treatments-xml/data/38/6A/BE/386ABE42FFA9FFB3FF06ABF52C1EFC00.xml
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<document id="C5E0C123DCFC36E3E0552F4051F0D2A8" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFA9FFB3FF06ABF52C1EFC00" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanopsis tarahomii Marin, Sheibani &amp; Sari 2014" docType="treatment" docVersion="3" lastPageNumber="207" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="204" updateTime="1702658426235" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="4C61B125C72A103BB2B804E81F5ED6EE">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:namePart id="0F0F22B1EDDDDC0289E9FAEF06413899">Anker, Arthur</mods:namePart>
<mods:affiliation id="F6B3F7F75FA4F29EEF5A04121065ED94">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<heading id="EB34B838FFA9FFB6FF06ABF52D35FA77" bold="true" box="[151,789,1464,1491]" fontSize="11" level="1" pageId="25" pageNumber="204" reason="1">
<taxonomicName id="77C374D7FFA9FFB6FF06ABF52D35FA77" ID-CoL="85MDJ" authority="Marin, Sheibani &amp; Sari, 2014" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[151,789,1464,1491]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFA9FFB6FF06ABF52D35FA77" bold="true" box="[151,789,1464,1491]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF06ABF52FBDFA76" bold="true" box="[151,413,1464,1491]" italics="true" pageId="25" pageNumber="204">Athanopsis tarahomii</emphasis>
<bibRefCitation id="D45272A5FFA9FFB6FE35ABF52D35FA77" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[420,789,1464,1490]" pageId="25" pageNumber="204" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">Marin, Sheibani &amp; Sari, 2014</bibRefCitation>
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<paragraph id="B07C0F54FFA9FFB6FF06AB912EDAFA52" blockId="25.[151,789,1464,1527]" box="[151,250,1500,1527]" pageId="25" pageNumber="204">
(
<figureCitation id="28F813D1FFA9FFB6FF0EAB912ED1FA52" box="[159,241,1500,1527]" captionStart="FIGURE 14" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[170,1417,181,1884]" captionTargetId="figure-19@27.[170,1417,181,1884]" captionTargetPageId="27" captionText="FIGURE 14. Athanopsis tarahomii Marin, Sheibani &amp; Sari, 2014, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350819" httpUri="https://zenodo.org/record/10350819/files/figure.png" pageId="25" pageNumber="204">Fig. 14</figureCitation>
)
</paragraph>
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<subSubSection id="F8D95CDFFFA9FFB6FF06A8572F1BF9D5" pageId="25" pageNumber="204" type="reference_group">
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<treatmentCitationGroup id="90D3287AFFA9FFB6FF06A8572ABBF997" box="[151,1179,1562,1586]" pageId="25" pageNumber="204">
<treatmentCitation id="31622945FFA9FFB6FF06A8572C7CF997" ID-DOI="http://doi.org/10.5281/zenodo.6131289" ID-Zenodo-Dep="6131289" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[151,604,1562,1586]" httpUri="http://treatment.plazi.org/id/C03687869A40FFF1FF40FF0DFA5CA63E" page="404" pageId="25" pageNumber="204" year="2014">
<taxonomicName id="77C374D7FFA9FFB6FF06A8572C7CF997" ID-CoL="85MDJ" authority="Marin et al. 2014: 404" authorityName="Marin" authorityPageNumber="404" authorityYear="2014" box="[151,604,1562,1586]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFA9FFB6FF06A8572F52F997" box="[151,370,1562,1586]" italics="true" pageId="25" pageNumber="204">Athanopsis tarahomii</emphasis>
<bibRefCitation id="D45272A5FFA9FFB6FEE9A8572C7CF997" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[376,604,1562,1586]" pageId="25" pageNumber="204" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFA9FFB6FE2CA8562FD0F997" box="[445,496,1562,1586]" italics="true" pageId="25" pageNumber="204">et al.</emphasis>
2014: 404
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(part., not “Coloration” on p. 409), figs. 46 (not fig. 7).
