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<document id="1D299DEE78BF43EAC07987D1405333B3" ID-CLB-Dataset="28520" ID-DOI="10.11646/zootaxa.4559.1.3" ID-GBIF-Dataset="becdb1cf-be93-48ca-abc4-898d0f84d0bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="2585159" ID-ZooBank="F051C9DE-CCB7-454E-8184-B6240049F004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550815179529" checkinUser="plazi" docAuthor="Hlaváč, Peter, Bregović, Petra &amp; Jalžić, Branko" docDate="2019" docId="5110AB180402FFC2FF56270DA40EA867" docLanguage="en" docName="zootaxa.4559.1.3.pdf" docOrigin="Zootaxa 4559 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumastocephalus bilandzijae Hlaváč &amp; Bregović &amp; Jalžić 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="91" masterDocId="AD29D3600403FFC0FFC12371A56DA903" masterDocTitle="Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)" masterLastPageNumber="110" masterPageNumber="90" pageNumber="91" updateTime="1698699937030" updateUser="ExternalLinkService">
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<mods:title id="98BA8CF2F7AF77193AFFA6E101184720">Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="6778B0FB1D24589756B1ABECF68D2739">Hlaváč, Peter</mods:namePart>
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<mods:namePart id="C72C8DDD371F50492DED5FB9A6A36F8D">Bregović, Petra</mods:namePart>
</mods:name>
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<mods:namePart id="1EE0000AEA479AAFD4463478CE83654D">Jalžić, Branko</mods:namePart>
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<mods:title id="24F7AECB7051B14AB431DE6AD0C126D7">Zootaxa</mods:title>
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<mods:date id="1FDFCEF7BBDBEFA25CD194FA62E0A578">2019</mods:date>
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<mods:number id="729638F8EB5A5D7145B600137EDE01E5">2019-02-19</mods:number>
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<mods:number id="849D921826446EA9276102AA1D1B1E1D">4559</mods:number>
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<mods:identifier id="CE9D7D5E8C455BCD6979ED5E80B0C812" type="DOI">10.11646/zootaxa.4559.1.3</mods:identifier>
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<mods:identifier id="41291BADF0882DBE5C1E73606195D0BB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5110AB180402FFC2FF56270DA40EA867" ID-DOI="http://doi.org/10.5281/zenodo.5934380" ID-GBIF-Taxon="154007273" ID-Zenodo-Dep="5934380" LSID="urn:lsid:plazi:treatment:5110AB180402FFC2FF56270DA40EA867" httpUri="http://treatment.plazi.org/id/5110AB180402FFC2FF56270DA40EA867" lastPageId="2" lastPageNumber="91" pageId="1" pageNumber="91">
<subSubSection id="91A349850402FFC1FF56270DA6DFAD95" box="[151,946,1148,1174]" pageId="1" pageNumber="91" type="nomenclature">
<paragraph id="D9061A0E0402FFC1FF56270DA6DFAD95" blockId="1.[151,946,1148,1207]" box="[151,946,1148,1174]" pageId="1" pageNumber="91">
<heading id="824EAD620402FFC1FF56270DA6DFAD95" bold="true" box="[151,946,1148,1174]" fontSize="11" level="1" pageId="1" pageNumber="91" reason="1">
<emphasis id="EBCDC61C0402FFC1FF56270DA6DFAD95" bold="true" box="[151,946,1148,1174]" pageId="1" pageNumber="91">
<taxonomicName id="1EB9618D0402FFC1FF56270DA60EAD95" ID-CoL="8YBQL" authority="Hlavac, Bregovic &amp; Jalzic" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[151,867,1148,1174]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="bilandzijae" status="sp. nov.">
<emphasis id="EBCDC61C0402FFC1FF56270DA77BAD95" bold="true" box="[151,534,1148,1174]" italics="true" pageId="1" pageNumber="91">Thaumastocephalus bilandzijae</emphasis>
Hlaváč, Bregović &amp; Jalžić
</taxonomicName>
,
<taxonomicNameLabel id="F0FE7B670402FFC1FCAE270DA6DFAD95" box="[879,946,1148,1174]" pageId="1" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91A349850402FFC1FF5627EFA475ADB4" box="[151,280,1182,1207]" pageId="1" pageNumber="91" type="description">
<paragraph id="D9061A0E0402FFC1FF5627EFA475ADB4" blockId="1.[151,946,1148,1207]" box="[151,280,1182,1207]" pageId="1" pageNumber="91">
(
