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<document ID-DOI="http://doi.org/10.5281/zenodo.5070224" ID-GBIF-Dataset="899b95a0-13b7-42aa-959e-4ad7678fb8a9" ID-GBIF-Taxon="183319411" ID-ISSN="1982-0224" ID-Zenodo-Dep="5070224" LSID="urn:lsid:zoobank.org:act:005062EC-37AE-4834-A166-569CC61DCE4F" checkinTime="1614362464116" checkinUser="carolina" docAuthor="Lima, Flávio C. T. &amp; Moreira, Cristiano R." docDate="2003" docId="512387A3FFBFFFD5BA87281FCF2EFADC" docLanguage="en" docName="NeotropIchthyol.1.1.21-33.pdf" docOrigin="Neotropical Ichthyology 1 (1)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252003000100003&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:7EA6CBAC9B92AC526D6A0354C904CC04.3:NeotropIchthyol.2014-2015.journal_article" docStyleId="7EA6CBAC9B92AC526D6A0354C904CC04" docStyleName="NeotropIchthyol.2014-2015.journal_article" docStyleVersion="3" docTitle="Hyphessobrycon eilyos Lima &amp; Moreira 2003, new species" docType="treatment" docVersion="7" lastPageId="6" lastPageNumber="27" masterDocId="AD1AFFDBFFBBFFD3B9452A5CCC22FFB3" masterDocTitle="Three new species of Hyphessobrycon (Characiformes: Characidae) from the upper rio Araguaia basin in Brazil" masterLastPageNumber="33" masterPageNumber="21" pageId="4" pageNumber="25" updateTime="1635476462862" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of Hyphessobrycon (Characiformes: Characidae) from the upper rio Araguaia basin in Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lima, Flávio C. T.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moreira, Cristiano R.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2003</mods:date>
<mods:detail type="pubDate">
<mods:number>2003-09-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>21</mods:start>
<mods:end>33</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252003000100003&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252003000100003</mods:identifier>
<mods:identifier type="GBIF-Dataset">899b95a0-13b7-42aa-959e-4ad7678fb8a9</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4566752</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5070224" ID-GBIF-Taxon="183319411" ID-Zenodo-Dep="5070224" LSID="urn:lsid:zoobank.org:act:005062EC-37AE-4834-A166-569CC61DCE4F" httpUri="http://treatment.plazi.org/id/512387A3FFBFFFD5BA87281FCF2EFADC" lastPageId="6" lastPageNumber="27" pageId="4" pageNumber="25">
<subSubSection box="[962,1359,578,603]" pageId="4" pageNumber="25" type="nomenclature">
<paragraph blockId="4.[962,1359,578,634]" box="[962,1359,578,603]" pageId="4" pageNumber="25">
<heading box="[962,1359,578,603]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="25" reason="2">
<emphasis bold="true" box="[962,1359,578,603]" pageId="4" pageNumber="25">
<taxonomicName authority="Lima &amp; Moreira, 2003" authorityName="Lima &amp; Moreira" authorityYear="2003" box="[962,1213,579,603]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos" status="new species">
<emphasis bold="true" box="[962,1213,579,603]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1227,1359,578,602]" pageId="4" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1114,1207,611,634]" pageId="4" pageNumber="25" type="description">
<paragraph blockId="4.[962,1359,578,634]" box="[1114,1207,611,634]" pageId="4" pageNumber="25">
<figureCitation box="[1114,1207,611,634]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[151,190,1922,1946]" captionStartId-1="6.[159,197,764,788]" captionTargetBox-0="[261,1347,1467,1896]" captionTargetBox-1="[206,758,177,710]" captionTargetId-0="figure-12@5.[261,1347,1467,1896]" captionTargetId-1="figure-10@6.[191,762,161,715]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 6. Hyphessobrycon eilyos, holotype, MZUSP 75126, 22.3 mm SL, Brazil, Mato Grosso, município de Alto Araguaia, ribeirão do Sapo." captionText-1="Fig. 7. Hyphessobrycon eilyos, paratype, MZUSP 41406.Upper and lower jaws, lateral view, left side. Scale bar 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.4566768" figureDoi-1="http://doi.org/10.5281/zenodo.4566770" httpUri-0="https://zenodo.org/record/4566768/files/figure.png" httpUri-1="https://zenodo.org/record/4566770/files/figure.png" pageId="4" pageNumber="25">Figs. 6-7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="25" type="materials_examined">
<paragraph blockId="4.[839,1482,674,794]" pageId="4" pageNumber="25">
<materialsCitation ID-GBIF-Occurrence="3313891304" collectingDate="2001-05-21" collectionCode="R, T" collectorName="de Alto Araguaia" country="Brazil" latitude="-17.519722" location="Mato Grosso" longLatPrecision="21" longitude="-53.259167" pageId="4" pageNumber="25" specimenCode="MZUSP 75126" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis bold="true" box="[839,947,674,698]" pageId="4" pageNumber="25">
<typeStatus box="[839,942,674,698]" pageId="4" pageNumber="25">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[951,1108,675,698]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="25">MZUSP 75126</specimenCode>
(
<quantity box="[1120,1212,675,698]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="4" pageNumber="25" unit="mm" value="22.3">22.3 mm</quantity>
SL):
<collectingCountry box="[1268,1334,675,698]" name="Brazil" pageId="4" pageNumber="25">Brazil</collectingCountry>
,
<collectingRegion box="[1342,1478,675,698]" country="Brazil" name="Mato Grosso" pageId="4" pageNumber="25">Mato Grosso</collectingRegion>
, município
<collectorName box="[954,1140,707,730]" pageId="4" pageNumber="25">de Alto Araguaia</collectorName>
, ribeirão do Sapo, km 464.04 of Ferronorte railroad,
<geoCoordinate box="[1042,1165,739,762]" degrees="17" direction="south" minutes="31" orientation="latitude" pageId="4" pageNumber="25" precision="15" seconds="11" value="-17.519722">17°3111”S</geoCoordinate>
,
<geoCoordinate box="[1174,1309,739,762]" degrees="53" direction="west" minutes="15" orientation="longitude" pageId="4" pageNumber="25" precision="15" seconds="33" value="-53.259167">53°1533”W</geoCoordinate>
; C.
<collectionCode box="[1346,1366,739,762]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="25">R</collectionCode>
. Moreira &amp; F.C.
