309 lines
31 KiB
XML
309 lines
31 KiB
XML
<document id="19A9F3CD6B0CBDBA6FA496F3739FDE23" ID-DOI="10.1649/072.065.0203" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102619" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630154770511" checkinUser="diego" docAuthor="Ratcliffe, Brett C." docDate="2011" docId="03E887B2FF9353593DC2FB82BFEFFEB1" docLanguage="en" docName="072.065.0203.pdf.imf" docOrigin="The Coleopterists Bulletin 65 (2)" docSource="http://dx.doi.org/10.1649/072.065.0203" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Phileurus zonianis Ratcliffe 2011, new species" docType="treatment" docVersion="3" lastPageNumber="129" masterDocId="FFD1FFCAFF91535D3F25FFABBD7AFFED" masterDocTitle="A Review of the Species ofPhileurusLatreille (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with Quadridentate Protibiae and Description of a New Species from Panama" masterLastPageNumber="129" masterPageNumber="125" pageNumber="127" updateTime="1699621880442" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="C28318AE5B7A22F62CE5FBEF7AA7A3B6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C856770EE5137BDE3CA49A473780C447">
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<mods:title id="6BB6935E08AA10B2234F7C375C6CEF53">A Review of the Species ofPhileurusLatreille (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) with Quadridentate Protibiae and Description of a New Species from Panama</mods:title>
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</mods:titleInfo>
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<mods:name id="9F6B2BD23B3734548A0D282800442B03" type="personal">
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<mods:role id="EE46A5FFFC6DE5A44E5C563566D2124D">
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<mods:roleTerm id="2664087DF7B4EA85AFE12E7BF18CEB97">Author</mods:roleTerm>
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<mods:namePart id="ADEA6161486A21F47E82E4AB4C7E81D2">Ratcliffe, Brett C.</mods:namePart>
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<mods:typeOfResource id="2379FBAF02794DE38D30DA90B7B72168">text</mods:typeOfResource>
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<mods:title id="862934693ECDD6072E0C8B389037306B">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="E3BE68060E4472361679DAC04632917A">
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<mods:date id="79182B1670DD7D6A96CC2FF57ED2B24C">2011</mods:date>
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<mods:detail id="85221200C9D20227D52D81E988A6F110" type="pubDate">
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<mods:number id="E8C7EE11386B9DD31A83FA7471B0F0C7">2011-06-30</mods:number>
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<mods:detail id="7473EC4297AF3661669434BF496ED8F2" type="volume">
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<mods:number id="4EDF3618C74F47FB1D897E89B06362D9">65</mods:number>
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</mods:detail>
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<mods:detail id="BDE65B43A954C3C8A1BAF26F07C5D64A" type="issue">
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<mods:number id="306F43CD6068B43E66E00EA97842C6CE">2</mods:number>
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<mods:start id="3BEF741DA64224E2E4227DE7C4E7DBF1">125</mods:start>
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<mods:end id="44B3C03F219A9D27EE6B7E040DEE9845">129</mods:end>
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<mods:location id="2763FBFE208431DA8BDCAA9077B628EF">
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<mods:url id="17A21934AEEF5D7A12F956594C463CF6">http://dx.doi.org/10.1649/072.065.0203</mods:url>
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<mods:classification id="9FA2487B311CC2F4BF451E54A76263F3">journal article</mods:classification>
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<mods:identifier id="B2ACBC395611355B32FE71FF56853CAA" type="DOI">10.1649/072.065.0203</mods:identifier>
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<mods:identifier id="D9C2C015F2313D88CC585108207B516E" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="89A2EFE898126F344F8A610059E32497" type="Zenodo-Dep">10102619</mods:identifier>
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</mods:mods>
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<treatment id="03E887B2FF9353593DC2FB82BFEFFEB1" LSID="urn:lsid:plazi:treatment:03E887B2FF9353593DC2FB82BFEFFEB1" httpUri="http://treatment.plazi.org/id/03E887B2FF9353593DC2FB82BFEFFEB1" lastPageId="4" lastPageNumber="129" pageId="2" pageNumber="127">
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<subSubSection id="C35B652FFF93535F3DC2FB82BE96FBB4" pageId="2" pageNumber="127" type="nomenclature">
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<paragraph id="8BFE36A4FF93535F3DC2FB82BE96FBB4" blockId="2.[743,1156,1065,1114]" pageId="2" pageNumber="127">
