres, gonocoxites and gonostyli. 11 Genitalia at focal level of aedeagus and gonostyli. 12–14 Variation in shapes of gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425551" httpUri="https://zenodo.org/record/4425551/files/figure.png" pageId="3" pageNumber="432">Fig. 11</figureCitation>
); southeastern
<collectingCountry id="A1F976102813FFBAFCF0FA42FC6FFA29" box="[891,938,1437,1458]" name="United States of America" pageId="3" pageNumber="432">USA</collectingCountry>
species,
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to
<collectingRegion id="1B2AF8622813FFBAFBF0FA42FB06FA28" box="[1147,1219,1437,1459]" country="United States of America" name="Florida" pageId="3" pageNumber="432">Florida</collectingRegion>
, west in
<collectingRegion id="1B2AF8622813FFBAFA9DFA42FA95FA29" box="[1302,1360,1437,1458]" country="United States of America" name="Texas" pageId="3" pageNumber="432">Texas</collectingRegion>
,
<collectingRegion id="1B2AF8622813FFBAFAD3FA42FF3AFA4B" country="United States of America" name="Arkansas" pageId="3" pageNumber="432">Arkansas</collectingRegion>
and
<collectingRegion id="1B2AF8622813FFBAFEA5FA65FE48FA4B" box="[302,397,1466,1488]" country="United States of America" name="Louisiana" pageId="3" pageNumber="432">Louisiana</collectingRegion>
.............................................................
<taxonomicName id="1EEE4D032813FFBAFBF9FA64FA59FA54" ID-CoL="8B6QM" authority="Dow &amp; Turner" authorityName="Dow &amp; Turner" authorityYear="1976" box="[1138,1436,1466,1488]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="laciniastyla">
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.
<emphasis id="EB9AEA922813FFBAFB05FA65FA59FA54" bold="true" box="[1166,1436,1466,1488]" pageId="3" pageNumber="432">
<emphasis id="EB9AEA922813FFBAFB05FA65FB3AFA4B" bold="true" box="[1166,1279,1466,1488]" italics="true" pageId="3" pageNumber="432">laciniastyla</emphasis>
Dow &amp; Turner
</emphasis>
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</materialsCitation>
</paragraph>
<paragraph id="D95136802813FFBAFF1CFA07FA59F9DE" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CFA07FA59F9DE" pageId="3" pageNumber="432">
- Gonostylus broad distally with elongate, curved outer prong with pointed tip and short rounded inner prong (
<figureCitation id="41D52A052813FFBAFA92FA07FA57FA76" box="[1305,1426,1495,1518]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="3" pageNumber="432">Figs. 25–26</figureCitation>
); aedeagus broad with elongate, triangular relatively narrow apical section (
<figureCitation id="41D52A052813FFBAFC21FA2AFBD2F990" box="[938,1047,1525,1547]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="3" pageNumber="432">Fig. 25–26</figureCitation>
); wide ranging Nearctic species,
<collectingRegion id="1B2AF8622813FFBAFADFFA2AFE4FF9B3" isEnumeration="true" pageId="3" pageNumber="432">Alaska, British Columbia</collectingRegion>
to
<collectingRegion id="1B2AF8622813FFBAFE20F9CCFD8FF9B3" box="[427,586,1554,1576]" country="Canada" name="New Brunswick" pageId="3" pageNumber="432">New Brunswick</collectingRegion>
, south to
<collectingRegion id="1B2AF8622813FFBAFD27F9CDFC5EF9B3" box="[684,923,1554,1576]" isEnumeration="true" pageId="3" pageNumber="432">California, New Mexico</collectingRegion>
, east in
<collectingRegion id="1B2AF8622813FFBAFC66F9CDFE51F9DD" isEnumeration="true" pageId="3" pageNumber="432">Utah, Wyoming, Wisconsin, Nebraska, New York, Connecticut</collectingRegion>
, south to
<collectingRegion id="1B2AF8622813FFBAFE64F9EFFD0CF9DD" box="[495,713,1584,1606]" isEnumeration="true" pageId="3" pageNumber="432">Maryland and Virginia</collectingRegion>
.....................................
<taxonomicName id="1EEE4D032813FFBAFB01F9EFFA59F9DE" ID-CoL="8B6S4" authority="Dow &amp; Turner" authorityName="Dow &amp; Turner" authorityYear="1976" box="[1162,1436,1583,1605]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="uncistyla">
<emphasis id="EB9AEA922813FFBAFB01F9EFFB5FF9DE" bold="true" box="[1162,1178,1584,1605]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB2DF9F0FA59F9DE" bold="true" box="[1190,1436,1583,1605]" pageId="3" pageNumber="432">
<emphasis id="EB9AEA922813FFBAFB2DF9F0FB3AF9DE" bold="true" box="[1190,1279,1583,1605]" italics="true" pageId="3" pageNumber="432">uncistyla</emphasis>
Dow &amp; Turner
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CF992FA59F9E4" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CF992FA59F9F9" blockId="3.[151,1437,616,2016]" box="[151,1436,1613,1635]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CF992FA59F9F9" box="[151,1436,1613,1635]" pageId="3" pageNumber="432">
9. Gonostylus with well-developed inner and outer prongs (
<figureCitation id="41D52A052813FFBAFC8DF992FC8CF9F9" box="[774,841,1613,1635]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="3" pageNumber="432">Figs. 3</figureCitation>
,
<figureCitation id="41D52A052813FFBAFCDFF992FC57F9F8" box="[852,914,1613,1635]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">69–70</figureCitation>
)........................................ 10
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CF9B4FA59F9E4" blockId="3.[151,1437,616,2016]" box="[151,1436,1642,1664]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CF9B4FA59F9E4" box="[151,1436,1642,1664]" pageId="3" pageNumber="432">
- Gonostylus with elongate outer prong, and rounded or without an inner prong (
<figureCitation id="41D52A052813FFBAFC53F9B5FBEDF9E4" box="[984,1064,1642,1664]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">Figs. 20</figureCitation>
,
<figureCitation id="41D52A052813FFBAFBB9F9B5FBAAF91B" box="[1074,1135,1642,1664]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">59–60</figureCitation>
)...................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBAFF1CF957FA58F8B4" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CF957FA59F920" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CF957FA59F920" pageId="3" pageNumber="432">
10. Gonostylus moderately long, slender, apex with inner prong longer than outer prong (
<figureCitation id="41D52A052813FFBAFB95F957FB9DF905" box="[1054,1112,1672,1694]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="3" pageNumber="432">Fig. 3</figureCitation>
); fore femur yellow without dark bands, with 4–9 ventral spines (
<figureCitation id="41D52A052813FFBAFD99F97AFD88F920" box="[530,589,1701,1723]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="3" pageNumber="432">Fig. 4</figureCitation>
); large species, wing length
<quantity id="1E169B652813FFBAFCD4F97AFC2FF921" box="[863,1002,1701,1723]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.35" metricValueMin="1.95" pageId="3" pageNumber="432" unit="mm" value="2.15" valueMax="2.35" valueMin="1.95">1.95–2.35 mm</quantity>
...............