<collectionCode box="[881,899,771,794]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="25">T</collectionCode>
. Lima,
<date box="[971,1108,771,794]" pageId="4" pageNumber="25" value="2001-05-21">
<collectingDate box="[971,1108,771,794]" pageId="4" pageNumber="25" value="2001-05-21">21 May 2001</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4566764" ID-Zenodo-Dep="4566764" httpUri="https://zenodo.org/record/4566764/files/figure.png" pageId="4" pageNumber="25" startId="4.[159,198,1130,1154]" targetBox="[158,795,190,1100]" targetPageId="4">
<paragraph blockId="4.[159,802,1130,1346]" pageId="4" pageNumber="25">
<emphasis bold="true" box="[159,227,1130,1154]" pageId="4" pageNumber="25">Fig. 5.</emphasis>
Upper rio Araguaia basin, showing sites of occurrence of
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[202,536,1163,1186]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[202,536,1163,1186]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon langeanii</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[565,683,1163,1186]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[565,683,1163,1186]" italics="true" pageId="4" pageNumber="25">H. eilyos</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="weitzmanorum">
<emphasis italics="true" pageId="4" pageNumber="25">H. weitzmanorum</emphasis>
</taxonomicName>
(diamonds),
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[523,687,1195,1218]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[523,687,1195,1218]" italics="true" pageId="4" pageNumber="25">H. langeanii</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="weitzmanorum">
<emphasis italics="true" pageId="4" pageNumber="25">H. weitzmanorum</emphasis>
</taxonomicName>
(dots),
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[401,536,1227,1250]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[401,536,1227,1250]" italics="true" pageId="4" pageNumber="25">H. langeanii</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[590,686,1227,1250]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[590,686,1227,1250]" italics="true" pageId="4" pageNumber="25">H. eilyos</emphasis>
</taxonomicName>
(triangle), or
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[188,322,1259,1282]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[188,322,1259,1282]" italics="true" pageId="4" pageNumber="25">H. langeanii</emphasis>
</taxonomicName>
(squares). 1. Ribeirão do Sapo, type locality of
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[185,276,1291,1314]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[185,276,1291,1314]" italics="true" pageId="4" pageNumber="25">H. eilyos</emphasis>
</taxonomicName>
. 2. Córrego Mosquito, type locality of
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[673,801,1291,1314]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[673,801,1291,1314]" italics="true" pageId="4" pageNumber="25">H. langeanii</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[204,394,1323,1346]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="weitzmanorum">
<emphasis box="[204,394,1323,1346]" italics="true" pageId="4" pageNumber="25">H. weitzmanorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="4.[159,801,1443,1530]" pageId="4" pageNumber="25">
<emphasis bold="true" box="[159,458,1443,1467]" pageId="4" pageNumber="25">Geographic distribution.</emphasis>
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[469,772,1443,1466]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[469,772,1443,1466]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon langeanii</emphasis>
</taxonomicName>
<materialsCitation ID-GBIF-Occurrence="3313891338" country="Brazil" location="upper rio Araguaia" pageId="4" pageNumber="25" specimenCount="1" stateProvince="Mato Grosso">
is known from several streams in the
<location LSID="urn:lsid:plazi:treatment:512387A3FFBFFFD5BA87281FCF2EFADC:DC55606EFFBFFFD7BB562F9FCEFEFA69" box="[531,732,1475,1498]" country="Brazil" name="upper rio Araguaia" pageId="4" pageNumber="25" stateProvince="Mato Grosso">upper rio Araguaia</location>
basin, states of
<collectingRegion box="[252,390,1507,1530]" country="Brazil" name="Mato Grosso" pageId="4" pageNumber="25">Mato Grosso</collectingRegion>
and
<collectingRegion box="[439,504,1507,1530]" country="Brazil" name="Goias" pageId="4" pageNumber="25">Goiás</collectingRegion>
,
<collectingCountry box="[514,579,1507,1530]" name="Brazil" pageId="4" pageNumber="25">Brazil</collectingCountry>
(
<figureCitation box="[593,657,1507,1530]" captionStart="Fig" captionStartId="4.[159,198,1130,1154]" captionTargetBox="[158,795,190,1100]" captionTargetId="figure-9@4.[147,801,183,1107]" captionTargetPageId="4" captionText="Fig. 5. Upper rio Araguaia basin, showing sites of occurrence of Hyphessobrycon langeanii, H. eilyos, and H. weitzmanorum (diamonds), H. langeanii and H. weitzmanorum (dots), H. langeanii and H. eilyos (triangle), or H. langeanii (squares). 1. Ribeirão do Sapo, type locality of H.eilyos. 2. Córrego Mosquito, type locality of H. langeanii and H. weitzmanorum." figureDoi="http://doi.org/10.5281/zenodo.4566764" httpUri="https://zenodo.org/record/4566764/files/figure.png" pageId="4" pageNumber="25">Fig. 5</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="4.[159,802,1571,1978]" pageId="4" pageNumber="25">
<emphasis bold="true" box="[159,352,1571,1595]" pageId="4" pageNumber="25">Ecological notes.</emphasis>
We observed
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[512,802,1571,1594]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[512,802,1571,1594]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon langeanii</emphasis>
</taxonomicName>
in schools, generally at midwater, sometimes associated with an unidentified
<taxonomicName box="[331,429,1635,1658]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis box="[331,429,1635,1658]" italics="true" pageId="4" pageNumber="25">Astyanax</emphasis>
</taxonomicName>
species. Habitats occupied by the species ranged from relatively large, deep, clearwater streams partially covered with aquatic vegetation (i.e., ribeirão do Sapo, córrego Gordura) to shallow, small streams with flooded areas (i.e., córrego Mosquito).