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<emphasis id="B935EAB6FF93535F3DC2FB82B9FEFBD3" bold="true" box="[743,1156,1065,1087]" pageId="2" pageNumber="127">
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<taxonomicName id="4C414D27FF93535F3DC2FB82B97BFBD3" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2011" box="[743,1025,1065,1087]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="zonianis" status="sp. nov.">
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<emphasis id="B935EAB6FF93535F3DC2FB82BEDBFBD2" bold="true" box="[743,929,1065,1087]" italics="true" pageId="2" pageNumber="127">Phileurus zonianis</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20657CDFF93535F3B2BFB82B9FEFBD3" box="[1038,1156,1065,1086]" pageId="2" pageNumber="127" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="137A2A21FF93535F3CAFFBEFBE9EFBB4" box="[906,996,1092,1114]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[175,218,1058,1077]" captionStartId-1="2.[731,763,1710,1729]" captionStartId-2="3.[175,207,998,1017]" captionTargetBox-0="[151,638,191,1024]" captionTargetBox-1="[707,1194,1481,1682]" captionTargetBox-2="[189,597,215,958]" captionTargetId-0="figure-487@1.[151,638,183,1034]" captionTargetId-1="figure-714@2.[707,1194,1470,1686]" captionTargetId-2="figure-647@3.[151,637,183,974]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figs. 1–7. Parameres of 1) Phileurus didymus; 2) Phileurus toulgoeti; 3) Phileurus valgus; 4) Phileurus voirinae; 5–7) Phileurus zonianis." captionText-1="Fig. 8. Distribution of Phileurus zonianis." captionText-2="Fig. 9. Phileurus zonianis, holotype." figureDoi-0="http://doi.org/10.5281/zenodo.10102621" figureDoi-1="http://doi.org/10.5281/zenodo.10102623" figureDoi-2="http://doi.org/10.5281/zenodo.10102625" httpUri-0="https://zenodo.org/record/10102621/files/figure.png" httpUri-1="https://zenodo.org/record/10102623/files/figure.png" httpUri-2="https://zenodo.org/record/10102625/files/figure.png" pageId="2" pageNumber="127">Figs. 5–9</figureCitation>
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)
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C35B652FFF93535E3DFEFBC4BC28FA85" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127" type="description">
|
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<paragraph id="8BFE36A4FF93535E3DFEFBC4BC8CFB25" blockId="2.[707,1194,1135,1422]" lastBlockId="3.[151,639,1070,1729]" lastPageId="3" lastPageNumber="128" pageId="2" pageNumber="127">
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<emphasis id="B935EAB6FF93535F3DFEFBC4BEE8FB69" bold="true" box="[731,914,1135,1156]" pageId="2" pageNumber="127">Ty p e M a t e r i a l.</emphasis>
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<materialsCitation id="3B293CF9FF93535E3C87FBC4BC8CFB25" collectingDate="1956-06-02" collectingDateMax="1995-06-03" collectingDateMin="1956-06-02" collectionCode="T, BARRO, COLORADO" collectorName="H. Wolda & Ft. Sherman & B. Ratcliffe & M. Jameson & Coll. H. Wolda & Col. H. Voris" country="Panama" county="One" lastPageId="3" lastPageNumber="128" location="One" municipality="Two" pageId="2" pageNumber="127" specimenCount="10" stateProvince="Colon" typeStatus="holotype">
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H o l o t y p e m a l e l a b e l e d “
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<collectingCountry id="F3567634FF93535F3DEBFB22BE40FB73" box="[718,826,1161,1182]" name="Panama" pageId="2" pageNumber="127">PANAMA</collectingCountry>
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||
: Canal Zone, Barro Colorado Isl.,
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<date id="FFFF1064FF93535F3DE6FB0FBE73FB54" box="[707,777,1188,1209]" pageId="2" pageNumber="127" value="1976-05">
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||
<collectingDate id="EFBBE98CFF93535F3DE6FB0FBE73FB54" box="[707,777,1188,1209]" pageId="2" pageNumber="127" value="1976-05">V-1976</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="26B45372FF93535F3C36FB0FBE16FB54" box="[787,876,1188,1209]" pageId="2" pageNumber="127">H. Wolda</collectorName>
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||
” and with my red
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||
<typeStatus id="54FA8806FF93535F3B07FB0FB90BFB54" box="[1058,1137,1188,1209]" pageId="2" pageNumber="127" type="holotype">holotype</typeStatus>
|
||
label.
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||
<typeStatus id="54FA8806FF93535F3DE6FB15BE68FB3E" box="[707,786,1214,1235]" pageId="2" pageNumber="127" type="allotype">Allotype</typeStatus>
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||
female with same data and my red
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<typeStatus id="54FA8806FF93535F3BA5FB15BF91FB03" pageId="2" pageNumber="127" type="allotype">allotype</typeStatus>
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label.