<taxonomicName id="1EEE4D032813FFBAFB25F97AFA59F920" ID-CoL="8B6RF" authority="(Coquillett)" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[1198,1436,1700,1723]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="pulverea">
<emphasis id="EB9AEA922813FFBAFB25F97AFA59F920" bold="true" box="[1198,1436,1700,1723]" pageId="3" pageNumber="432">
<emphasis id="EB9AEA922813FFBAFB25F97AFB7BF921" bold="true" box="[1198,1214,1701,1722]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB41F97BFADAF921" bold="true" box="[1226,1311,1700,1722]" italics="true" pageId="3" pageNumber="432">pulverea</emphasis>
(Coquillett)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBAFF1CF91CFA58F8B4" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CF91CFA58F8B4" pageId="3" pageNumber="432">
- Gonostylus very short with very broad claw-like apex, outer prong very long, greatly curved, inner prong shorter, broad, blade-like (
<figureCitation id="41D52A052813FFBAFEC5F93FFE01F96D" box="[334,452,1760,1782]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">Figs. 69–70</figureCitation>
); fore femur yellowish or light brown with brown basal band, with or without poorly developed narrow subapical brown band and 2–4 ventral spines (
<figureCitation id="41D52A052813FFBAFD14F922FD22F888" box="[671,743,1789,1811]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="3" pageNumber="432">Fig. 67</figureCitation>
); medium-size species, wing length
<quantity id="1E169B652813FFBAFBCFF922FB0AF889" box="[1092,1231,1789,1811]" metricMagnitude="-3" metricUnit="m" metricValue="1.715" metricValueMax="1.81" metricValueMin="1.62" pageId="3" pageNumber="432" unit="mm" value="1.715" valueMax="1.81" valueMin="1.62">1.62–1.81 mm</quantity>
....................................................................................................
<emphasis id="EB9AEA922813FFBAFB5BF8C4FA58F8B4" bold="true" box="[1232,1437,1818,1840]" pageId="3" pageNumber="432">
<taxonomicName id="1EEE4D032813FFBAFB5BF8C4FA9EF8AB" ID-CoL="9HWFM" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1232,1371,1818,1840]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="titanochela" status="sp. nov.">
<emphasis id="EB9AEA922813FFBAFB5BF8C4FB25F8AB" bold="true" box="[1232,1248,1819,1840]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB67F8C5FA9EF8AB" bold="true" box="[1260,1371,1818,1840]" italics="true" pageId="3" pageNumber="432">titanochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92813FFBAFAEDF8C5FA58F8B4" box="[1382,1437,1818,1839]" pageId="3" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252813FFBDFF1CF8E7FA58FF51" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432">
<paragraph id="D95136802813FFBAFF1CF8E7FA59F81C" blockId="3.[151,1437,616,2016]" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBAFF1CF8E7FA59F81C" pageId="3" pageNumber="432">
11. Fore femur with 4 ventral spines; sternite 9 long with a narrow deep apico-medial excavation that lacks a fringe of setae; gonostylus very short, broad with slender, curved outer prong (
<figureCitation id="41D52A052813FFBAFC80F88AFC97F8F0" box="[779,850,1877,1899]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">Fig. 20</figureCitation>
); fore, mid tibiae yellowish brown with darker brown narrow dorsal stripes that extend 4/5 of their total lengths (
<figureCitation id="41D52A052813FFBAFD40F8ADFCD4F813" box="[715,785,1906,1928]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="3" pageNumber="432">Fig. 21</figureCitation>
)................................
<taxonomicName id="1EEE4D032813FFBAFB15F8ACFA59F81C" ID-CoL="8B6RW" authority="Dow &amp; Turner" authorityName="Dow &amp; Turner" authorityYear="1976" box="[1182,1436,1906,1928]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="spicata">
<emphasis id="EB9AEA922813FFBAFB15F8ACFA59F81C" bold="true" box="[1182,1436,1906,1928]" pageId="3" pageNumber="432">
<emphasis id="EB9AEA922813FFBAFB15F8ACFB6BF813" bold="true" box="[1182,1198,1907,1928]" italics="true" pageId="3" pageNumber="432">B</emphasis>
.