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</paragraph>
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<collectingRegion id="7207C1B6FFA9FFB6FF06A8742E9DF9F5" box="[151,189,1593,1616]" country="Maldives" name="Uthuru" pageId="25" pageNumber="204">Not</collectingRegion>
<taxonomicName id="77C374D7FFA9FFB6FF55A8742FBFF9F4" ID-CoL="85MDJ" box="[196,415,1593,1617]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFA9FFB6FF55A8742FBFF9F4" box="[196,415,1593,1617]" italics="true" pageId="25" pageNumber="204">Athanopsis tarahomii</emphasis>
</taxonomicName>
<treatmentCitation id="31622945FFA9FFB6FE28A8742C80F9F4" ID-DOI="http://doi.org/10.5281/zenodo.6131289" ID-Zenodo-Dep="6131289" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[441,672,1593,1617]" httpUri="http://treatment.plazi.org/id/C03687869A40FFF1FF40FF0DFA5CA63E" page="408" pageId="25" pageNumber="204" year="2014">
<bibRefCitation id="D45272A5FFA9FFB6FE28A8742C80F9F4" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[441,672,1593,1617]" pageId="25" pageNumber="204" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFA9FFB6FE6FA8772C11F9F4" box="[510,561,1593,1617]" italics="true" pageId="25" pageNumber="204">et al.</emphasis>
2014: 408
</bibRefCitation>
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, fig. 7, 409 (“Coloration”) [=
<taxonomicName id="77C374D7FFA9FFB6FC42A8742BB8F9F4" ID-CoL="85LT8" authority="(Coutiere, 1896)" baseAuthorityName="Coutiere" baseAuthorityYear="1896" box="[979,1432,1593,1617]" class="Malacostraca" family="Alpheidae" genus="Amphibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="jousseaumei">
<emphasis id="82B7D346FFA9FFB6FC42A8742AC7F9F4" box="[979,1255,1593,1617]" italics="true" pageId="25" pageNumber="204">Amphibetaeus jousseaumei</emphasis>
(
<bibRefCitation id="D45272A5FFA9FFB6FB64A8742BB1F9F4" author="Coutiere, H." box="[1269,1425,1593,1617]" pageId="25" pageNumber="204" pagination="313 - 317" refId="ref21438" refString="Coutiere, H. (1896) Note sur un nouvel Alpheide, Betaeus Jousseaumei [Crust.]. Bulletin de la Societe Entomologique de France, 13, 313 - 317. https: // doi. org / 10.3406 / bsef. 1896.21892" type="journal article" year="1896">Coutière, 1896</bibRefCitation>
)
</taxonomicName>
, see below].
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</paragraph>
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<emphasis id="82B7D346FFA9FFB6FF06A8DA2F5AF914" bold="true" box="[151,378,1687,1713]" pageId="25" pageNumber="204">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFA9FFB6FE1BA8DA2B5BF95C" collectingDate="2023-01-20" collectionCode="FLMNH, KAUST" collectorName="A. Anker" country="Saudi Arabia" elevation="1" latitude="22.340612" location="Thuwal" longLatPrecision="1" longitude="39.08753" municipality="King Abdullah Monument" pageId="25" pageNumber="204" specimenCode="UF 71425" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA9FFB6FE1BA8DA2FF6F914" box="[394,470,1687,1713]" count="1" pageId="25" pageNumber="204" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA9FFB6FD95A8DA2C7FF914" box="[516,607,1687,1713]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="25" pageNumber="204" unit="mm" value="4.6">4.6 mm</quantity>
), 1 ovig. female (cl
<quantity id="773BA2B1FFA9FFB6FCD5A8DA2DBFF914" box="[836,927,1687,1713]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="204" unit="mm" value="4.7">4.7 mm</quantity>
),
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<specimenCode id="E065A72FFFA9FFB6FB8CA8D52AB2F917" box="[1053,1170,1687,1714]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="25" pageNumber="204" type="Museum">UF 71425</specimenCode>
,
