186 lines
22 KiB
XML
186 lines
22 KiB
XML
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<mods:title id="6D644A39F6E323ACB34198B6188126B4">Diversity and morphology of Eocene and Oligocene Mordellidae (Coleoptera)</mods:title>
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<mods:namePart id="DE8C7BCBBDE30A78284EFB4E6ADCFF37">Batelka, Jan</mods:namePart>
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<mods:namePart id="5258ED74AF64A5AB385CE28B5D5413A4">Rosová, Kateřina</mods:namePart>
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<mods:namePart id="EA0587A6B979D761AD38330D590977E2">Prokop, Jakub</mods:namePart>
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<mods:title id="789BE24A2F0C920184B81614F12805C1">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DF6FD65350ADA59F4BC5E7C8C4E76B39">2023</mods:date>
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<paragraph id="8B8CCF4EF26D8C67FF35FC4BC1E4FBBA" blockId="4.[178,733,1019,1075]" pageId="4" pageNumber="455">
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<heading id="D0C47822F26D8C67FF35FC4BC1E4FBBA" centered="true" fontSize="10" level="2" pageId="4" pageNumber="455" reason="3">
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<emphasis id="B947135CF26D8C67FF35FC4BC1E4FBBA" bold="true" pageId="4" pageNumber="455">
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<taxonomicName id="4C33B4CDF26D8C67FF35FC4BC163FBBA" authority="(Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust, 2018)" authorityName="Batelka & Rosová & Prokop" authorityYear="2023" baseAuthorityName="Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust" baseAuthorityYear="2018" class="Insecta" family="Mordellidae" genus="Baltimorda" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="succinea" status="comb. nov.">
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<emphasis id="B947135CF26D8C67FF35FC4BC2A5FB9A" bold="true" box="[178,405,1019,1043]" italics="true" pageId="4" pageNumber="455">Baltimorda succinea</emphasis>
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(
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<bibRefCitation id="EFA2B2BFF26D8C67FE23FC4CC17CFBBA" author="BAO T. & WALCZYNSKA K. S. & BOJARSKI B. & JARZEMBOWSKI E. & WANG B. & RUST J." pageId="4" pageNumber="455" pagination="31 - 36" refId="ref19861" refString="BAO T., WALCZYNSKA K. S., BOJARSKI B., JARZEMBOWSKI E., WANG B. & RUST J. 2018: A new species of tumbling flower beetle (Coleoptera: Mordellidae) from Baltic amber. Palaontologische Zeitschrift 93 (14): 31 - 36." type="journal article" year="2018">Bao, Walczyńska, Bojarski, Jarzembowski, Wang & Rust, 2018</bibRefCitation>
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)
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</taxonomicName>
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||
<taxonomicNameLabel id="A274AE27F26D8C67FDDDFBACC1E4FBBA" box="[602,724,1052,1075]" pageId="4" pageNumber="455" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3299CC5F26D8C67FF10FBF2C14EFBF9" pageId="4" pageNumber="455" type="reference_group">
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<paragraph id="8B8CCF4EF26D8C67FF10FBF2C14EFBF9" blockId="4.[151,760,1090,1136]" pageId="4" pageNumber="455">
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<treatmentCitationGroup id="AB23E860F26D8C67FF10FBF2C14EFBF9" pageId="4" pageNumber="455">
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<treatmentCitation id="0A92E95FF26D8C67FF10FBF2C205FBF9" author="BAO T. & WALCZYNSKA K. S. & BOJARSKI B. & JARZEMBOWSKI E. & WANG B. & RUST J." page="32" pageId="4" pageNumber="455" year="2018">
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<taxonomicName id="4C33B4CDF26D8C67FF10FBF2C205FBF9" authority="Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust, 2018: 32" authorityName="Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust" authorityPageNumber="32" authorityYear="2018" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="succinea">
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<emphasis id="B947135CF26D8C67FF10FBF2C201FBDC" box="[151,305,1090,1109]" italics="true" pageId="4" pageNumber="455">Tomoxia succinea</emphasis>
