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<document id="4E3B600EBE2EA29F808DC1569C4A7786" ID-CLB-Dataset="28520" ID-DOI="10.11646/zootaxa.4559.1.3" ID-GBIF-Dataset="becdb1cf-be93-48ca-abc4-898d0f84d0bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="2585159" ID-ZooBank="F051C9DE-CCB7-454E-8184-B6240049F004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550815179529" checkinUser="plazi" docAuthor="Hlaváč, Peter, Bregović, Petra &amp; Jalžić, Branko" docDate="2019" docId="5110AB180404FFC8FF562609A152AAC8" docLanguage="en" docName="zootaxa.4559.1.3.pdf" docOrigin="Zootaxa 4559 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumastocephalus troglavi Hlaváč &amp; Bregović &amp; Jalžić 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="98" masterDocId="AD29D3600403FFC0FFC12371A56DA903" masterDocTitle="Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)" masterLastPageNumber="110" masterPageNumber="90" pageNumber="97" updateTime="1698699937030" updateUser="ExternalLinkService">
<mods:mods id="A0E4F819EC979DEC21D6BBC69C39649A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C2B62B1103960BD79824762B9EAB4F24">
<mods:title id="10AB3194C720E5864EB91127EAB51696">Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="6CB93FAFF4D3FCD606495E116451963B" type="personal">
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<mods:namePart id="807C34461B7AD16A1B3924824208A4B8">Hlaváč, Peter</mods:namePart>
</mods:name>
<mods:name id="6D25661DA2980CFDCD3C9FD5414468A1" type="personal">
<mods:role id="BB63CE131011474F4E9D9E9B6D2F902A">
<mods:roleTerm id="C676ACC119B6DD7B9013E36E228AB268">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23EEBE2E6BF587643A26B3EB2355F84C">Bregović, Petra</mods:namePart>
</mods:name>
<mods:name id="43E7B6F7ED917F5B281A3CE3F69E7B54" type="personal">
<mods:role id="6B40A7C01B38E8B91DAF8FC1E3665EC2">
<mods:roleTerm id="CB5752515FA6F4B3CC0158A6BCE8A3B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8651C5F86A5438B9B9181AF267DDB0BD">Jalžić, Branko</mods:namePart>
</mods:name>
<mods:typeOfResource id="A209EB250BF3E638C4F0F84A99A64582">text</mods:typeOfResource>
<mods:relatedItem id="470E00420D201790D601C453D3611C02" type="host">
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<mods:title id="BBEBB5DE135D74900B4B92D090488041">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4E99DEFC148B19D8F329702EAF835603">
<mods:date id="DAEDE1DE57D6339E8F3E32303FDB15A3">2019</mods:date>
<mods:detail id="934A8AF737E52F75F9281D4A26CD69E3" type="pubDate">
<mods:number id="3D227887371344A42D6CEB6D567D21AA">2019-02-19</mods:number>
</mods:detail>
<mods:detail id="7D1B9960943D7965B69B3506605D7168" type="volume">
<mods:number id="EF931F159AA3B5FEB545B9C6419BD346">4559</mods:number>
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<mods:number id="9EA79EE9A5F38AF87660ED5E34799F3B">1</mods:number>
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<mods:classification id="A69E1F2F83272087950FE4E7602F6A76">journal article</mods:classification>
<mods:identifier id="3DB88BF75EB2882B9FCA2DA9282A332B" type="CLB-Dataset">28520</mods:identifier>
<mods:identifier id="11F606ED6B2166C394F2FCA2335FA77A" type="DOI">10.11646/zootaxa.4559.1.3</mods:identifier>
<mods:identifier id="BE3F656A02271E70A5FE71DA3AABA9A1" type="GBIF-Dataset">becdb1cf-be93-48ca-abc4-898d0f84d0bd</mods:identifier>
<mods:identifier id="3ECB2C963BC834F83B31D601BC5CC4F1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="34D204ED18E62ECA349E0D5EB044ABEE" type="Zenodo-Dep">2585159</mods:identifier>
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<treatment id="5110AB180404FFC8FF562609A152AAC8" ID-DOI="http://doi.org/10.5281/zenodo.5934390" ID-GBIF-Taxon="154007271" ID-Zenodo-Dep="5934390" LSID="urn:lsid:plazi:treatment:5110AB180404FFC8FF562609A152AAC8" httpUri="http://treatment.plazi.org/id/5110AB180404FFC8FF562609A152AAC8" lastPageId="8" lastPageNumber="98" pageId="7" pageNumber="97">