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||
<collectingCounty id="629F4E28FF93535F3C0EFB72BE28FB03" box="[811,850,1241,1262]" pageId="2" pageNumber="127">One</collectingCounty>
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||
male
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||
<typeStatus id="54FA8806FF93535F3CA9FB72BEA6FB03" box="[908,988,1241,1262]" pageId="2" pageNumber="127" type="paratype">paratype</typeStatus>
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||
labeled “
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||
<collectingCountry id="F3567634FF93535F3B12FB72B9E1FB03" box="[1079,1179,1241,1262]" name="Panama" pageId="2" pageNumber="127">PANAMA</collectingCountry>
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||
- Canal Zone, Barro Colorado Island,
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||
<date id="FFFF1064FF93535F3B0EFB58B9DFFAE5" box="[1067,1189,1267,1288]" pageId="2" pageNumber="127" value="1956-06-02">
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||
<collectingDate id="EFBBE98CFF93535F3B0EFB58B9DFFAE5" box="[1067,1189,1267,1288]" pageId="2" pageNumber="127" value="1956-06-02">2 June 1956</collectingDate>
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||
</date>
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||
, Carl W. and Marian E. Rettenmeyer, No.”.
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||
<collectingMunicipality id="6B9AACDEFF93535F3B5BFAA5B9D3FACE" box="[1150,1193,1294,1315]" pageId="2" pageNumber="127">Two</collectingMunicipality>
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||
male
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||
<typeStatus id="54FA8806FF93535F3DDFFA83BE2FFAD0" box="[762,853,1320,1341]" pageId="2" pageNumber="127" type="paratype">paratypes</typeStatus>
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||
labeled “
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||
<collectingCountry id="F3567634FF93535F3C9DFA83B95BFAD0" box="[952,1057,1320,1341]" name="Panama" pageId="2" pageNumber="127">PANAMA</collectingCountry>
|
||
:
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||
<collectingRegion id="4985F846FF93535F3B08FA83B9DCFAD0" box="[1069,1190,1320,1341]" country="Panama" name="Colon" pageId="2" pageNumber="127">Colón Prov.</collectingRegion>
|
||
,
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<collectorName id="26B45372FF93535F3DE6FAE8BE44FAB5" box="[707,830,1347,1368]" pageId="2" pageNumber="127">Ft. Sherman</collectorName>
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||
, Pavon Hill,
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||
<date id="FFFF1064FF93535F3CE3FAE8B955FAB5" box="[966,1071,1347,1368]" pageId="2" pageNumber="127" value="1995-06-03">
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||
<collectingDate id="EFBBE98CFF93535F3CE3FAE8B955FAB5" box="[966,1071,1347,1368]" pageId="2" pageNumber="127" value="1995-06-03">VI-3-1995</collectingDate>
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||
</date>
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||
, at light,
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||
<collectorName id="26B45372FF93535F3BB6FAE8BE66FA9E" pageId="2" pageNumber="127">B. Ratcliffe</collectorName>
|
||
and
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||
<collectorName id="26B45372FF93535F3C73FAF5BEAEFA9E" box="[854,980,1374,1395]" pageId="2" pageNumber="127">M. Jameson</collectorName>
|
||
”.
|
||
<location id="8E9E607FFF93535F3CD4FAF5B960FA9E" LSID="urn:lsid:plazi:treatment:03E887B2FF9353593DC2FB82BFEFFEB1:8E9E607FFF93535F3CD4FAF5B960FA9E" box="[1009,1050,1374,1395]" country="Panama" county="One" municipality="Two" name="One" pageId="2" pageNumber="127" stateProvince="Colon">One</location>
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||
female
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||
<typeStatus id="54FA8806FF93535F3B50FAF5BF91FA60" pageId="2" pageNumber="127" type="paratype">paratype</typeStatus>
|
||
labeled “
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||
<collectingCountry id="F3567634FF93535F3C6FFAD3BED4FA60" box="[842,942,1400,1421]" name="Panama" pageId="2" pageNumber="127">PANAMA</collectingCountry>
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||
C. Z., Barro Colorado I.,
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||
<date id="FFFF1064FF92535E3FB2FB84BC6CFBA9" box="[151,278,1071,1092]" pageId="3" pageNumber="128" value="1985-01-16" valueMax="1985-01-27" valueMin="1985-01-16">
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||
<collectingDate id="EFBBE98CFF92535E3FB2FB84BC6CFBA9" box="[151,278,1071,1092]" pageId="3" pageNumber="128" value="1985-01-16" valueMax="1985-01-27" valueMin="1985-01-16">16-27-I-1985</collectingDate>
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||
</date>
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,
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<collectorName id="26B45372FF92535E3E3BFB84BC03FBA9" box="[286,377,1071,1092]" pageId="3" pageNumber="128">H. Wolda</collectorName>
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||
, L.