<emphasis id="EB9AEA922813FFBAFB31F8ADFB3AF81C" bold="true" box="[1210,1279,1906,1927]" italics="true" pageId="3" pageNumber="432">spicata</emphasis>
Dow &amp; Turner
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D95136802813FFBDFF1CF84FFA58FF51" blockId="3.[151,1437,616,2016]" lastBlockId="4.[151,1437,151,1464]" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432">
<keyLead id="621F95B52813FFBDFF1CF84FFA58FF51" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432">
- Fore femur with 6–8 ventral spines (
<figureCitation id="41D52A052813FFBAFDC9F84FFD4FF83E" box="[578,650,1935,1958]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">Fig. 56</figureCitation>
); sternite 9 moderately long with narrow shallow U-shaped apico-medial excavation that lacks a fringe of setae (
<figureCitation id="41D52A052813FFBAFDABF872FD5DF858" box="[544,664,1965,1987]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">Figs. 59–60</figureCitation>
); gonostylus elongate with very long slightly curved outer prong (
<figureCitation id="41D52A052813FFBAFAA9F872FAAEF858" box="[1314,1387,1965,1987]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="3" pageNumber="432">Fig. 60</figureCitation>
), but lacks an inner prong; fore tibia yellowish to light brown with broad central and narrower apical brown bands, mid tibia brown or medium brown with pale narrow subapical band or with poorly defined basal band and broader well developed pale subapical band (
<figureCitation id="41D52A052814FFBDFE95FF6AFEA1FF50" box="[286,356,181,203]" captionStart="FIGURES 56–60" captionStartId="29.[151,264,1862,1886]" captionTargetBox="[166,1403,190,1826]" captionTargetId="figure-16@29.[151,1436,181,1839]" captionTargetPageId="29" captionText="FIGURES 56–60. Bezzia leptostyla n. sp. Male. 56 Thorax and legs. 57 Hind tibial spines. 58 Wing. 59 Genitalia at focal level of parameres, gonocoxites and gonostyli. 60 Genitalia at focal level of parameres, aedeagus, right gonocoxite and gonostylus." figureDoi="http://doi.org/10.5281/zenodo.4425571" httpUri="https://zenodo.org/record/4425571/files/figure.png" pageId="4" pageNumber="433">Fig. 56</figureCitation>
).........................................................................
<emphasis id="EB9AEA922814FFBDFB6AFF6AFA58FF51" bold="true" box="[1249,1437,180,202]" pageId="4" pageNumber="433">
<taxonomicName id="1EEE4D032814FFBDFB6AFF6AFA9FFF51" ID-CoL="9HRXW" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1249,1370,180,202]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="leptostyla" status="sp. nov.">
<emphasis id="EB9AEA922814FFBDFB6AFF6AFB34FF51" bold="true" box="[1249,1265,181,202]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB76FF6BFA9FFF51" bold="true" box="[1277,1370,180,202]" italics="true" pageId="4" pageNumber="433">leptostyla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92814FFBDFAEDFF6AFA58FF51" box="[1382,1437,181,202]" pageId="4" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252814FFBDFF1CFF0DFA59FEE1" pageId="4" pageNumber="433">
<paragraph id="D95136802814FFBDFF1CFF0DFA58FEB9" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFF0DFA58FEB9" pageId="4" pageNumber="433">
12. Femora, tibiae brown, hind leg darkest with only a poorly developed narrow light brown apical band on forefemur, mid-, hind femur and all tibiae without pale bands (
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); mid-, hind femur without ventral spines; spermathecae subequal-size, very small (larger
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, smaller
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long)................................................
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Femora, tibiae yellowish to dark brown with yellowish or light brown bands on some or all legs (
<figureCitation id="41D52A052814FFBDFBF2FEF5FB79FEA4" box="[1145,1212,298,320]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 4</figureCitation>
,
<figureCitation id="41D52A052814FFBDFB4EFEF5FB1AFEDB" box="[1221,1247,298,320]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">16</figureCitation>
,
<figureCitation id="41D52A052814FFBDFB62FEF5FAC6FEA4" box="[1257,1283,298,319]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">21</figureCitation>
,
<figureCitation id="41D52A052814FFBDFA87FEF5FAE3FEA4" box="[1292,1318,298,319]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">34</figureCitation>
,
<figureCitation id="41D52A052814FFBDFAA4FEF5FA8CFEA4" box="[1327,1353,298,319]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="4" pageNumber="433">40</figureCitation>
,
<figureCitation id="41D52A052814FFBDFAD9FEF5FAA9FEDB" box="[1362,1388,298,320]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="4" pageNumber="433">54</figureCitation>
,
<figureCitation id="41D52A052814FFBDFAFEFEF5FA55FEDB" box="[1397,1424,298,320]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">64</figureCitation>
); mid-, hind femur usually with 1 or more ventral spines; spermathecae unequal-size, medium-size to very large (larger
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, smaller
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long) (
<figureCitation id="41D52A052814FFBDFDBAFEBAFDB0FEE0" box="[561,629,357,379]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 6</figureCitation>
,
<figureCitation id="41D52A052814FFBDFDF4FEBAFD5CFEE1" box="[639,665,357,378]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">18</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD28FEBAFD78FEE1" box="[675,701,357,378]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">23</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD4CFEBAFD24FEE1" box="[711,737,357,378]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">30</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD60FEBAFCC0FEE0" box="[747,773,357,379]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">36</figureCitation>
,
<figureCitation id="41D52A052814FFBDFC84FEBAFCECFEE1" box="[783,809,357,378]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="4" pageNumber="433">42</figureCitation>
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<figureCitation id="41D52A052814FFBDFCB8FEBAFC88FEE0" box="[819,845,357,379]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="4" pageNumber="433">55</figureCitation>
,
<figureCitation id="41D52A052814FFBDFCDCFEBAFCB4FEE0" box="[855,881,357,379]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">66</figureCitation>
,
<figureCitation id="41D52A052814FFBDFCF0FEBAFC51FEE0" box="[891,916,357,379]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="4" pageNumber="433">71</figureCitation>
)........................................ 13
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13. Hind tibia brown with broad to very broad (0.3–0.7 length of tibia) pale yellow subapical or apical band, without large stout apical spines (
<figureCitation id="41D52A052814FFBDFEE4FE40FE7CFE2E" box="[367,441,415,437]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="4" pageNumber="433">Fig. 54</figureCitation>
); spermathecae unequal-size with greatly tapered neck regions, the larger very slender (
<figureCitation id="41D52A052814FFBDFA95FE40FAADFE2E" box="[1310,1384,415,437]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="4" pageNumber="433">Fig. 55</figureCitation>
); AR 1.42–1.56..............................................................................