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<bibRefCitation id="EFA2B2BFF26D8C67FEB0FBF2C205FBF9" author="BAO T. & WALCZYNSKA K. S. & BOJARSKI B. & JARZEMBOWSKI E. & WANG B. & RUST J." pageId="4" pageNumber="455" pagination="31 - 36" refId="ref19861" refString="BAO T., WALCZYNSKA K. S., BOJARSKI B., JARZEMBOWSKI E., WANG B. & RUST J. 2018: A new species of tumbling flower beetle (Coleoptera: Mordellidae) from Baltic amber. Palaontologische Zeitschrift 93 (14): 31 - 36." type="journal article" year="2018">Bao, Walczyńska, Bojarski, Jarzembowski, Wang & Rust, 2018: 32</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="548871ECF26D8C67FEBAFBEDC2BCFBF9" box="[317,396,1117,1136]" pageId="4" pageNumber="455" type="holotype">Holotype</typeStatus>
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and
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<typeStatus id="548871ECF26D8C67FE31FBEDC133FBF9" box="[438,515,1117,1136]" pageId="4" pageNumber="455" type="paratype">paratype</typeStatus>
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: Baltic amber.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3299CC5F26D8C67FF10FB38C67DFD9C" pageId="4" pageNumber="455" type="discussion">
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<paragraph id="8B8CCF4EF26D8C67FF10FB38C6ADFE3C" blockId="4.[151,760,1160,2047]" lastBlockId="4.[826,1437,190,533]" pageId="4" pageNumber="455">
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<emphasis id="B947135CF26D8C67FF10FB38C2E6FB16" bold="true" box="[151,470,1160,1183]" pageId="4" pageNumber="455">Comments on identification.</emphasis>
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The examination of figures and description of
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<taxonomicName id="4C33B4CDF26D8C67FEE7FB18C12FFB36" authorityName="Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust" authorityYear="2018" box="[352,543,1192,1215]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="succinea">
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<emphasis id="B947135CF26D8C67FEE7FB18C12FFB36" box="[352,543,1192,1215]" italics="true" pageId="4" pageNumber="455">Tomoxia succinea</emphasis>
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||
</taxonomicName>
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||
provided by
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||
<bibRefCitation id="EFA2B2BFF26D8C67FD28FB18C3CFFB56" author="BAO T. & WALCZYNSKA K. S. & BOJARSKI B. & JARZEMBOWSKI E. & WANG B. & RUST J." pageId="4" pageNumber="455" pagination="31 - 36" refId="ref19861" refString="BAO T., WALCZYNSKA K. S., BOJARSKI B., JARZEMBOWSKI E., WANG B. & RUST J. 2018: A new species of tumbling flower beetle (Coleoptera: Mordellidae) from Baltic amber. Palaontologische Zeitschrift 93 (14): 31 - 36." type="journal article" year="2018">BAO et al. (2018)</bibRefCitation>
|
||
indicates that this species is not associated with the genus
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<taxonomicName id="4C33B4CDF26D8C67FE8DFB58C255FB76" box="[266,357,1256,1279]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B947135CF26D8C67FE8DFB58C255FB76" box="[266,357,1256,1279]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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</taxonomicName>
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, because this placement is based on erroneous arguments and poor knowledge of the generic delimitations within the
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<taxonomicName id="4C33B4CDF26D8C67FE22FA98C12BFAB6" authorityName="Latreille" authorityYear="1802" box="[421,539,1320,1343]" class="Insecta" family="Mordellidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="tribe" tribe="Mordellini">Mordellini</taxonomicName>
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. The diagnosis they provide is as follows: ‘Diagnosis (emended). The diagnostic characters for the genus