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<collectionCode id="ED50AE61FF92535E3EBFFB84BCD1FBA9" box="[410,427,1071,1092]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="128">T</collectionCode>
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||
., at ground level”. One female
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||
<typeStatus id="54FA8806FF92535E3FC6FBE2BC4FFBB3" box="[227,309,1097,1118]" pageId="3" pageNumber="128" type="paratype">paratype</typeStatus>
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||
labeled “
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<collectionCode id="ED50AE61FF92535E3EBEFBE2BF07FBB3" box="[411,637,1097,1118]" pageId="3" pageNumber="128">BARRO COLORADO</collectionCode>
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||
C. Z., Lights: Weir/SM-I. III,
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||
<date id="FFFF1064FF92535E3EE8FBCFBF45FB94" box="[461,575,1124,1145]" pageId="3" pageNumber="128" value="1977-03-24">
|
||
<collectingDate id="EFBBE98CFF92535E3EE8FBCFBF45FB94" box="[461,575,1124,1145]" pageId="3" pageNumber="128" value="1977-03-24">24-III-1977</collectingDate>
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||
</date>
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||
,
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<collectorName id="26B45372FF92535E3D6BFBCFBD88FB7E" pageId="3" pageNumber="128">Coll. H. Wolda</collectorName>
|
||
”. One female
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||
<typeStatus id="54FA8806FF92535E3EA4FBD5BCABFB7E" box="[385,465,1150,1171]" pageId="3" pageNumber="128" type="paratype">paratype</typeStatus>
|
||
labeled “Panama, Barro Colo.
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||
<date id="FFFF1064FF92535E3E2EFB32BCE8FB43" box="[267,402,1177,1198]" pageId="3" pageNumber="128" value="1965-08-31">
|
||
<collectingDate id="EFBBE98CFF92535E3E2EFB32BCE8FB43" box="[267,402,1177,1198]" pageId="3" pageNumber="128" value="1965-08-31">Aug. 31, 1965</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26B45372FF92535E3EB9FB32BF6FFB43" box="[412,533,1177,1198]" pageId="3" pageNumber="128">Col. H. Voris</collectorName>
|
||
”. All
|
||
<typeStatus id="54FA8806FF92535E3D6BFB32BDB2FB25" pageId="3" pageNumber="128" type="paratype">paratypes</typeStatus>
|
||
with my yellow
|
||
<typeStatus id="54FA8806FF92535E3E4EFB18BCC1FB25" box="[363,443,1203,1224]" pageId="3" pageNumber="128" type="paratype">paratype</typeStatus>
|
||
label.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF3E662CFF93535F3DFEF905B92CF92C" ID-DOI="http://doi.org/10.5281/zenodo.10102623" ID-Zenodo-Dep="10102623" box="[731,1110,1710,1729]" httpUri="https://zenodo.org/record/10102623/files/figure.png" pageId="2" pageNumber="127" startId="2.[731,763,1710,1729]" targetBox="[707,1194,1481,1682]" targetPageId="2">
|
||
<paragraph id="8BFE36A4FF93535F3DFEF905B92CF92C" blockId="2.[731,1110,1710,1729]" box="[731,1110,1710,1729]" pageId="2" pageNumber="127">
|
||
<emphasis id="B935EAB6FF93535F3DFEF905BE6DF92C" bold="true" box="[731,791,1710,1729]" pageId="2" pageNumber="127">Fig. 8.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C414D27FF93535F3C97F905B92BF92C" authorityName="Ratcliffe" authorityYear="2011" box="[946,1105,1710,1729]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="zonianis">
|
||
<emphasis id="B935EAB6FF93535F3C97F905B92BF92C" box="[946,1105,1710,1729]" italics="true" pageId="2" pageNumber="127">Phileurus zonianis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3E662CFF92535E3F8AFC4DBC84FC14" ID-DOI="http://doi.org/10.5281/zenodo.10102625" ID-Zenodo-Dep="10102625" box="[175,510,997,1017]" httpUri="https://zenodo.org/record/10102625/files/figure.png" pageId="3" pageNumber="128" startId="3.[175,207,998,1017]" targetBox="[189,597,215,958]" targetPageId="3">
|
||
<paragraph id="8BFE36A4FF92535E3F8AFC4DBC84FC14" blockId="3.[175,510,997,1017]" box="[175,510,997,1017]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3F8AFC4DBD96FC14" bold="true" box="[175,236,998,1017]" pageId="3" pageNumber="128">Fig. 9.</emphasis>
|
||
<taxonomicName id="4C414D27FF92535E3E24FC4EBCDAFC15" authorityName="Ratcliffe" authorityYear="2011" box="[257,416,997,1016]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="zonianis">
|
||
<emphasis id="B935EAB6FF92535E3E24FC4EBCDAFC15" box="[257,416,997,1016]" italics="true" pageId="3" pageNumber="128">Phileurus zonianis</emphasis>
|
||
</taxonomicName>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFE36A4FF92535E3F8AFB65BC28FA85" blockId="3.[151,639,1070,1729]" pageId="3" pageNumber="128">
|
||
<materialsCitation id="3B293CF9FF92535E3F8AFB65BF00FB13" collectorName="Lincoln, NE" location="Museum" municipality="University" pageId="3" pageNumber="128" specimenCount="2" stateProvince="Nebraska" typeStatus="holotype">
|
||
<typeStatus id="54FA8806FF92535E3F8AFB65BC7DFB0E" box="[175,263,1230,1251]" pageId="3" pageNumber="128" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54FA8806FF92535E3E19FB65BCF2FB0E" box="[316,392,1230,1251]" pageId="3" pageNumber="128" type="allotype">allotype</typeStatus>
|
||
deposited at the
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||
<collectingMunicipality id="6B9AACDEFF92535E3D11FB65BDC1FB13" pageId="3" pageNumber="128">University</collectingMunicipality>
|
||
of
|
||
<collectingRegion id="4985F846FF92535E3FC1FB42BC04FB13" box="[228,382,1257,1278]" country="United States of America" name="Nebraska" pageId="3" pageNumber="128">Nebraska State</collectingRegion>
|
||
<location id="8E9E607FFF92535E3EA2FB42BCA7FB13" LSID="urn:lsid:plazi:treatment:03E887B2FF9353593DC2FB82BFEFFEB1:8E9E607FFF92535E3EA2FB42BCA7FB13" box="[391,477,1257,1278]" municipality="University" name="Museum" pageId="3" pageNumber="128" stateProvince="Nebraska">Museum</location>
|
||
(
|
||
<collectorName id="26B45372FF92535E3ECAFB42BF09FB13" box="[495,627,1257,1278]" pageId="3" pageNumber="128">Lincoln, NE</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B293CF9FF92535E3FB2FAA8BC34FA85" collectorName="Battleground, WA & Brett C. Ratcliffe & Lincoln, NE" country="United States of America" location="Washington" pageId="3" pageNumber="128" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
|
||
<typeStatus id="54FA8806FF92535E3FB2FAA8BD89FAF5" box="[151,243,1283,1304]" pageId="3" pageNumber="128" type="paratype">Paratypes</typeStatus>
|
||
deposited at the
|
||
<collectingCountry id="F3567634FF92535E3E84FAA8BCAEFAF5" box="[417,468,1283,1304]" name="United States of America" pageId="3" pageNumber="128">U. S.</collectingCountry>
|
||
National Collection (
|
||
<collectingRegion id="4985F846FF92535E3FEEFAB5BC3AFADE" box="[203,320,1310,1331]" country="United States of America" name="Washington" pageId="3" pageNumber="128">Washington</collectingRegion>
|
||
, D. C.), Ron McPeak collection (
|
||
<collectorName id="26B45372FF92535E3FBBFA93BC28FAA0" box="[158,338,1336,1357]" pageId="3" pageNumber="128">Battleground, WA</collectorName>
|
||
), and
|
||
<collectorName id="26B45372FF92535E3EABFA93BF4CFAA0" box="[398,566,1336,1357]" pageId="3" pageNumber="128">Brett C. Ratcliffe</collectorName>
|
||
collection (
|
||
<collectorName id="26B45372FF92535E3FEEFAF8BC3DFA85" box="[203,327,1363,1384]" pageId="3" pageNumber="128">Lincoln, NE</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B652FFF92535E3F8AFAC5B9F4FB13" pageId="3" pageNumber="128" type="materials_examined">
|
||
<paragraph id="8BFE36A4FF92535E3F8AFAC5B9EEFC82" blockId="3.[151,639,1070,1729]" lastBlockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3F8AFAC5BC75FA6E" bold="true" box="[175,271,1390,1411]" pageId="3" pageNumber="128">
|
||
<typeStatus id="54FA8806FF92535E3F8AFAC5BC71FA6E" box="[175,267,1390,1411]" pageId="3" pageNumber="128" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male.Length
|
||
<quantity id="4CB99B41FF92535E3EBDFAC5BC90FA6E" box="[408,490,1390,1411]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="3" pageNumber="128" unit="mm" value="18.1">18.1 mm</quantity>
|
||
, width 13.0 mm. Color black.
|
||
<emphasis id="B935EAB6FF92535E3E36FA23BC32FA70" bold="true" box="[275,328,1416,1437]" pageId="3" pageNumber="128">Head</emphasis>
|
||
: Hollow of vertex deep, smooth inside. Frons rugose. Short horn present on each side of head; horn conical, stout, erect. Clypeus triangular, apex acute and sharply reflexed, surface transversely rugopunctate; clypeal carina extending from clypeal apex to base of each tubercle absent. Interocular width equals 3.3 transverse eye diameters; antenna with 10 segments, club a little longer than segments 2–7. Mandibles narrow, arcuate on lateral edge, apex subacute.
|
||
<emphasis id="B935EAB6FF92535E3E4DF9DCBCB4F961" bold="true" box="[360,462,1655,1676]" pageId="3" pageNumber="128">Pronotum</emphasis>
|
||
: Median, longitudinal furrow narrow, deep, extending from near base to just past middle where it joins small, transverse fovea. Furrow with moderately large punctures, fovea with transverse rugae. Strong tubercle present immediately behind anterior margin. Disc either side of furrow moderately densely punctate, punctures moderately large, becoming slightly denser along lateral margins; anterior angles with large, transverse, mostly confluent punctures. Base with complete marginal bead.
|
||
<emphasis id="B935EAB6FF92535E3CA1FED7BEBCFE7C" bold="true" box="[900,966,380,401]" pageId="3" pageNumber="128">Elytra</emphasis>
|
||
: Rows of punctures equidistant from one another, in furrows; punctures large, ocellate-umbilicate, separated by 1–2 puncture diameters from each other. Intervals convex, shiny, all subequal in height and width, with sparse, small punctures.
|
||
<emphasis id="B935EAB6FF92535E3C6BFDAABED7FDFB" bold="true" box="[846,941,513,534]" pageId="3" pageNumber="128">Pygidium</emphasis>
|
||
: Surface with punctures moderately large, ocellate, setigerous; setae minute to short, tawny. In lateral view, surface evenly and strongly convex.
|
||
<emphasis id="B935EAB6FF92535E3C68FDFABE01FD8B" bold="true" box="[845,891,593,614]" pageId="3" pageNumber="128">Legs</emphasis>
|
||
: Protibia quadridentate, apical 2 teeth obliquely truncate and closer to one another than to other teeth. Apex of metatibia with 2 large teeth subequal in size. Apex of basal tarsomere of metatarsus extended into long, slender spine.
|
||
<emphasis id="B935EAB6FF92535E3B6BFD10B9F7FD3D" bold="true" box="[1102,1165,699,720]" pageId="3" pageNumber="128">Venter</emphasis>
|
||
: Prosternal process short, glabrous, apex broadly rounded, posterior surface with transverse furrow, base with subacute lobe.
|
||
<emphasis id="B935EAB6FF92535E3CB1FCA1BE84FCF2" bold="true" box="[916,1022,778,799]" pageId="3" pageNumber="128">Parameres</emphasis>
|
||
: Each paramere with a medially projecting lobe crossing opposing lobe, lobes turning downwards at apex; lobes crossing either left over right or right over left (
|
||
<figureCitation id="137A2A21FF92535E3B15FCF1B9F3FC82" box="[1072,1161,858,880]" captionStart="Figs" captionStartId="1.[175,218,1058,1077]" captionTargetBox="[151,638,191,1024]" captionTargetId="figure-487@1.[151,638,183,1034]" captionTargetPageId="1" captionText="Figs. 1–7. Parameres of 1) Phileurus didymus; 2) Phileurus toulgoeti; 3) Phileurus valgus; 4) Phileurus voirinae; 5–7) Phileurus zonianis." figureDoi="http://doi.org/10.5281/zenodo.10102621" httpUri="https://zenodo.org/record/10102621/files/figure.png" pageId="3" pageNumber="128">Figs. 5–6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE36A4FF92535E3DE0FCDEB964FBC4" blockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3DE0FCDEBE59FC67" bold="true" box="[709,803,885,906]" pageId="3" pageNumber="128">
|
||
<typeStatus id="54FA8806FF92535E3DE0FCDEBE65FC67" box="[709,799,885,906]" pageId="3" pageNumber="128" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. Length
|
||
<quantity id="4CB99B41FF92535E3CC3FCDEB932FC67" box="[998,1096,885,906]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="128" unit="mm" value="19.1">19.1 mm</quantity>
|
||
, width 13.0 mm. As
|
||
<typeStatus id="54FA8806FF92535E3C65FC24BEEDFC49" box="[832,919,911,932]" pageId="3" pageNumber="128" type="holotype">holotype</typeStatus>
|
||
except in the following respects:
|
||
<emphasis id="B935EAB6FF92535E3C22FC01BE47FC52" bold="true" box="[775,829,938,959]" pageId="3" pageNumber="128">Head</emphasis>
|
||
: Hollow of vertex with a few moderately large punctures, an erect tubercle on each side of head instead of a short horn, and interocular width equals 3.6 transverse eye diameters.
|
||
<emphasis id="B935EAB6FF92535E3B79FC51B9F6FBE2" bold="true" box="[1116,1164,1018,1039]" pageId="3" pageNumber="128">Legs</emphasis>
|
||
: Left front leg missing tarsomeres 4–5.
|
||
</paragraph>
|
||
<paragraph id="8BFE36A4FF92535E3DE0FB84B9EBFB7E" blockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3DE0FB84BE48FBA9" bold="true" box="[709,818,1071,1092]" pageId="3" pageNumber="128">
|
||
<typeStatus id="54FA8806FF92535E3DE0FB84BE54FBA9" box="[709,814,1071,1092]" pageId="3" pageNumber="128" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Males (3). Length
|
||
<quantity id="4CB99B41FF92535E3CD9FB84B9EAFBA9" box="[1020,1168,1070,1092]" metricMagnitude="-2" metricUnit="m" metricValue="2.735" metricValueMax="2.88" metricValueMin="2.59" pageId="3" pageNumber="128" unit="mm" value="27.35" valueMax="28.8" valueMin="25.9">25.9–28.8 mm</quantity>
|
||
, width
|
||
<quantity id="4CB99B41FF92535E3DC2FBE2BE0BFBB3" box="[743,881,1097,1119]" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.3599999999999999" metricValueMin="1.23" pageId="3" pageNumber="128" unit="mm" value="12.95" valueMax="13.6" valueMin="12.3">12.3–13.6 mm</quantity>
|
||
. The male
|
||
<typeStatus id="54FA8806FF92535E3CF0FBE2B957FBB3" box="[981,1069,1097,1118]" pageId="3" pageNumber="128" type="paratype">paratypes</typeStatus>
|
||
do not differ significantly from the
|
||
<typeStatus id="54FA8806FF92535E3CB6FBCFBE9CFB94" box="[915,998,1124,1145]" pageId="3" pageNumber="128" type="holotype">holotype</typeStatus>
|
||
. One specimen has the pronotal surface slightly more densely punctate.
|
||
</paragraph>
|
||
<paragraph id="8BFE36A4FF92535E3DE0FB32B9F4FB13" blockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
Females (3). Length
|
||
<quantity id="4CB99B41FF92535E3CAEFB32B962FB43" box="[907,1048,1176,1198]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.85" metricValueMin="1.83" pageId="3" pageNumber="128" unit="mm" value="18.4" valueMax="18.5" valueMin="18.3">18.3–18.5 mm</quantity>
|
||
, width 13.0–
|
||
<quantity id="4CB99B41FF92535E3D88FB1FBE7DFB24" box="[685,775,1204,1225]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="128" unit="mm" value="13.1">13.1 mm</quantity>
|
||
. The female
|
||
<typeStatus id="54FA8806FF92535E3CA0FB1FBE9AFB24" box="[901,992,1204,1225]" pageId="3" pageNumber="128" type="paratype">paratypes</typeStatus>
|
||
do not differs significantly from the
|
||
<typeStatus id="54FA8806FF92535E3C4DFB65BECDFB0E" box="[872,951,1230,1251]" pageId="3" pageNumber="128" type="allotype">allotype</typeStatus>
|
||
. One specimen has the interocular width of 3.4 transverse eye diameters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B652FFF92535E3DE0FAA8BE90FA6E" pageId="3" pageNumber="128" type="etymology">
|
||
<paragraph id="8BFE36A4FF92535E3DE0FAA8BE90FA6E" blockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3DE0FAA8BE42FAF5" bold="true" box="[709,824,1283,1304]" pageId="3" pageNumber="128">Etymology.</emphasis>
|
||
A Zonian is a dweller, usually a person, associated with the
|
||
<collectingCountry id="F3567634FF92535E3C89FAB5BE81FADE" box="[940,1019,1310,1331]" name="Panama" pageId="3" pageNumber="128">Panama</collectingCountry>
|
||
Canal Zone, a political entity that existed between 1903 and the absorption of the Canal Zone into the
|
||
<collectingCountry id="F3567634FF92535E3B07FAF8BF82FA6E" name="Panama" pageId="3" pageNumber="128">Republic of Panama</collectingCountry>
|
||
between 1980 and 2000.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B652FFF92535E3DE0FA23BE63FA00" pageId="3" pageNumber="128" type="distribution">
|
||
<paragraph id="8BFE36A4FF92535E3DE0FA23BE63FA00" blockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3DE0FA23BE32FA70" bold="true" box="[709,840,1416,1437]" pageId="3" pageNumber="128">Distribution.</emphasis>
|
||
<taxonomicName id="4C414D27FF92535E3C70FA23B958FA70" authorityName="Ratcliffe" authorityYear="2011" box="[853,1058,1416,1437]" class="Insecta" family="Dynastidae" genus="Phileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="zonianis">
|
||
<emphasis id="B935EAB6FF92535E3C70FA23B958FA70" box="[853,1058,1416,1437]" italics="true" pageId="3" pageNumber="128">Phileurus zonianis</emphasis>
|
||
</taxonomicName>
|
||
is known only from Barro Colorado Island in the area of the former Canal Zone in the
|
||
<collectingCountry id="F3567634FF92535E3CF5FA16B9EEFA3F" box="[976,1172,1469,1490]" name="Panama" pageId="3" pageNumber="128">Republic of Panama</collectingCountry>
|
||
(
|
||
<figureCitation id="137A2A21FF92535E3D91FA73BF8BFA00" box="[692,753,1496,1517]" captionStart="Fig" captionStartId="2.[731,763,1710,1729]" captionTargetBox="[707,1194,1481,1682]" captionTargetId="figure-714@2.[707,1194,1470,1686]" captionTargetPageId="2" captionText="Fig. 8. Distribution of Phileurus zonianis." figureDoi="http://doi.org/10.5281/zenodo.10102623" httpUri="https://zenodo.org/record/10102623/files/figure.png" pageId="3" pageNumber="128">Fig. 8</figureCitation>
|
||
,
|
||
<figureCitation id="137A2A21FF92535E3DDAFA73BE77FA00" box="[767,781,1496,1517]" captionStart="Fig" captionStartId="3.[175,207,998,1017]" captionTargetBox="[189,597,215,958]" captionTargetId="figure-647@3.[151,637,183,974]" captionTargetPageId="3" captionText="Fig. 9. Phileurus zonianis, holotype." figureDoi="http://doi.org/10.5281/zenodo.10102625" httpUri="https://zenodo.org/record/10102625/files/figure.png" pageId="3" pageNumber="128">9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B652FFF9253593DE0FA59BC34FEE1" lastPageId="4" lastPageNumber="129" pageId="3" pageNumber="128" type="diagnosis">
|
||
<paragraph id="8BFE36A4FF92535E3DE0FA59B916F92C" blockId="3.[685,1172,194,1730]" pageId="3" pageNumber="128">
|
||
<emphasis id="B935EAB6FF92535E3DE0FA59BE54F9EA" bold="true" box="[709,814,1522,1543]" pageId="3" pageNumber="128">Diagnosis.</emphasis>
|
||
This species is distinguished by the smooth hollow of the vertex in males, pronotum densely punctate either side of median furrow, elytral intervals all subequal in height and with sparse, small punctures, apex of the metatibia with two strong teeth, prosternal process glabrous, and form of the parameres where the median lobe of each paramere angles downwards (
|
||
<figureCitation id="137A2A21FF92535E3B24F907B91AF92C" box="[1025,1120,1708,1730]" captionStart="Figs" captionStartId="1.[175,218,1058,1077]" captionTargetBox="[151,638,191,1024]" captionTargetId="figure-487@1.[151,638,183,1034]" captionTargetPageId="1" captionText="Figs. 1–7. Parameres of 1) Phileurus didymus; 2) Phileurus toulgoeti; 3) Phileurus valgus; 4) Phileurus voirinae; 5–7) Phileurus zonianis." figureDoi="http://doi.org/10.5281/zenodo.10102621" httpUri="https://zenodo.org/record/10102621/files/figure.png" pageId="3" pageNumber="128">Figs. 5–6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFE36A4FF9553593FE0FF69BC34FEE1" blockId="4.[173,661,194,348]" pageId="4" pageNumber="129">
|
||
This species was overlooked in my treatment of the Dynastinae of
|
||
<collectingCountry id="F3567634FF9553593EB0FF76BF72FF1F" box="[405,520,221,242]" name="Costa Rica" pageId="4" pageNumber="129">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3567634FF9553593D61FF76BFE9FF1F" box="[580,659,221,242]" name="Panama" pageId="4" pageNumber="129">Panama</collectingCountry>
|
||
(Ratcliffe 2003).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35B652FFF9553593FE0FEB9BFEFFEB1" pageId="4" pageNumber="129" type="biology_ecology">
|
||
<paragraph id="8BFE36A4FF9553593FE0FEB9BFEFFEB1" blockId="4.[173,661,194,348]" pageId="4" pageNumber="129">
|
||
<emphasis id="B935EAB6FF9553593FE0FEB9BC6CFECA" bold="true" box="[197,278,274,295]" pageId="4" pageNumber="129">Biology.</emphasis>
|
||
Other than being attracted to lights, nothing is known of the life history of this species. All of the specimens were taken in lowland rainforest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |