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<document id="CC80AE64918CE70E96194CBDC18402A7" ID-CLB-Dataset="28528" ID-DOI="10.11646/zootaxa.4556.1.1" ID-GBIF-Dataset="9ffb3429-ba6c-4d23-a88e-781dde361e56" ID-ISSN="1175-5326" ID-Zenodo-Dep="2585204" ID-ZooBank="7C55D5CB-5A0F-4DCE-A5A7-755339CF45F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1551120081848" checkinUser="plazi" docAuthor="Qiu, Jian-Yue, Xu, Hao &amp; Chen, Li" docDate="2019" docId="03CA3425FF97A421C2E9F98C103614E2" docLanguage="en" docName="zootaxa.4556.1.1.pdf" docOrigin="Zootaxa 4556 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Macronotops biserratus Qiu &amp; Xu &amp; Chen 2019, new species" docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FFF34C5DFF90A429C27EFF3E140B123D" masterDocTitle="A revision of the rare flower beetle genus Macronotops Krikken (Coleoptera: Scarabaeidae: Cetoniinae) from Asia with biological notes" masterLastPageNumber="65" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1550769681392" originalUpdateUser="plazi" pageNumber="8" updateTime="1698700146227" updateUser="ExternalLinkService">
<mods:mods id="813D50BAA7641FD47EFE5378BE1B1AFA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1524D52FB8A3FE8D1A4BB866B5160A0E">A revision of the rare flower beetle genus Macronotops Krikken (Coleoptera: Scarabaeidae: Cetoniinae) from Asia with biological notes</mods:title>
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<mods:namePart id="068BD33EB5DF4E85DD5A5D2233329235">Qiu, Jian-Yue</mods:namePart>
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<mods:namePart id="D0CDDB174DAA51B2732D29C988E2AC21">Xu, Hao</mods:namePart>
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<mods:namePart id="5DD60CA62475AD602997675F1DDBFC38">Chen, Li</mods:namePart>
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<mods:title id="8F86A4B3870ECCEA96C7AEA358547895">Zootaxa</mods:title>
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<mods:date id="7F48CD6CAE2846DB8728E5D7C61AC89F">2019</mods:date>
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<mods:number id="48D99FBDD0DC846ECB1A9708ECE044DE">2019-02-18</mods:number>
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<mods:identifier id="49613D3126BC4ED8819F4DCB6CE55681" type="DOI">10.11646/zootaxa.4556.1.1</mods:identifier>
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<treatment id="03CA3425FF97A421C2E9F98C103614E2" ID-DOI="http://doi.org/10.5281/zenodo.5934003" ID-GBIF-Taxon="152633810" ID-Zenodo-Dep="5934003" LSID="urn:lsid:plazi:treatment:03CA3425FF97A421C2E9F98C103614E2" httpUri="http://treatment.plazi.org/id/03CA3425FF97A421C2E9F98C103614E2" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C379D6B8FF97A42EC2E9F98C172914F0" box="[151,802,1714,1741]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BDC8533FF97A42EC2E9F98C172914F0" blockId="7.[151,802,1714,1774]" box="[151,802,1714,1741]" pageId="7" pageNumber="8">
<heading id="D094325FFF97A42EC2E9F98C172914F0" bold="true" box="[151,802,1714,1741]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B9175921FF97A42EC2E9F98C172914F0" bold="true" box="[151,802,1714,1741]" pageId="7" pageNumber="8">
<taxonomicName id="4C63FEB0FF97A42EC2E9F98C168014F0" ID-CoL="3X78L" authority="Qiu, Xu &amp; Chen" authorityName="Qiu &amp; Xu &amp; Chen" authorityYear="2019" box="[151,651,1714,1741]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="biserratus" status="sp. nov.">
<emphasis id="B9175921FF97A42EC2E9F98C15BD14F1" bold="true" box="[151,438,1714,1740]" italics="true" pageId="7" pageNumber="8">Macronotops biserratus</emphasis>
Qiu, Xu &amp; Chen
</taxonomicName>
<taxonomicNameLabel id="A224E45AFF97A42EC0ECF98D172914F0" box="[658,802,1715,1741]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C379D6B8FF97A42EC2E9F9EB16E714D3" box="[151,748,1749,1774]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BDC8533FF97A42EC2E9F9EB16E714D3" blockId="7.[151,802,1714,1774]" box="[151,748,1749,1774]" pageId="7" pageNumber="8">
(
<figureCitation id="135899B6FF97A42EC2E1F9EB14E714D3" box="[159,236,1749,1774]" captionStart="FIGURES 1–23" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 1–23. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 2–3, M. dianensis new species; 4–5, M. fulvoguttatus; 6–7, M. fulvopilosus; 8–9, M. miksici new species; 10–11, M. nigropubescens; 12–13, M. olivaceofuscus; 14–15, M. sexmaculatus; 16–17, M. vuilleti; 18–19, M. curvimarginatus; 20–21, M. medogensis new species; 22–23, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="7" pageNumber="8">Figs. 1</figureCitation>
,
<figureCitation id="135899B6FF97A42EC284F9E8151214D3" box="[250,281,1750,1774]" captionStart="FIGURES 24–47" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 24–47. Protibia of Macronotops (male and female). 24, M. biserratus new species; 25–26, M. dianensis new species; 27–28, M. fulvoguttatus; 29–30, M. fulvopilosus; 31–32, M. miksici new species; 33–34, M. nigropubescens; 35–36, M. olivaceofuscus; 37–38, M. sexmaculatus; 39–40, M. vuilleti; 41, Macronotops sp.; 42–43, M. curvimarginatus; 44–45, M. medogensis new species; 46–47, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="7" pageNumber="8">24</figureCitation>
,
<figureCitation id="135899B6FF97A42EC35BF9E8154814D3" box="[293,323,1750,1774]" captionStart="FIGURES 48–65" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 48–65. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 49–50, M. dianensis new species; 51–52, M. fulvoguttatus; 53–54, M. fulvopilosus; 55–56, M. miksici new species; 57–58, M. nigropubescens;59–60, M. olivaceofuscus; 61–62, M. sexmaculatus; 63–64, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="7" pageNumber="8">48</figureCitation>
,
<figureCitation id="135899B6FF97A42EC330F9E8159C14D3" box="[334,407,1750,1774]" captionStart="FIGURES 72–98" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 72–98. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 72–73, M. biserratus new species; 74–75, M. dianensis new species; 76–79, M. fulvoguttatus; 80–83, M. fulvopilosus; 84–85, M. miksici new species; 86–89, M. nigropubescens; 90–91, M. olivaceofuscus; 92–93, M. sexmaculatus; 94–97, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="7" pageNumber="8">72–73</figureCitation>
,
<figureCitation id="135899B6FF97A42EC3DCF9E815C714D3" box="[418,460,1750,1774]" captionStart="FIGURES 111–122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111–122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="7" pageNumber="8">111</figureCitation>
,
<figureCitation id="135899B6FF97A42EC3A6F9E8160F14D3" box="[472,516,1750,1774]" captionStart="FIGURES 122–134" captionStartId="17.[151,264,1757,1780]" captionTargetBox="[168,1424,207,1720]" captionTargetId="figure@17.[151,1436,193,1737]" captionTargetPageId="17" captionText="FIGURES 122–134. Male genitalia of Macronotops (lateral view). 123, M. biserratus new species; 124, M. dianensis new species; 125, M. fulvoguttatus; 126, M. fulvopilosus; 127, M. miksici new species; 128, M. nigropubescens; 129, M. olivaceofuscus; 130, M. sexmaculatus; 131, M. vuilleti; 132, M. curvimarginatus; 133, M. medogensis new species; 134, M. ovaliceps." httpUri="https://zenodo.org/record/2585222/files/figure.png" pageId="7" pageNumber="8">123</figureCitation>
,
<figureCitation id="135899B6FF97A42EC06EF9EB167E14D3" box="[528,629,1749,1774]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="7" pageNumber="8">135–136</figureCitation>
,
<figureCitation id="135899B6FF97A42EC0FEF9E816A614D3" box="[640,685,1750,1774]" captionStart="FIGURES 203–214" captionStartId="50.[151,264,1945,1968]" captionTargetBox="[173,1415,188,1914]" captionTargetId="figure@50.[151,1436,185,1924]" captionTargetPageId="50" captionText="FIGURES 203–214. Habitus of Macronotops (male in left, female in right; oblique view). 203, M. biserratus new species; 204, Macronotops sp.; 205–206, M. dianensis new species; 207–208, M. fulvoguttatus. 209–210, M. fulvopilosus; 211–212, M. miksici new species; 213–214, M. nigropubescens." httpUri="https://zenodo.org/record/2585254/files/figure.png" pageId="7" pageNumber="8">203</figureCitation>
,
<figureCitation id="135899B6FF97A42EC0C6F9EB16E814D3" box="[696,739,1749,1774]" captionStart="FIGURES 258–260" captionStartId="55.[151,264,1245,1268]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@55.[151,1436,193,1224]" captionTargetPageId="55" captionText="FIGURES 258–260. Known distribution of Macronotops. 258, Distribution for the whole genus; 259, map of SE Xizang, China; 260, map of W Sichuan, China. Syncretic markers indicate identical or very close localities; type localities of the known species are underlined, that of M. olivaceofuscus is missing because the precise type locality is unknown." httpUri="https://zenodo.org/record/2585264/files/figure.png" pageId="7" pageNumber="8">258</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="B9175921FF97A42EC2E9F82315AC150B" bold="true" box="[151,423,1821,1846]" pageId="7" pageNumber="8">
Type material.
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<collectingCountry id="F374C5A3FF97A42EC32FF82315AA150B" box="[337,417,1821,1846]" name="Laos" pageId="7" pageNumber="8">LAOS</collectingCountry>
:
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(
<emphasis id="B9175921FF97A42EC3A7F823164D150B" bold="true" box="[473,582,1821,1846]" pageId="7" pageNumber="8">
<typeStatus id="54D83B91FF97A42EC3A7F823164D150B" box="[473,582,1821,1846]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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, SWU),
<date id="FFDDA3F3FF97A42EC0D1F8201705150B" box="[687,782,1822,1846]" pageId="7" pageNumber="8" value="2014-06">
<collectingDate id="EF995A1BFF97A42EC0D1F8201705150B" box="[687,782,1822,1846]" pageId="7" pageNumber="8" value="2014-06">VI.2014</collectingDate>
</date>
,
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,
<collectingRegion id="49A74BD1FF97A42EC190F8201054150B" box="[1006,1119,1821,1846]" country="Laos" name="Houaphan" pageId="7" pageNumber="8">Hua Phan</collectingRegion>
Prov.,
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(
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):
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(
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),
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<collectingDate id="EF995A1BFF97A42EC3F7F87C15E31567" box="[393,488,1858,1882]" pageId="7" pageNumber="8" value="2014-06">VI.2014</collectingDate>
</date>
,
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,
<collectingRegion id="49A74BD1FF97A42EC0BEF87C17251567" box="[704,814,1857,1882]" country="Laos" name="Houaphan" pageId="7" pageNumber="8">Hua Phan</collectingRegion>
Prov.,
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;
<materialsCitation id="3B0B8F6EFF97A42EC195F87C160B1543" ID-GBIF-Occurrence="2004970361" collectingDate="2010-06-10" collectingDateMax="2010-06-21" collectingDateMin="2010-06-10" collectionCode="KSCJ" country="Laos" location="St. Jakl" municipality="Phou Pan" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Houa Phan" typeStatus="holotype">
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(
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),
<date id="FFDDA3F3FF97A42EC607F87C112F1567" box="[1145,1316,1858,1882]" pageId="7" pageNumber="8" value="2010-06-10" valueMax="2010-06-21" valueMin="2010-06-10">
<collectingDate id="EF995A1BFF97A42EC607F87C112F1567" box="[1145,1316,1858,1882]" pageId="7" pageNumber="8" value="2010-06-10" valueMax="2010-06-21" valueMin="2010-06-10">10–21.VI.2010</collectingDate>
</date>
,
<collectingMunicipality id="6BB81F49FF97A42EC74FF87F11971564" box="[1329,1436,1857,1882]" pageId="7" pageNumber="8">Phou Pan</collectingMunicipality>
(Mt.),
<collectingRegion id="49A74BD1FF97A42EC2A1F85815501543" box="[223,347,1893,1918]" inferredBySuffix="true" pageId="7" pageNumber="8">Houa Phan</collectingRegion>
Prov.,
<location id="8EBCD3E8FF97A42EC3D4F858160B1543" LSID="urn:lsid:plazi:treatment:03CA3425FF97A421C2E9F98C103614E2:8EBCD3E8FF97A42EC3D4F858160B1543" box="[426,512,1893,1918]" country="Laos" municipality="Phou Pan" name="St. Jakl" pageId="7" pageNumber="8" stateProvince="Houa Phan">St. Jakl</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC8533FF97A421C2B9F8B714C41641" blockId="7.[151,1437,1821,2026]" lastBlockId="8.[151,1437,151,1760]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B9175921FF97A42EC2B9F8B71602159F" bold="true" box="[199,521,1929,1954]" pageId="7" pageNumber="8">
<typeStatus id="54D83B91FF97A42EC2B9F8B7153F159F" box="[199,308,1929,1954]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
(male). General:
</emphasis>
Body length 18.0 mm; width 8.0 mm, widest at humeral umbone, gradually narrowed backward. Body brown. Surface with fulvous, brown, light yellow and black setae, and yellow tomentous maculae.
<emphasis id="B9175921FF97A42EC37AF8EF154515D7" bold="true" box="[260,334,2001,2026]" pageId="7" pageNumber="8">Head:</emphasis>
Dorsal surface with dense, setiferous punctures; setae fulvous, short. Clypeus with week green metallic reflections; anterior margin nearly straight and slightly raised; clypeolateral ridge not distinct. Frons with a slightly raised longitudinal ridge (
<figureCitation id="135899B6FF98A421C06CFF82165812E9" box="[530,595,188,213]" captionStart="FIGURES 1–23" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 1–23. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 2–3, M. dianensis new species; 4–5, M. fulvoguttatus; 6–7, M. fulvopilosus; 8–9, M. miksici new species; 10–11, M. nigropubescens; 12–13, M. olivaceofuscus; 14–15, M. sexmaculatus; 16–17, M. vuilleti; 18–19, M. curvimarginatus; 20–21, M. medogensis new species; 22–23, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
). Maxillary palpus and labial palpus brown. Antenna brown, antennal club long, about 1.5 times length of antennomeres 2–7 combined; inner side of antennomere 8 with short, dense, fulvous setae (
<figureCitation id="135899B6FF98A421C2A0FE3A15141321" box="[222,287,260,285]" captionStart="FIGURES 1–23" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 1–23. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 2–3, M. dianensis new species; 4–5, M. fulvoguttatus; 6–7, M. fulvopilosus; 8–9, M. miksici new species; 10–11, M. nigropubescens; 12–13, M. olivaceofuscus; 14–15, M. sexmaculatus; 16–17, M. vuilleti; 18–19, M. curvimarginatus; 20–21, M. medogensis new species; 22–23, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
). Ventral surface clad with long, orange setae (
<figureCitation id="135899B6FF98A421C14FFE3A179B1320" box="[817,912,260,285]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 136</figureCitation>
).
<emphasis id="B9175921FF98A421C1D9FE3A10221320" bold="true" box="[935,1065,260,285]" pageId="8" pageNumber="9">Pronotum:</emphasis>
Widest at base, basomedian area depressed. Lateral margin curved; posterior margin distinctly protruded. Surface clad with short, reddish-brown setae except for basolateral area; basolateral area clad with long, black and light yellow setae (
<figureCitation id="135899B6FF98A421C758FE7211861358" box="[1318,1421,332,357]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 135</figureCitation>
).
<emphasis id="B9175921FF98A421C2E9FE51151313B5" bold="true" box="[151,280,367,392]" pageId="8" pageNumber="9">Scutellum:</emphasis>
Surface clad with short, sparse, black setae.
<emphasis id="B9175921FF98A421C176FE51176713B5" bold="true" box="[776,876,367,392]" pageId="8" pageNumber="9">Elytron:</emphasis>
With posthumeral macula, lateral macula, median macula, and distal macula; median macula and distal macula small. Surface densely clad with setae; setae light yellow on maculae; setae black behind humeral umbone and on lateral and distal declivity; setae dark brown on juxtascutellar area; setae long, fulvous on disc; setae fulvous along lateral margins; humeral umbone glabrous (
<figureCitation id="135899B6FF98A421C710FEE214C91025" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 135</figureCitation>
).
<emphasis id="B9175921FF98A421C29AFEC1159B1025" bold="true" box="[228,400,511,536]" pageId="8" pageNumber="9">Mesepimeron:</emphasis>
Dark brown, clad with light yellow and dark brown setae.
<emphasis id="B9175921FF98A421C6DAFEC115351000" bold="true" pageId="8" pageNumber="9">Metepisternum and metepimeron:</emphasis>
Dark brown, clad with dark brown and black setae.
<emphasis id="B9175921FF98A421C1E7FD1A10021000" bold="true" box="[921,1033,548,573]" pageId="8" pageNumber="9">Sternum:</emphasis>
Preprosternum dark brown, almost glabrous. Mesosternum black, clad with long, sparse, light yellow setae. Mesometasternal process short, glabrous, brown, apex rounded; mesometasternal suture depressed, clad with long, yellow setae. Metasternum black, with long, sparse, yellow and brown setae; middle portion glabrous (
<figureCitation id="135899B6FF98A421C10BFDB117DC1095" box="[885,983,655,680]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 136</figureCitation>
).
<emphasis id="B9175921FF98A421C191FDB110671095" bold="true" box="[1007,1132,655,680]" pageId="8" pageNumber="9">Pygidium:</emphasis>
Dark brown. Surface clad with short, dense, fulvous and black setae; setae fulvous in middle (
<figureCitation id="135899B6FF98A421C1D4FD8A100110F1" box="[938,1034,692,717]" captionStart="FIGURES 72–98" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 72–98. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 72–73, M. biserratus new species; 74–75, M. dianensis new species; 76–79, M. fulvoguttatus; 80–83, M. fulvopilosus; 84–85, M. miksici new species; 86–89, M. nigropubescens; 90–91, M. olivaceofuscus; 92–93, M. sexmaculatus; 94–97, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="8" pageNumber="9">Figs. 72</figureCitation>
,
<figureCitation id="135899B6FF98A421C667FD8A104F10F0" box="[1049,1092,692,717]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">135</figureCitation>
).
<emphasis id="B9175921FF98A421C623FD8A10D010F0" bold="true" box="[1117,1243,692,717]" pageId="8" pageNumber="9">Abdomen:</emphasis>
Without median groove. Six abdominal sternites visible; sternites II–V clad with yellow setae; setae on sternite II long, dense; setae on sternites III–V short, sparse, but longer on sides; setae near posterior margin of sternite VI long, orange; sternite VII with sparse, short, fulvous setae on sides. Sternites II–V with a small, indistinct yellow macula close to posterior margin on each side; densely clad with fulvous and black setae in dorsal and lateral portion (
<figureCitation id="135899B6FF98A421C752FC7A11861160" box="[1324,1421,836,861]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 136</figureCitation>
).
<emphasis id="B9175921FF98A421C2E9FC5914D311BD" bold="true" box="[151,216,871,896]" pageId="8" pageNumber="9">Legs:</emphasis>
Slender, simple. Metacoxa brown; clad with long, orange and yellow setae. Femora and tibia reddish-brown; clad with long, orange and yellow setae. Tiny teeth between 3 large teeth of protibia distinct (
<figureCitation id="135899B6FF98A421C6C0FCB2111B1198" box="[1214,1296,908,933]" captionStart="FIGURES 24–47" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 24–47. Protibia of Macronotops (male and female). 24, M. biserratus new species; 25–26, M. dianensis new species; 27–28, M. fulvoguttatus; 29–30, M. fulvopilosus; 31–32, M. miksici new species; 33–34, M. nigropubescens; 35–36, M. olivaceofuscus; 37–38, M. sexmaculatus; 39–40, M. vuilleti; 41, Macronotops sp.; 42–43, M. curvimarginatus; 44–45, M. medogensis new species; 46–47, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="8" pageNumber="9">Fig. 25</figureCitation>
); spine near the middle of outer margin of mesotibia sharp, but blunt in metatibia; metatibia with a row of dense, brush-like, orange setae along inner margin (
<figureCitation id="135899B6FF98A421C067FCEA16B611D0" box="[537,701,980,1005]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Figs. 135–136</figureCitation>
); dorsal tooth of metatibia longer than basitarsus of metatarsus, outer tooth small and short (
<figureCitation id="135899B6FF98A421C3A5FCC91625162D" box="[475,558,1015,1040]" captionStart="FIGURES 48–65" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 48–65. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 49–50, M. dianensis new species; 51–52, M. fulvoguttatus; 53–54, M. fulvopilosus; 55–56, M. miksici new species; 57–58, M. nigropubescens;59–60, M. olivaceofuscus; 61–62, M. sexmaculatus; 63–64, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="8" pageNumber="9">Fig. 48</figureCitation>
). Tarsi reddish-brown; basitarsus of metatarsus with a cluster of sparse, long setae (
<figureCitation id="135899B6FF98A421C29FFB22153E1609" box="[225,309,1052,1077]" captionStart="FIGURES 48–65" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 48–65. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 49–50, M. dianensis new species; 51–52, M. fulvoguttatus; 53–54, M. fulvopilosus; 55–56, M. miksici new species; 57–58, M. nigropubescens;59–60, M. olivaceofuscus; 61–62, M. sexmaculatus; 63–64, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="8" pageNumber="9">Fig. 48</figureCitation>
).
<emphasis id="B9175921FF98A421C330FB2215CD1608" bold="true" box="[334,454,1052,1077]" pageId="8" pageNumber="9">Genitalia:</emphasis>
Parameres narrow, long; apex rounded, slightly expanded. Interparameral split wide; base with membrane, distinctly divided in medial. Median lobe ribbon-like; distinctly longer than paramere (
<figureCitation id="135899B6FF98A421C710FB0114B41641" captionStart="FIGURES 111–122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111–122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="8" pageNumber="9">Fig. 111</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FF98A421C2B9FBB9106616F9" pageId="8" pageNumber="9" type="description">
<paragraph id="8BDC8533FF98A421C2B9FBB915A9169D" blockId="8.[151,1437,151,1760]" box="[199,418,1159,1184]" pageId="8" pageNumber="9">
<emphasis id="B9175921FF98A421C2B9FBB9152E169D" bold="true" box="[199,293,1159,1184]" pageId="8" pageNumber="9">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BDC8533FF98A421C2B9FB92106616F9" blockId="8.[151,1437,151,1760]" box="[199,1133,1196,1221]" pageId="8" pageNumber="9">
<emphasis id="B9175921FF98A421C2B9FB92154616F8" bold="true" box="[199,333,1196,1221]" pageId="8" pageNumber="9">Variability.</emphasis>
Pygidium of the male
<typeStatus id="54D83B91FF98A421C032FB9316A716F8" box="[588,684,1197,1221]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
with a distinct yellow macula (
<figureCitation id="135899B6FF98A421C673FB92105516F9" box="[1037,1118,1196,1221]" captionStart="FIGURES 72–98" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 72–98. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 72–73, M. biserratus new species; 74–75, M. dianensis new species; 76–79, M. fulvoguttatus; 80–83, M. fulvopilosus; 84–85, M. miksici new species; 86–89, M. nigropubescens; 90–91, M. olivaceofuscus; 92–93, M. sexmaculatus; 94–97, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="8" pageNumber="9">Fig. 73</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FF98A421C2B9FBF1176A17D9" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BDC8533FF98A421C2B9FBF11693174A" blockId="8.[151,1437,151,1760]" pageId="8" pageNumber="9">
<emphasis id="B9175921FF98A421C2B9FBF1162916D5" bold="true" box="[199,546,1231,1256]" pageId="8" pageNumber="9">Differential diagnosis (male).</emphasis>
Length of clypeus almost equal to width. Antenna brown (
<figureCitation id="135899B6FF98A421C6BCFBF1110816D5" box="[1218,1283,1231,1256]" captionStart="FIGURES 1–23" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 1–23. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 2–3, M. dianensis new species; 4–5, M. fulvoguttatus; 6–7, M. fulvopilosus; 8–9, M. miksici new species; 10–11, M. nigropubescens; 12–13, M. olivaceofuscus; 14–15, M. sexmaculatus; 16–17, M. vuilleti; 18–19, M. curvimarginatus; 20–21, M. medogensis new species; 22–23, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
). Elytra dark brown, with posthumeral maculae, median maculae, lateral maculae and distal maculae (
<figureCitation id="135899B6FF98A421C600FBCA10D51730" box="[1150,1246,1268,1293]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 135</figureCitation>
). Pygidium with numerous black setae (
<figureCitation id="135899B6FF98A421C3E4FA2915E6170D" box="[410,493,1303,1328]" captionStart="FIGURES 72–98" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 72–98. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 72–73, M. biserratus new species; 74–75, M. dianensis new species; 76–79, M. fulvoguttatus; 80–83, M. fulvopilosus; 84–85, M. miksici new species; 86–89, M. nigropubescens; 90–91, M. olivaceofuscus; 92–93, M. sexmaculatus; 94–97, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="8" pageNumber="9">Fig. 72</figureCitation>
). Abdominal sternites II–V with a small or indistinct, yellow maculae on each side (
<figureCitation id="135899B6FF98A421C2E1FA02150E1768" box="[159,261,1340,1365]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Fig. 136</figureCitation>
); male abdomen without median groove. Protibia with tiny teeth (
<figureCitation id="135899B6FF98A421C673FA0210681769" box="[1037,1123,1340,1365]" captionStart="FIGURES 24–47" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 24–47. Protibia of Macronotops (male and female). 24, M. biserratus new species; 25–26, M. dianensis new species; 27–28, M. fulvoguttatus; 29–30, M. fulvopilosus; 31–32, M. miksici new species; 33–34, M. nigropubescens; 35–36, M. olivaceofuscus; 37–38, M. sexmaculatus; 39–40, M. vuilleti; 41, Macronotops sp.; 42–43, M. curvimarginatus; 44–45, M. medogensis new species; 46–47, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="8" pageNumber="9">Fig. 24</figureCitation>
); dorsal tooth of metatibia longer than basitarsus of metatarsus (
<figureCitation id="135899B6FF98A421C049FA6116821745" box="[567,649,1375,1400]" captionStart="FIGURES 48–65" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 48–65. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 49–50, M. dianensis new species; 51–52, M. fulvoguttatus; 53–54, M. fulvopilosus; 55–56, M. miksici new species; 57–58, M. nigropubescens;59–60, M. olivaceofuscus; 61–62, M. sexmaculatus; 63–64, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="8" pageNumber="9">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="8BDC8533FF98A421C2B9FABA176A17D9" blockId="8.[151,1437,151,1760]" pageId="8" pageNumber="9">
This new species is similar to
<taxonomicName id="4C63FEB0FF98A421C05AFABB169317A0" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[548,664,1412,1437]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FF98A421C05AFABB169317A0" box="[548,664,1412,1437]" italics="true" pageId="8" pageNumber="9">M. vuilleti</emphasis>
</taxonomicName>
, but can be separated by male protibia with several tiny teeth (
<figureCitation id="135899B6FF98A421C710FABA14BF17FD" captionStart="FIGURES 24–47" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 24–47. Protibia of Macronotops (male and female). 24, M. biserratus new species; 25–26, M. dianensis new species; 27–28, M. fulvoguttatus; 29–30, M. fulvopilosus; 31–32, M. miksici new species; 33–34, M. nigropubescens; 35–36, M. olivaceofuscus; 37–38, M. sexmaculatus; 39–40, M. vuilleti; 41, Macronotops sp.; 42–43, M. curvimarginatus; 44–45, M. medogensis new species; 46–47, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="8" pageNumber="9">Fig. 24</figureCitation>
), numerous black setae on pygidium (
<figureCitation id="135899B6FF98A421C009FA9916C117FD" box="[631,714,1447,1472]" captionStart="FIGURES 72–98" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 72–98. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 72–73, M. biserratus new species; 74–75, M. dianensis new species; 76–79, M. fulvoguttatus; 80–83, M. fulvopilosus; 84–85, M. miksici new species; 86–89, M. nigropubescens; 90–91, M. olivaceofuscus; 92–93, M. sexmaculatus; 94–97, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="8" pageNumber="9">Fig. 72</figureCitation>
), distinctly longer legs (
<figureCitation id="135899B6FF98A421C194FA99108417FD" box="[1002,1167,1447,1472]" captionStart="FIGURES 135–136" captionStartId="19.[151,264,1266,1289]" captionTargetBox="[182,1373,208,1232]" captionTargetId="figure@19.[151,1436,193,1245]" captionTargetPageId="19" captionText="FIGURES 135–136. Habitus of Macronotops biserratus new species (dorsal and ventral view). Male holotype." httpUri="https://zenodo.org/record/2585224/files/figure.png" pageId="8" pageNumber="9">Figs. 135–136</figureCitation>
), elongated parameres, wider median lobe, and distinctly divided membrane (
<figureCitation id="135899B6FF98A421C08AFAF2175917D9" box="[756,850,1484,1509]" captionStart="FIGURES 111–122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111–122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="8" pageNumber="9">Fig. 111</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FF98A421C2B9FAD115EB146D" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BDC8533FF98A421C2B9FAD115EB146D" blockId="8.[151,1437,151,1760]" pageId="8" pageNumber="9">
<emphasis id="B9175921FF98A421C2B9FAD115451435" bold="true" box="[199,334,1519,1544]" pageId="8" pageNumber="9">Etymology.</emphasis>
The species name “
<taxonomicName id="4C63FEB0FF98A421C048FAD116A31435" authorityName="Qiu &amp; Xu &amp; Chen" authorityYear="2019" box="[566,680,1519,1544]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis id="B9175921FF98A421C048FAD116A31435" box="[566,680,1519,1544]" italics="true" pageId="8" pageNumber="9">biserratus</emphasis>
</taxonomicName>
” is a combination of the Latin words “
<emphasis id="B9175921FF98A421C615FAD1108B1435" box="[1131,1152,1519,1544]" italics="true" pageId="8" pageNumber="9">bi</emphasis>
” meaning “double”, and “
<emphasis id="B9175921FF98A421C2DDF92B14F41411" box="[163,255,1557,1580]" italics="true" pageId="8" pageNumber="9">serratus</emphasis>
” meaning “serrated”, since this species is similar to
<taxonomicName id="4C63FEB0FF98A421C121F92B17DF1410" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[863,980,1556,1581]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FF98A421C121F92B17DF1410" box="[863,980,1556,1581]" italics="true" pageId="8" pageNumber="9">M. vuilleti</emphasis>
</taxonomicName>
, but the protibia bear distinct tiny teeth between the three large teeth.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FF98A421C2B9F96215AE1449" box="[199,421,1628,1653]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BDC8533FF98A421C2B9F96215AE1449" blockId="8.[151,1437,151,1760]" box="[199,421,1628,1653]" pageId="8" pageNumber="9">
<emphasis id="B9175921FF98A421C2B9F962156B1448" bold="true" box="[199,352,1628,1653]" pageId="8" pageNumber="9">Distribution.</emphasis>
<collectingCountry id="F374C5A3FF98A421C319F96315AA1449" box="[359,417,1629,1652]" name="Laos" pageId="8" pageNumber="9">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FF98A421C2B9F941103614E2" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BDC8533FF98A421C2B9F941103614E2" blockId="8.[151,1437,151,1760]" pageId="8" pageNumber="9">
<emphasis id="B9175921FF98A421C2B9F941153014A5" bold="true" box="[199,315,1663,1688]" pageId="8" pageNumber="9">Remarks.</emphasis>
This species occurs sympatrically with
<taxonomicName id="4C63FEB0FF98A421C081F9BF177814A5" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[767,883,1663,1688]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FF98A421C081F9BF177814A5" box="[767,883,1663,1688]" italics="true" pageId="8" pageNumber="9">M. vuilleti</emphasis>
</taxonomicName>
at Mount Phu Pane,
<collectingCountry id="F374C5A3FF98A421C61CF9BE109014AA" box="[1122,1179,1664,1687]" name="Laos" pageId="8" pageNumber="9">Laos</collectingCountry>
, and these two species are very similar, but several morphological differences (see differential diagnosis section) are observed, especially in tiny teeth and male genitalia which are stable diagnostic characters of this genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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