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<taxonomicName id="4C33B4CDF26D8C67FE6AFAD8C178FAF6" box="[493,584,1384,1407]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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<emphasis id="B947135CF26D8C67FE6AFAD8C178FAF6" box="[493,584,1384,1407]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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</taxonomicName>
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when compared to the other members of the tribe
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<taxonomicName id="4C33B4CDF26D8C67FE7CFA38C15DFA16" authorityName="Latreille" authorityYear="1802" box="[507,621,1416,1439]" class="Insecta" family="Mordellidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="tribe" tribe="Mordellini">Mordellini</taxonomicName>
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are: (1) eyes finely granulated; (2) scutellum quadrilateral or irregular shaped; (3) hind tibia with a fine dorsal ridge and a lateral ridge; basal tarsomere of hindleg also with a fine dorsal ridge (SMITH 1882,
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<bibRefCitation id="EFA2B2BFF26D8C67FEEDF9B8C128F996" author="LILJEBLAD E." box="[362,536,1544,1567]" pageId="4" pageNumber="455" pagination="1 - 229" refId="ref21721" refString="LILJEBLAD E. 1945: Monograph of the family Mordellidae (Coleoptera) of North America, north of Mexico. Miscellaneous Publications, Museum of Zoology, University of Michigan 62: 1 - 229." type="journal article" year="1945">LILJEBLAD 1945</bibRefCitation>
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, FRANCISCOLO 1965, JACKMAN & LU 2001).’ While points (1) and (2) do not provide any information relevant to the definition of the genus (e.g., eyes with small facets are present in all three genera included in the informal
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<taxonomicName id="4C33B4CDF26D8C67FE64F938C10EF916" box="[483,574,1672,1695]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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<emphasis id="B947135CF26D8C67FE64F938C10EF916" box="[483,574,1672,1695]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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</taxonomicName>
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complex (
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<bibRefCitation id="EFA2B2BFF26D8C67FD2DF938C3FEF936" author="HORAK J." pageId="4" pageNumber="455" pagination="51 - 58" refId="ref21235" refString="HORAK J. 2007: First records of genera Tomoxioda Ermisch and Paratomoxioda Ermisch (Coleoptera: Mordellidae) from Palaearctic region. Studies and Reports of District Museum Prague-East, Taxonomical Series 3 (1 - 2): 51 - 58." type="book chapter" year="2007">HORÁK 2007</bibRefCitation>
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) and in many other
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<taxonomicName id="4C33B4CDF26D8C67FE1BF918C122F936" authorityName="Latreille" authorityYear="1802" box="[412,530,1704,1727]" class="Insecta" family="Mordellidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="tribe" tribe="Mordellini">Mordellini</taxonomicName>
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), point (3) is incorrect
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<emphasis id="B947135CF26D8C67FF10F978C3EBF956" box="[151,219,1736,1759]" italics="true" pageId="4" pageNumber="455">per se</emphasis>
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because in
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<taxonomicName id="4C33B4CDF26D8C67FEE4F978C28EF956" box="[355,446,1736,1759]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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<emphasis id="B947135CF26D8C67FEE4F978C28EF956" box="[355,446,1736,1759]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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</taxonomicName>
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, a dorsolateral ridge (dorsal ridge sensu
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<bibRefCitation id="EFA2B2BFF26D8C67FE95F958C281F976" author="BAO T. & WALCZYNSKA K. S. & BOJARSKI B. & JARZEMBOWSKI E. & WANG B. & RUST J." box="[274,433,1768,1791]" pageId="4" pageNumber="455" pagination="31 - 36" refId="ref19861" refString="BAO T., WALCZYNSKA K. S., BOJARSKI B., JARZEMBOWSKI E., WANG B. & RUST J. 2018: A new species of tumbling flower beetle (Coleoptera: Mordellidae) from Baltic amber. Palaontologische Zeitschrift 93 (14): 31 - 36." type="journal article" year="2018">BAO et al. 2018</bibRefCitation>
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) is present on the metatibia, but absent on the first metatarsomere.According to the generic key to the
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<taxonomicName id="4C33B4CDF26D8C67FE8EF898C254F8B6" box="[265,356,1832,1855]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B947135CF26D8C67FE8EF898C254F8B6" box="[265,356,1832,1855]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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||
</taxonomicName>
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complex provided by
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<bibRefCitation id="EFA2B2BFF26D8C67FDD1F898C1C4F8B6" author="HORAK J." box="[598,756,1832,1855]" pageId="4" pageNumber="455" pagination="51 - 58" refId="ref21235" refString="HORAK J. 2007: First records of genera Tomoxioda Ermisch and Paratomoxioda Ermisch (Coleoptera: Mordellidae) from Palaearctic region. Studies and Reports of District Museum Prague-East, Taxonomical Series 3 (1 - 2): 51 - 58." type="book chapter" year="2007">HORÁK (2007)</bibRefCitation>
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, the genus
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<taxonomicName id="4C33B4CDF26D8C67FE8AF8F8C25AF8D6" box="[269,362,1864,1887]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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<emphasis id="B947135CF26D8C67FE8AF8F8C25AF8D6" box="[269,362,1864,1887]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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</taxonomicName>
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is defined as follows: ‘Hind tibiae besides preapical ridge with fine dorsolateral ridge. First tarsomere of posterior tarsus
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<emphasis id="B947135CF26D8C67FE4DF838C1C1F816" bold="true" box="[458,753,1928,1951]" pageId="4" pageNumber="455">without dorsolateral ridge</emphasis>
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. Eyes very finely facetted, with interfacetal setae.’ In the description of eyes of
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<taxonomicName id="4C33B4CDF26D8C67FEF9F878C10BF856" authorityName="Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust" authorityYear="2018" box="[382,571,1992,2015]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="succinea">
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<emphasis id="B947135CF26D8C67FEF9F878C10BF856" box="[382,571,1992,2015]" italics="true" pageId="4" pageNumber="455">Tomoxia succinea</emphasis>
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</taxonomicName>
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the authors stated those are ‘large, ellipsoidal, finely faceted and granulated, apparently not reaching occiput, hairy.’ This term likely indicates the presence of interfacetal setae, which are present in
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<taxonomicName id="4C33B4CDF26D8C67FCD1FF4EC081FE9C" box="[854,945,254,277]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
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<emphasis id="B947135CF26D8C67FCD1FF4EC081FE9C" box="[854,945,254,277]" italics="true" pageId="4" pageNumber="455">Tomoxia</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="4C33B4CDF26D8C67FC65FF4EC768FE9C" authorityName="Ermisch" authorityYear="1950" box="[994,1112,254,277]" class="Insecta" family="Mordellidae" genus="Tomoxioda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947135CF26D8C67FC65FF4EC768FE9C" box="[994,1112,254,277]" italics="true" pageId="4" pageNumber="455">Tomoxioda</emphasis>
|
||
</taxonomicName>
|
||
, but absent in