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Hind tibia brown with (
<figureCitation id="41D52A052814FFBDFE4BFE05FDC1FE74" box="[448,516,474,496]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 4</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD86FE05FDE2FE74" box="[525,551,474,495]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">21</figureCitation>
,
<figureCitation id="41D52A052814FFBDFDBBFE05FD8CFE6B" box="[560,585,474,496]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">34</figureCitation>
) or without narrow, pale yellow subapical band (
<figureCitation id="41D52A052814FFBDFB92FE05FBADFE6B" box="[1049,1128,474,496]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">Figs. 16</figureCitation>
,
<figureCitation id="41D52A052814FFBDFBF9FE05FB49FE74" box="[1138,1164,474,495]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">28</figureCitation>
,
<figureCitation id="41D52A052814FFBDFB1EFE05FB6AFE74" box="[1173,1199,474,495]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="4" pageNumber="433">40</figureCitation>
,
<figureCitation id="41D52A052814FFBDFB33FE05FB16FE6B" box="[1208,1235,474,496]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">64</figureCitation>
), apex usually with 1 or more large apical spines (
<figureCitation id="41D52A052814FFBDFE7BFE28FDF7FD96" box="[496,562,503,525]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 1</figureCitation>
,
<figureCitation id="41D52A052814FFBDFDB5FE27FD9DFD96" box="[574,600,504,525]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">22</figureCitation>
,
<figureCitation id="41D52A052814FFBDFDE9FE28FDB8FD96" box="[610,637,503,525]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">65</figureCitation>
); spermathecae with or without tapered neck regions, the larger stout or moderately slender; AR 1.00–1.65................................................................................ 14
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14. Hind femur and tibia brown to dark brown without yellowish or light brown bands (
<figureCitation id="41D52A052814FFBDFB83FDEDFB9DFDD3" box="[1032,1112,562,584]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">Figs. 16</figureCitation>
,
<figureCitation id="41D52A052814FFBDFBE9FDEDFBB9FDDC" box="[1122,1148,562,583]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">28</figureCitation>
,
<figureCitation id="41D52A052814FFBDFB0DFDEDFB5AFDD3" box="[1158,1183,562,584]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="4" pageNumber="433">40</figureCitation>
).................. 15
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<paragraph id="D95136802814FFBDFF1CFD8FFA52FDFD" blockId="4.[151,1437,151,1464]" box="[151,1431,591,614]" pageId="4" pageNumber="433">
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- Hind femur and tibia brown with yellowish or light brown bands on both or only on one of these segments (
<figureCitation id="41D52A052814FFBDFB66FD90FAF4FDFE" box="[1261,1329,591,613]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 4</figureCitation>
,
<figureCitation id="41D52A052814FFBDFAB0FD8FFA90FDFE" box="[1339,1365,592,613]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">21</figureCitation>
,
<figureCitation id="41D52A052814FFBDFAD4FD8FFABCFDFD" box="[1375,1401,592,614]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">34</figureCitation>
)18
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<keyStep id="621A2E252814FFBDFF1CFDB2FA59FD40" pageId="4" pageNumber="433">
<paragraph id="D95136802814FFBDFF1CFDB2FA58FD04" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
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15. Hind tarsomere 1 dark brown (
<figureCitation id="41D52A052814FFBDFD81FDB2FD94FD18" box="[522,593,621,643]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="4" pageNumber="433">Fig. 40</figureCitation>
); spermathecae with slightly tapered neck regions (
<figureCitation id="41D52A052814FFBDFBB5FDB2FB41FD18" box="[1086,1156,621,643]" captionStart="FIGURES 37–42" captionStartId="22.[151,264,1961,1985]" captionTargetBox="[179,1416,192,1927]" captionTargetId="figure-21@22.[161,1426,181,1938]" captionTargetPageId="22" captionText="FIGURES 37–42. Bezzia brunneipedia, n. sp. Male, 37–38, Female, 39–42. 37, 40 Thorax and legs. 38 Genitalia at focal level of parameres, aedeagus, gonostyli and gonocoxites. 39 Head. 41 Wing. 42 Abdominal segments 8–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425565" httpUri="https://zenodo.org/record/4425565/files/figure.png" pageId="4" pageNumber="433">Fig. 42</figureCitation>
).......................................................................................................