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[449,747,1763,1786]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[449,747,1763,1786]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon langeanii</emphasis>
</taxonomicName>
was collected syntopically with
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[448,635,1795,1818]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="weitzmanorum">
<emphasis box="[448,635,1795,1818]" italics="true" pageId="4" pageNumber="25">H. weitzmanorum</emphasis>
</taxonomicName>
at córrego Gordura, córrego Boiadeiro, córrego do Sapinho, and córrego do Mosquito, with
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[337,436,1859,1882]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[337,436,1859,1882]" italics="true" pageId="4" pageNumber="25">H. eilyos</emphasis>
</taxonomicName>
at córrego do Rancho, and with both species at the ribeirão do Sapo. The occurrence of
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[159,445,1923,1946]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[159,445,1923,1946]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon langeanii</emphasis>
</taxonomicName>
in a broad number of sites, even in highly degraded, silted streams such as córrego Bandeira,
</paragraph>
<paragraph blockId="4.[839,1482,834,1146]" pageId="4" pageNumber="25">
<materialsCitation ID-GBIF-Occurrence="3313891343" collectingDate="2001-05-22" collectionCode="R, T" country="Brazil" county="All" latitude="-17.27" location="Veado" longLatPrecision="21" longitude="-53.414448" municipality="Rancho" pageId="4" pageNumber="25" specimenCode="MZUSP 73344, MZUSP 73363" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<emphasis bold="true" box="[839,958,834,858]" pageId="4" pageNumber="25">
<typeStatus box="[839,953,834,858]" pageId="4" pageNumber="25">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty box="[962,996,835,858]" pageId="4" pageNumber="25">All</collectingCounty>
localities in
<collectingCountry box="[1125,1192,835,858]" name="Brazil" pageId="4" pageNumber="25">Brazil</collectingCountry>
,
<collectingRegion box="[1200,1337,835,858]" country="Brazil" name="Mato Grosso" pageId="4" pageNumber="25">Mato Grosso</collectingRegion>
, município
<collectorName pageId="4" pageNumber="25">de Alto Araguaia</collectorName>
:
<specimenCode box="[1003,1165,867,890]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="25">MZUSP 73344</specimenCode>
(11, 15.0-
<quantity box="[1273,1369,867,890]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="4" pageNumber="25" unit="mm" value="23.3">23.3 mm</quantity>
SL), same data as holotype.
<specimenCode box="[1020,1181,899,922]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="25">MZUSP 73363</specimenCode>
(1,
<quantity box="[1219,1312,899,922]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="4" pageNumber="25" unit="mm" value="22.1">22.1 mm</quantity>
SL), córrego do
<collectingMunicipality box="[839,920,931,954]" pageId="4" pageNumber="25">Rancho</collectingMunicipality>
, below lagoa do
<location LSID="urn:lsid:plazi:treatment:512387A3FFBFFFD5BA87281FCF2EFADC:DC55606EFFBFFFD7BD7829FFC8A2FC09" box="[1085,1152,931,954]" country="Brazil" county="All" latitude="-17.27" longLatPrecision="21" longitude="-53.414448" municipality="Rancho" name="Veado" pageId="4" pageNumber="25" stateProvince="Mato Grosso">Veado</location>
,
<geoCoordinate box="[1160,1284,931,954]" degrees="17" direction="south" minutes="16" orientation="latitude" pageId="4" pageNumber="25" precision="15" seconds="12" value="-17.27">17°1612”S</geoCoordinate>
,
<geoCoordinate box="[1292,1426,931,954]" degrees="53" direction="west" minutes="24" orientation="longitude" pageId="4" pageNumber="25" precision="15" seconds="52" value="-53.414448">53°2452”W</geoCoordinate>
; C.
<collectionCode box="[1457,1478,931,954]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="25">R</collectionCode>
.
<collectorName box="[839,921,963,986]" pageId="4" pageNumber="25">Moreira</collectorName>
&amp; F.C.
<collectionCode box="[991,1008,963,986]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="25">T</collectionCode>
.
<collectorName box="[1015,1072,963,986]" pageId="4" pageNumber="25">Lima</collectorName>
,
<date box="[1080,1215,963,986]" pageId="4" pageNumber="25" value="2001-05-22">
<collectingDate box="[1080,1215,963,986]" pageId="4" pageNumber="25" value="2001-05-22">22 May 2001</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3313891339" collectingDate="1989-03-08" collectionCode="MZUSP, MCZ, USNM, MNRJ" collectorName="R Pezzi da Silva &amp; F. Langeani" country="Portugal" latitude="-17.266666" location="Portugal" longLatPrecision="1279" longitude="-53.4" pageId="4" pageNumber="25" specimenCode="MZUSP 41406, MCZ 162371, USNM 371923, MNRJ 24781" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1222,1376,963,986]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="25">MZUSP 41406</specimenCode>
(179, 5 cs,
<quantity box="[839,982,995,1018]" metricMagnitude="-2" metricUnit="m" metricValue="1.9649999999999999" metricValueMax="2.4699999999999998" metricValueMin="1.46" pageId="4" pageNumber="25" unit="mm" value="19.65" valueMax="24.7" valueMin="14.6">14.6-24.7 mm</quantity>
SL);
<specimenCode box="[1036,1173,995,1018]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="25" type="Museum">MCZ 162371</specimenCode>
(5,
<quantity box="[1209,1352,995,1018]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.36" metricValueMin="2.04" pageId="4" pageNumber="25" unit="mm" value="22.0" valueMax="23.6" valueMin="20.4">20.4-23.6 mm</quantity>
SL);
<specimenCode collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="25" type="Museum">USNM 371923</specimenCode>
(5,
<quantity box="[946,1083,1027,1050]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" metricValueMax="2.31" metricValueMin="1.95" pageId="4" pageNumber="25" unit="mm" value="21.3" valueMax="23.1" valueMin="19.5">19.5-23.1 mm</quantity>
SL);
<specimenCode box="[1133,1266,1027,1050]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="4" pageNumber="25">MNRJ 24781</specimenCode>
(5,
<quantity box="[1299,1435,1027,1050]" metricMagnitude="-2" metricUnit="m" metricValue="2.185" metricValueMax="2.21" metricValueMin="2.16" pageId="4" pageNumber="25" unit="mm" value="21.85" valueMax="22.1" valueMin="21.6">21.6-22.1mm</quantity>
SL), córrego do Rancho (headwaters), swamp near the road, approx.
<geoCoordinate box="[839,929,1091,1114]" degrees="17" direction="south" minutes="16" orientation="latitude" pageId="4" pageNumber="25" precision="925" value="-17.266666">17°16S</geoCoordinate>
,
<geoCoordinate box="[941,1041,1091,1114]" degrees="53" direction="west" minutes="24" orientation="longitude" pageId="4" pageNumber="25" precision="925" value="-53.4">53°24W</geoCoordinate>
; L.