<subSubSection id="91A349850404FFC7FF562609A6E6AC91" box="[151,907,1400,1426]" pageId="7" pageNumber="97" type="nomenclature">
<paragraph id="D9061A0E0404FFC7FF562609A6E6AC91" blockId="7.[151,907,1400,1459]" box="[151,907,1400,1426]" pageId="7" pageNumber="97">
<heading id="824EAD620404FFC7FF562609A6E6AC91" bold="true" box="[151,907,1400,1426]" fontSize="11" level="1" pageId="7" pageNumber="97" reason="1">
<emphasis id="EBCDC61C0404FFC7FF562609A6E6AC91" bold="true" box="[151,907,1400,1426]" pageId="7" pageNumber="97">
<taxonomicName id="1EB9618D0404FFC7FF562609A656AC91" ID-CoL="8YBQS" authority="Hlavac, Bregovic &amp; Jalzic" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[151,827,1400,1426]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="97" phylum="Arthropoda" rank="species" species="troglavi" status="sp. nov.">
<emphasis id="EBCDC61C0404FFC7FF562609A482AC91" bold="true" box="[151,495,1400,1426]" italics="true" pageId="7" pageNumber="97">Thaumastocephalus troglavi</emphasis>
Hlaváč, Bregović &amp; Jalžić
</taxonomicName>
,
<taxonomicNameLabel id="F0FE7B670404FFC7FC892609A6E6AC91" box="[840,907,1400,1426]" pageId="7" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91A349850404FFC7FF5626EBA474ACB0" box="[151,281,1434,1459]" pageId="7" pageNumber="97" type="description">
<paragraph id="D9061A0E0404FFC7FF5626EBA474ACB0" blockId="7.[151,907,1400,1459]" box="[151,281,1434,1459]" pageId="7" pageNumber="97">
(
<figureCitation id="4182068B0404FFC7FF5E26EBA58AACB0" box="[159,231,1434,1459]" captionStart="FIGURES 5–9" captionStartId="5.[151,264,1906,1929]" captionTargetBox="[152,1435,193,1885]" captionTargetId="figure@5.[151,1436,193,1885]" captionTargetPageId="5" captionText="FIGURES 5–9. 5—Thaumatocephalus slavkoi sp. n., aedeagus, dorsal view (scale: 0.1 mm); 6—Thaumatocephalus troglavi sp. n., aedeagus, dorsal view (scale: 0.2 mm); Thaumatocephalus dahnae sp. n., 7—aedeagus, dorsal view (scale: 0.1 mm); 8— protibiae (scale: 0.1 mm); 9—maxillary palpomere (scale: 0.1 mm)." httpUri="https://zenodo.org/record/2585163/files/figure.png" pageId="7" pageNumber="97">Figs 6</figureCitation>
,
<figureCitation id="4182068B0404FFC7FF3226EBA47DACB0" box="[243,272,1434,1459]" captionStart="FIGURES 25–27" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[210,1382,193,1921]" captionTargetId="figure@15.[210,1382,193,1921]" captionTargetPageId="15" captionText="FIGURES 25–27. 25—entrance to Sistem Velika -Velika (cave), type locality (photo Petra Bregović); 26—Thaumatocephalus troglavi sp. n., habitus in situ (photo Petra Bregović); 27—topographical map of the type locality (the finding place of the T. troglavi sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585175/files/figure.png" pageId="7" pageNumber="97">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="91A349850404FFC7FF562693A07AACF8" box="[151,1303,1506,1531]" pageId="7" pageNumber="97" type="etymology">
<paragraph id="D9061A0E0404FFC7FF562693A07AACF8" blockId="7.[151,1437,1506,2035]" box="[151,1303,1506,1531]" pageId="7" pageNumber="97">
<emphasis id="EBCDC61C0404FFC7FF562693A473ACF8" bold="true" box="[151,286,1506,1531]" pageId="7" pageNumber="97">Etymology.</emphasis>
The species is named after the mountain peak Troglav near the cave Sistem Velika-Velika.
</paragraph>
</subSubSection>
<subSubSection id="91A349850404FFC7FF062577A646AF88" pageId="7" pageNumber="97" type="materials_examined">
<paragraph id="D9061A0E0404FFC7FF062577A646AF88" blockId="7.[151,1437,1506,2035]" pageId="7" pageNumber="97">
<emphasis id="EBCDC61C0404FFC7FF062577A64FAF1C" bold="true" box="[199,802,1542,1567]" pageId="7" pageNumber="97">
Material studied.