<collectingCountry id="C8D44FC4FFA9FFB6FB0FA8DA2B14F914" box="[1182,1332,1687,1713]" name="Saudi Arabia" pageId="25" pageNumber="204">Saudi Arabia</collectingCountry>
,
<location id="B51C598FFFA9FFB6FAD1A8DA2BB8F914" LSID="urn:lsid:plazi:treatment:386ABE42FFA9FFB3FF06ABF52C1EFC00:B51C598FFFA9FFB6FAD1A8DA2BB8F914" box="[1344,1432,1687,1713]" country="Saudi Arabia" latitude="22.340612" longLatPrecision="1" longitude="39.08753" municipality="King Abdullah Monument" name="Thuwal" pageId="25" pageNumber="204">Thuwal</location>
, near
<collectionCode id="D6D29791FFA9FFB6FF5CA8F62F0AF970" box="[205,298,1723,1749]" pageId="25" pageNumber="204">KAUST</collectionCode>
<collectingMunicipality id="5018952EFFA9FFB6FEA1A8F62C79F973" box="[304,601,1723,1750]" pageId="25" pageNumber="204">King Abdullah Monument</collectingMunicipality>
,
<geoCoordinate id="D5F76993FFA9FFB6FDF2A8F62D22F970" box="[611,770,1723,1749]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="25" pageNumber="204" precision="1" seconds="26.2" value="22.340612">22°2026.2”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFA9FFB6FC84A8F62D93F973" box="[789,947,1723,1750]" degrees="39" direction="east" minutes="05" orientation="longitude" pageId="25" pageNumber="204" precision="1" seconds="15.1" value="39.08753">39°0515.1”E</geoCoordinate>
, shallow subtidal sandflat with seagrass and rubble patches, depth
<quantity id="773BA2B1FFA9FFB6FE1CA8922C20F95F" box="[397,512,1759,1786]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="25" pageNumber="204" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">
<elevation id="3BEEE867FFA9FFB6FE1CA8922C20F95F" box="[397,512,1759,1786]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="25" pageNumber="204" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.51.5 m</elevation>
</quantity>
, suction pump, in echiuran burrow, leg.
<collectorName id="1D366A82FFA9FFB6FC50A8922A0AF95C" box="[961,1066,1759,1785]" pageId="25" pageNumber="204">A. Anker</collectorName>
,
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<collectingDate id="D439D07CFFA9FFB6FBA4A8922A93F95C" box="[1077,1203,1759,1785]" pageId="25" pageNumber="204" value="2023-01-20">20.01.2023</collectingDate>
</date>
[fcn AA-22-493]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF56A94E2F57F850" pageId="25" pageNumber="204" type="description">
<paragraph id="B07C0F54FFA9FFB6FF56A94E2C69F8E4" blockId="25.[151,1436,1687,2037]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56A94E2F71F8B8" bold="true" box="[199,337,1795,1821]" pageId="25" pageNumber="204">Description</emphasis>
. See
<bibRefCitation id="D45272A5FFA9FFB6FE00A94E2C4AF8B8" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[401,618,1795,1821]" pageId="25" pageNumber="204" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFA9FFB6FE4FA9492C37F8B8" box="[478,535,1795,1821]" italics="true" pageId="25" pageNumber="204">et al.</emphasis>
(2014)
</bibRefCitation>
for original description and illustrations; colour photographs of the
<collectingRegion id="7207C1B6FFA9FFB6FAFEA94E2E9FF8E4" country="Eritrea" name="Semenawi Keyih Bahri" pageId="25" pageNumber="204">Red Sea</collectingRegion>
specimens are provided in
<figureCitation id="28F813D1FFA9FFB6FE62A96A2C65F8E4" box="[499,581,1831,1857]" captionStart="FIGURE 14" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[170,1417,181,1884]" captionTargetId="figure-19@27.[170,1417,181,1884]" captionTargetPageId="27" captionText="FIGURE 14. Athanopsis tarahomii Marin, Sheibani &amp; Sari, 2014, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350819" httpUri="https://zenodo.org/record/10350819/files/figure.png" pageId="25" pageNumber="204">Fig. 14</figureCitation>
.
