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: 6
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, 42;
<treatmentCitation id="0A8D0659FFE0095E6554FACFCEDEFAAB" author="VIENNA P." page="115" pageId="119" pageNumber="112" year="1980">
<bibRefCitation id="EFBD5DB9FFE0095E6554FACFCEE1FAAB" author="VIENNA P." pageId="119" pageNumber="112" refId="ref122194" refString="VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp." type="book" year="1980">
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(1980)
</bibRefCitation>
: 115
</treatmentCitation>
, 117;
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<bibRefCitation id="EFBD5DB9FFE0095E6073FAA5CF82FAAB" author="MAZUR S." box="[292,424,1346,1365]" pageId="119" pageNumber="112" refId="ref118463" refString="MAZUR S. 1981 a: Histeridae - Gnilikowate (Insecta: Coleoptera). Fauna Polski, Tom 9. Polish Entomological Society, Warszawa, 206 pp (In Polish)" type="book" year="1981">MAZUR (1981a)</bibRefCitation>
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<treatmentCitation id="0A8D0659FFE0095E60BCFAA5CCA4FAAB" author="MAZUR S." box="[491,654,1346,1365]" page="103" pageId="119" pageNumber="112" year="1984">
<bibRefCitation id="EFBD5DB9FFE0095E60BCFAA5CC4FFAAB" author="MAZUR S." box="[491,613,1346,1365]" pageId="119" pageNumber="112" pagination="1 - 376" refId="ref118519" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3 - 4): 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
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<bibRefCitation id="EFBD5DB9FFE0095E63C1FAA6CD24FAAB" author="OHARA M." box="[662,782,1345,1365]" pageId="119" pageNumber="112" pagination="1 - 238" refId="ref119358" refString="OHARA M. 1994: A revision of the superfamily Histeroidea of Japan (Coleoptera). Insecta Matsumurana (N. S.) 51: 1 - 238." type="journal article" year="1994">ÔHARA (1994)</bibRefCitation>
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<bibRefCitation id="EFBD5DB9FFE0095E6269FAA5CA3FFAAB" author="DOWNIE N. M. &amp; ARNETT R. H. JR." box="[830,1045,1346,1365]" pageId="119" pageNumber="112" refId="ref114280" refString="DOWNIE N. M. &amp; ARNETT R. H. JR. 1996: The beetles of northeastern North America. Volume I: Introduction; suborders Archostemata, Adephaga, and Polyphaga, thru superfamily Cantharoidea. The Sandhill Crane Press, Gainesvilles, FL, xiv + 879 pp." type="book" year="1996">DOWNIE &amp; ARNETT (1996)</bibRefCitation>
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, 612;
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<bibRefCitation id="EFBD5DB9FFE0095E61ECFABACF1CFA8E" author="MAZUR S." box="[187,310,1373,1392]" pageId="119" pageNumber="112" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
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<bibRefCitation id="EFBD5DB9FFE0095E603DFABACC40FA8E" author="BOUSQUET Y. &amp; LAPLANTE S." box="[362,618,1373,1392]" pageId="119" pageNumber="112" pagination="1 - 190" refId="ref112971" refString="BOUSQUET Y. &amp; LAPLANTE S. 1999: Les Coleopteres Histerides du Quebec. Fabreries, Supplement 8: 1 - 190." type="journal article" year="1999">BOUSQUET &amp; LAPLANTE (1999)</bibRefCitation>
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, 142;
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<bibRefCitation id="EFBD5DB9FFE0095E639FFABACD6EFA8E" author="MAZUR S." box="[712,836,1373,1392]" pageId="119" pageNumber="112" pagination="17 - 66" refId="ref118660" refString="MAZUR S. 2001: Review of the Histeridae (Coleoptera) of Mexico. Dugesiana 8: 17 - 66." type="journal article" year="2001">MAZUR (2001)</bibRefCitation>
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<treatmentCitation id="0A8D0659FFE0095E62DAFABACEDFFA74" author="KOVARIK P. W. &amp; CATERINO M. S." page="221" pageId="119" pageNumber="112" year="2001">
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<bibRefCitation id="EFBD5DB9FFE0095E6346FA9FCCA7FA74" author="MAZUR S." box="[529,653,1399,1419]" pageId="119" pageNumber="112" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR (2004)</bibRefCitation>
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81.