<materialsCitation id="69D110530404FFC7FE6F2577A64FAF1C" ID-GBIF-Occurrence="2005761077" box="[430,802,1542,1567]" collectionCode="VI" collectorName="Biokovo, M &amp; Sistem Velika-Velika" country="Croatia" location="CROATIA" pageId="7" pageNumber="97" specimenCount="1" typeStatus="holotype">
<typeStatus id="0602A4AC0404FFC7FE6F2577A73CAF1C" box="[430,593,1542,1567]" pageId="7" pageNumber="97" type="holotype">HOLOTYPE</typeStatus>
: Ƌ:
<collectingCountry id="A1AE5A9E0404FFC7FD572577A671AF1C" box="[662,796,1542,1567]" name="Croatia" pageId="7" pageNumber="97">CROATIA</collectingCountry>
:
</materialsCitation>
</emphasis>
<collectorName id="744C7FD80404FFC7FCF32577A6ABAF1D" box="[818,966,1542,1567]" pageId="7" pageNumber="97">Biokovo, M</collectorName>
. Troglav,
<collectorName id="744C7FD80404FFC7FB882577A053AF1C" box="[1097,1342,1542,1567]" pageId="7" pageNumber="97">Sistem Velika-Velika</collectorName>
(cave),
<date id="AD073CCE0404FFC7FF56255AA474AF40" box="[151,281,1579,1603]" pageId="7" pageNumber="97" value="2017-06-22">
22.VI
<materialsCitation id="69D110530404FFC7FF19255AA474AF40" ID-GBIF-Occurrence="2005761082" box="[216,281,1579,1603]" collectionCode="CNHM" collectorName="A. Kirin" country="Croatia" pageId="7" pageNumber="97" specimenCount="1" typeStatus="holotype">.2017</materialsCitation>
</date>
,
<collectorName id="744C7FD80404FFC7FEE4255AA4EEAF40" box="[293,387,1578,1603]" pageId="7" pageNumber="97">A. Kirin</collectorName>
lgt. (CNHM).
<materialsCitation id="69D110530404FFC7FDEF255AA64AAF88" ID-GBIF-Occurrence="2005761074" collectingDate="2016-01-23" collectingDateMax="2016-05-15" collectingDateMin="2016-01-23" collectionCode="CNHM, CPH" collectorName="V. Sudar &amp; M. Lukic" country="Croatia" county="Gradska" location="Other" municipality="Zupa" pageId="7" pageNumber="97" specimenCode="PARATYPES 1" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<specimenCode id="891FB2750404FFC7FDEF255AA7B6AF40" box="[558,731,1579,1603]" collectionCode="CNHM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="7" pageNumber="97" type="Museum">PARATYPES 1</specimenCode>
♂,
<specimenCount id="CFBFD1870404FFC7FCC3255AA645AF40" box="[770,808,1579,1603]" pageId="7" pageNumber="97" type="female">2♀</specimenCount>
: same data as holotype (
<collectionCode id="BFA882CB0404FFC7FBFF255AA1FBAF40" box="[1086,1174,1579,1603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="7" pageNumber="97" type="Museum">CNHM</collectionCode>
,
<collectionCode id="BFA882CB0404FFC7FB63255AA1B0AF40" box="[1186,1245,1579,1603]" country="USA" lsid="urn:lsid:biocol.org:col:15612" name="University of the Pacific" pageId="7" pageNumber="97" type="Herbarium">CPH</collectionCode>
).
<location id="DC664CD50404FFC7FB30255BA05CAF40" LSID="urn:lsid:plazi:treatment:5110AB180404FFC8FF562609A152AAC8:DC664CD50404FFC7FB30255BA05CAF40" box="[1265,1329,1578,1603]" country="Croatia" county="Gradska" municipality="Zupa" name="Other" pageId="7" pageNumber="97">Other</location>
material:
<specimenCount id="CFBFD1870404FFC7FF56253EA5D0AF64" box="[151,189,1615,1639]" pageId="7" pageNumber="97" type="male">1♂</specimenCount>
:
<collectingCountry id="A1AE5A9E0404FFC7FF0B253EA425AF64" box="[202,328,1615,1639]" name="Croatia" pageId="7" pageNumber="97">CROATIA</collectingCountry>
:
<location id="DC664CD50404FFC7FE95253FA4DAAF64" LSID="urn:lsid:plazi:treatment:5110AB180404FFC8FF562609A152AAC8:DC664CD50404FFC7FE95253FA4DAAF64" box="[340,439,1614,1639]" country="Croatia" county="Gradska" municipality="Zupa" name="Biokovo" pageId="7" pageNumber="97">Biokovo</location>
,
<collectingMunicipality id="396280740404FFC7FE05253FA76DAF64" box="[452,512,1614,1639]" pageId="7" pageNumber="97">Župa</collectingMunicipality>
,
<collectingCounty id="306762820404FFC7FDCC253FA707AF64" box="[525,618,1614,1639]" pageId="7" pageNumber="97">Gradska</collectingCounty>
spila (cave),
<date id="AD073CCE0404FFC7FCC4253FA617AF64" box="[773,890,1614,1639]" pageId="7" pageNumber="97" value="2016-05-15">
<collectingDate id="BD43C5260404FFC7FCC4253FA617AF64" box="[773,890,1614,1639]" pageId="7" pageNumber="97" value="2016-05-15">15.V.2016</collectingDate>
</date>
,
<collectorName id="744C7FD80404FFC7FC46253EA685AF64" box="[903,1000,1614,1639]" pageId="7" pageNumber="97">V. Sudar</collectorName>
lgt (
<collectionCode id="BFA882CB0404FFC7FBDE253EA131AF64" box="[1055,1116,1615,1639]" country="USA" lsid="urn:lsid:biocol.org:col:15612" name="University of the Pacific" pageId="7" pageNumber="97" type="Herbarium">CPH</collectionCode>
).