<collectorName box="[1077,1117,1091,1114]" name="R Pezzi da Silva" pageId="4" pageNumber="25">P.S.</collectorName>
<collectingCountry box="[1124,1215,1091,1114]" name="Portugal" pageId="4" pageNumber="25">Portugal</collectingCountry>
&amp;
<collectorName box="[1251,1380,1091,1114]" pageId="4" pageNumber="25">F. Langeani</collectorName>
,
<date pageId="4" pageNumber="25" value="1989-03-08">
<collectingDate pageId="4" pageNumber="25" value="1989-03-08">8 March 1989</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="25" type="diagnosis">
<paragraph blockId="4.[839,1483,1186,1562]" pageId="4" pageNumber="25">
<emphasis bold="true" box="[839,956,1186,1210]" pageId="4" pageNumber="25">Diagnosis.</emphasis>
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[964,1212,1187,1210]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[964,1212,1187,1210]" italics="true" pageId="4" pageNumber="25">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
is distinguished from its congeners by the absence of humeral and caudal spots; by the presence of numerous dark chromatophores on the lateral surface of the body, with the chromatophores intensely concentrated on the ventral region from the pelvic-fin origin to the end of the caudal-fin base; dorsal, adipose, and caudal fins with carmine red pigmentation in life; the presence of 7- 11 maxillary teeth; 6 or 7 horizontal scale rows between the dorsal-fin origin and the lateral line; and 15 to 19 branched anal-fin rays. Detailed comparisons between
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[1336,1433,1475,1498]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="25" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[1336,1433,1475,1498]" italics="true" pageId="4" pageNumber="25">H. eilyos</emphasis>
</taxonomicName>
and congeners sharing a general dark color pattern are presented in the “Discussion”, below.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="27" pageId="4" pageNumber="25" type="description">
<paragraph blockId="4.[839,1482,1602,1978]" lastBlockId="5.[151,795,195,1434]" lastPageId="5" lastPageNumber="26" pageId="4" pageNumber="25">
<emphasis bold="true" box="[877,1020,1602,1626]" pageId="4" pageNumber="25">Description.</emphasis>
Morphometric data of the
<typeStatus box="[1335,1431,1603,1626]" pageId="4" pageNumber="25">holotype</typeStatus>
and
<typeStatus box="[839,948,1635,1658]" pageId="4" pageNumber="25">paratypes</typeStatus>
are presented in
<tableCitation box="[1161,1253,1635,1658]" captionStart="Table 2" captionStartId="6.[839,904,194,218]" captionTargetBox="[839,1455,354,1082]" captionTargetId="graphics-50@6.[839,1482,341,1099]" captionText="Table 2. Morphometric data for Hyphessobrycon eilyos. Measurements based on the holotype (MZUSP 75126) and 61 paratypes (MZUSP 41406, 56 ex.; MZUSP 73344, 4 ex.; MZUSP 73363, 1 ex.)." httpUri="http://table.plazi.org/id/8DF5663DFFBDFFD5BA022A9EC80BFE89" pageId="4" pageNumber="25" tableUuid="8DF5663DFFBDFFD5BA022A9EC80BFE89">Table 2</tableCitation>
. Body compressed, moderately deep, greatest body depth immediately anterior to dorsal-fin origin. Dorsal profile of head slightly convex from upper lip to vertical through middle of the orbit; straight to slightly concave from latter point to tip of supraoccipital spine. Predorsal profile of body convex, dorsal-fin base straight to slightly convex, posteroventrally inclined. Body profile straight to convex from end of dorsal-fin base to adipose fin; slightly concave between latter point and origin of dorsal most procurrent caudal-fin ray. Ventral profile of head and body convex from lower lip to pelvic-fin origin. Body profile straight from pelvic-fin insertion to anal-fin origin. Body profile along anal-fin base posterodorsally slanted and slightly concave to slightly convex. Ventral profile of caudal peduncle slightly concave.
</paragraph>
<paragraph blockId="5.[151,795,195,1434]" pageId="5" pageNumber="26">
Jaws unequal, mouth terminal, anteroventral end of dentary protruding slightly. Maxilla extends posteriorly to under middle of orbit. Premaxillary teeth in two rows (
<figureCitation box="[717,780,387,410]" captionStart="Fig" captionStartId="6.[159,197,764,788]" captionTargetBox="[206,758,177,710]" captionTargetId="figure-10@6.[191,762,161,715]" captionTargetPageId="6" captionText="Fig. 7. Hyphessobrycon eilyos, paratype, MZUSP 41406.Upper and lower jaws, lateral view, left side. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4566770" httpUri="https://zenodo.org/record/4566770/files/figure.png" pageId="5" pageNumber="26">Fig. 7</figureCitation>
). Outer row with 3(7) uni- to tricuspid teeth. Inner row with 6(7) tri- to pentacuspid teeth. Maxilla with 7(2), 8(3), 9(1), or 11(1), uni- to tricuspid teeth. Dentary with 4(3), 5(3), or 6(1) large tricuspid teeth followed by 9(1), 11(3), 12(2), or 14(1) smaller uni- to tricuspid teeth.
</paragraph>
<paragraph blockId="5.[151,795,195,1434]" pageId="5" pageNumber="26">Scales cycloid, with few radii. Lateral line incompletely pored, with 3(1), 4(7), 5(18), 6*(13), or 8(1) perforated scales. Lateral series scales including perforated scales 28(1), 29(2), 30*(10), 31(5), 32(1), or 33(1). Horizontal scale rows between dorsal-fin origin and lateral line 6*(26) or 7(16), not including scale of predorsal series situated just anterior to first dorsalfin ray. Horizontal scale rows between lateral line and pelvicfin origin 4*(43) or 5(3). Predorsal scales 9(1), 10*(8), 11(7), 12(4), or 13(1). Circumpeduncular scales 11(1), 12(4), 13(7), or 15(1). Single row of 5-6 scales covering basis of anteriormost anal-fin rays.</paragraph>
<paragraph blockId="5.[151,795,195,1434]" lastBlockId="5.[832,1475,195,378]" pageId="5" pageNumber="26">
Dorsal-fin rays ii,8(3), or ii,9*(61). Dorsal-fin origin at middle of standard length. Base of last dorsal-fin ray at vertical through just before anal-fin origin. First dorsal-fin pterygiophore inserting behind neural spine of 9th(7) vertebra. Adipose fin typically present, but reduced in some specimens, and absent in 52 of
<specimenCount box="[374,534,1091,1114]" pageId="5" pageNumber="26" type="generic">215 specimens</specimenCount>
examined. Unbranched anal-fin rays iii*(52) or iv(12). Branched anal fin rays 15(1), 16(4), 17*(28), 18(23), or 19(8). First anal-fin pterygiophore inserting behind hemal spine of 15th(1), or 16th(6) vertebra. Pectoral-fin rays i,7(1), 9(4), 10(27), 11*(29), or 12(3). Specimens up to
<quantity box="[208,300,1251,1274]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="5" pageNumber="26" unit="mm" value="14.9">14.9 mm</quantity>
SL retaining larval pectoral fin anatomy. Tip of pectoral fin reaching vertical through pelvic-fin origin. Pelvicfin rays i,5(7), 6*(56), or 9(1). Tip of pelvic fin reaching origin of anal fin. Caudal fin forked, upper and lower lobes rounded, and similar in size. Principal caudal-fin rays 10+9(4). Eight (1), 10(1), or 12(2) dorsal procurrent caudal-fin rays, and 9(1), 10(2), or 11(1) ventral procurrent caudal-fin rays. First gill arch with 6(7) epibranchial; 1(7) on cartilage between epibranchial and ceratobranchial, 10(7) ceratobranchial, and 2(7) hypobranchial gill-rakers. Four (7) branchiostegal rays, 3(7) on anterior ceratohyal, and 1(7) on posterior ceratohyal Vertebrae 32(4), 33(2), or 34(1). Supraneurals 4(5), or 5(2).