<figureCitation id="4182068B0402FFC1FF5E27EFA588ADB4" box="[159,229,1182,1207]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[270,1322,195,1885]" captionTargetId="figure@4.[269,1322,195,1885]" captionTargetPageId="4" captionText="FIGURES 1–4. 1—Thaumatocephalus bilandzijae sp. n., aedeagus, dorsal view (scale: 0.1 mm); 2—Thaumatocephalus kirini sp. n., aedeagus, dorsal view (scale: 0.2 mm); 3—Thaumatocephalus marsici sp. n., aedeagus, dorsal view (scale: 0.2 mm); 4—Thaumatocephalus rujnicensis sp. n., aedeagus, dorsal view (scale: 0.1 mm)." httpUri="https://zenodo.org/record/2585161/files/figure.png" pageId="1" pageNumber="91">Figs 1</figureCitation>
,
<figureCitation id="4182068B0402FFC1FF3227EEA463ADB4" box="[243,270,1183,1207]" captionStart="FIGURES 10–12" captionStartId="6.[151,264,1792,1815]" captionTargetBox="[185,1399,411,1792]" captionTargetId="figure@6.[164,1423,411,1771]" captionTargetPageId="6" captionText="FIGURES 10–12. 10—entrance to Peć u Čulinovim raljevinama (cave), type locality (photo Marko Lukić); 11— Thaumatocephalus bilandzijae sp. n., habitus in situ (photo Alen Kirin); 12—topographical map of the type locality (the finding place of the T. bilandzijae sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585165/files/figure.png" pageId="1" pageNumber="91">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="91A349850402FFC1FF562797A6A3AC20" pageId="1" pageNumber="91" type="etymology">
<paragraph id="D9061A0E0402FFC1FF562797A6A3AC20" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="91">
<emphasis id="EBCDC61C0402FFC1FF562797A473ADFC" bold="true" box="[151,286,1254,1279]" pageId="1" pageNumber="91">Etymology.</emphasis>
The species is named after biospeleologist Helena Bilandžija, a member of the Croatian Biospeleological Society (CBSS) and collector of the
<typeStatus id="0602A4AC0402FFC1FD32267BA638AC20" box="[755,853,1290,1315]" pageId="1" pageNumber="91" type="holotype">holotype</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="91A349850402FFC1FF06265FA4BCAC8C" pageId="1" pageNumber="91" type="materials_examined">
<paragraph id="D9061A0E0402FFC1FF06265FA4BCAC8C" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="91">
<emphasis id="EBCDC61C0402FFC1FF06265FA661AC44" bold="true" box="[199,780,1326,1351]" pageId="1" pageNumber="91">
Material studied.
<materialsCitation id="69D110530402FFC1FE62265FA661AC44" ID-GBIF-Occurrence="2005761081" box="[419,780,1326,1351]" collectingDate="2009-01-06" collectionCode="CNHM" collectorName="H. Bilandzija" country="Croatia" county="Culinovim" location="Dugopolje" municipality="Pec" pageId="1" pageNumber="91" specimenCount="1" typeStatus="holotype">
<typeStatus id="0602A4AC0402FFC1FE62265FA72BAC44" box="[419,582,1326,1351]" pageId="1" pageNumber="91" type="holotype">HOLOTYPE</typeStatus>
: Ƌ:
<collectingCountry id="A1AE5A9E0402FFC1FD41265FA665AC44" box="[640,776,1326,1351]" name="Croatia" pageId="1" pageNumber="91">CROATIA</collectingCountry>
:
</materialsCitation>
</emphasis>
<location id="DC664CD50402FFC1FCD6265FA6FDAC44" LSID="urn:lsid:plazi:treatment:5110AB180402FFC2FF56270DA40EA867:DC664CD50402FFC1FCD6265FA6FDAC44" box="[791,912,1326,1351]" country="Croatia" county="Culinovim" municipality="Pec" name="Dugopolje" pageId="1" pageNumber="91">Dugopolje</location>
,
<location id="DC664CD50402FFC1FC61265FA68CAC44" LSID="urn:lsid:plazi:treatment:5110AB180402FFC2FF56270DA40EA867:DC664CD50402FFC1FC61265FA68CAC44" box="[928,993,1326,1351]" country="Croatia" county="Culinovim" municipality="Pec" name="Kolic" pageId="1" pageNumber="91">Kolić</location>
,
<collectingMunicipality id="396280740402FFC1FC31265EA175AC44" box="[1008,1048,1327,1351]" pageId="1" pageNumber="91">Peć</collectingMunicipality>
u
<collectingCounty id="306762820402FFC1FBFC265FA1D8AC44" box="[1085,1205,1326,1351]" pageId="1" pageNumber="91">Čulinovim</collectingCounty>
raljevinama (cave),
<date id="AD073CCE0402FFC1FF562623A59BAC68" box="[151,246,1362,1387]" pageId="1" pageNumber="91" value="2009-01-06">
<collectingDate id="BD43C5260402FFC1FF562623A59BAC68" box="[151,246,1362,1387]" pageId="1" pageNumber="91" value="2009-01-06">6.I.2009</collectingDate>
</date>
,
<collectorName id="744C7FD80402FFC1FEC32622A4FAAC68" box="[258,407,1362,1387]" pageId="1" pageNumber="91">H. Bilandžija</collectorName>
lgt. (CNHM).