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<taxonomicName id="1EEE4D032814FFBDFB31FD55FA9FFD3B" ID-CoL="9HV5F" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1210,1370,650,672]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="brunneipedia" status="sp. nov.">
<emphasis id="EB9AEA922814FFBDFB31FD55FB0FFD04" bold="true" box="[1210,1226,650,671]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB5DFD55FA9FFD3B" bold="true" box="[1238,1370,650,672]" italics="true" pageId="4" pageNumber="433">brunneipedia</emphasis>
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<paragraph id="D95136802814FFBDFF1CFD77FA59FD40" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
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- Hind tarsomere 1 yellowish to light brown (
<figureCitation id="41D52A052814FFBDFD03FD78FD09FD26" box="[648,716,679,701]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 4</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD5DFD78FD35FD26" box="[726,752,679,701]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">16</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD71FD77FCD1FD26" box="[762,788,680,701]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">21</figureCitation>
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<figureCitation id="41D52A052814FFBDFC95FD77FCFDFD26" box="[798,824,680,701]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">28</figureCitation>
,
<figureCitation id="41D52A052814FFBDFCC9FD77FC99FD26" box="[834,860,680,701]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">34</figureCitation>
,
<figureCitation id="41D52A052814FFBDFCEDFD78FC44FD26" box="[870,897,679,701]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">64</figureCitation>
); spermathecae with broad neck regions, non-tapered or slightly to moderately tapered (
<figureCitation id="41D52A052814FFBDFD85FD1AFD97FD40" box="[526,594,709,731]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 6</figureCitation>
,
<figureCitation id="41D52A052814FFBDFDD7FD1AFDB3FD41" box="[604,630,709,730]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">18</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD0BFD1AFD5FFD41" box="[640,666,709,730]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">23</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD2FFD1AFD7BFD41" box="[676,702,709,730]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">30</figureCitation>
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<figureCitation id="41D52A052814FFBDFD43FD1AFD27FD40" box="[712,738,709,731]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">36</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD67FD1AFCC3FD40" box="[748,774,709,731]" captionStart="FIGURES 49–55" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[184,1412,207,1911]" captionTargetId="figure-16@27.[170,1418,181,1919]" captionTargetPageId="27" captionText="FIGURES 49–55. Bezzia huberti n. sp. Male, 49–52, Female, 53–55. 49 Head, thorax and legs. 50 Wing. 51 Hind tibial spines. 52 Genitalia at focal level of sternite 9, parameres, aedeagus and gonostyli. 53 Head. 54 Thorax and legs. 55 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425569" httpUri="https://zenodo.org/record/4425569/files/figure.png" pageId="4" pageNumber="433">55</figureCitation>
,
<figureCitation id="41D52A052814FFBDFC9BFD1AFCEFFD40" box="[784,810,709,731]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">66</figureCitation>
,
<figureCitation id="41D52A052814FFBDFCBFFD1AFC88FD40" box="[820,845,709,731]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="4" pageNumber="433">71</figureCitation>
).............................................. 16
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<keyStep id="621A2E252814FFBDFF1CFD3DFA59FCD4" pageId="4" pageNumber="433">
<paragraph id="D95136802814FFBDFF1CFD3DFA58FC8E" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
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16. Antennal flagellum moderately long to very long (length
<quantity id="1E169B652814FFBDFC98FD3DFC66FD63" box="[787,931,738,760]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" metricValueMax="1.52" metricValueMin="1.22" pageId="4" pageNumber="433" unit="mm" value="1.37" valueMax="1.52" valueMin="1.22">1.22–1.52 mm</quantity>
); AR 1.51–1.65; fore femur with 5–8 ventral spines (
<figureCitation id="41D52A052814FFBDFF62FD20FEF4FC8E" box="[233,305,767,789]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">Fig. 64</figureCitation>
); spermathecae with broad, non-tapered necks (
<figureCitation id="41D52A052814FFBDFD70FD20FC84FC8E" box="[763,833,767,789]" captionStart="FIGURES 61–66" captionStartId="32.[151,264,1849,1873]" captionTargetBox="[164,1425,192,1803]" captionTargetId="figure-21@32.[151,1436,181,1825]" captionTargetPageId="32" captionText="FIGURES 61–66. Bezzia marylandensis n. sp. Male, 61–63, Female, 64–66. 61 Head. 62 Thorax and legs. 63 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 64 Thorax and legs. 65 Mid tibial apical spines. 66 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425573" httpUri="https://zenodo.org/record/4425573/files/figure.png" pageId="4" pageNumber="433">Fig. 66</figureCitation>
).............................
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<taxonomicName id="1EEE4D032814FFBDFB25FCDFFA9FFC8E" ID-CoL="9HTYF" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1198,1370,767,789]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="marylandensis" status="sp. nov.">
<emphasis id="EB9AEA922814FFBDFB25FCDFFB7BFC8E" bold="true" box="[1198,1214,768,789]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB41FD20FA9FFC8E" bold="true" box="[1226,1370,767,789]" italics="true" pageId="4" pageNumber="433">marylandensis</emphasis>
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,
<taxonomicNameLabel id="F0A957E92814FFBDFAEDFCDFFA58FC8E" box="[1382,1437,768,789]" pageId="4" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="D95136802814FFBDFF1CFCC2FA59FCD4" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFCC2FA59FCD4" pageId="4" pageNumber="433">
- Antennal flagellum very short to moderately long (length
<quantity id="1E169B652814FFBDFC9DFCC2FC60FCA9" box="[790,933,797,819]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" metricValueMax="1.28" metricValueMin="0.86" pageId="4" pageNumber="433" unit="mm" value="1.07" valueMax="1.28" valueMin="0.86">0.86–1.28 mm</quantity>
); AR 1.21–1.72; fore femur with 2–8 ventral spines (
<figureCitation id="41D52A052814FFBDFF62FCE5FEFCFCCB" box="[233,313,826,848]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">Figs. 16</figureCitation>
,
<figureCitation id="41D52A052814FFBDFEC8FCE5FE98FCD4" box="[323,349,826,847]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">21</figureCitation>
,
<figureCitation id="41D52A052814FFBDFEECFCE5FE47FCCB" box="[359,386,826,848]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">28</figureCitation>
); spermathecae with slightly or moderately tapered necks (
<figureCitation id="41D52A052814FFBDFC33FCE5FBCDFCD4" box="[952,1032,826,848]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">Figs. 23</figureCitation>
,
<figureCitation id="41D52A052814FFBDFB99FCE5FBE9FCCB" box="[1042,1068,826,848]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">36</figureCitation>
,
<figureCitation id="41D52A052814FFBDFBBDFCE5FB8AFCCB" box="[1078,1103,826,848]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="4" pageNumber="433">71</figureCitation>
)......................... 17
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<keyStep id="621A2E252814FFBDFF1CFC87FA59FC79" pageId="4" pageNumber="433">
<paragraph id="D95136802814FFBDFF1CFC87FA59FC11" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<materialsCitation id="69863CDD2814FFBDFF1CFC87FA59FC11" ID-GBIF-Occurrence="3042855405" country="United States of America" location="Maryland" pageId="4" pageNumber="433" specimenCount="1" stateProvince="Maryland">
<keyLead id="621F95B52814FFBDFF1CFC87FA59FC11" pageId="4" pageNumber="433">
17. Medium-size to large species (wing length
<quantity id="1E169B652814FFBDFD09FC88FCD4FCF6" box="[642,785,855,877]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.23" metricValueMin="1.57" pageId="4" pageNumber="433" unit="mm" value="1.9" valueMax="2.23" valueMin="1.57">1.57–2.23 mm</quantity>
;
<figureCitation id="41D52A052814FFBDFC90FC88FCA4FCF5" box="[795,865,855,878]" captionStart="FIGURES 15–19" captionStartId="10.[151,264,1912,1936]" captionTargetBox="[166,1424,198,1873]" captionTargetId="figure-21@10.[151,1436,181,1889]" captionTargetPageId="10" captionText="FIGURES 15–19. Bezzia laciniastyla. Female. 15 head. 16 thorax and legs. 17 wing. 18 Abdominal segments 7–9 and spermathecae. 19 Abdominal segments 8–10 and smaller spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4425555" httpUri="https://zenodo.org/record/4425555/files/figure.png" pageId="4" pageNumber="433">Fig. 17</figureCitation>
); AR 1.25–1.72; southeastern
<collectingCountry id="A1F976102814FFBDFB0DFC87FB70FCF6" box="[1158,1205,856,877]" name="United States of America" pageId="4" pageNumber="433">USA</collectingCountry>
species,
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,
<collectingRegion id="1B2AF8622814FFBDFAFDFC88FED7FC10" country="United States of America" name="Virginia" pageId="4" pageNumber="433">Virginia</collectingRegion>
south to
<collectingRegion id="1B2AF8622814FFBDFEE0FCAAFE76FC10" box="[363,435,885,907]" country="United States of America" name="Florida" pageId="4" pageNumber="433">Florida</collectingRegion>
, west in
<collectingRegion id="1B2AF8622814FFBDFD8DFCAAFD85FC11" box="[518,576,885,906]" country="United States of America" name="Texas" pageId="4" pageNumber="433">Texas</collectingRegion>
,
<collectingRegion id="1B2AF8622814FFBDFDC3FCAAFD64FC10" box="[584,673,885,907]" country="United States of America" name="Arkansas" pageId="4" pageNumber="433">Arkansas</collectingRegion>
and
<collectingRegion id="1B2AF8622814FFBDFD5BFCAAFCEAFC10" box="[720,815,885,907]" country="United States of America" name="Louisiana" pageId="4" pageNumber="433">Louisiana</collectingRegion>
..........................
<taxonomicName id="1EEE4D032814FFBDFBF9FCAAFA59FC11" ID-CoL="8B6QM" authority="Dow &amp; Turner" authorityName="Dow &amp; Turner" authorityYear="1976" box="[1138,1436,884,907]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="laciniastyla">
<emphasis id="EB9AEA922814FFBDFBF9FCAAFA59FC11" bold="true" box="[1138,1436,884,907]" pageId="4" pageNumber="433">
<emphasis id="EB9AEA922814FFBDFBF9FCAAFB47FC11" bold="true" box="[1138,1154,885,906]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB05FCABFB3AFC11" bold="true" box="[1166,1279,884,906]" italics="true" pageId="4" pageNumber="433">laciniastyla</emphasis>
Dow &amp; Turner
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</paragraph>
<paragraph id="D95136802814FFBDFF1CFC4CFA59FC79" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFC4CFA59FC79" pageId="4" pageNumber="433">
- Large to very large species (wing length
<quantity id="1E169B652814FFBDFDEEFC4DFD37FC33" box="[613,754,914,936]" metricMagnitude="-3" metricUnit="m" metricValue="2.415" metricValueMax="2.86" metricValueMin="1.97" pageId="4" pageNumber="433" unit="mm" value="2.415" valueMax="2.86" valueMin="1.97">1.97–2.86 mm</quantity>
;
<figureCitation id="41D52A052814FFBDFD77FC4DFC87FC33" box="[764,834,914,936]" captionStart="FIGURES 25–30" captionStartId="17.[151,264,1944,1968]" captionTargetBox="[196,1401,192,1909]" captionTargetId="figure-16@17.[170,1417,181,1920]" captionTargetPageId="17" captionText="FIGURES 25–30. Bezzia uncistyla. Male, 25–26, Female, 27–30 (from type locality, Taos, New Mexico). 25 Genitalia at focal level of sternite 9, parameres, and gonostyli (from New York); 26 Genitalia at focal level of sternite 9, aedeagus and gonostyli (from Utah). 27 Head. 28 Thorax and legs. 29 Wing. 30 Abdominal segments 7–9 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425559" httpUri="https://zenodo.org/record/4425559/files/figure.png" pageId="4" pageNumber="433">Fig. 29</figureCitation>
); AR 1.21–1.57; wide ranging Nearctic species,
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to
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, south to
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, east to
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, south to
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...........................................................
<taxonomicName id="1EEE4D032814FFBDFB00FC12FA59FC79" ID-CoL="8B6S4" authority="Dow &amp; Turner" authorityName="Dow &amp; Turner" authorityYear="1976" box="[1163,1436,972,995]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="uncistyla">
<emphasis id="EB9AEA922814FFBDFB00FC12FA59FC79" bold="true" box="[1163,1436,972,995]" pageId="4" pageNumber="433">
<emphasis id="EB9AEA922814FFBDFB00FC12FB5EFC79" bold="true" box="[1163,1179,973,994]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB2CFC13FAC5FC79" bold="true" box="[1191,1280,972,994]" italics="true" pageId="4" pageNumber="433">uncistyla</emphasis>
Dow &amp; Turner
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<paragraph id="D95136802814FFBDFF1CFC35FA59FB86" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFC35FA59FB86" pageId="4" pageNumber="433">
18. Hind tibia without large apical spines (
<figureCitation id="41D52A052814FFBDFDD7FC35FD60FB9B" box="[604,677,1002,1024]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">Fig. 34</figureCitation>
); antennal flagellum very short to short (length
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); spermathecae with greatly tapered necks (
<figureCitation id="41D52A052814FFBDFE67FBD8FDF9FB86" box="[492,572,1031,1053]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">Figs. 36</figureCitation>
,
<figureCitation id="41D52A052814FFBDFDCDFBD7FD9AFB85" box="[582,607,1032,1054]" captionStart="FIGURES 67–71" captionStartId="34.[151,264,1932,1956]" captionTargetBox="[308,1371,612,1898]" captionTargetId="figure-231@34.[208,1380,597,1908]" captionTargetPageId="34" captionText="FIGURES 67–71. Bezzia titanochela n. sp. Male, 67–70, Female, 71. 67 Thorax and legs. 68 Mid tibial apical spines. 69 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 70 Genitalia at focal level of parameres, aedeagus, gonocoxites and gonostyli." figureDoi="http://doi.org/10.5281/zenodo.4425575" httpUri="https://zenodo.org/record/4425575/files/figure.png" pageId="4" pageNumber="433">71</figureCitation>
).................................................................. 19
</keyLead>
</paragraph>
<paragraph id="D95136802814FFBDFF1CFBFAFA59FBCC" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFBFAFA59FBCC" pageId="4" pageNumber="433">
- Hind tibia with 3 large apical spines; antennal flagellum short to moderately long (length
<quantity id="1E169B652814FFBDFBCCFBFAFB12FBA1" box="[1095,1239,1061,1083]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.35" metricValueMin="1.01" pageId="4" pageNumber="433" unit="mm" value="1.18" valueMax="1.35" valueMin="1.01">1.01–1.35 mm</quantity>
); spermathecae with slightly to non-tapered neck regions (
<figureCitation id="41D52A052814FFBDFDC3FB9DFD49FBC3" box="[584,652,1090,1112]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Figs. 6</figureCitation>
,
<figureCitation id="41D52A052814FFBDFD1DFB9DFD6AFBC3" box="[662,687,1090,1112]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">23</figureCitation>
)........................................................... 20
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</keyStep>
<keyStep id="621A2E252814FFBDFF1CFBBFFA58FB56" pageId="4" pageNumber="433">
<paragraph id="D95136802814FFBDFF1CFBBFFA58FB09" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFBBFFA58FB09" pageId="4" pageNumber="433">
19. Smaller species (wing length
<quantity id="1E169B652814FFBDFE75FB80FD49FBEE" box="[510,652,1119,1141]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.76" metricValueMin="1.39" pageId="4" pageNumber="433" unit="mm" value="1.575" valueMax="1.76" valueMin="1.39">1.39–1.76 mm</quantity>
;
<figureCitation id="41D52A052814FFBDFD1DFB80FD18FBEE" box="[662,733,1119,1141]" captionStart="FIGURES 31–36" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[172,1386,193,1917]" captionTargetId="figure-21@20.[166,1421,181,1920]" captionTargetPageId="20" captionText="FIGURES 31–36. Bezzia amblystyla, n. sp. Male, 31–33, Female, 34–36. 31, 34 Thorax and legs. 32, 35 Wing. 33 Genitalia at focal level of sternite 9, parameres, aedeagus, gonocoxites and gonostyli. 36 Abdominal segments 7–10 and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425563" httpUri="https://zenodo.org/record/4425563/files/figure.png" pageId="4" pageNumber="433">Fig. 35</figureCitation>
); antennal flagellum very short (length
<quantity id="1E169B652814FFBDFBD2FBBFFB2DFBEE" box="[1113,1256,1120,1141]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" metricValueMax="8.7" metricValueMin="7.7" pageId="4" pageNumber="433" unit="mm" value="0.82" valueMax="0.87" valueMin="0.77">0.77–0.87 mm</quantity>
); palpal segment 3 moderately long (palpal ratio 2.75–4.14)...................................................
<emphasis id="EB9AEA922814FFBDFB5EFBA2FA58FB09" bold="true" box="[1237,1437,1148,1170]" pageId="4" pageNumber="433">
<taxonomicName id="1EEE4D032814FFBDFB5EFBA2FA9FFB09" ID-CoL="9HSM5" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1237,1370,1148,1170]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="amblystyla" status="sp. nov.">
<emphasis id="EB9AEA922814FFBDFB5EFBA2FB20FB09" bold="true" box="[1237,1253,1149,1170]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB7AFBA3FA9FFB09" bold="true" box="[1265,1370,1148,1170]" italics="true" pageId="4" pageNumber="433">amblystyla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92814FFBDFAEDFBA2FA58FB09" box="[1382,1437,1149,1170]" pageId="4" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="D95136802814FFBDFF1CFB44FA58FB56" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFB44FA58FB56" pageId="4" pageNumber="433">
- Larger species (wing length
<quantity id="1E169B652814FFBDFE77FB45FD4BFB2B" box="[508,654,1178,1200]" metricMagnitude="-3" metricUnit="m" metricValue="1.785" metricValueMax="1.83" metricValueMin="1.74" pageId="4" pageNumber="433" unit="mm" value="1.785" valueMax="1.83" valueMin="1.74">1.74–1.83 mm</quantity>
); antennal flagellum short (length
<quantity id="1E169B652814FFBDFC6FFB45FBB3FB2B" box="[996,1142,1178,1200]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" metricValueMax="1.09" metricValueMin="1.03" pageId="4" pageNumber="433" unit="mm" value="1.06" valueMax="1.09" valueMin="1.03">1.03–1.09 mm</quantity>
); palpal segment 3 very short (palpal ratio 2.50–2.60)................................................................