</paragraph>
<paragraph blockId="5.[832,1475,418,1082]" pageId="5" pageNumber="26">
<emphasis bold="true" box="[832,1016,418,442]" pageId="5" pageNumber="26">Color in alcohol</emphasis>
. Ground color cream, tanner in specimens stored for long time in ethanol. Guanine present on opercle and infraorbitals. Dark chromatophores densely concentrated on dorsal surface of head, and anterior surface of lower jaw. Dark chromatophores scattered on remaining portions of head. Dorsal midline with dense concentration of small dark chromatophores. Dorso and ventrolateral portions of body with dense concentration of dark chromatophores, uniformly distributed. Humeral spot absent (apparent humeral spot in
<figureCitation box="[832,893,707,730]" captionStart="Fig" captionStartId="5.[151,190,1922,1946]" captionTargetBox="[261,1347,1467,1896]" captionTargetId="figure-12@5.[261,1347,1467,1896]" captionTargetPageId="5" captionText="Fig. 6. Hyphessobrycon eilyos, holotype, MZUSP 75126, 22.3 mm SL, Brazil, Mato Grosso, município de Alto Araguaia, ribeirão do Sapo." figureDoi="http://doi.org/10.5281/zenodo.4566768" httpUri="https://zenodo.org/record/4566768/files/figure.png" pageId="5" pageNumber="26">Fig. 6</figureCitation>
is actually a result of the reduction of the musculature in that portion of the body wall). Dense concentration of deep-lying dark chromatophores along ventral midline, more concentrated from pelvic-fin insertion to posterior portion of anal-fin base. Chromatophores on caudal-fin base absent, resulting in a light area. Caudal fin with dark chromatophores concentrated on middle rays and distal margin. Dorsal fin with concentration of dark chromatophores along distal margin. Anal fin with heavy concentration of dark chromatophores, mainly on interradial fin membrane. Pectoral, pelvic, and adipose fins with dense concentration of dark chromatophores.
</paragraph>
<paragraph blockId="5.[832,1475,1123,1274]" pageId="5" pageNumber="26">
<emphasis bold="true" box="[832,992,1123,1147]" pageId="5" pageNumber="26">Color in life.</emphasis>
Description based on photograph of the
<typeStatus box="[832,926,1155,1178]" pageId="5" pageNumber="26">holotype</typeStatus>
(MZUSP 75126). Ground color dark gray. Sides of head, and body silvery. Dorsal and adipose fins bright red. Caudal fin bright red, except for distal portion of caudal-fin lobes.
</paragraph>
<paragraph blockId="5.[832,1474,1315,1434]" pageId="5" pageNumber="26">
<emphasis bold="true" box="[832,1115,1315,1339]" pageId="5" pageNumber="26">Geographic distribution.</emphasis>
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[1121,1369,1315,1338]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[1121,1369,1315,1338]" italics="true" pageId="5" pageNumber="26">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
is known from córrego do Rancho and ribeirão do Sapo, both tributaries of the upper rio Araguaia basin in the state of
<collectingRegion box="[1328,1469,1379,1402]" country="Brazil" name="Mato Grosso" pageId="5" pageNumber="26">Mato Grosso</collectingRegion>
,
<collectingCountry box="[832,894,1411,1434]" name="Brazil" pageId="5" pageNumber="26">Brazil</collectingCountry>
(
<figureCitation box="[905,965,1411,1434]" captionStart="Fig" captionStartId="4.[159,198,1130,1154]" captionTargetBox="[158,795,190,1100]" captionTargetId="figure-9@4.[147,801,183,1107]" captionTargetPageId="4" captionText="Fig. 5. Upper rio Araguaia basin, showing sites of occurrence of Hyphessobrycon langeanii, H. eilyos, and H. weitzmanorum (diamonds), H. langeanii and H. weitzmanorum (dots), H. langeanii and H. eilyos (triangle), or H. langeanii (squares). 1. Ribeirão do Sapo, type locality of H.eilyos. 2. Córrego Mosquito, type locality of H. langeanii and H. weitzmanorum." figureDoi="http://doi.org/10.5281/zenodo.4566764" httpUri="https://zenodo.org/record/4566764/files/figure.png" pageId="5" pageNumber="26">Fig. 5</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4566768" ID-Zenodo-Dep="4566768" httpUri="https://zenodo.org/record/4566768/files/figure.png" pageId="5" pageNumber="26" startId="5.[151,190,1922,1946]" targetBox="[261,1347,1467,1896]" targetPageId="5">
<paragraph blockId="5.[151,1473,1922,1977]" pageId="5" pageNumber="26">
<emphasis bold="true" box="[151,223,1922,1946]" pageId="5" pageNumber="26">Fig. 6.</emphasis>
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[231,479,1923,1946]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="26" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[231,479,1923,1946]" italics="true" pageId="5" pageNumber="26">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
, holotype, MZUSP 75126, 22.3 mm SL, Brazil, Mato Grosso, município de Alto Araguaia, ribeirão do Sapo.
</paragraph>
</caption>
<caption ID-Table-UUID="8DF5663DFFBDFFD5BA022A9EC80BFE89" httpUri="http://table.plazi.org/id/8DF5663DFFBDFFD5BA022A9EC80BFE89" pageId="6" pageNumber="27" startId="6.[839,904,194,218]" targetBox="[839,1455,354,1082]" targetIsTable="true" targetPageId="6">
<paragraph blockId="6.[839,1482,194,314]" pageId="6" pageNumber="27">
<emphasis bold="true" box="[839,934,194,218]" pageId="6" pageNumber="27">Table 2.</emphasis>
Morphometric data for
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[1216,1474,195,218]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[1216,1474,195,218]" italics="true" pageId="6" pageNumber="27">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
. Measurements based on the holotype (MZUSP 75126) and 61 paratypes (MZUSP 41406, 56 ex.; MZUSP 73344, 4 ex.; MZUSP 73363, 1 ex.).
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="27">
<table box="[839,1455,354,1082]" gridcols="5" gridrows="25" pageId="6" pageNumber="27">
<tr box="[839,1455,354,375]" gridrow="0" pageId="6" pageNumber="27" rowspan-0="1">
<th box="[1150,1272,354,375]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="27">holotype n</th>
<th box="[1293,1387,354,375]" gridcol="3" gridrow="0" pageId="6" pageNumber="27">range</th>
<th box="[1405,1455,354,375]" gridcol="4" gridrow="0" pageId="6" pageNumber="27">mean</th>
</tr>
<tr box="[839,1455,398,419]" gridrow="1" pageId="6" pageNumber="27">
<th box="[839,1129,398,419]" gridcol="0" gridrow="1" pageId="6" pageNumber="27">Standard length (mm)</th>
<td box="[1150,1193,398,419]" gridcol="1" gridrow="1" pageId="6" pageNumber="27">22.3</td>
<td box="[1247,1272,398,419]" gridcol="2" gridrow="1" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,398,419]" gridcol="3" gridrow="1" pageId="6" pageNumber="27">15.8-24.7</td>
<td box="[1405,1455,398,419]" gridcol="4" gridrow="1" pageId="6" pageNumber="27">-</td>
</tr>
<tr box="[839,1455,427,448]" gridrow="2" pageId="6" pageNumber="27">
<th box="[839,1455,427,448]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="6" pageNumber="27">
<emphasis bold="true" box="[839,1129,427,448]" pageId="6" pageNumber="27">Percents of standard length</emphasis>
</th>
</tr>
<tr box="[839,1455,456,477]" gridrow="3" pageId="6" pageNumber="27">
<th box="[839,1129,456,477]" gridcol="0" gridrow="3" pageId="6" pageNumber="27">Depth at dorsal-fin origin</th>
<td box="[1150,1193,456,477]" gridcol="1" gridrow="3" pageId="6" pageNumber="27">40.0</td>
<td box="[1247,1272,456,477]" gridcol="2" gridrow="3" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,456,477]" gridcol="3" gridrow="3" pageId="6" pageNumber="27">32.0-40.7</td>
<td box="[1405,1455,456,477]" gridcol="4" gridrow="3" pageId="6" pageNumber="27">37.2</td>
</tr>
<tr box="[839,1455,485,506]" gridrow="4" pageId="6" pageNumber="27">
<th box="[839,1129,485,506]" gridcol="0" gridrow="4" pageId="6" pageNumber="27">Snout to dorsal fin origin</th>
<td box="[1150,1193,485,506]" gridcol="1" gridrow="4" pageId="6" pageNumber="27">54.1</td>
<td box="[1247,1272,485,506]" gridcol="2" gridrow="4" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,485,506]" gridcol="3" gridrow="4" pageId="6" pageNumber="27">50.6-58.5</td>
<td box="[1405,1455,485,506]" gridcol="4" gridrow="4" pageId="6" pageNumber="27">53.6</td>
</tr>
<tr box="[839,1455,514,535]" gridrow="5" pageId="6" pageNumber="27">
<th box="[839,1129,514,535]" gridcol="0" gridrow="5" pageId="6" pageNumber="27">Snout to pectoral-fin origin</th>
<td box="[1150,1193,514,535]" gridcol="1" gridrow="5" pageId="6" pageNumber="27">33.8</td>
<td box="[1247,1272,514,535]" gridcol="2" gridrow="5" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,514,535]" gridcol="3" gridrow="5" pageId="6" pageNumber="27">30.5-36.6</td>
<td box="[1405,1455,514,535]" gridcol="4" gridrow="5" pageId="6" pageNumber="27">33.0</td>
</tr>
<tr box="[839,1455,542,563]" gridrow="6" pageId="6" pageNumber="27">
<th box="[839,1129,542,563]" gridcol="0" gridrow="6" pageId="6" pageNumber="27">Snout to pelvic-fin origin</th>
<td box="[1150,1193,542,563]" gridcol="1" gridrow="6" pageId="6" pageNumber="27">51.4</td>
<td box="[1247,1272,542,563]" gridcol="2" gridrow="6" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,542,563]" gridcol="3" gridrow="6" pageId="6" pageNumber="27">47.4-55.9</td>
<td box="[1405,1455,542,563]" gridcol="4" gridrow="6" pageId="6" pageNumber="27">50.2</td>
</tr>
<tr box="[839,1455,571,592]" gridrow="7" pageId="6" pageNumber="27">
<th box="[839,1129,571,592]" gridcol="0" gridrow="7" pageId="6" pageNumber="27">Snout to anal-fin origin</th>
<td box="[1150,1193,571,592]" gridcol="1" gridrow="7" pageId="6" pageNumber="27">64.8</td>
<td box="[1247,1272,571,592]" gridcol="2" gridrow="7" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,571,592]" gridcol="3" gridrow="7" pageId="6" pageNumber="27">61.9-69.0</td>
<td box="[1405,1455,571,592]" gridcol="4" gridrow="7" pageId="6" pageNumber="27">64.7</td>
</tr>
<tr box="[839,1455,600,621]" gridrow="8" pageId="6" pageNumber="27">
<th box="[839,1129,600,621]" gridcol="0" gridrow="8" pageId="6" pageNumber="27">Caudal peduncle depth</th>
<td box="[1150,1193,600,621]" gridcol="1" gridrow="8" pageId="6" pageNumber="27">12.2</td>
<td box="[1247,1272,600,621]" gridcol="2" gridrow="8" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,600,621]" gridcol="3" gridrow="8" pageId="6" pageNumber="27">9.2-13.0</td>
<td box="[1405,1455,600,621]" gridcol="4" gridrow="8" pageId="6" pageNumber="27">11.3</td>
</tr>
<tr box="[839,1455,629,650]" gridrow="9" pageId="6" pageNumber="27">
<th box="[839,1129,629,650]" gridcol="0" gridrow="9" pageId="6" pageNumber="27">Caudal peduncle length</th>
<td box="[1150,1193,629,650]" gridcol="1" gridrow="9" pageId="6" pageNumber="27">12.6</td>
<td box="[1247,1272,629,650]" gridcol="2" gridrow="9" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,629,650]" gridcol="3" gridrow="9" pageId="6" pageNumber="27">11.7-17.4</td>
<td box="[1405,1455,629,650]" gridcol="4" gridrow="9" pageId="6" pageNumber="27">15.1</td>
</tr>
<tr box="[839,1455,658,679]" gridrow="10" pageId="6" pageNumber="27">
<th box="[839,1129,658,679]" gridcol="0" gridrow="10" pageId="6" pageNumber="27">Pectoral-fin length</th>
<td box="[1150,1193,658,679]" gridcol="1" gridrow="10" pageId="6" pageNumber="27">21.0</td>
<td box="[1247,1272,658,679]" gridcol="2" gridrow="10" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,658,679]" gridcol="3" gridrow="10" pageId="6" pageNumber="27">14.0-22.6</td>