<specimenCount id="CFBFD1870404FFC7FBAE253EA1F9AF64" box="[1135,1172,1615,1639]" pageId="7" pageNumber="97" type="female">1♀</specimenCount>
: same data as previous
<typeStatus id="0602A4AC0404FFC7FF562502A59AAF88" box="[151,247,1651,1675]" pageId="7" pageNumber="97" type="paratype">paratype</typeStatus>
but collected on
<date id="AD073CCE0404FFC7FE792503A74EAF88" box="[440,547,1650,1675]" pageId="7" pageNumber="97" value="2016-01-23">
<collectingDate id="BD43C5260404FFC7FE792503A74EAF88" box="[440,547,1650,1675]" pageId="7" pageNumber="97" value="2016-01-23">23.I.2016</collectingDate>
</date>
by
<collectorName id="744C7FD80404FFC7FD8F2502A7D8AF88" box="[590,693,1650,1675]" pageId="7" pageNumber="97">M. Lukić</collectorName>
(
<collectionCode id="BFA882CB0404FFC7FD042502A64DAF88" box="[709,800,1651,1675]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="7" pageNumber="97" type="Museum">CNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91A349850404FFC7FF0625E7A0E0AE3D" pageId="7" pageNumber="97" type="discussion">
<paragraph id="D9061A0E0404FFC7FF0625E7A0E0AE3D" blockId="7.[151,1437,1506,2035]" pageId="7" pageNumber="97">
<emphasis id="EBCDC61C0404FFC7FF0625E7A456AFAC" bold="true" box="[199,315,1686,1711]" pageId="7" pageNumber="97">Remarks.</emphasis>
The specimens from Gradska spila are tentatively attributed to this species as the male has practically identical aedeagus, but due to the following reasons we did not include them in the
<typeStatus id="0602A4AC0404FFC7FB0925CAA195AFD0" box="[1224,1272,1723,1747]" pageId="7" pageNumber="97">type</typeStatus>
material od
<taxonomicName id="1EB9618D0404FFC7FA4625CDA582AFF4" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="97" phylum="Arthropoda" rank="species" species="troglavi">
<emphasis id="EBCDC61C0404FFC7FA4625CDA582AFF4" italics="true" pageId="7" pageNumber="97">T. troglavi</emphasis>
</taxonomicName>
: 1) ecological differences of habitat between Sistem Velika-Velika and Gradska spila (see details further in text), and 2) different fauna composition in the highest part of Biokovo Mountain (see Discussion). More male specimens from both caves will be needed to check the identity of specimens from Gradska spila with
<taxonomicName id="1EB9618D0404FFC7FAD22456A0EBAE3C" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[1299,1414,1830,1855]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="97" phylum="Arthropoda" rank="species" species="troglavi">
<emphasis id="EBCDC61C0404FFC7FAD22456A0EBAE3C" box="[1299,1414,1830,1855]" italics="true" pageId="7" pageNumber="97">T. troglavi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="91A349850404FFC8FF06243BA6F6A88B" lastPageId="8" lastPageNumber="98" pageId="7" pageNumber="97" type="description">
<paragraph id="D9061A0E0404FFC8FF06243BA63EA867" blockId="7.[151,1437,1506,2035]" lastBlockId="8.[151,1437,151,971]" lastPageId="8" lastPageNumber="98" pageId="7" pageNumber="97">
<emphasis id="EBCDC61C0404FFC7FF06243BA434AE60" bold="true" box="[199,345,1866,1891]" pageId="7" pageNumber="97">Description.</emphasis>
Body shiny (
<figureCitation id="4182068B0404FFC7FE37243BA721AE60" box="[502,588,1866,1891]" captionStart="FIGURES 25–27" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[210,1382,193,1921]" captionTargetId="figure@15.[210,1382,193,1921]" captionTargetPageId="15" captionText="FIGURES 25–27. 25—entrance to Sistem Velika -Velika (cave), type locality (photo Petra Bregović); 26—Thaumatocephalus troglavi sp. n., habitus in situ (photo Petra Bregović); 27—topographical map of the type locality (the finding place of the T. troglavi sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585175/files/figure.png" pageId="7" pageNumber="97">Fig. 26</figureCitation>
), reddish-brown, covered with dense, uneven setation, with long setae on posterior part of elytra; legs, antennae and maxillary palpi of the same colour, length
<quantity id="1E41B7EB0404FFC7FB92241EA1D4AE85" box="[1107,1209,1903,1927]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="7" pageNumber="97" unit="mm" value="2.07">2.07 mm</quantity>
, maximum width of elytra
<quantity id="1E41B7EB0404FFC7FF1F24E3A42BAEA9" box="[222,326,1938,1963]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="7" pageNumber="97" unit="mm" value="0.69">0.69 mm</quantity>