</paragraph>
<paragraph id="B07C0F54FFA9FFB6FF56A9062F57F850" blockId="25.[151,1436,1687,2037]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56A9062F46F8C0" bold="true" box="[199,358,1867,1893]" pageId="25" pageNumber="204">Colour in life</emphasis>
. Background translucent whitish; carapace with small red patch between eyes, numerous white, star-shaped chromatophores covering most of dorsal surface, and broad, red patch on each flank, at about middle of carapace; most of pleon densely covered with white chromatophores, ventral pleura of third pleonite and most of fourth pleonite deep red; chelipeds and remaining appendages hyaline whitish; eggs yolky yellow (
<figureCitation id="28F813D1FFA9FFB6FA9DA9FA2B7FF874" box="[1292,1375,1975,2001]" captionStart="FIGURE 14" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[170,1417,181,1884]" captionTargetId="figure-19@27.[170,1417,181,1884]" captionTargetPageId="27" captionText="FIGURE 14. Athanopsis tarahomii Marin, Sheibani &amp; Sari, 2014, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350819" httpUri="https://zenodo.org/record/10350819/files/figure.png" pageId="25" pageNumber="204">Fig. 14</figureCitation>
). See also remarks below.
</paragraph>
</subSubSection>
<caption id="E4BC5FDCFFAAFFB5FF06A9212DECF869" ID-DOI="http://doi.org/10.5281/zenodo.10350817" ID-Zenodo-Dep="10350817" httpUri="https://zenodo.org/record/10350817/files/figure.png" pageId="26" pageNumber="205" startId="26.[151,250,1900,1924]" targetBox="[151,1436,181,1876]" targetPageId="26" targetType="figure">
<paragraph id="B07C0F54FFAAFFB5FF06A9212DECF869" blockId="26.[151,1437,1900,1996]" pageId="26" pageNumber="205">
<emphasis id="82B7D346FFAAFFB5FF06A9212F00F821" bold="true" box="[151,288,1900,1924]" pageId="26" pageNumber="205">FIGURE 13.</emphasis>
<taxonomicName id="77C374D7FFAAFFB5FEB7A9212CE8F821" authority="Berggren, 1991" authorityName="Berggren" authorityYear="1991" box="[294,712,1900,1925]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="205" phylum="Arthropoda" rank="species" species="rubricinctuta">
<emphasis id="82B7D346FFAAFFB5FEB7A9212C03F821" box="[294,547,1900,1924]" italics="true" pageId="26" pageNumber="205">Athanopsis rubricinctuta</emphasis>
<bibRefCitation id="D45272A5FFAAFFB5FDB8A9202CE8F821" author="Berggren, M." box="[553,712,1900,1925]" pageId="26" pageNumber="205" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren, 1991</bibRefCitation>
</taxonomicName>
, male (cl 5.4 mm) from Masirah Island, Oman (FLMNH UF 71437) [a, b]; male (cl 5.8 mm) from the same locality (FLMNH UF 71443) [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, shrimp in life (different individual), lateral (left side). Photographs by the author.
</paragraph>
</caption>
<caption id="E4BC5FDCFFABFFB4FF06A9382EFEF85C" ID-DOI="http://doi.org/10.5281/zenodo.10350819" ID-Zenodo-Dep="10350819" httpUri="https://zenodo.org/record/10350819/files/figure.png" pageId="27" pageNumber="206" startId="27.[151,250,1909,1933]" targetBox="[170,1417,181,1884]" targetPageId="27" targetType="figure">
<paragraph id="B07C0F54FFABFFB4FF06A9382EFEF85C" blockId="27.[151,1436,1909,2041]" pageId="27" pageNumber="206">
<emphasis id="82B7D346FFABFFB4FF06A9382F3FF828" bold="true" box="[151,287,1909,1933]" pageId="27" pageNumber="206">FIGURE 14.</emphasis>
<taxonomicName id="77C374D7FFABFFB4FEB5A9382D0EF828" authority="Marin, Sheibani &amp; Sari, 2014" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[292,814,1909,1933]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="206" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFABFFB4FEB5A9382FDEF828" box="[292,510,1909,1933]" italics="true" pageId="27" pageNumber="206">Athanopsis tarahomii</emphasis>
<bibRefCitation id="D45272A5FFABFFB4FD95A9382D0EF828" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[516,814,1909,1933]" pageId="27" pageNumber="206" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">Marin, Sheibani &amp; Sari, 2014</bibRefCitation>
</taxonomicName>
, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author.
</paragraph>
</caption>
<subSubSection id="F8D95CDFFFACFFB3FF56AEDA2C6BFF14" box="[199,587,150,177]" pageId="28" pageNumber="207" type="materials_examined">
<paragraph id="B07C0F54FFACFFB3FF56AEDA2C6BFF14" blockId="28.[151,1437,150,933]" box="[199,587,150,177]" pageId="28" pageNumber="207">
<emphasis id="82B7D346FFACFFB3FF56AEDA2F40FF15" bold="true" box="[199,352,150,177]" pageId="28" pageNumber="207">
<typeStatus id="6F78B1F6FFACFFB3FF56AEDA2F22FF14" box="[199,258,151,177]" pageId="28" pageNumber="207">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="C8D44FC4FFACFFB3FEFFAEDA2C2BFF14" box="[366,523,151,177]" name="Qeshm Island" pageId="28" pageNumber="207">Qeshm Island</collectingCountry>
,
<collectingCountry id="C8D44FC4FFACFFB3FD86AEDA2C68FF14" box="[535,584,151,177]" name="Iran" pageId="28" pageNumber="207">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFACFFB3FF56AEF72FFEFF5C" pageId="28" pageNumber="207" type="distribution">
<paragraph id="B07C0F54FFACFFB3FF56AEF72FFEFF5C" blockId="28.[151,1437,150,933]" pageId="28" pageNumber="207">
<emphasis id="82B7D346FFACFFB3FF56AEF72F79FF71" bold="true" box="[199,345,186,212]" pageId="28" pageNumber="207">Distribution</emphasis>
. Currently known only from the
<typeStatus id="6F78B1F6FFACFFB3FD5AAEF62CDBFF70" box="[715,763,187,213]" pageId="28" pageNumber="207">type</typeStatus>
locality in
<collectingCountry id="C8D44FC4FFACFFB3FCEBAEF62D87FF70" box="[890,935,187,213]" name="Iran" pageId="28" pageNumber="207">Iran</collectingCountry>
(
<bibRefCitation id="D45272A5FFACFFB3FC24AEF62A5BFF70" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[949,1147,187,213]" pageId="28" pageNumber="207" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFACFFB3FB93AEF62A13FF70" box="[1026,1075,187,213]" italics="true" pageId="28" pageNumber="207">et al</emphasis>
. 2014
</bibRefCitation>
) and the Red Sea coast of
<collectingCountry id="C8D44FC4FFACFFB3FF06AE922F08FF5C" box="[151,296,223,249]" name="Saudi Arabia" pageId="28" pageNumber="207">Saudi Arabia</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFACFFB3FF56AF4E2C3BFE2C" pageId="28" pageNumber="207" type="biology_ecology">
<paragraph id="B07C0F54FFACFFB3FF56AF4E2C3BFE2C" blockId="28.[151,1437,150,933]" pageId="28" pageNumber="207">
<emphasis id="82B7D346FFACFFB3FF56AF4E2F06FEB8" bold="true" box="[199,294,259,285]" pageId="28" pageNumber="207">Ecology</emphasis>
. The two Saudi Arabian specimens were extracted from the burrow of a large echiuran (probably
<taxonomicName id="77C374D7FFACFFB3FF06AF6A2F74FEE4" box="[151,340,295,321]" class="Polychaeta" family="Thalassematidae" genus="Ochetostoma" kingdom="Animalia" order="Echiuroidea" pageId="28" pageNumber="207" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="82B7D346FFACFFB3FF06AF6A2F0BFEE4" box="[151,299,295,321]" italics="true" pageId="28" pageNumber="207">Ochetostoma</emphasis>
sp.