<td box="[1405,1455,658,679]" gridcol="4" gridrow="10" pageId="6" pageNumber="27">20.1</td>
</tr>
<tr box="[839,1455,686,707]" gridrow="11" pageId="6" pageNumber="27">
<th box="[839,1129,686,707]" gridcol="0" gridrow="11" pageId="6" pageNumber="27">Pelvic-fin length</th>
<td box="[1150,1193,686,707]" gridcol="1" gridrow="11" pageId="6" pageNumber="27">18.3</td>
<td box="[1247,1272,686,707]" gridcol="2" gridrow="11" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,686,707]" gridcol="3" gridrow="11" pageId="6" pageNumber="27">14.1-20.0</td>
<td box="[1405,1455,686,707]" gridcol="4" gridrow="11" pageId="6" pageNumber="27">17.3</td>
</tr>
<tr box="[839,1455,715,736]" gridrow="12" pageId="6" pageNumber="27">
<th box="[839,1129,715,736]" gridcol="0" gridrow="12" pageId="6" pageNumber="27">Dorsal-fin base length</th>
<td box="[1150,1193,715,736]" gridcol="1" gridrow="12" pageId="6" pageNumber="27">14.7</td>
<td box="[1247,1272,715,736]" gridcol="2" gridrow="12" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,715,736]" gridcol="3" gridrow="12" pageId="6" pageNumber="27">12.4-16.4</td>
<td box="[1405,1455,715,736]" gridcol="4" gridrow="12" pageId="6" pageNumber="27">14.3</td>
</tr>
<tr box="[839,1455,744,765]" gridrow="13" pageId="6" pageNumber="27">
<th box="[839,1129,744,765]" gridcol="0" gridrow="13" pageId="6" pageNumber="27">Dorsal fin height</th>
<td box="[1150,1193,744,765]" gridcol="1" gridrow="13" pageId="6" pageNumber="27">28.7</td>
<td box="[1247,1272,744,765]" gridcol="2" gridrow="13" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,744,765]" gridcol="3" gridrow="13" pageId="6" pageNumber="27">23.7-31.5</td>
<td box="[1405,1455,744,765]" gridcol="4" gridrow="13" pageId="6" pageNumber="27">28.4</td>
</tr>
<tr box="[839,1455,773,794]" gridrow="14" pageId="6" pageNumber="27">
<th box="[839,1129,773,794]" gridcol="0" gridrow="14" pageId="6" pageNumber="27">Anal-fin base length</th>
<td box="[1150,1193,773,794]" gridcol="1" gridrow="14" pageId="6" pageNumber="27">27.5</td>
<td box="[1247,1272,773,794]" gridcol="2" gridrow="14" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,773,794]" gridcol="3" gridrow="14" pageId="6" pageNumber="27">23.1-29.3</td>
<td box="[1405,1455,773,794]" gridcol="4" gridrow="14" pageId="6" pageNumber="27">25.8</td>
</tr>
<tr box="[839,1455,802,823]" gridrow="15" pageId="6" pageNumber="27">
<th box="[839,1129,802,823]" gridcol="0" gridrow="15" pageId="6" pageNumber="27">Anal-fin lobe length</th>
<td box="[1150,1193,802,823]" gridcol="1" gridrow="15" pageId="6" pageNumber="27">19.5</td>
<td box="[1247,1272,802,823]" gridcol="2" gridrow="15" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,802,823]" gridcol="3" gridrow="15" pageId="6" pageNumber="27">14.2-22.5</td>
<td box="[1405,1455,802,823]" gridcol="4" gridrow="15" pageId="6" pageNumber="27">19.6</td>
</tr>
<tr box="[839,1455,830,851]" gridrow="16" pageId="6" pageNumber="27">
<th box="[839,1129,830,851]" gridcol="0" gridrow="16" pageId="6" pageNumber="27">Eye to dorsal-fin origin</th>
<td box="[1150,1193,830,851]" gridcol="1" gridrow="16" pageId="6" pageNumber="27">36.4</td>
<td box="[1247,1272,830,851]" gridcol="2" gridrow="16" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,830,851]" gridcol="3" gridrow="16" pageId="6" pageNumber="27">32.7-38.9</td>
<td box="[1405,1455,830,851]" gridcol="4" gridrow="16" pageId="6" pageNumber="27">35.8</td>
</tr>
<tr box="[839,1455,859,880]" gridrow="17" pageId="6" pageNumber="27">
<th box="[839,1129,859,880]" gridcol="0" gridrow="17" pageId="6" pageNumber="27">Dorsal-fin origin to caudal-fin</th>
<td box="[1150,1193,859,880]" gridcol="1" gridrow="17" pageId="6" pageNumber="27">51.9</td>
<td box="[1247,1272,859,880]" gridcol="2" gridrow="17" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,859,880]" gridcol="3" gridrow="17" pageId="6" pageNumber="27">47.8-55.5</td>
<td box="[1405,1455,859,880]" gridcol="4" gridrow="17" pageId="6" pageNumber="27">52.4</td>
</tr>
<tr box="[839,1455,888,909]" gridrow="18" pageId="6" pageNumber="27">
<th box="[839,1455,888,909]" colspan="5" colspanRight="4" gridcol="0" gridrow="18" pageId="6" pageNumber="27">base</th>
</tr>
<tr box="[839,1455,917,938]" gridrow="19" pageId="6" pageNumber="27">
<th box="[839,1129,917,938]" gridcol="0" gridrow="19" pageId="6" pageNumber="27">Bony head length</th>
<td box="[1150,1193,917,938]" gridcol="1" gridrow="19" pageId="6" pageNumber="27">34.3</td>
<td box="[1247,1272,917,938]" gridcol="2" gridrow="19" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,917,938]" gridcol="3" gridrow="19" pageId="6" pageNumber="27">30.4-37.6</td>
<td box="[1405,1455,917,938]" gridcol="4" gridrow="19" pageId="6" pageNumber="27">32.8</td>
</tr>
<tr box="[839,1455,945,966]" gridrow="20" pageId="6" pageNumber="27" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th box="[839,1129,945,966]" gridcol="0" gridrow="20" pageId="6" pageNumber="27">
<emphasis bold="true" box="[839,1089,945,966]" pageId="6" pageNumber="27">Percents of head length</emphasis>
</th>
</tr>
<tr box="[839,1455,974,995]" gridrow="21" pageId="6" pageNumber="27">
<th box="[839,1129,974,995]" gridcol="0" gridrow="21" pageId="6" pageNumber="27">Horizontal eye diameter</th>
<td box="[1150,1193,974,995]" gridcol="1" gridrow="21" pageId="6" pageNumber="27">35.4</td>
<td box="[1247,1272,974,995]" gridcol="2" gridrow="21" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,974,995]" gridcol="3" gridrow="21" pageId="6" pageNumber="27">35.2-44.4</td>
<td box="[1405,1455,974,995]" gridcol="4" gridrow="21" pageId="6" pageNumber="27">39.9</td>
</tr>
<tr box="[839,1455,1003,1024]" gridrow="22" pageId="6" pageNumber="27">
<th box="[839,1129,1003,1024]" gridcol="0" gridrow="22" pageId="6" pageNumber="27">Snout length</th>