<emphasis id="EB9AEA922814FFBDFB5BFB67FA58FB56" bold="true" box="[1232,1437,1207,1229]" pageId="4" pageNumber="433">
<taxonomicName id="1EEE4D032814FFBDFB5BFB67FA9EFB56" ID-CoL="9HWFM" authority="Grogan, 2020" authorityName="Grogan" authorityYear="2020" box="[1232,1371,1207,1229]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="titanochela" status="sp. nov.">
<emphasis id="EB9AEA922814FFBDFB5BFB67FB25FB56" bold="true" box="[1232,1248,1208,1229]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB67FB68FA9EFB56" bold="true" box="[1260,1371,1207,1229]" italics="true" pageId="4" pageNumber="433">titanochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="F0A957E92814FFBDFAEDFB67FA58FB56" box="[1382,1437,1208,1229]" pageId="4" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="621A2E252814FFBDFF1CFB0AFA59FA2C" pageId="4" pageNumber="433">
<paragraph id="D95136802814FFBDFF1CFB0AFA59FAD9" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFB0AFA59FAD9" pageId="4" pageNumber="433">
20. Fore femur with 5–8 ventral spines; fore femur yellow without brown bands, mid-, hind femora yellow with or without narrow brown apical band (
<figureCitation id="41D52A052814FFBDFE2EFB2DFE27FA93" box="[421,482,1266,1288]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Fig. 4</figureCitation>
); fore tibia yellow with poorly developed brown apical band, mid tibia brown with broad yellow subapical band (
<figureCitation id="41D52A052814FFBDFEF6FAD0FE72FABD" box="[381,439,1295,1318]" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Fig. 4</figureCitation>
); palpal segment 3 moderately stout (palpal ratio 2.82–3.54); spermathecae with non-tapered necks (
<figureCitation id="41D52A052814FFBDFAFEFAD0FF2AFAD8" captionStart="FIGURES 1–6" captionStartId="6.[151,264,1932,1956]" captionTargetBox="[231,1392,192,1896]" captionTargetId="figure-21@6.[185,1403,181,1908]" captionTargetPageId="6" captionText="FIGURES 1–6. Bezzia pulverea. Male, 1–3, Female, 4–6. 1 Apical mid-tibial spines. 2 Genitalia at focal level of aedeagus, gonocoxites and gonostyli. 3 Genitalia at focal level of sternite 9 and parameres. 4 Thorax and legs. 5 Wing. 6 Abdomen, ventral view of spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4425549" httpUri="https://zenodo.org/record/4425549/files/figure.png" pageId="4" pageNumber="433">Fig. 6</figureCitation>
)...............................................................................
<taxonomicName id="1EEE4D032814FFBDFB25FAF2FA59FAD9" ID-CoL="8B6RF" authority="(Coquillett)" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[1198,1436,1324,1346]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="pulverea">
<emphasis id="EB9AEA922814FFBDFB25FAF2FA59FAD9" bold="true" box="[1198,1436,1324,1346]" pageId="4" pageNumber="433">
<emphasis id="EB9AEA922814FFBDFB25FAF2FB7BFAD9" bold="true" box="[1198,1214,1325,1346]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB41FAF3FADAFAD9" bold="true" box="[1226,1311,1324,1346]" italics="true" pageId="4" pageNumber="433">pulverea</emphasis>
(Coquillett)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D95136802814FFBDFF1CFA94FA59FA2C" blockId="4.[151,1437,151,1464]" pageId="4" pageNumber="433">
<keyLead id="621F95B52814FFBDFF1CFA94FA59FA2C" pageId="4" pageNumber="433">
- Fore femur with 2–5 ventral spines; fore-, mid femora yellow with basal and subapical brown bands, hind femur brown with broad subapical yellow band (
<figureCitation id="41D52A052814FFBDFD8BFAB8FD8DFAE5" box="[512,584,1383,1406]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">Fig. 21</figureCitation>
); fore-, mid tibiae light brown with brown stripes on proximal 4/5 of their dorsal surfaces (
<figureCitation id="41D52A052814FFBDFF62FA5AFEF4FA00" box="[233,305,1413,1435]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">Fig. 21</figureCitation>
); palpal segment 3 moderately slender (palpal ratio 3.40–4.50); spermathecae with slightly tapered necks (
<figureCitation id="41D52A052814FFBDFAA4FA5AFAB0FA00" box="[1327,1397,1413,1435]" captionStart="FIGURES 20–24" captionStartId="14.[151,264,1811,1835]" captionTargetBox="[164,1425,186,1775]" captionTargetId="figure-21@14.[151,1436,181,1787]" captionTargetPageId="14" captionText="FIGURES 20–24. Bezzia spicata. Male allotype, 20 Genitalia at focal level of parameres, gonocoxites and gonostyli. Female 21–24. 21 Thorax and legs. 22 Hind tibial spines. 23 Abdominal segments 7–10 and spermathecae. 24 Abdominal segments 8–10 with posteriorly folded anterior prongs of segment 9." figureDoi="http://doi.org/10.5281/zenodo.4425557" httpUri="https://zenodo.org/record/4425557/files/figure.png" pageId="4" pageNumber="433">Fig. 23</figureCitation>
).................................................................................
<taxonomicName id="1EEE4D032814FFBDFB15FA7DFA59FA2C" ID-CoL="8B6RW" authority="Dow &amp; Turner" authorityName="Dow &amp; Turner" authorityYear="1976" box="[1182,1436,1442,1464]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="spicata">
<emphasis id="EB9AEA922814FFBDFB15FA7DFA59FA2C" bold="true" box="[1182,1436,1442,1464]" pageId="4" pageNumber="433">
<emphasis id="EB9AEA922814FFBDFB15FA7DFB6BFA2C" bold="true" box="[1182,1198,1442,1463]" italics="true" pageId="4" pageNumber="433">B</emphasis>
.
<emphasis id="EB9AEA922814FFBDFB31FA7DFB3AFA2C" bold="true" box="[1210,1279,1442,1463]" italics="true" pageId="4" pageNumber="433">spicata</emphasis>
Dow &amp; Turner
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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