<materialsCitation id="69D110530402FFC1FDFE2622A4A0AC8C" ID-GBIF-Occurrence="2005761079" collectingDate="2016-11-05" collectionCode="CNHM, CPH" collectorName="A. Kirin" country="Croatia" county="Culinovim" location="Dugopolje" municipality="Pec" pageId="1" pageNumber="91" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="0602A4AC0402FFC1FDFE2622A7B4AC68" box="[575,729,1363,1387]" pageId="1" pageNumber="91" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="CFBFD1870402FFC1FD2A2622A662AC68" box="[747,783,1363,1387]" pageId="1" pageNumber="91" type="female">2♀</specimenCount>
): same locality as the holotype but collected on
<date id="AD073CCE0402FFC1FAEA2623A0F1AC68" box="[1323,1436,1362,1387]" pageId="1" pageNumber="91" value="2016-11-05">
<collectingDate id="BD43C5260402FFC1FAEA2623A0F1AC68" box="[1323,1436,1362,1387]" pageId="1" pageNumber="91" value="2016-11-05">5.XI.2016</collectingDate>
</date>
by
<collectorName id="744C7FD80402FFC1FF7B2606A47AAC8C" box="[186,279,1398,1423]" pageId="1" pageNumber="91">A. Kirin</collectorName>
(
<collectionCode id="BFA882CB0402FFC1FEE72606A412AC8C" box="[294,383,1399,1423]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="1" pageNumber="91" type="Museum">CNHM</collectionCode>
,
<collectionCode id="BFA882CB0402FFC1FE482606A4A8AC8C" box="[393,453,1399,1423]" country="USA" lsid="urn:lsid:biocol.org:col:15612" name="University of the Pacific" pageId="1" pageNumber="91" type="Herbarium">CPH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91A349850402FFC1FF0626EBA17BAE3C" pageId="1" pageNumber="91" type="description">
<paragraph id="D9061A0E0402FFC1FF0626EBA7D3AE19" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="91">
<emphasis id="EBCDC61C0402FFC1FF0626EBA434ACB0" bold="true" box="[199,345,1434,1459]" pageId="1" pageNumber="91">Description.</emphasis>
Body shiny (
<figureCitation id="4182068B0402FFC1FE3F26EBA738ACB0" box="[510,597,1434,1459]" captionStart="FIGURES 10–12" captionStartId="6.[151,264,1792,1815]" captionTargetBox="[185,1399,411,1792]" captionTargetId="figure@6.[164,1423,411,1771]" captionTargetPageId="6" captionText="FIGURES 10–12. 10—entrance to Peć u Čulinovim raljevinama (cave), type locality (photo Marko Lukić); 11— Thaumatocephalus bilandzijae sp. n., habitus in situ (photo Alen Kirin); 12—topographical map of the type locality (the finding place of the T. bilandzijae sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585165/files/figure.png" pageId="1" pageNumber="91">Fig. 11</figureCitation>
), reddish-brown, covered with very sparse, uneven, golden setae; legs, antennae and maxillary palpi of same colour, length
<quantity id="1E41B7EB0402FFC1FD2126CEA62BACD4" box="[736,838,1471,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="1" pageNumber="91" unit="mm" value="1.82">1.82 mm</quantity>
, maximum width of elytra
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. Head 1.22 times as long as wide, 0.84 times narrower than pronotum, rostrum 2.17 times as wide as long. Antennae about
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long, scape twice as long as pedicel, antennomeres III–VII of same length, VIII slightly shorter than VII and as long as IX, antennomere X about as long as VII, terminal antennomere four times as long as X, relative length of antennomeres: 1.00 / 0.50 / 0.33 / 0.33 / 0.33 / 0.33 / 0.25 / 0.25 / 0.30 / 1.17. Pronotum 1.16 times as long as wide. Elytra 1.32 times as wide as long and 1.28 times as long as pronotum. Abdomen 1.31 times as long as elytra. Legs long, protibiae modified, depressed in apical half, with median, well-defined sharp spine, mesotibiae narrowed and slightly bent at apex, metatibiae simple. Aedeagus (
<figureCitation id="4182068B0402FFC1FD1B25CBA676AFD0" box="[730,795,1722,1747]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[270,1322,195,1885]" captionTargetId="figure@4.[269,1322,195,1885]" captionTargetPageId="4" captionText="FIGURES 1–4. 1—Thaumatocephalus bilandzijae sp. n., aedeagus, dorsal view (scale: 0.1 mm); 2—Thaumatocephalus kirini sp. n., aedeagus, dorsal view (scale: 0.2 mm); 3—Thaumatocephalus marsici sp. n., aedeagus, dorsal view (scale: 0.2 mm); 4—Thaumatocephalus rujnicensis sp. n., aedeagus, dorsal view (scale: 0.1 mm)." httpUri="https://zenodo.org/record/2585161/files/figure.png" pageId="1" pageNumber="91">Fig. 1</figureCitation>
)
<quantity id="1E41B7EB0402FFC1FCEC25CAA6FCAFD1" box="[813,913,1723,1747]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="91" unit="mm" value="0.33">0.33 mm</quantity>
long, with basal bulb about 1.54 times as long as apical part, right paramere more robust than left, both with three preapical setae, internal sac with two asymmetric structures, dorsal diaphragm present.
</paragraph>
<paragraph id="D9061A0E0402FFC1FF062457A17BAE3C" blockId="1.[151,1437,1254,2035]" box="[199,1046,1830,1855]" pageId="1" pageNumber="91">
<emphasis id="EBCDC61C0402FFC1FF062457A4DAAE3C" bold="true" box="[199,439,1830,1855]" pageId="1" pageNumber="91">Sexual dimorphism.</emphasis>
Females lacking modification of pro- and mesotibiae.
</paragraph>
</subSubSection>
<subSubSection id="91A349850402FFC1FF06243BA463AEA8" pageId="1" pageNumber="91" type="diagnosis">
<paragraph id="D9061A0E0402FFC1FF06243BA463AEA8" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="91">
<emphasis id="EBCDC61C0402FFC1FF06243BA4BDAE60" bold="true" box="[199,464,1866,1891]" pageId="1" pageNumber="91">Differential diagnosis.</emphasis>
<taxonomicName id="1EB9618D0402FFC1FE1B243DA71EAE60" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[474,627,1866,1891]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="bilandzijae">
<emphasis id="EBCDC61C0402FFC1FE1B243DA71EAE60" box="[474,627,1866,1891]" italics="true" pageId="1" pageNumber="91">T. bilandzijae</emphasis>
</taxonomicName>
is readily separated from all other known species of the genus by: 1) small size, length under
<quantity id="1E41B7EB0402FFC1FE6B241FA77CAE84" box="[426,529,1902,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="91" unit="mm" value="1.85">1.85 mm</quantity>
, 2) antennae short, only
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long, and 3) scape which is only twice as long as pedicel.
</paragraph>
</subSubSection>
<subSubSection id="91A349850402FFC2FF0624C7A40EA867" lastPageId="2" lastPageNumber="92" pageId="1" pageNumber="91" type="distribution">
<paragraph id="D9061A0E0402FFC2FF0624C7A40EA867" blockId="1.[151,1437,1254,2035]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="92" pageId="1" pageNumber="91">
<emphasis id="EBCDC61C0402FFC1FF0624C7A49EAECC" bold="true" box="[199,499,1974,1999]" pageId="1" pageNumber="91">Distribution and habitat.</emphasis>
<taxonomicName id="1EB9618D0402FFC1FE3B24C9A7FEAECC" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[506,659,1974,1999]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="bilandzijae">
<emphasis id="EBCDC61C0402FFC1FE3B24C9A7FEAECC" box="[506,659,1974,1999]" italics="true" pageId="1" pageNumber="91">T. bilandzijae</emphasis>
</taxonomicName>
is known only from the cave Peć u Čulinovim raljevinama (
<figureCitation id="4182068B0402FFC1FA8124C7A5DBAEF0" captionStart="FIGURES 10–12" captionStartId="6.[151,264,1792,1815]" captionTargetBox="[185,1399,411,1792]" captionTargetId="figure@6.[164,1423,411,1771]" captionTargetPageId="6" captionText="FIGURES 10–12. 10—entrance to Peć u Čulinovim raljevinama (cave), type locality (photo Marko Lukić); 11— Thaumatocephalus bilandzijae sp. n., habitus in situ (photo Alen Kirin); 12—topographical map of the type locality (the finding place of the T. bilandzijae sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585165/files/figure.png" pageId="1" pageNumber="91">Figs 10, 12</figureCitation>
), situated at
<quantity id="1E41B7EB0402FFC1FE8324ABA4E5AEF1" box="[322,392,2010,2035]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="1" pageNumber="91" unit="m" value="425.0">425 m</quantity>
a.s.l., near the village Kolić in the municipality Dugopolje (
<figureCitation id="4182068B0402FFC1FBE524ABA11BAEF0" box="[1060,1142,2010,2035]" captionStart="FIGURE 30" captionStartId="16.[151,250,1943,1965]" captionTargetBox="[150,1436,846,1922]" captionTargetId="figure@16.[150,1436,846,1922]" captionTargetPageId="16" captionText="FIGURE 30. Distribution map of the species of the genus Thaumastocephalus mentioned in this work." httpUri="https://zenodo.org/record/2585179/files/figure.png" pageId="1" pageNumber="91">Fig. 30</figureCitation>
, red square). The cave has two separate chambers, and is about
<quantity id="1E41B7EB0401FFC2FDF023E6A707A9AC" box="[561,618,151,176]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="92" unit="m" value="25.0">25 m</quantity>
long and
<quantity id="1E41B7EB0401FFC2FD1923E6A67CA9AC" box="[728,785,151,176]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="92" unit="m" value="15.0">15 m</quantity>
deep (
<bibRefCitation id="BD2867FF0401FFC2FC9A23E6A172A9B3" author="Jalzic, B. &amp; Bedek, J. &amp; Bilandzija, H. &amp; Cvitanovic, H. &amp; Drazina, T. &amp; Gottstein, S. &amp; Kljakovic Gaspic, F. &amp; Lukic, M. &amp; Ozimec, R. &amp; Pavlek, M. &amp; Slapnik, R. &amp; Stamol, V." box="[859,1055,151,176]" pageId="2" pageNumber="92" refId="ref10811" refString="Jalzic, B., Bedek, J., Bilandzija, H., Cvitanovic, H., Drazina, T., Gottstein, S., Kljakovic Gaspic, F., Lukic, M., Ozimec, R., Pavlek, M., Slapnik, R. &amp; Stamol, V. (2010) Atlas spiljskih tipskih lokaliteta faune Republike Hrvatske. Fol. 1. Hrvatsko biospeleolosko drustvo, Drzavni zavod za zastitu prirode, Zagreb, 261 pp." type="book" year="2010">
Jalžić
<emphasis id="EBCDC61C0401FFC2FC6223E8A6BEA9B3" box="[931,979,151,176]" italics="true" pageId="2" pageNumber="92">et al</emphasis>
. 2010
</bibRefCitation>
). The air temperature measured in the cave on
<date id="AD073CCE0401FFC2FEE823CDA4B7A9D7" box="[297,474,188,213]" pageId="2" pageNumber="92" value="2011-06-25">25th June 2011</date>
was 13 °C. The cave is the
<typeStatus id="0602A4AC0401FFC2FCF123CCA60DA9D6" box="[816,864,189,213]" pageId="2" pageNumber="92">type</typeStatus>