<td box="[1150,1193,1003,1024]" gridcol="1" gridrow="22" pageId="6" pageNumber="27">19.7</td>
<td box="[1247,1272,1003,1024]" gridcol="2" gridrow="22" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,1003,1024]" gridcol="3" gridrow="22" pageId="6" pageNumber="27">19.7-27.2</td>
<td box="[1405,1455,1003,1024]" gridcol="4" gridrow="22" pageId="6" pageNumber="27">22.7</td>
</tr>
<tr box="[839,1455,1032,1053]" gridrow="23" pageId="6" pageNumber="27">
<th box="[839,1129,1032,1053]" gridcol="0" gridrow="23" pageId="6" pageNumber="27">Least interorbital width</th>
<td box="[1150,1193,1032,1053]" gridcol="1" gridrow="23" pageId="6" pageNumber="27">26.8</td>
<td box="[1247,1272,1032,1053]" gridcol="2" gridrow="23" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,1032,1053]" gridcol="3" gridrow="23" pageId="6" pageNumber="27">23.3-33.6</td>
<td box="[1405,1455,1032,1053]" gridcol="4" gridrow="23" pageId="6" pageNumber="27">27.7</td>
</tr>
<tr box="[839,1455,1061,1082]" gridrow="24" pageId="6" pageNumber="27">
<th box="[839,1129,1061,1082]" gridcol="0" gridrow="24" pageId="6" pageNumber="27">Upper jaw length</th>
<td box="[1150,1193,1061,1082]" gridcol="1" gridrow="24" pageId="6" pageNumber="27">45.2</td>
<td box="[1247,1272,1061,1082]" gridcol="2" gridrow="24" pageId="6" pageNumber="27">62</td>
<td box="[1293,1387,1061,1082]" gridcol="3" gridrow="24" pageId="6" pageNumber="27">33.9-51.3</td>
<td box="[1405,1455,1061,1082]" gridcol="4" gridrow="24" pageId="6" pageNumber="27">46.7</td>
</tr>
</table>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4566770" ID-Zenodo-Dep="4566770" httpUri="https://zenodo.org/record/4566770/files/figure.png" pageId="6" pageNumber="27" startId="6.[159,197,764,788]" targetBox="[206,758,177,710]" targetPageId="6">
<paragraph blockId="6.[159,804,764,819]" pageId="6" pageNumber="27">
<emphasis bold="true" box="[159,224,764,788]" pageId="6" pageNumber="27">Fig. 7.</emphasis>
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[229,463,764,787]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[229,463,764,787]" italics="true" pageId="6" pageNumber="27">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
, paratype, MZUSP 41406. Upper and lower jaws, lateral view, left side. Scale bar 1mm.
</paragraph>
</caption>
<paragraph blockId="6.[159,803,888,1232]" pageId="6" pageNumber="27">
<emphasis bold="true" box="[159,331,888,912]" pageId="6" pageNumber="27">Ecological notes.</emphasis>
At the ribeirão do Sapo(
<figureCitation box="[567,623,888,911]" captionStart="Fig" captionStartId="6.[159,198,1922,1946]" captionTargetBox="[159,802,1455,1896]" captionTargetId="figure-31@6.[159,803,1455,1896]" captionTargetPageId="6" captionText="Fig. 8. Type locality of Hyphessobrycon eilyos, ribeirão do Sapo, Mato Grosso, município de Alto Araguaia, Brazil." figureDoi="http://doi.org/10.5281/zenodo.4566772" httpUri="https://zenodo.org/record/4566772/files/figure.png" pageId="6" pageNumber="27">Fig. 8</figureCitation>
), the type-locality,
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[159,391,920,943]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[159,391,920,943]" italics="true" pageId="6" pageNumber="27">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
was only collected in backwaters of the stream. These backwaters were characteristically tea-colored, contrasting with the clear water of the mainstream, and were choked with vegetable debris from the adjacent riparian forest.Our limited collecting activity at the Córrego do Rancho did not allow us to ascertain what microhabitat was occupied by
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="eilyos">
<emphasis italics="true" pageId="6" pageNumber="27">Hyphessobrycon eilyos</emphasis>
</taxonomicName>
, but presumably the species is associated with large flooded areas with abundant aquatic vegetation present in that stream. For remarks on its syntopy with
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[497,625,1177,1200]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[497,625,1177,1200]" italics="true" pageId="6" pageNumber="27">H. langeanii</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="weitzmanorum">
<emphasis italics="true" pageId="6" pageNumber="27">H. weitzmanorum</emphasis>
</taxonomicName>
, see “Ecological notes” under
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[535,660,1209,1232]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="langeanii">
<emphasis box="[535,660,1209,1232]" italics="true" pageId="6" pageNumber="27">H. langeanii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="27" type="etymology">
<paragraph blockId="6.[159,802,1272,1392]" pageId="6" pageNumber="27">
<emphasis bold="true" box="[159,279,1272,1296]" pageId="6" pageNumber="27">Etymology.</emphasis>
From the Greek
<taxonomicName authorityName="Lima &amp; Moreira" authorityYear="2003" box="[442,499,1273,1296]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="eilyos">
<emphasis box="[442,499,1273,1296]" italics="true" pageId="6" pageNumber="27">eilyos</emphasis>
</taxonomicName>
, den, lurking-place, in allusion to the habitat (backwaters choked with vegetal matter) occupied by the new species (and according to local information, shared with anacondas,
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[336,525,1369,1392]" class="Reptilia" family="Boidae" genus="Eunectes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="27" phylum="Chordata" rank="species" species="murinus">
<emphasis box="[336,525,1369,1392]" italics="true" pageId="6" pageNumber="27">Eunectes murinus</emphasis>
</taxonomicName>
). A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>