</taxonomicName>
) on a shallow sandflat with large patches of seagrass and some fragmented rubble, at a depth of about
<specimenCount id="A6C5C4DDFFACFFB3FF4DAF062F2DFEC0" box="[220,269,331,357]" count="1" pageId="28" pageNumber="207" type="generic">
<quantity id="773BA2B1FFACFFB3FF4DAF062F29FEC0" box="[220,265,331,357]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="207" unit="m" value="1.0">1 m</quantity>
.
</specimenCount>
The Iranian
<typeStatus id="6F78B1F6FFACFFB3FE0BAF062FEAFEC0" box="[410,458,331,357]" pageId="28" pageNumber="207">type</typeStatus>
specimens were also found in association with echiurans, in a “cobble/rocky area under large boulder” (
<bibRefCitation id="D45272A5FFACFFB3FED7AF222C2FFE2C" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[326,527,367,393]" pageId="28" pageNumber="207" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFACFFB3FE02AF222FE5FE2C" box="[403,453,367,393]" italics="true" pageId="28" pageNumber="207">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFACFFB3FF56AFDE2C1EFC00" pageId="28" pageNumber="207" type="discussion">
<paragraph id="B07C0F54FFACFFB3FF56AFDE2CFEFD98" blockId="28.[151,1437,150,933]" pageId="28" pageNumber="207">
<emphasis id="82B7D346FFACFFB3FF56AFDE2F14FE08" bold="true" box="[199,308,403,429]" pageId="28" pageNumber="207">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFACFFB3FED0AFDE2C11FE08" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[321,561,403,429]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFACFFB3FED0AFDE2C11FE08" box="[321,561,403,429]" italics="true" pageId="28" pageNumber="207">Athanopsis tarahomii</emphasis>
</taxonomicName>
was hitherto known only from
<specimenCount id="A6C5C4DDFFACFFB3FC1BAFDE2DCAFE08" box="[906,1002,403,429]" count="2" pageId="28" pageNumber="207" type="generic" typeStatus="type">two type</specimenCount>
specimens from
<collectingCountry id="C8D44FC4FFACFFB3FB36AFDE2B63FE08" box="[1191,1347,403,429]" name="Qeshm Island" pageId="28" pageNumber="207">Qeshm Island</collectingCountry>
,
<collectingCountry id="C8D44FC4FFACFFB3FADCAFDE2B5DFE08" box="[1357,1405,403,429]" name="Iran" pageId="28" pageNumber="207">Iran</collectingCountry>
, in the Strait of
<collectingCountry id="C8D44FC4FFACFFB3FE8EAFFA2F5EFE74" box="[287,382,439,465]" name="Iran" pageId="28" pageNumber="207">Hormuz</collectingCountry>
. The present Saudi Arabian material of
<taxonomicName id="77C374D7FFACFFB3FCA2AFFA2D9FFE74" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[819,959,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFACFFB3FCA2AFFA2D9FFE74" box="[819,959,439,465]" italics="true" pageId="28" pageNumber="207">A. tarahomii</emphasis>
</taxonomicName>
significantly extends its previously known range from the Arabian-Persian Gulf to the central Red Sea. The two Red Sea specimens agree well with the
<typeStatus id="6F78B1F6FFACFFB3FAFDAF962BBCFE50" box="[1388,1436,475,501]" pageId="28" pageNumber="207">type</typeStatus>
material, including the very characteristic descendant and distally notched rostrum, although not as pronounced as in the
<typeStatus id="6F78B1F6FFACFFB3FF4CAC6E2F2DFD98" box="[221,269,547,573]" pageId="28" pageNumber="207">type</typeStatus>
specimens (
<bibRefCitation id="D45272A5FFACFFB3FE09AC6E2C40FD98" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[408,608,547,573]" pageId="28" pageNumber="207" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFACFFB3FE74AC6E2C36FD98" box="[485,534,547,573]" italics="true" pageId="28" pageNumber="207">et al</emphasis>
. 2014
</bibRefCitation>
: fig. 4d, e).