|
||
<taxonomicName id="4C33B4CDF26D8C67FB77FF4EC6A6FE9C" authorityName="Ermisch" authorityYear="1954" box="[1264,1430,254,277]" class="Insecta" family="Mordellidae" genus="Paratomoxioda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947135CF26D8C67FB77FF4EC6A6FE9C" box="[1264,1430,254,277]" italics="true" pageId="4" pageNumber="455">Paratomoxioda</emphasis>
|
||
</taxonomicName>
|
||
. The combination of eyes with small facets covered with setae and presence of dorsolateral ridges on hind tibia and first metatarsomere conforms with the diagnosis of the genus
|
||
<taxonomicName id="4C33B4CDF26D8C67FC05FECEC7DEFE1C" authority="Ermisch, 1968" authorityName="Ermisch" authorityYear="1968" box="[898,1262,382,405]" class="Insecta" family="Mordellidae" genus="Klapperichimorda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947135CF26D8C67FC05FECEC776FE1C" box="[898,1094,382,405]" italics="true" pageId="4" pageNumber="455">Klapperichimorda</emphasis>
|
||
<bibRefCitation id="EFA2B2BFF26D8C67FBC9FECEC7DEFE1C" author="ERMISCH K." box="[1102,1262,382,405]" pageId="4" pageNumber="455" pagination="279 - 292" refId="ref20313" refString="ERMISCH K. 1968: Neue Mordellini aus der chinensischen Provinz Fukien. Reichenbachia 10: 279 - 292." type="journal article" year="1968">Ermisch, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
, which contains four
|
||
<collectingRegion id="49F701ACF26D8C67FCEAFE2EC0F4FE3C" box="[877,964,414,437]" country="Democratic Republic of the Congo" name="Orientale" pageId="4" pageNumber="455">Oriental</collectingRegion>
|
||
species (
|
||
<bibRefCitation id="EFA2B2BFF26D8C67FBA5FE2EC78DFE3C" author="ERMISCH K." box="[1058,1213,414,437]" pageId="4" pageNumber="455" pagination="279 - 292" refId="ref20313" refString="ERMISCH K. 1968: Neue Mordellini aus der chinensischen Provinz Fukien. Reichenbachia 10: 279 - 292." type="journal article" year="1968">ERMISCH 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA2B2BFF26D8C67FB41FE2EC67FFE3C" author="HORAK J." box="[1222,1359,414,437]" pageId="4" pageNumber="455" pagination="153 - 164" refId="ref21176" refString="HORAK J. 1996: Revision of some Oriental Mordellini with description of three new species. Part 2. Acta Societatis Zoologicae Bohemicae 60: 153 - 164." type="journal article" year="1996">HORÁK 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA2B2BFF26D8C67FADFFE2EC6A2FE3C" author="HORAK J." box="[1368,1426,414,437]" pageId="4" pageNumber="455" pagination="79 - 104" refId="ref21325" refString="HORAK J. 2020: Mordellidae. Pp. 79 - 104. In: IWAN D. & LOBL I. (eds): Catalogue of Palaearctic Coleoptera. Volume 5. Tenebrionoidea. Revised and updated second edition. Brill, Leiden, Boston, xxvi + 945 pp." type="book chapter" year="2020">2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8CCF4EF26D8C67FCDDFE0EC67DFD9C" blockId="4.[826,1437,190,533]" pageId="4" pageNumber="455">
|
||
Because of the uncertain generic identity of the
|
||
<typeStatus id="548871ECF26D8C67FAC6FE0EC6ADFE5C" box="[1345,1437,446,469]" pageId="4" pageNumber="455" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="548871ECF26D8C67FCEEFE6EC0F5FE7C" box="[873,965,478,501]" pageId="4" pageNumber="455" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C33B4CDF26D8C67FC68FE6EC780FE7C" authorityName="Bao, Walczynska, Bojarski, Jarzembowski, Wang & Rust" authorityYear="2018" box="[1007,1200,478,501]" class="Insecta" family="Mordellidae" genus="Tomoxia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="succinea">
|
||
<emphasis id="B947135CF26D8C67FC68FE6EC780FE7C" box="[1007,1200,478,501]" italics="true" pageId="4" pageNumber="455">Tomoxia succinea</emphasis>
|
||
</taxonomicName>
|
||
, this species is transferred here into
|
||
<taxonomicName id="4C33B4CDF26D8C67FC60FE4EC750FD9C" box="[999,1120,510,533]" class="Insecta" family="Mordellidae" genus="Baltimorda" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947135CF26D8C67FC60FE4EC750FD9C" box="[999,1120,510,533]" italics="true" pageId="4" pageNumber="455">Baltimorda</emphasis>
|
||
</taxonomicName>
|
||
new collective group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |