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<document id="6442365141212DED166CAFA0983B9E41" ID-CLB-Dataset="28520" ID-DOI="10.11646/zootaxa.4559.1.3" ID-GBIF-Dataset="becdb1cf-be93-48ca-abc4-898d0f84d0bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="2585159" ID-ZooBank="F051C9DE-CCB7-454E-8184-B6240049F004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550815179529" checkinUser="plazi" docAuthor="Hlaváč, Peter, Bregović, Petra &amp; Jalžić, Branko" docDate="2019" docId="5110AB180401FFC2FF5622C9A67CAF28" docLanguage="en" docName="zootaxa.4559.1.3.pdf" docOrigin="Zootaxa 4559 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumastocephalus kirini Hlaváč &amp; Bregović &amp; Jalžić 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="92" masterDocId="AD29D3600403FFC0FFC12371A56DA903" masterDocTitle="Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)" masterLastPageNumber="110" masterPageNumber="90" pageNumber="92" updateTime="1698699937030" updateUser="ExternalLinkService">
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<mods:title id="9BF210E8B21658C451086CD52DE01980">Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="1A71E59FDDB9F7571C31A136D69121AA">Hlaváč, Peter</mods:namePart>
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<mods:namePart id="6369BBA9B62FF76A4EBD03D4F513EAB2">Bregović, Petra</mods:namePart>
</mods:name>
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<mods:namePart id="C6AA0C26374199F036D13D6404F75BDF">Jalžić, Branko</mods:namePart>
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<treatment id="5110AB180401FFC2FF5622C9A67CAF28" ID-DOI="http://doi.org/10.5281/zenodo.5934382" ID-GBIF-Taxon="154007276" ID-Zenodo-Dep="5934382" LSID="urn:lsid:plazi:treatment:5110AB180401FFC2FF5622C9A67CAF28" httpUri="http://treatment.plazi.org/id/5110AB180401FFC2FF5622C9A67CAF28" lastPageNumber="92" pageId="2" pageNumber="92">
<subSubSection id="91A349850401FFC2FF5622C9A61CA8D1" box="[151,881,440,466]" pageId="2" pageNumber="92" type="nomenclature">
<paragraph id="D9061A0E0401FFC2FF5622C9A61CA8D1" blockId="2.[151,881,440,499]" box="[151,881,440,466]" pageId="2" pageNumber="92">
<heading id="824EAD620401FFC2FF5622C9A61CA8D1" bold="true" box="[151,881,440,466]" fontSize="11" level="1" pageId="2" pageNumber="92" reason="1">
<emphasis id="EBCDC61C0401FFC2FF5622C9A61CA8D1" bold="true" box="[151,881,440,466]" pageId="2" pageNumber="92">
<taxonomicName id="1EB9618D0401FFC2FF5622C9A64FA8D1" ID-CoL="8YBQN" authority="Hlavac, Bregovic &amp; Jalzic" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[151,802,440,466]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="kirini" status="sp. nov.">
<emphasis id="EBCDC61C0401FFC2FF5622C9A4B8A8D1" bold="true" box="[151,469,440,466]" italics="true" pageId="2" pageNumber="92">Thaumastocephalus kirini</emphasis>
Hlaváč, Bregović &amp; Jalžić
</taxonomicName>
,
<taxonomicNameLabel id="F0FE7B670401FFC2FCEF22C9A61CA8D1" box="[814,881,440,466]" pageId="2" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="91A349850401FFC2FF5622ABA474A8F0" box="[151,281,474,499]" pageId="2" pageNumber="92" type="description">
<paragraph id="D9061A0E0401FFC2FF5622ABA474A8F0" blockId="2.[151,881,440,499]" box="[151,281,474,499]" pageId="2" pageNumber="92">
(
<figureCitation id="4182068B0401FFC2FF5E22ABA58AA8F0" box="[159,231,474,499]" captionStart="FIGURES 14" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[270,1322,195,1885]" captionTargetId="figure@4.[269,1322,195,1885]" captionTargetPageId="4" captionText="FIGURES 14. 1—Thaumatocephalus bilandzijae sp. n., aedeagus, dorsal view (scale: 0.1 mm); 2—Thaumatocephalus kirini sp. n., aedeagus, dorsal view (scale: 0.2 mm); 3—Thaumatocephalus marsici sp. n., aedeagus, dorsal view (scale: 0.2 mm); 4—Thaumatocephalus rujnicensis sp. n., aedeagus, dorsal view (scale: 0.1 mm)." httpUri="https://zenodo.org/record/2585161/files/figure.png" pageId="2" pageNumber="92">Figs 2</figureCitation>
,
<figureCitation id="4182068B0401FFC2FF3222AAA47DA8F0" box="[243,272,475,499]" captionStart="FIGURES 1315" captionStartId="11.[151,264,1784,1807]" captionTargetBox="[210,1377,193,1764]" captionTargetId="figure@11.[210,1377,193,1764]" captionTargetPageId="11" captionText="FIGURES 1315. 13—entrance to Jamica u docima (cave), type locality (photo Branko Jalžić); 14—Thaumatocephalus kirini sp. n., habitus in situ (photo Boris Krstinić); 15—topographical map of the type locality (the finding place of the T. kirini sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585167/files/figure.png" pageId="2" pageNumber="92">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="91A349850401FFC2FF562153A6A0AB5C" pageId="2" pageNumber="92" type="etymology">
<paragraph id="D9061A0E0401FFC2FF562153A6A0AB5C" blockId="2.[151,1437,546,1579]" pageId="2" pageNumber="92">
<emphasis id="EBCDC61C0401FFC2FF562153A473AB38" bold="true" box="[151,286,546,571]" pageId="2" pageNumber="92">Etymology.</emphasis>
This species is named after speleologist Alen Kirin, a member of the Croatian Biospeleological Society (CBSS) and collector of the first specimen from the
<typeStatus id="0602A4AC0401FFC2FCFA2136A606AB5C" box="[827,875,583,607]" pageId="2" pageNumber="92">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="91A349850401FFC2FF06211BA421ABEC" pageId="2" pageNumber="92" type="materials_examined">
<paragraph id="D9061A0E0401FFC2FF06211BA421ABEC" blockId="2.[151,1437,546,1579]" pageId="2" pageNumber="92">
<emphasis id="EBCDC61C0401FFC2FF06211BA793AB80" bold="true" box="[199,766,618,643]" pageId="2" pageNumber="92">
Material studied.
<materialsCitation id="69D110530401FFC2FE5D211BA793AB80" ID-GBIF-Occurrence="2005761070" box="[412,766,618,643]" collectingDate="2015-06-17" collectionCode="CNHM" collectorName="B. Jalzic" country="Croatia" county="Rujnica" location="Metkovic" municipality="Desne" pageId="2" pageNumber="92" specimenCount="1" typeStatus="holotype">
<typeStatus id="0602A4AC0401FFC2FE5D211BA752AB80" box="[412,575,618,643]" pageId="2" pageNumber="92" type="holotype">HOLOTYPE</typeStatus>
: Ƌ:
<collectingCountry id="A1AE5A9E0401FFC2FDB2211BA794AB80" box="[627,761,618,643]" name="Croatia" pageId="2" pageNumber="92">CROATIA</collectingCountry>
:
</materialsCitation>
</emphasis>
<location id="DC664CD50401FFC2FCC7211BA619AB80" LSID="urn:lsid:plazi:treatment:5110AB180401FFC2FF5622C9A67CAF28:DC664CD50401FFC2FCC7211BA619AB80" box="[774,884,618,643]" country="Croatia" county="Rujnica" municipality="Desne" name="Metkovic" pageId="2" pageNumber="92">Metković</location>
,
<collectingMunicipality id="396280740401FFC2FC41211AA6A4AB81" box="[896,969,619,642]" pageId="2" pageNumber="92">Desne</collectingMunicipality>
,
<collectingCounty id="306762820401FFC2FC14211BA142AB80" box="[981,1071,618,643]" pageId="2" pageNumber="92">Rujnica</collectingCounty>
, Varda. Jamica u docima (cave),
<date id="AD073CCE0401FFC2FF5621FFA474ABA4" box="[151,281,654,679]" pageId="2" pageNumber="92" value="2015-06-17">
<collectingDate id="BD43C5260401FFC2FF5621FFA474ABA4" box="[151,281,654,679]" pageId="2" pageNumber="92" value="2015-06-17">
17.
<bibRefCitation id="BD2867FF0401FFC2FF7721FFA474ABA4" author="Bregovic, P. &amp; Delic, T. &amp; Jalzic, B." box="[182,281,654,679]" pageId="2" pageNumber="92" pagination="40 - 49" refId="ref9929" refString="Bregovic, P., Delic, T. &amp; Jalzic, B. (2015) Doprinos poznavanju rasprostranjenosti rijetke faune podzemnih kornjasa Biokova i ponovno istrazivanje Pretnerove jame (The Distribution of Rare Subterranean Beetles from Biokovo Mountain and the Re- Exploration of Pretnerova Jama). Subterranea Croatica, 19, 40 - 49." type="journal article" year="2015">VI.2015</bibRefCitation>
</collectingDate>
</date>
,
<collectorName id="744C7FD80401FFC2FEE521FEA4E8ABA4" box="[292,389,654,679]" pageId="2" pageNumber="92">B. Jalžić</collectorName>
lgt. (CNHM).
<materialsCitation id="69D110530401FFC2FDED21FEA409ABC8" ID-GBIF-Occurrence="2005761076" collectingDate="2015-06-17" collectionCode="IV" collectorName="B. Jalzic" country="Croatia" county="Rujnica" location="Metkovic" municipality="Desne" pageId="2" pageNumber="92" specimenCount="13" specimenCount-female="13" typeStatus="paratype">
<typeStatus id="0602A4AC0401FFC2FDED21FEA7AAABA4" box="[556,711,655,679]" pageId="2" pageNumber="92" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="CFBFD1870401FFC2FD1921FEA791ABA4" box="[728,764,655,679]" pageId="2" pageNumber="92" type="female">7♀</specimenCount>
):
<specimenCount id="CFBFD1870401FFC2FCD321FEA655ABA4" box="[786,824,655,679]" pageId="2" pageNumber="92" type="female">2♀</specimenCount>
: same data as holotype;
<specimenCount id="CFBFD1870401FFC2FB8821FEA102ABA4" box="[1097,1135,655,679]" pageId="2" pageNumber="92" type="female">4♀</specimenCount>
: same data as
<typeStatus id="0602A4AC0401FFC2FACE21FFA01CABA4" box="[1295,1393,654,679]" pageId="2" pageNumber="92" type="holotype">holotype</typeStatus>
but collected on
<date id="AD073CCE0401FFC2FEE721C2A409ABC8" box="[294,356,691,715]" pageId="2" pageNumber="92" value="2010-04-30">
30.
<collectionCode id="BFA882CB0401FFC2FE8721C2A409ABC8" box="[326,356,691,715]" pageId="2" pageNumber="92">IV</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="69D110530401FFC2FEA521C2A425ABEC" ID-GBIF-Occurrence="2005761068" collectingDate="2016-09-16" collectionCode="CNHM, CPH" collectorName="A. Kirin &amp; B. Jalzic" country="Croatia" county="Rujnica" location="Metkovic" municipality="Desne" pageId="2" pageNumber="92" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
.2010 by
<collectorName id="744C7FD80401FFC2FE0C21C2A746ABC8" box="[461,555,690,715]" pageId="2" pageNumber="92">A. Kirin</collectorName>
(
<collectionCode id="BFA882CB0401FFC2FDFB21C2A7FEABC8" box="[570,659,691,715]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="2" pageNumber="92" type="Museum">CNHM</collectionCode>
,
<collectionCode id="BFA882CB0401FFC2FD5F21C2A7B4ABC8" box="[670,729,691,715]" country="USA" lsid="urn:lsid:biocol.org:col:15612" name="University of the Pacific" pageId="2" pageNumber="92" type="Herbarium">CPH</collectionCode>
);
<specimenCount id="CFBFD1870401FFC2FD2C21C2A67CABC8" box="[749,785,691,715]" pageId="2" pageNumber="92" type="female">1♀</specimenCount>
: same data as holotype but collected on
<date id="AD073CCE0401FFC2FB1921C3A035ABC8" box="[1240,1368,690,715]" pageId="2" pageNumber="92" value="2016-09-16">
<collectingDate id="BD43C5260401FFC2FB1921C3A035ABC8" box="[1240,1368,690,715]" pageId="2" pageNumber="92" value="2016-09-16">16.IX.2016</collectingDate>
</date>
by
<collectorName id="744C7FD80401FFC2FA4221C2A5BAABEC" pageId="2" pageNumber="92">B. Jalžić</collectorName>
(
<collectionCode id="BFA882CB0401FFC2FF2421A6A42DABEC" box="[229,320,727,751]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="2" pageNumber="92" type="Museum">CNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91A349850401FFC2FF06218BA6F6ADC0" pageId="2" pageNumber="92" type="description">
<paragraph id="D9061A0E0401FFC2FF06218BA403AD9D" blockId="2.[151,1437,546,1579]" pageId="2" pageNumber="92">
<emphasis id="EBCDC61C0401FFC2FF06218BA434AA10" bold="true" box="[199,345,762,787]" pageId="2" pageNumber="92">Description.</emphasis>
Body shiny (
<figureCitation id="4182068B0401FFC2FE3C218BA739AA10" box="[509,596,762,787]" captionStart="FIGURES 1315" captionStartId="11.[151,264,1784,1807]" captionTargetBox="[210,1377,193,1764]" captionTargetId="figure@11.[210,1377,193,1764]" captionTargetPageId="11" captionText="FIGURES 1315. 13—entrance to Jamica u docima (cave), type locality (photo Branko Jalžić); 14—Thaumatocephalus kirini sp. n., habitus in situ (photo Boris Krstinić); 15—topographical map of the type locality (the finding place of the T. kirini sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585167/files/figure.png" pageId="2" pageNumber="92">Fig. 14</figureCitation>
), reddish-brown, covered with dense, uneven setae, with long setae on posterior part of elytra, setae lacking on disc of elytra; legs, antennae and maxillary palpi slightly lighter, length
<quantity id="1E41B7EB0401FFC2FF562033A591AA59" box="[151,252,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" pageId="2" pageNumber="92" unit="mm" value="1.96">1.96 mm</quantity>
, maximum width of elytra
<quantity id="1E41B7EB0401FFC2FDE82033A7E2AA59" box="[553,655,834,859]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="92" unit="mm" value="0.68">0.68 mm</quantity>
. Head 1.32 times as long as wide, 0.79 times narrower than pronotum, rostrum 3.00 times as wide as long. Antennae about
<quantity id="1E41B7EB0401FFC2FCE22016A6E3AA7D" box="[803,910,871,895]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="92" unit="mm" value="0.82">0.82 mm</quantity>
long, scape 2.57 times as long as pedicel, antennomere III slightly longer than IV, antennomeres IVVII of same length, VIII slightly shorter than VII and as long as IX, antennomere X about as long as VII, terminal antennomere 3.10 times as long as X, relative length of antennomeres: 1.00 / 0.39 / 0.28 / 0.25 / 0.25 / 0.25 / 0.25 / 0.22 / 0.22 / 0.25 / 0.78. Pronotum 1.10 times as long as wide. Elytra 1.22 times as wide as long and 1.40 times as long as pronotum. Abdomen 1.14 times as long as elytra. Legs long, protibiae slightly modified, slightly depressed in apical half, median spine minuscule, mesotibiae simple, metatibiae simple. Aedeagus (
<figureCitation id="4182068B0401FFC2FD81274FA7EEAD54" box="[576,643,1086,1111]" captionStart="FIGURES 14" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[270,1322,195,1885]" captionTargetId="figure@4.[269,1322,195,1885]" captionTargetPageId="4" captionText="FIGURES 14. 1—Thaumatocephalus bilandzijae sp. n., aedeagus, dorsal view (scale: 0.1 mm); 2—Thaumatocephalus kirini sp. n., aedeagus, dorsal view (scale: 0.2 mm); 3—Thaumatocephalus marsici sp. n., aedeagus, dorsal view (scale: 0.2 mm); 4—Thaumatocephalus rujnicensis sp. n., aedeagus, dorsal view (scale: 0.1 mm)." httpUri="https://zenodo.org/record/2585161/files/figure.png" pageId="2" pageNumber="92">Fig. 2</figureCitation>
)
<quantity id="1E41B7EB0401FFC2FD53274EA798AD55" box="[658,757,1087,1111]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="92" unit="mm" value="0.49">0.49 mm</quantity>
long, with basal bulb about 1.15 times as long as apical part, right paramere more robust than left, both with three preapical setae, internal sac with one spiral structure, dorsal diaphragm present.
</paragraph>
<paragraph id="D9061A0E0401FFC2FF0627DBA6F6ADC0" blockId="2.[151,1437,546,1579]" box="[199,923,1194,1219]" pageId="2" pageNumber="92">
<emphasis id="EBCDC61C0401FFC2FF0627DBA4DAADC0" bold="true" box="[199,439,1194,1219]" pageId="2" pageNumber="92">Sexual dimorphism.</emphasis>
Females lacking modification of protibiae.
</paragraph>
</subSubSection>
<subSubSection id="91A349850401FFC2FF0627BFA485AC50" pageId="2" pageNumber="92" type="diagnosis">
<paragraph id="D9061A0E0401FFC2FF0627BFA485AC50" blockId="2.[151,1437,546,1579]" pageId="2" pageNumber="92">
<emphasis id="EBCDC61C0401FFC2FF0627BFA4A2ADE4" bold="true" box="[199,463,1230,1255]" pageId="2" pageNumber="92">Differential diagnosis.</emphasis>
<taxonomicName id="1EB9618D0401FFC2FE1627BEA742ADE4" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[471,559,1230,1255]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="kirini">
<emphasis id="EBCDC61C0401FFC2FE1627BEA742ADE4" box="[471,559,1230,1255]" italics="true" pageId="2" pageNumber="92">T. kirini</emphasis>
</taxonomicName>
is readily separated from all other known species of the genus by: 1) frontal tibiae at most with minuscule tooth, 2) body length under
<quantity id="1E41B7EB0401FFC2FCDA2782A630AC09" box="[795,861,1267,1291]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="92" unit="mm" value="2.0">2 mm</quantity>
, 3) terminal antennomere short, less than 4 times as long as antenommere X, 4) elytra long, more than 1.3 times as long as pronotum, and 5) short rostrum which is about 3 times as wide as long.
</paragraph>
</subSubSection>
<subSubSection id="91A349850401FFC2FF06262FA67CAF28" pageId="2" pageNumber="92" type="distribution">
<paragraph id="D9061A0E0401FFC2FF06262FA67CAF28" blockId="2.[151,1437,546,1579]" pageId="2" pageNumber="92">
<emphasis id="EBCDC61C0401FFC2FF06262FA49DAC74" bold="true" box="[199,496,1374,1399]" pageId="2" pageNumber="92">Distribution and habitat.</emphasis>
<taxonomicName id="1EB9618D0401FFC2FE36262EA720AC74" authorityName="Hlaváč &amp; Bregović &amp; Jalžić" authorityYear="2019" box="[503,589,1374,1399]" class="Insecta" family="Staphylinidae" genus="Thaumastocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="kirini">
<emphasis id="EBCDC61C0401FFC2FE36262EA720AC74" box="[503,589,1374,1399]" italics="true" pageId="2" pageNumber="92">T. kirini</emphasis>
</taxonomicName>
is known only from the cave Jamica u docima (
<figureCitation id="4182068B0401FFC2FBA4262FA189AC74" box="[1125,1252,1374,1399]" captionStart="FIGURES 1315" captionStartId="11.[151,264,1784,1807]" captionTargetBox="[210,1377,193,1764]" captionTargetId="figure@11.[210,1377,193,1764]" captionTargetPageId="11" captionText="FIGURES 1315. 13—entrance to Jamica u docima (cave), type locality (photo Branko Jalžić); 14—Thaumatocephalus kirini sp. n., habitus in situ (photo Boris Krstinić); 15—topographical map of the type locality (the finding place of the T. kirini sp. n. is marked with red dots)." httpUri="https://zenodo.org/record/2585167/files/figure.png" pageId="2" pageNumber="92">Figs 13, 15</figureCitation>
), situated at
<quantity id="1E41B7EB0401FFC2FAB3262FA5C0AC99" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="2" pageNumber="92" unit="m" value="345.0">345 m</quantity>
a.s.l. near peak Varda (
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) and village Desne (
<figureCitation id="4182068B0401FFC2FD2B26F3A653AC98" box="[746,830,1410,1435]" captionStart="FIGURE 30" captionStartId="16.[151,250,1943,1965]" captionTargetBox="[150,1436,846,1922]" captionTargetId="figure@16.[150,1436,846,1922]" captionTargetPageId="16" captionText="FIGURE 30. Distribution map of the species of the genus Thaumastocephalus mentioned in this work." httpUri="https://zenodo.org/record/2585179/files/figure.png" pageId="2" pageNumber="92">Fig. 30</figureCitation>
, violet star). The cave is a vertical shaft,
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deep and
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long. The air temperature measured in the cave on
<date id="AD073CCE0401FFC2FCF726D7A687ACBC" box="[822,1002,1446,1471]" pageId="2" pageNumber="92" value="2010-04-30">30th April 2010</date>
was 10.1 °C and the relative humidity was 98 %. The following fauna is also recorded from the cave: Isopoda—
<taxonomicName id="1EB9618D0401FFC2FC1226BBA010ACE0" authority="(Strouhal, 1938)" authorityName="Strouhal" authorityYear="1938" baseAuthorityName="Strouhal" baseAuthorityYear="1938" box="[979,1405,1482,1507]" class="Malacostraca" family="Trichoniscidae" genus="Alpioniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="verheoffi">
<emphasis id="EBCDC61C0401FFC2FC1226BBA1D2ACE0" box="[979,1215,1482,1507]" italics="true" pageId="2" pageNumber="92">Alpioniscus verheoffi</emphasis>
(Strouhal, 1938)
</taxonomicName>
as noted by
<bibRefCitation id="BD2867FF0401FFC2FEC0269FA4A1AF04" author="Bedek, J. &amp; Taiti, S. &amp; Gottstein, S." box="[257,460,1518,1543]" pageId="2" pageNumber="92" pagination="237 - 354" refId="ref9760" refString="Bedek, J., Taiti, S. &amp; Gottstein, S. (2011) Catalogue and atlas of cave-dwelling terrestrial isopods (Crustacea: Oniscidea) from Croatia. Natura Croatica, 20 (2), 237 - 354." type="journal article" year="2011">
Bedek
<emphasis id="EBCDC61C0401FFC2FE91269EA4ECAF04" box="[336,385,1518,1543]" italics="true" pageId="2" pageNumber="92">et al</emphasis>
. 2011
</bibRefCitation>
; Coleoptera—
<taxonomicName id="1EB9618D0401FFC2FDAE269FA780AF04" box="[623,749,1518,1543]" class="Insecta" family="Carabidae" genus="Neotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C0401FFC2FDAE269FA780AF04" box="[623,749,1518,1543]" italics="true" pageId="2" pageNumber="92">Neotrechus</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="1EB9618D0401FFC2FCE5269EA6D8AF05" authorityName="Jeannel" authorityYear="1910" box="[804,949,1519,1542]" class="Insecta" family="Leiodidae" genus="Speonesiotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="EBCDC61C0401FFC2FCE5269EA6D8AF05" box="[804,949,1519,1542]" italics="true" pageId="2" pageNumber="92">Speonesiotes</emphasis>
</taxonomicName>
sp. (det. B. Jalžić), and
<taxonomicName id="1EB9618D0401FFC2FB0B269EA457AF28" authority="(Reitter, 1881)" authorityName="Reitter" authorityYear="1881" baseAuthorityName="Reitter" baseAuthorityYear="1881" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="scapularis">
<emphasis id="EBCDC61C0401FFC2FB0B269EA0F1AF04" box="[1226,1436,1518,1543]" italics="true" pageId="2" pageNumber="92">Bryaxis scapularis</emphasis>
(Reitter, 1881)
</taxonomicName>
as noted by Bekchiev and
<bibRefCitation id="BD2867FF0401FFC2FDA92563A661AF28" author="Hlavac, P. &amp; Lakota, J. &amp; Ceplik, D." box="[616,780,1554,1579]" pageId="2" pageNumber="92" pagination="137 - 140" refId="ref10527" refString="Hlavac, P., Lakota, J. &amp; Ceplik, D. (2016) Endogean and cavernicolous Coleoptera of the Balkans. XVII. A new species of the genus Speluncarius Reitter, 1886 (Coleoptera, Carabidae, Harpalinae, Pterostichini) from Croatia. Zootaxa, 4161 (1), 137 - 140. https: // doi. org / 10.11646 / zootaxa. 4161.1.10" type="journal article" year="2016">Hlaváč (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>