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<document id="0674F745CAAFCF5F6408ACC38B32CBE0" ID-DOI="10.11646/zootaxa.3974.3.2" ID-GBIF-Dataset="67110af0-00a8-4fab-9025-cf54a705ba3e" ID-ISSN="1175-5326" ID-Zenodo-Dep="243714" ID-ZooBank="59DD0D8D-1F66-4F25-B8C1-57D94734BC39" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461219698134" checkinUser="plazi" docAuthor="Schmidt, Olga" docDate="2015" docId="03E88788182BFFB57CCEFC102C3CFE10" docLanguage="en" docName="zt03974p340.pdf" docOrigin="Zootaxa 3974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chesiadini" docType="treatment" docVersion="7" lastPageNumber="331" masterDocId="FFD1FFF01829FFB67C59FE562E71FFBD" masterDocTitle="One century after: a reappraisal of the gnathos (sensu Pierce, 1914) in Larentiinae (Lepidoptera: Geometridae)" masterLastPageNumber="340" masterPageNumber="328" pageNumber="330" updateTime="1698603722042" updateUser="plazi">
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<mods:title id="F86680760CAF3675DB8236F054783D0D">One century after: a reappraisal of the gnathos (sensu Pierce, 1914) in Larentiinae (Lepidoptera: Geometridae)</mods:title>
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<mods:namePart id="C6DD436F2B435C9F53DEA4F7722D6C9C">Schmidt, Olga</mods:namePart>
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<treatment id="03E88788182BFFB57CCEFC102C3CFE10" ID-DOI="http://doi.org/10.5281/zenodo.6121732" ID-GBIF-Taxon="119631500" ID-Zenodo-Dep="6121732" LSID="urn:lsid:plazi:treatment:03E88788182BFFB57CCEFC102C3CFE10" httpUri="http://treatment.plazi.org/id/03E88788182BFFB57CCEFC102C3CFE10" lastPageId="3" lastPageNumber="331" pageId="2" pageNumber="330">
<subSubSection id="C35B6515182BFFB47CCEFC102A1AF8CE" pageId="2" pageNumber="330" type="nomenclature">
<paragraph id="8BFE369E182BFFB47CCEFC102F19FDDD" blockId="2.[151,360,582,608]" box="[151,360,582,608]" pageId="2" pageNumber="330">
<heading id="D0B681F2182BFFB47CCEFC102F19FDDD" bold="true" box="[151,360,582,608]" fontSize="11" level="1" pageId="2" pageNumber="330" reason="1">
<emphasis id="B935EA8C182BFFB47CCEFC102F19FDDD" bold="true" box="[151,360,582,608]" pageId="2" pageNumber="330">
Tribe
<taxonomicName id="4C414D1D182BFFB47CBBFC102F19FDDD" box="[226,360,582,608]" class="Insecta" family="Geometridae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="330" phylum="Arthropoda" rank="tribe" tribe="Chesiadini">Chesiadini</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFE369E182BFFB47CCEFCDB2A54FD53" blockId="2.[151,1436,653,750]" pageId="2" pageNumber="330">
<taxonomicName id="4C414D1D182BFFB47CCEFCDB2F06FD1B" box="[151,375,653,678]" class="Insecta" family="Geometridae" genus="Aplocera" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="330" phylum="Arthropoda" rank="species" species="efformata">
<emphasis id="B935EA8C182BFFB47CCEFCDB2F06FD1B" bold="true" box="[151,375,653,678]" italics="true" pageId="2" pageNumber="330">Aplocera efformata</emphasis>
</taxonomicName>
(
<figureCitation id="137A2A1B182BFFB47DDCFCDB2FB7FD1B" box="[389,454,653,678]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[165,1436,802,1713]" captionTargetId="figure@2.[150,1436,799,1726]" captionTargetPageId="2" captionText="FIGURES 1 2. Male genitalia apically: 1. Aplocera efformata. 2. Chesias linogriseata. Gnathos is shown in grey." httpUri="https://zenodo.org/record/243715/files/figure.png" pageId="2" pageNumber="330">Fig. 1</figureCitation>
). The gnathos arms are thin, less pronounced than in other chesiadine genera and partly fused with the tegumen. The base of the large anal tube with its long sclerotized subscaphium is connected to the gnathos and membrane of the fultura superior (which is ventrad of the anal tube).
</paragraph>
<caption id="DF3E6616182BFFB47CCEF8822B6EF957" box="[151,1311,1748,1771]" httpUri="https://zenodo.org/record/243715/files/figure.png" pageId="2" pageNumber="330" targetBox="[165,1436,802,1713]" targetPageId="2">
<paragraph id="8BFE369E182BFFB47CCEF8822B6EF957" blockId="2.[151,1311,1748,1771]" box="[151,1311,1748,1771]" pageId="2" pageNumber="330">
<emphasis id="B935EA8C182BFFB47CCEF8822F48F957" bold="true" box="[151,313,1748,1771]" pageId="2" pageNumber="330">FIGURES 12.</emphasis>
Male genitalia apically:
<emphasis id="B935EA8C182BFFB47E6CF8832C38F957" bold="true" box="[565,585,1749,1770]" pageId="2" pageNumber="330">1.</emphasis>
<taxonomicName id="4C414D1D182BFFB47E16F8822D64F957" box="[591,789,1748,1770]" class="Insecta" family="Geometridae" genus="Aplocera" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="330" phylum="Arthropoda" rank="species" species="efformata">
<emphasis id="B935EA8C182BFFB47E16F8822D64F957" box="[591,789,1748,1770]" italics="true" pageId="2" pageNumber="330">Aplocera efformata</emphasis>
</taxonomicName>
.
<emphasis id="B935EA8C182BFFB47F78F8832D45F957" bold="true" box="[801,820,1749,1770]" pageId="2" pageNumber="330">2.</emphasis>
<taxonomicName id="4C414D1D182BFFB47F63F8822A7AF957" box="[826,1035,1748,1770]" class="Insecta" family="Geometridae" genus="Chesias" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="330" phylum="Arthropoda" rank="species" species="linogriseata">
<emphasis id="B935EA8C182BFFB47F63F8822A7AF957" box="[826,1035,1748,1770]" italics="true" pageId="2" pageNumber="330">Chesias linogriseata</emphasis>
</taxonomicName>
. Gnathos is shown in grey.
</paragraph>
</caption>
<paragraph id="8BFE369E182BFFB47C9EF9442A1AF8CE" blockId="2.[151,1437,1810,2014]" pageId="2" pageNumber="330">
<taxonomicName id="4C414D1D182BFFB47C9EF9442FCDF896" box="[199,444,1810,1835]" class="Insecta" family="Geometridae" genus="Chesias" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="330" phylum="Arthropoda" rank="species" species="linogrisearia">
<emphasis id="B935EA8C182BFFB47C9EF9442FCDF896" bold="true" box="[199,444,1810,1835]" italics="true" pageId="2" pageNumber="330">Chesias linogrisearia</emphasis>
</taxonomicName>
(
<figureCitation id="137A2A1B182BFFB47D92F9442C7FF897" box="[459,526,1810,1835]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1748,1771]" captionTargetBox="[165,1436,802,1713]" captionTargetId="figure@2.[150,1436,799,1726]" captionTargetPageId="2" captionText="FIGURES 1 2. Male genitalia apically: 1. Aplocera efformata. 2. Chesias linogriseata. Gnathos is shown in grey." httpUri="https://zenodo.org/record/243715/files/figure.png" pageId="2" pageNumber="330">Fig. 2</figureCitation>
). The gnathos arms are distinct, thin, long and weakly sclerotized, and connected to the base of the uncus laterally. The base of the medium sized anal tube with its distinct subscaphium is connected to the gnathos and membrane of the fultura superior (which is ventrad of the anal tube).
</paragraph>
</subSubSection>
<subSubSection id="C35B6515182BFFB57C9EF92B2C3CFE10" lastPageId="3" lastPageNumber="331" pageId="2" pageNumber="330" type="reference_group">
<paragraph id="8BFE369E182BFFB57C9EF92B2D5EFF69" blockId="2.[151,1437,1810,2014]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="331" pageId="2" pageNumber="330">
<taxonomicName id="4C414D1D182BFFB47C9EF92B2FCDF82B" box="[199,444,1917,1942]" class="Insecta" family="Geometridae" genus="Docirava" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="330" phylum="Arthropoda" rank="species" species="mundulata">
<emphasis id="B935EA8C182BFFB47C9EF92B2FCDF82B" bold="true" box="[199,444,1917,1942]" italics="true" pageId="2" pageNumber="330">Docirava mundulata</emphasis>
</taxonomicName>
(
<figureCitation id="137A2A1B182BFFB47D88F92B2C6BF82B" box="[465,538,1917,1942]" captionStart="FIGURES 3 4" captionStartId="3.[151,264,1443,1466]" captionTargetBox="[158,1436,495,1409]" captionTargetId="figure@3.[151,1436,483,1422]" captionTargetPageId="3" captionText="FIGURES 3 4. Male genitalia apically: 3. Docirava mundulata. 4. Lithostege amoenata. Gnathos is shown in grey." httpUri="https://zenodo.org/record/243716/files/figure.png" pageId="2" pageNumber="330">Fig. 3</figureCitation>
). The gnathos arms are distinct, sclerotized, stout, broadening and dentate proximally. The base of the anal tube with its broad sclerotized subscaphium is connected to the gnathos and membrane of the fultura superior (which is ventrad of the anal tube).
<emphasis id="B935EA8C182BFFB47F98F9932A44F863" bold="true" box="[961,1077,1989,2014]" pageId="2" pageNumber="330">Remarks.</emphasis>
The
<emphasis id="B935EA8C182BFFB4782DF9932A9EF863" box="[1140,1263,1989,2014]" italics="true" pageId="2" pageNumber="330">dervenaria</emphasis>
species-group, including
<taxonomicName id="4C414D1D182AFFB57D55FECF2FDDFF0D" box="[268,428,151,176]" class="Insecta" family="Geometridae" genus="Docirava" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="331" phylum="Arthropoda" rank="species" species="mundulata">
<emphasis id="B935EA8C182AFFB57D55FECF2FDDFF0D" box="[268,428,151,176]" italics="true" pageId="3" pageNumber="331">D. mundulata</emphasis>
</taxonomicName>
, was only tentatively transferred to
<taxonomicName id="4C414D1D182AFFB57F05FECF2DB7FF0D" ID-CoL="927BT" box="[860,966,153,176]" class="Insecta" family="Geometridae" genus="Docirava" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="B935EA8C182AFFB57F05FECF2DB7FF0D" box="[860,966,153,176]" italics="true" pageId="3" pageNumber="331">Docirava</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD04B6F182AFFB57F83FEC12ABDFF0D" author="Karsholt" box="[986,1228,151,176]" pageId="3" pageNumber="331" refString="Karsholt, O., Nieukerken, E. J. van &amp; de Jong, Y. S. D. M. (2013) Lepidoptera, Moths. Fauna Europaea. Version 2.6.2. Available from: http: // www. faunaeur. org (Accessed 29 August 2013)" type="journal article" year="2013">
Karsholt
<emphasis id="B935EA8C182AFFB57811FECF2AF4FF0D" box="[1096,1157,151,176]" italics="true" pageId="3" pageNumber="331">et al.</emphasis>
2013
</bibRefCitation>
, Hausmann, pers. comm.) but is questionable (
<bibRefCitation id="EFD04B6F182AFFB57D8CFEEA2D52FF69" author="Hausmann" box="[469,803,188,213]" pageId="3" pageNumber="331" refString="Hausmann, A. &amp; Viidalepp, J. (2012) Larentiinae I. In: Hausmann, A. (Ed.), The Geometrid Moths of Europe. Apollo Books, Stenstrup, pp. 1 - 743. [25 plates., 147 text-figs, 268 maps.]" type="book chapter" year="2012">Hausmann &amp; Viidalepp 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BFE369E182AFFB57C9EFE892C3CFE10" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="331">
<taxonomicName id="4C414D1D182AFFB57C9EFE892FC9FF4A" ID-CoL="3VLK4" box="[199,440,223,248]" class="Insecta" family="Geometridae" genus="Lithostege" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="331" phylum="Arthropoda" rank="species" species="amoenata">
<emphasis id="B935EA8C182AFFB57C9EFE892FC9FF4A" bold="true" box="[199,440,223,248]" italics="true" pageId="3" pageNumber="331">Lithostege amoenata</emphasis>
</taxonomicName>
(
<figureCitation id="137A2A1B182AFFB57D90FE892C7CFF45" box="[457,525,223,248]" captionStart="FIGURES 3 4" captionStartId="3.[151,264,1443,1466]" captionTargetBox="[158,1436,495,1409]" captionTargetId="figure@3.[151,1436,483,1422]" captionTargetPageId="3" captionText="FIGURES 3 4. Male genitalia apically: 3. Docirava mundulata. 4. Lithostege amoenata. Gnathos is shown in grey." httpUri="https://zenodo.org/record/243716/files/figure.png" pageId="3" pageNumber="331">Fig. 4</figureCitation>
). The gnathos arms are distinct, weakly sclerotized and thickened at the base of the uncus. The base of the medium sized anal tube with its rather broad sclerotized subscaphium is connected to the gnathos and membrane of the fultura superior (which is ventrad of the anal tube).
<emphasis id="B935EA8C182AFFB5781AFF712AC6FEFD" bold="true" box="[1091,1207,295,320]" pageId="3" pageNumber="331">Remarks.</emphasis>
<bibRefCitation id="EFD04B6F182AFFB57899FF712BEDFEFD" box="[1216,1436,295,320]" pageId="3" pageNumber="331" refString="Rajaei, H. S., Stuning, D. &amp; Viidalepp, J. (2011) A review of the species of Lithostege Hubner, [1825] 1816 (Lepidoptera: Geometridae, Larentiinae), occurring in Iran and adjacent countries, with description of two new species from Iran and Pakistan. Zootaxa, 3105, 1 - 46." type="journal article">
Rajaei
<emphasis id="B935EA8C182AFFB57949FF7F2B3BFEFD" box="[1296,1354,295,320]" italics="true" pageId="3" pageNumber="331">et al.</emphasis>
(2011)
</bibRefCitation>
mentioned that, “Chesiadine genera have an additional rib-like sclerite in male genitalia, from below the uncus to the hemitranstilla (partially fused to the lateral margins of tegumen in some
<taxonomicName id="4C414D1D182AFFB57878FF392AF9FE35" ID-CoL="922J5" box="[1057,1160,367,392]" class="Insecta" family="Geometridae" genus="Aplocera" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="B935EA8C182AFFB57878FF392AF9FE35" box="[1057,1160,367,392]" italics="true" pageId="3" pageNumber="331">Aplocera</emphasis>
</taxonomicName>
species)”. This rib-like sclerite is the gnathos described above.
</paragraph>
</subSubSection>
</treatment>
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