. Head 1.25 times as long as wide, 0.71 times narrower than pronotum, rostrum 2.14 times as wide as long. Antennae about
<quantity id="1E41B7EB0404FFC7FE6B24C6A760AECD" box="[426,525,1975,1999]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="97" unit="mm" value="0.87">0.87 mm</quantity>
long, scape 2.38 times as long as pedicel, antennomere III slightly longer than IV, IV–VII of same length, VIII slightly shorter than VII and as long as IX, X 1.25 times as long as IX, terminal antennomere 3.78 times as long as X, relative length of antennomeres: 1.00 / 0.42 / 0.26 / 0.24 / 0.24 / 0.24/ 0.24 / 0.21 / 0.21 / 0.24 / 0.89. Pronotum 1.07 as long as wide. Elytra 1.31 times as wide as long and 1.17 times as long as pronotum. Abdomen 1.37 times as long as elytra. Legs long, protibiae slightly modified, slightly depressed in apical half, median spine absent, mesotibiae simple, metatibiae simple. Aedeagus (
<figureCitation id="4182068B040BFFC8FBF02275A11EA81E" box="[1073,1139,260,285]" captionStart="FIGURES 5–9" captionStartId="5.[151,264,1906,1929]" captionTargetBox="[152,1435,193,1885]" captionTargetId="figure@5.[151,1436,193,1885]" captionTargetPageId="5" captionText="FIGURES 5–9. 5—Thaumatocephalus slavkoi sp. n., aedeagus, dorsal view (scale: 0.1 mm); 6—Thaumatocephalus troglavi sp. n., aedeagus, dorsal view (scale: 0.2 mm); Thaumatocephalus dahnae sp. n., 7—aedeagus, dorsal view (scale: 0.1 mm); 8— protibiae (scale: 0.1 mm); 9—maxillary palpomere (scale: 0.1 mm)." httpUri="https://zenodo.org/record/2585163/files/figure.png" pageId="8" pageNumber="98">Fig. 6</figureCitation>
)
<quantity id="1E41B7EB040BFFC8FB432275A188A81F" box="[1154,1253,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="8" pageNumber="98" unit="mm" value="0.59">0.59 mm</quantity>
long, with basal bulb about 1.5 times as long as apical part, right paramere smaller than left, both with three preapical setae, internal sac with two asymmetric structures, dorsal diaphragm present.
</paragraph>
<paragraph id="D9061A0E040BFFC8FF06221EA6F6A88B" blockId="8.[151,1437,151,971]" box="[199,923,367,392]" pageId="8" pageNumber="98">
<emphasis id="EBCDC61C040BFFC8FF06221EA4DAA88B" bold="true" box="[199,439,367,392]" pageId="8" pageNumber="98">Sexual dimorphism.</emphasis>
Females lacking modification of protibiae.
</paragraph>
</subSubSection>
<subSubSection id="91A34985040BFFC8FF0622E5A6D0A8CC" pageId="8" pageNumber="98" type="diagnosis">
<paragraph id="D9061A0E040BFFC8FF0622E5A6D0A8CC" blockId="8.[151,1437,151,971]" pageId="8" pageNumber="98">
<emphasis id="EBCDC61C040BFFC8FF0622E5A4BFA8AE" bold="true" box="[199,466,404,429]" pageId="8" pageNumber="98">Differential diagnosis.</emphasis>
<taxonomicName id="1EB9618D040BFFC8FE1F22E4A73AA8AE" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[478,599,404,429]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="troglavi">
<emphasis id="EBCDC61C040BFFC8FE1F22E4A73AA8AE" box="[478,599,404,429]" italics="true" pageId="8" pageNumber="98">T. troglavi</emphasis>
</taxonomicName>
is readily separated from all other known species of the genus by: 1) protibiae at most with minuscule tooth and 2) body length over
<quantity id="1E41B7EB040BFFC8FCA122C6A6D4A8CC" box="[864,953,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="98" unit="mm" value="2.5">2.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="91A34985040BFFC8FF0622ADA152AAC8" pageId="8" pageNumber="98" type="distribution">
<paragraph id="D9061A0E040BFFC8FF0622ADA152AAC8" blockId="8.[151,1437,151,971]" pageId="8" pageNumber="98">
<emphasis id="EBCDC61C040BFFC8FF0622ADA499A8F6" bold="true" box="[199,500,476,501]" pageId="8" pageNumber="98">Distribution and habitat.</emphasis>
<taxonomicName id="1EB9618D040BFFC8FE3D22ACA703A8F6" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[508,622,476,501]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="troglavi">
<emphasis id="EBCDC61C040BFFC8FE3D22ACA703A8F6" box="[508,622,476,501]" italics="true" pageId="8" pageNumber="98">T. troglavi</emphasis>
</taxonomicName>
is known from the cave system Velika-Velika (
<figureCitation id="4182068B040BFFC8FB4F22ADA07FA8F7" box="[1166,1298,476,501]" captionStart="FIGURES 25–27" captionStartId="15.[151,264,1942,1965]" captionTargetBox="[210,1382,193,1921]" captionTargetId="figure@15.[210,1382,193,1921]" captionTargetPageId="15" captionText="FIGURES 25–27. 25—entrance to Sistem Velika -Velika (cave), type locality (photo Petra Bregović); 26—Thaumatocephalus troglavi sp. n., habitus in situ (photo Petra Bregović); 27—topographical map of the type locality (the finding place of the T. troglavi sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585175/files/figure.png" pageId="8" pageNumber="98">Figs 25, 27</figureCitation>
), situated at
<quantity id="1E41B7EB040BFFC8FF56228EA580AB14" box="[151,237,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.375" pageId="8" pageNumber="98" unit="m" value="1375.0">1375 m</quantity>
a.s.l. near the peak M. Troglav on Biokovo Mountain (
<figureCitation id="4182068B040BFFC8FCA4228EA6D4AB1B" box="[869,953,511,536]" captionStart="FIGURE 30" captionStartId="16.[151,250,1943,1965]" captionTargetBox="[150,1436,846,1922]" captionTargetId="figure@16.[150,1436,846,1922]" captionTargetPageId="16" captionText="FIGURE 30. Distribution map of the species of the genus Thaumastocephalus mentioned in this work." httpUri="https://zenodo.org/record/2585179/files/figure.png" pageId="8" pageNumber="98">Fig. 30</figureCitation>
, dark blue circle). The cave is
<quantity id="1E41B7EB040BFFC8FADB228EA00FAB14" box="[1306,1378,511,536]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="98" unit="m" value="135.0">135 m</quantity>
long and
<quantity id="1E41B7EB040BFFC8FF062156A46FAB3C" box="[199,258,551,575]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="98" unit="m" value="48.0">48 m</quantity>
deep, and it was discovered during 1
<superScript id="2ECCB746040BFFC8FD6B2152A7D9AB32" attach="left" box="[682,692,547,561]" fontSize="6" pageId="8" pageNumber="98">st</superScript>
Biospeleological Expedition—Biokovo 2017 (
<bibRefCitation id="BD2867FF040BFFC8FB062157A0FCAB3C" author="Sudar, V. &amp; Kuharic, N. &amp; Bregovic, P. &amp; Kirin, A." box="[1223,1425,550,575]" pageId="8" pageNumber="98" pagination="2 - 20" refId="ref11954" refString="Sudar, V., Kuharic, N., Bregovic, P. &amp; Kirin, A. (2017) Prva biospeleoloska ekspedicija-Biokovo 2017 (1 st Biospeleological Expedition-Biokovo 2017). Subterranea Croatica, 23, 2 - 20." type="journal article" year="2017">
Sudar
<emphasis id="EBCDC61C040BFFC8FAD32159A028AB3C" box="[1298,1349,550,575]" italics="true" pageId="8" pageNumber="98">et al</emphasis>
. 2017
</bibRefCitation>
). The air temperature measured in the cave on
<date id="AD073CCE040BFFC8FD5F213AA656AB60" box="[670,827,587,611]" pageId="8" pageNumber="98" value="2017-06-01">1st June 2017</date>
was 2.2 °C and the relative humidity was 99 %. The following fauna is also recorded in the cave: Diplopoda—
<taxonomicName id="1EB9618D040BFFC8FC85211FA1AAAB84" authority="Mrsic, 1992" authorityName="Mrsic" authorityYear="1992" box="[836,1223,622,647]" class="Diplopoda" family="Biokoviellidae" genus="Biokoviella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="mauriesi">
<emphasis id="EBCDC61C040BFFC8FC85211FA15DAB84" box="[836,1072,622,647]" italics="true" pageId="8" pageNumber="98">Biokoviella mauriesi</emphasis>
Mršić, 1992
</taxonomicName>
(det. T. Dražina); Coleoptera—
<taxonomicName id="1EB9618D040BFFC8FEEC21E5A4F6ABA8" box="[301,411,660,683]" class="Insecta" family="Staphylinidae" genus="Pygoxyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C040BFFC8FEEC21E5A4F6ABA8" box="[301,411,660,683]" italics="true" pageId="8" pageNumber="98">Pygoxyon</emphasis>
</taxonomicName>
sp. (det. P. Hlaváč) and
<taxonomicName id="1EB9618D040BFFC8FD7021E3A1A5ABA8" authority="Muller, 1934" authorityName="Muller" authorityYear="1934" box="[689,1224,658,683]" class="Insecta" family="Leiodidae" genus="Speoplanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="subSpecies" species="giganteus" subSpecies="biocovensis">
<emphasis id="EBCDC61C040BFFC8FD7021E3A142ABA8" box="[689,1071,658,683]" italics="true" pageId="8" pageNumber="98">Speoplanes giganteus biocovensis</emphasis>
Müller, 1934
</taxonomicName>
(det. P. Bregović). The second possible location of
<taxonomicName id="1EB9618D040BFFC8FDC521C9A715ABCC" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[516,632,694,719]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="troglavi">
<emphasis id="EBCDC61C040BFFC8FDC521C9A77CABCC" box="[516,529,696,719]" italics="true" pageId="8" pageNumber="98">T</emphasis>
.
<emphasis id="EBCDC61C040BFFC8FDE021C7A715ABCC" box="[545,632,694,719]" italics="true" pageId="8" pageNumber="98">troglavi</emphasis>
</taxonomicName>
is the cave Gradska spila (see above Remarks) situated at
<quantity id="1E41B7EB040BFFC8FAD021C7A034ABCC" box="[1297,1369,694,719]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="98" unit="m" value="750.0">750 m</quantity>
a.s.l., near the village Župa, on the slopes of Biokovo Mountain (
<figureCitation id="4182068B040BFFC8FCE621ABA615ABF0" box="[807,888,730,755]" captionStart="FIGURE 30" captionStartId="16.[151,250,1943,1965]" captionTargetBox="[150,1436,846,1922]" captionTargetId="figure@16.[150,1436,846,1922]" captionTargetPageId="16" captionText="FIGURE 30. Distribution map of the species of the genus Thaumastocephalus mentioned in this work." httpUri="https://zenodo.org/record/2585179/files/figure.png" pageId="8" pageNumber="98">Fig. 30</figureCitation>
, light blue circle). The cave is small,
<quantity id="1E41B7EB040BFFC8FAD621ABA009ABF1" box="[1303,1380,730,755]" metricMagnitude="1" metricUnit="m" metricValue="2.65" pageId="8" pageNumber="98" unit="m" value="26.5">26.5 m</quantity>
long and
<quantity id="1E41B7EB040BFFC8FF07218EA468AA14" box="[198,261,767,791]" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="98" unit="m" value="1.8">1.8 m</quantity>
deep. The air temperature measured in the cave on
<date id="AD073CCE040BFFC8FC84218FA69DAA14" box="[837,1008,766,792]" pageId="8" pageNumber="98" value="2016-05-15">15th May 2016</date>
was 10.1 °C and the relative humidity was 97.9 %. The following fauna is also recorded from the cave: Isopoda—
<taxonomicName id="1EB9618D040BFFC8FC352053A11AAA38" authorityName="Racovitza" authorityYear="1908" box="[1012,1143,802,827]" class="Malacostraca" family="Trichoniscidae" genus="Alpioniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C040BFFC8FC352053A11AAA38" box="[1012,1143,802,827]" italics="true" pageId="8" pageNumber="98">Alpioniscus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="1EB9618D040BFFC8FB6F2053A438AA5C" authority="(Strouhal, 1938)" authorityName="Strouhal" authorityYear="1938" baseAuthorityName="Strouhal" baseAuthorityYear="1938" class="Malacostraca" family="Trichoniscidae" genus="Alpioniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="verheoffi">
<emphasis id="EBCDC61C040BFFC8FB6F2053A0F1AA38" box="[1198,1436,802,827]" italics="true" pageId="8" pageNumber="98">Alpioniscus verheoffi</emphasis>
(Strouhal, 1938)
</taxonomicName>
(det. J. Bedek) and
<taxonomicName id="1EB9618D040BFFC8FD9B2036A162AA5C" authority="Strouhal, 1937" authorityName="Strouhal" authorityYear="1937" box="[602,1039,838,863]" class="Malacostraca" family="Oniscidae" genus="Oroniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="dalmaticus">
<emphasis id="EBCDC61C040BFFC8FD9B2036A638AA5C" box="[602,853,838,863]" italics="true" pageId="8" pageNumber="98">Oroniscus dalmaticus</emphasis>
Strouhal, 1937
</taxonomicName>
as noted by
<bibRefCitation id="BD2867FF040BFFC8FB7B2037A0F5AA5C" author="Bedek, J. &amp; Taiti, S. &amp; Gottstein, S." box="[1210,1432,838,863]" pageId="8" pageNumber="98" pagination="237 - 354" refId="ref9760" refString="Bedek, J., Taiti, S. &amp; Gottstein, S. (2011) Catalogue and atlas of cave-dwelling terrestrial isopods (Crustacea: Oniscidea) from Croatia. Natura Croatica, 20 (2), 237 - 354." type="journal article" year="2011">
Bedek
<emphasis id="EBCDC61C040BFFC8FAD12039A02AAA5C" box="[1296,1351,838,863]" italics="true" pageId="8" pageNumber="98">et al</emphasis>
. 2011
</bibRefCitation>
; Collembola—
<taxonomicName id="1EB9618D040BFFC8FEF4201DA787AA80" authority="Templeton, 1835" authorityName="Templeton" authorityYear="1835" box="[309,746,874,899]" class="Collembola" family="Entomobryidae" genus="Heteromurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Entomobryomorpha" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="EBCDC61C040BFFC8FEF4201DA773AA80" box="[309,542,874,899]" italics="true" pageId="8" pageNumber="98">Heteromurus nitidus</emphasis>