</paragraph>
<paragraph id="B07C0F54FFACFFB3FF56AC0A2C1EFC00" blockId="28.[151,1437,150,933]" pageId="28" pageNumber="207">
The colour pattern of
<taxonomicName id="77C374D7FFACFFB3FE57AC0A2C76FDC4" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[454,598,583,609]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFACFFB3FE57AC0A2C76FDC4" box="[454,598,583,609]" italics="true" pageId="28" pageNumber="207">A. tarahomii</emphasis>
</taxonomicName>
is herein illustrated for the first time (
<figureCitation id="28F813D1FFACFFB3FB89AC0A2A4CFDC4" box="[1048,1132,583,609]" captionStart="FIGURE 14" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[170,1417,181,1884]" captionTargetId="figure-19@27.[170,1417,181,1884]" captionTargetPageId="27" captionText="FIGURE 14. Athanopsis tarahomii Marin, Sheibani &amp; Sari, 2014, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350819" httpUri="https://zenodo.org/record/10350819/files/figure.png" pageId="28" pageNumber="207">Fig. 14</figureCitation>
) and is diagnostic for this species, despite possessing some similarities with the colour patterns of
<taxonomicName id="77C374D7FFACFFB3FC5DAC262A4DFD20" authorityName="Banner &amp; Banner" authorityYear="1981" box="[972,1133,619,645]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="82B7D346FFACFFB3FC5DAC262A4DFD20" box="[972,1133,619,645]" italics="true" pageId="28" pageNumber="207">A. brevirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFACFFB3FB37AC262B31FD21" authorityName="Anker" authorityYear="2011" box="[1190,1297,619,645]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="saurus">
<emphasis id="82B7D346FFACFFB3FB37AC262B31FD21" box="[1190,1297,619,645]" italics="true" pageId="28" pageNumber="207">A. saurus</emphasis>
</taxonomicName>
(cf.
<figureCitation id="28F813D1FFACFFB3FAD9AC262BB6FD20" box="[1352,1430,619,645]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner &amp; Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="28" pageNumber="207">Figs. 8</figureCitation>
,
<figureCitation id="28F813D1FFACFFB3FF06ACC22E87FD0C" box="[151,167,655,681]" captionStart="FIGURE 9" captionStartId="18.[151,250,1878,1902]" captionTargetBox="[151,1436,151,1860]" captionTargetId="figure-34@18.[151,1436,151,1860]" captionTargetPageId="18" captionText="FIGURE 9. Athanopsis saurus Anker, 2011, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [ae] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably Ochetostoma sp. (Thalassematidae), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350801" httpUri="https://zenodo.org/record/10350801/files/figure.png" pageId="28" pageNumber="207">9</figureCitation>
). The description of the colour pattern of
<taxonomicName id="77C374D7FFACFFB3FD12ACC22D32FD0C" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[643,786,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFACFFB3FD12ACC22D32FD0C" box="[643,786,655,681]" italics="true" pageId="28" pageNumber="207">A. tarahomii</emphasis>
</taxonomicName>
and the accompanying photograph in
<bibRefCitation id="D45272A5FFACFFB3FB57ACC22BB9FD0C" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[1222,1433,655,681]" pageId="28" pageNumber="207" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFACFFB3FA83ACC22B6BFD0C" box="[1298,1355,655,681]" italics="true" pageId="28" pageNumber="207">et al.</emphasis>
(2014
</bibRefCitation>
: fig. 7) are erroneous and actually refer to
<taxonomicName id="77C374D7FFACFFB3FDE2ACFE2A49FD68" authority="(Coutiere, 1896)" baseAuthorityName="Coutiere" baseAuthorityYear="1896" box="[627,1129,691,717]" class="Malacostraca" family="Alpheidae" genus="Amphibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="jousseaumei">