locality for Coleoptera—
<taxonomicName id="1EB9618D0401FFC2FB4D23CDA42FA9FB" authority="novaki (Muller, 1911)" authorityName="Muller" authorityYear="1911" baseAuthorityName="Muller" baseAuthorityYear="1911" class="Insecta" family="Carabidae" genus="Duvalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subSpecies" species="novaki" subSpecies="novaki">
<emphasis id="EBCDC61C0401FFC2FB4D23CDA0F1A9D6" box="[1164,1436,188,213]" italics="true" pageId="2" pageNumber="92">Duvalius novaki novaki</emphasis>
(
<bibRefCitation id="BD2867FF0401FFC2FF5F23AEA454A9FB" author="Muller, J." box="[158,313,223,248]" pageId="2" pageNumber="92" pagination="1 - 4" refId="ref11136" refString="Muller, J. [G.] (1911) Neue Hohlenkafer aus dem osterreichischen Karst. Wiener entomologische Zeitung, 30 (1), 1 - 4." type="journal article" year="1911">Müller, 1911</bibRefCitation>
)
</taxonomicName>
. The following fauna is also recorded from the cave: Araneae—
<taxonomicName id="1EB9618D0401FFC2FBF82390A440A81F" authority="(Kratochvil &amp; Miller, 1939)" authorityName="Kratochvil &amp; Miller" authorityYear="1939" baseAuthorityName="Kratochvil &amp; Miller" baseAuthorityYear="1939" class="Arachnida" family="Leptonetidae" genus="Barusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="maheni">
<emphasis id="EBCDC61C0401FFC2FBF82390A182A9FB" box="[1081,1263,223,248]" italics="true" pageId="2" pageNumber="92">Barusia maheni</emphasis>
(Kratochvíl &amp; Miller, 1939)
</taxonomicName>
and
<taxonomicName id="1EB9618D0401FFC2FEA62275A6A7A81F" authority="Absolon &amp; Kratochvil, 1932" authorityName="Absolon &amp; Kratochvil" authorityYear="1932" box="[359,970,260,285]" class="Arachnida" family="Linyphiidae" genus="Troglohyphantes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="EBCDC61C0401FFC2FEA62275A717A81E" box="[359,634,260,285]" italics="true" pageId="2" pageNumber="92">Troglohyphantes strandi</emphasis>
Absolon &amp; Kratochvíl, 1932
</taxonomicName>
as noted by
<bibRefCitation id="BD2867FF0401FFC2FB9E2275A0FAA81F" author="Deeleman-Reinhold, C. L." box="[1119,1431,260,285]" pageId="2" pageNumber="92" pagination="1 - 221" refId="ref10258" refString="Deeleman-Reinhold, C. L. (1978) Revision of the cave-dwelling and related spiders of the genus Troglohyphantes Joseph (Linyphiidae), with special reference to the yugoslav species. Slovenska Akademija Znanosti in Umetnosti, Razred za Prirodoslovne Fede, Classis IV, Historia Naturalis, 23 (6), 1 - 221." type="journal article" year="1978">Deeleman-Reinhold (1978)</bibRefCitation>
; Isopoda—
<taxonomicName id="1EB9618D0401FFC2FECD2256A4E2A843" authorityName="Racovitza" authorityYear="1908" box="[268,399,295,320]" class="Malacostraca" family="Trichoniscidae" genus="Alpioniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C0401FFC2FECD2256A4E2A843" box="[268,399,295,320]" italics="true" pageId="2" pageNumber="92">Alpioniscus</emphasis>
</taxonomicName>
sp. (det. J. Bedek); Coleoptera—
<taxonomicName id="1EB9618D0401FFC2FCC22258A054A843" authority="(Schaufuss, 1862)" authorityName="Schaufuss" authorityYear="1862" baseAuthorityName="Schaufuss" baseAuthorityYear="1862" box="[771,1337,295,320]" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="subSpecies" species="cavicola" subSpecies="modestus">
<emphasis id="EBCDC61C0401FFC2FCC22258A104A843" box="[771,1129,295,320]" italics="true" pageId="2" pageNumber="92">Laemostenus cavicola modestus</emphasis>
(Schaufuss, 1862)
</taxonomicName>
as noted by
<bibRefCitation id="BD2867FF0401FFC2FF7B223CA40DA867" author="Pretner, E." box="[186,352,332,356]" pageId="2" pageNumber="92" pagination="101 - 239" refId="ref11804" refString="Pretner, E. (1973) Koleopteroloska fauna pecina i jama Hrvatske. Krs Jugoslavije, 8, 101 - 239." type="journal article" year="1973">Pretner (1973)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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