Templeton, 1835
</taxonomicName>
,
<taxonomicName id="1EB9618D040BFFC8FD36201BA61CAA80" box="[759,881,874,899]" class="Insecta" family="Entomobryidae" genus="Orchesella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C040BFFC8FD36201BA61CAA80" box="[759,881,874,899]" italics="true" pageId="8" pageNumber="98">Orchesella</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="1EB9618D040BFFC8FC6B201BA155AA80" box="[938,1080,874,899]" class="Insecta" family="Orchesellidae" genus="Verhoeffiella" higherTaxonomySource="CoL" kingdom="Animalia" order="Dermaptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C040BFFC8FC6B201BA155AA80" box="[938,1080,874,899]" italics="true" pageId="8" pageNumber="98">Verhoeffiella</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="1EB9618D040BFFC8FB5C201DA471AAA4" authority="Stach, 1922" authorityName="Stach" authorityYear="1922" class="Insecta" family="Tomoceridae" genus="Tomocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="terrestralis">
<emphasis id="EBCDC61C040BFFC8FB5C201DA0F1AA80" box="[1181,1436,874,899]" italics="true" pageId="8" pageNumber="98">Tomocerus terrestralis</emphasis>
Stach, 1922
</taxonomicName>
(
<bibRefCitation id="BD2867FF040BFFC8FEEC20FFA735AAA4" author="Lukic, M. &amp; Deharveng, L." box="[301,600,910,935]" pageId="8" pageNumber="98" pagination="119 - 136" refId="ref11025" refString="Lukic, M. &amp; Deharveng, L. (2008) Biodiversity and distribution of cave Collembola (Hexapoda) on Biokovo Mt., Croatia. In: Pavicevic, D. &amp; Perreau, M. (Eds.), Advances in the Studies of the Fauna of the Balkan Peninsula-Papers Dedicated to the Memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, pp. 119 - 136." type="book chapter" year="2008">Lukić &amp; Deharveng 2008</bibRefCitation>
); Coleoptera—
<taxonomicName id="1EB9618D040BFFC8FCC420FFA1B8AAA4" authority="(L. Miller, 1861)" authorityName="L. Miller" authorityYear="1861" baseAuthorityName="L. Miller" baseAuthorityYear="1861" box="[773,1237,910,935]" class="Insecta" family="Carabidae" genus="Neotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="dalmatinus">
<emphasis id="EBCDC61C040BFFC8FCC420FFA165AAA4" box="[773,1032,910,935]" italics="true" pageId="8" pageNumber="98">Neotrechus dalmatinus</emphasis>
(L. Miller, 1861)
</taxonomicName>
,
<taxonomicName id="1EB9618D040BFFC8FB2320FEA01EAAA5" authorityName="Jeannel" authorityYear="1910" box="[1250,1395,911,934]" class="Insecta" family="Leiodidae" genus="Speonesiotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C040BFFC8FB2320FEA01EAAA5" box="[1250,1395,911,934]" italics="true" pageId="8" pageNumber="98">Speonesiotes</emphasis>
</taxonomicName>
sp. and
<taxonomicName id="1EB9618D040BFFC8FF0720C3A601AAC8" authority="Hlavac, Lakota &amp; Ceplik, 2016" authorityName="Hlavac, Lakota &amp; Ceplik" authorityYear="2016" box="[198,876,946,971]" class="Insecta" family="Carabidae" genus="Speluncarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="98" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis id="EBCDC61C040BFFC8FF0720C3A435AAC8" box="[198,344,946,971]" italics="true" pageId="8" pageNumber="98">Speluncarius</emphasis>
cf.
<emphasis id="EBCDC61C040BFFC8FE4220C3A76BAAC8" box="[387,518,946,971]" italics="true" pageId="8" pageNumber="98">biokovensis</emphasis>
<bibRefCitation id="BD2867FF040BFFC8FDCC20C3A601AAC8" author="Hlavac, P. &amp; Lakota, J. &amp; Ceplik, D." box="[525,876,946,971]" pageId="8" pageNumber="98" pagination="137 - 140" refId="ref10527" refString="Hlavac, P., Lakota, J. &amp; Ceplik, D. (2016) Endogean and cavernicolous Coleoptera of the Balkans. XVII. A new species of the genus Speluncarius Reitter, 1886 (Coleoptera, Carabidae, Harpalinae, Pterostichini) from Croatia. Zootaxa, 4161 (1), 137 - 140. https: // doi. org / 10.11646 / zootaxa. 4161.1.10" type="journal article" year="2016">Hlaváč, Lakota &amp; Čeplík, 2016</bibRefCitation>
</taxonomicName>
(det. P. Bregović).
</paragraph>
</subSubSection>
</treatment>
</document>
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