<emphasis id="82B7D346FFACFFB3FDE2ACFE2D84FD69" box="[627,932,691,717]" italics="true" pageId="28" pageNumber="207">Amphibetaeus jousseaumei</emphasis>
(
<bibRefCitation id="D45272A5FFACFFB3FC25ACFE2A41FD68" author="Coutiere, H." box="[948,1121,691,717]" pageId="28" pageNumber="207" pagination="313 - 317" refId="ref21438" refString="Coutiere, H. (1896) Note sur un nouvel Alpheide, Betaeus Jousseaumei [Crust.]. Bulletin de la Societe Entomologique de France, 13, 313 - 317. https: // doi. org / 10.3406 / bsef. 1896.21892" type="journal article" year="1896">Coutière, 1896</bibRefCitation>
)
</taxonomicName>
, which was reported in the same study. The present author is in the possession of a colour photograph showing two individuals (presumably type specimens) of the real
<taxonomicName id="77C374D7FFACFFB3FE5DACB62C79FCB0" authorityName="Marin, Sheibani &amp; Sari" authorityYear="2014" box="[460,601,763,789]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="tarahomii">
<emphasis id="82B7D346FFACFFB3FE5DACB62C79FCB0" box="[460,601,763,789]" italics="true" pageId="28" pageNumber="207">A. tarahomii</emphasis>
</taxonomicName>
from
<collectingCountry id="C8D44FC4FFACFFB3FD0CACB62D18FCB0" box="[669,824,763,789]" name="Qeshm Island" pageId="28" pageNumber="207">Qeshm Island</collectingCountry>
[sent by R. Sheibani, the second author in
<bibRefCitation id="D45272A5FFACFFB3FA88ACB62EC2FC9C" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." pageId="28" pageNumber="207" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFACFFB3FAF5ACB62BB5FCB0" box="[1380,1429,763,789]" italics="true" pageId="28" pageNumber="207">et al</emphasis>
. (2014)
</bibRefCitation>
well before the publication of their study), which confirms that the colour patterns of the Iranian and Saudi Arabian specimens are almost identical. Another error in
<bibRefCitation id="D45272A5FFACFFB3FC86AD0E2DCCFCF8" author="Marin, I. &amp; Sheibani, R. &amp; Sari, A." box="[791,1004,835,861]" pageId="28" pageNumber="207" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. &amp; Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
Marin
<emphasis id="82B7D346FFACFFB3FCF2AD0E2DBBFCF8" box="[867,923,835,861]" italics="true" pageId="28" pageNumber="207">et al.</emphasis>
(2014)
</bibRefCitation>
is the invalid designation of a
<typeStatus id="6F78B1F6FFACFFB3FAD5AD0E2BBDFCF8" box="[1348,1437,835,861]" pageId="28" pageNumber="207" type="neotype">neotype</typeStatus>
for
<taxonomicName id="77C374D7FFACFFB3FF51AD2A2F4AFC25" baseAuthorityName="Coutiere" baseAuthorityYear="1896" box="[192,362,871,897]" class="Malacostraca" family="Alpheidae" genus="Amphibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="28" pageNumber="207" phylum="Arthropoda" rank="species" species="jousseaumei">
<emphasis id="82B7D346FFACFFB3FF51AD2A2F4AFC25" box="[192,362,871,897]" italics="true" pageId="28" pageNumber="207">A. jousseaumei</emphasis>
</taxonomicName>
(SMF-43232, deposited in the Senckenberg Naturmuseum, Frankfurt) as a
<typeStatus id="6F78B1F6FFACFFB3FB54AD2A2B0CFC24" box="[1221,1324,871,897]" pageId="28" pageNumber="207" type="lectotype">lectotype</typeStatus>
exists for this species (see
<bibRefCitation id="D45272A5FFACFFB3FEC0ADC62C12FC00" author="Anker, A. &amp; Jeng, M. S." box="[337,562,907,933]" pageId="28" pageNumber="207" pagination="379 - 391" refId="ref20138" refString="Anker, A. &amp; Jeng, M. S. (2006) Richalpheus palmeri, n. gen., n. sp., an infaunal alpheid shrimp from the Philippines, with redescription of Amphibetaeus jousseaumei Coutiere, 1896 (Decapoda: Caridea). Journal of Crustacean Biology, 26, 379 - 391. https: // doi. org / 10.1651 / S- 2647.1" type="journal article" year="2006">Anker &amp; Jeng 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>