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</paragraph>
<paragraph id="8B932048FFE0095E6085FB6FCF39FB4B" blockId="119.[113,1090,1159,1419]" pageId="119" pageNumber="112">
<typeStatus id="54979EEAFFE0095E6085FB6FCFD4FB65" box="[466,510,1160,1179]" pageId="119" pageNumber="112">Type</typeStatus>
species:
<taxonomicName id="4C2C5BCBFFE0095E631CFB6FCD5AFB64" ID-CoL="9MR68" authority="Kugelann, 1792" authorityName="Kugelann" authorityYear="1792" box="[587,880,1159,1178]" class="Insecta" family="Histeridae" genus="Hister" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="112" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B958FC5AFFE0095E631CFB6FCCCBFB64" box="[587,737,1160,1178]" italics="true" pageId="119" pageNumber="112">Hister rotundatus</emphasis>
<bibRefCitation id="EFBD5DB9FFE0095E63B1FB60CD5AFB64" author="KUGELANN J. G." box="[742,880,1159,1178]" pageId="119" pageNumber="112" pagination="513 - 582" refId="ref117125" refString="KUGELANN J. G. 1792: Verzeichniss der in einigen Gegenden Preussens bis jetz entdeckten Kafer-Arten, nebst kurzen Nachrichten von denselben. Neustes Magazin Fur die Liebhaber der Entomologie, 1, 5: 513 - 582." type="journal article" year="1792">Kugelann, 1792</bibRefCitation>
</taxonomicName>
, designated by C. THOM- SON (1859: 75).
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection id="C33673C3FFE009516126FA7CCFACFC8A" lastPageId="120" lastPageNumber="117" pageId="119" pageNumber="112" type="diagnosis">
<paragraph id="8B932048FFE009516126FA7CCD26FE4D" blockId="119.[113,1089,1435,1619]" lastBlockId="120.[151,1127,219,1620]" lastPageId="120" lastPageNumber="117" pageId="119" pageNumber="112">
<emphasis id="B958FC5AFFE0095E6126FA7CCECCFA4D" bold="true" box="[113,230,1435,1459]" pageId="119" pageNumber="112">Diagnosis.</emphasis>
Cuticle brown to black, never metallic; frontal, supraorbital striae absent; pronotum flattened; pronotal hypomeron glabrous; pronotal foveae absent; punctures of pronotum often forming longitudinal rugae laterally (in
<taxonomicName id="4C2C5BCBFFE0095E631AFA3BCD7BFA0D" authorityName="Reichardt" authorityYear="1930" box="[589,849,1500,1523]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="119" pageNumber="112" phylum="Arthropoda" rank="species" species="kiritshenkoi">
<emphasis id="B958FC5AFFE0095E631AFA3BCD7BFA0D" box="[589,849,1500,1523]" italics="true" pageId="119" pageNumber="112">Gnathoncus kiritshenkoi</emphasis>
</taxonomicName>
pronotal disc medially with curious tubercle); elytra often imbricate, punctures on apical part of elytra often forming longitudinal rugae; marginal epipleural stria usually double; fourth dorsal elytral stria never connected with sutural stria; apical elytral stria often shortened. In most species there is a characteristic hooked appendix between fourth dorsal and sutural striae at elytral base; anterior ends of fourth dorsal elytral and sutural elytral striae form a small hook. Prosternum without pre-apical foveae; median fossa often present; carinal prosternal striae strongly convergent anteriorly, often united under sharp angle; lateral prosternal striae shortened, strongly convergent anteriorly; prosternal process flattened, broad; outer-lateral costa reaches prosternal process, its basal margin distinctly elevated; metaventrite of males often longitudinally concave; ninth tergite of male genitalia divided longitudinally.
</paragraph>
<paragraph id="8B932048FFEF095161C0FE5CCFACFC8A" blockId="120.[151,1127,219,1620]" pageId="120" pageNumber="117">
<emphasis id="B958FC5AFFEF095161C0FE5CCFBBFE2D" bold="true" box="[151,401,443,467]" pageId="120" pageNumber="117">Differential diagnosis.</emphasis>
<taxonomicName id="4C2C5BCBFFEF095160C2FE5ACC3CFE2A" authorityName=": THOMSON" authorityYear="1867" box="[405,534,445,468]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF095160C2FE5ACC3CFE2A" box="[405,534,445,468]" italics="true" pageId="120" pageNumber="117">Gnathoncus</emphasis>
</taxonomicName>
is most similar to the presumably related genera
<taxonomicName id="4C2C5BCBFFEF09516541FE5ACED9FE0A" authorityName=": ROSS" authorityYear="1940" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF09516541FE5ACED9FE0A" italics="true" pageId="120" pageNumber="117">Eremosaprinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C5BCBFFEF0951607EFE3ACFB8FE0A" authorityName="Marseul" authorityYear="1862" box="[297,402,477,500]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF0951607EFE3ACFB8FE0A" box="[297,402,477,500]" italics="true" pageId="120" pageNumber="117">Myrmetes</emphasis>
</taxonomicName>
sharing with them several putative synapomorphies: absent frontal and supraorbital striae, body without metallic luster and longitudinally divided ninth tergite of male genitalia.
<taxonomicName id="4C2C5BCBFFEF09516001FDFACFFDFDCA" authorityName=": THOMSON" authorityYear="1867" box="[342,471,541,564]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF09516001FDFACFFDFDCA" box="[342,471,541,564]" italics="true" pageId="120" pageNumber="117">Gnathoncus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2C5BCBFFEF09516334FDFACD2CFDCA" authorityName=": ROSS" authorityYear="1940" box="[611,774,541,564]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF09516334FDFACD2CFDCA" box="[611,774,541,564]" italics="true" pageId="120" pageNumber="117">Eremosaprinus</emphasis>
</taxonomicName>
chiefly by the presence of lateral prosternal striae (absent in
<taxonomicName id="4C2C5BCBFFEF095160FBFDDACC66FDAA" authorityName=": ROSS" authorityYear="1940" box="[428,588,573,596]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF095160FBFDDACC66FDAA" box="[428,588,573,596]" italics="true" pageId="120" pageNumber="117">Eremosaprinus</emphasis>
</taxonomicName>
) and shorter meso- and metatibiae. From
<taxonomicName id="4C2C5BCBFFEF095162AFFDDACA4AFDAA" authorityName="Marseul" authorityYear="1862" box="[1016,1120,573,596]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF095162AFFDDACA4AFDAA" box="[1016,1120,573,596]" italics="true" pageId="120" pageNumber="117">Myrmetes</emphasis>
</taxonomicName>
, it differs by the presence of sutural elytral stria (absent in
<taxonomicName id="4C2C5BCBFFEF095163A3FDBACD77FD8A" authorityName="Marseul" authorityYear="1862" box="[756,861,605,628]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF095163A3FDBACD77FD8A" box="[756,861,605,628]" italics="true" pageId="120" pageNumber="117">Myrmetes</emphasis>
</taxonomicName>
), punctate dorsal surface (impunctate, matt in
<taxonomicName id="4C2C5BCBFFEF09516025FD9ACFF1FD6A" authorityName="Marseul" authorityYear="1862" box="[370,475,637,660]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF09516025FD9ACFF1FD6A" box="[370,475,637,660]" italics="true" pageId="120" pageNumber="117">Myrmetes</emphasis>
</taxonomicName>
), outer margin of protibia with teeth (only short denticles present in
<taxonomicName id="4C2C5BCBFFEF095161B1FD7ACF65FD4A" authorityName="Marseul" authorityYear="1862" box="[230,335,669,692]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF095161B1FD7ACF65FD4A" box="[230,335,669,692]" italics="true" pageId="120" pageNumber="117">Myrmetes</emphasis>
</taxonomicName>
) and the characteristic hooked appendix between the fourth dorsal elytral and sutural elytral striae (absent in
<taxonomicName id="4C2C5BCBFFEF0951635DFD5ACC59FD2A" authorityName="Marseul" authorityYear="1862" box="[522,627,701,724]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF0951635DFD5ACC59FD2A" box="[522,627,701,724]" italics="true" pageId="120" pageNumber="117">Myrmetes</emphasis>
</taxonomicName>
). Superficially also similar the sole Palaearctic representative of the subgenus
<taxonomicName id="4C2C5BCBFFEF0951608FFD3ACC74FD0A" box="[472,606,733,756]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="subGenus" subGenus="Neosaprinus">
<emphasis id="B958FC5AFFEF0951608FFD3ACC74FD0A" box="[472,606,733,756]" italics="true" pageId="120" pageNumber="117">Neosaprinus</emphasis>
</taxonomicName>
of the genus
<taxonomicName id="4C2C5BCBFFEF095163B1FD3ACD7CFD0A" authorityName=": BICKHARDT" authorityYear="1916" box="[742,854,733,756]" class="Insecta" family="Histeridae" genus="Euspilotus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF095163B1FD3ACD7CFD0A" box="[742,854,733,756]" italics="true" pageId="120" pageNumber="117">Euspilotus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516236FD3ACA4AFD0A" baseAuthorityName="Marseul" baseAuthorityYear="1872" box="[865,1120,732,756]" class="Insecta" family="Histeridae" genus="Euspilotus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="perrisi" subGenus="Neosaprinus">
<emphasis id="B958FC5AFFEF09516236FD3ACD53FD0A" box="[865,889,733,756]" italics="true" pageId="120" pageNumber="117">E.</emphasis>
(
<emphasis id="B958FC5AFFEF095162D1FD3ACA26FD0A" box="[902,1036,733,756]" italics="true" pageId="120" pageNumber="117">Neosaprinus</emphasis>
)
<emphasis id="B958FC5AFFEF0951654EFD3ACA4AFD0A" box="[1049,1120,733,756]" italics="true" pageId="120" pageNumber="117">perrisi</emphasis>
</taxonomicName>
, but the species of
<taxonomicName id="4C2C5BCBFFEF09516037FD1ACFCBFCEA" authorityName=": THOMSON" authorityYear="1867" box="[352,481,765,788]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF09516037FD1ACFCBFCEA" box="[352,481,765,788]" italics="true" pageId="120" pageNumber="117">Gnathoncus</emphasis>
</taxonomicName>
differ from it by absent preapical foveae and flattened and broad prosternal process (distinctly convex in
<taxonomicName id="4C2C5BCBFFEF09516320FCFACD5CFCCA" baseAuthorityName="Marseul" baseAuthorityYear="1872" box="[631,886,796,820]" class="Insecta" family="Histeridae" genus="Euspilotus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="perrisi" subGenus="Neosaprinus">
<emphasis id="B958FC5AFFEF09516320FCFACCA4FCCA" box="[631,654,797,820]" italics="true" pageId="120" pageNumber="117">E.</emphasis>
(
<emphasis id="B958FC5AFFEF095163CCFCFACD0BFCCA" box="[667,801,797,820]" italics="true" pageId="120" pageNumber="117">Neosaprinus</emphasis>
)
<emphasis id="B958FC5AFFEF09516278FCFACD5CFCCA" box="[815,886,797,820]" italics="true" pageId="120" pageNumber="117">perrisi</emphasis>
</taxonomicName>
). Female specimens of
<taxonomicName id="4C2C5BCBFFEF095161C0FCDACFB1FCAA" baseAuthorityName="Marseul" baseAuthorityYear="1872" box="[151,411,828,852]" class="Insecta" family="Histeridae" genus="Euspilotus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="perrisi" subGenus="Neosaprinus">
<emphasis id="B958FC5AFFEF095161C0FCDACE85FCAA" box="[151,175,829,852]" italics="true" pageId="120" pageNumber="117">E.</emphasis>
(
<emphasis id="B958FC5AFFEF095161E9FCDACF6FFCAA" box="[190,325,829,852]" italics="true" pageId="120" pageNumber="117">Neosaprinus</emphasis>
)
<emphasis id="B958FC5AFFEF09516003FCDACFB1FCAA" box="[340,411,829,852]" italics="true" pageId="120" pageNumber="117">perrisi</emphasis>
</taxonomicName>
also possess curious pygidial sulci (
<figureCitation id="13173CCDFFEF09516249FCDCCD93FCAD" box="[798,953,827,851]" captionStart="Figs" captionStartId="56.[152,191,1557,1576]" captionTargetBox="[227,1045,933,1531]" captionTargetId="figure-254@56.[220,1054,928,1534]" captionTargetPageId="56" captionText="Figs. 162167.Euspilotus (Neosaprinus) perrisi (Marseul, 1872), variability of female pygidial sulci. Figs. 162165 after OLEXA (1975)." figureDoi="http://doi.org/10.5281/zenodo.4272181" httpUri="https://zenodo.org/record/4272181/files/figure.png" pageId="120" pageNumber="117">Figs. 162167</figureCitation>
) that are always absent in
<emphasis id="B958FC5AFFEF095161AAFCBACFACFC8A" box="[253,390,861,884]" italics="true" pageId="120" pageNumber="117">
<taxonomicName id="4C2C5BCBFFEF095161AAFCBACFABFC8A" authorityName=": THOMSON" authorityYear="1867" box="[253,385,861,884]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">Gnathoncus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FFEF095161C0FC9CCF1CFB6D" pageId="120" pageNumber="117" type="biology_ecology">
<paragraph id="8B932048FFEF095161C0FC9CCF1CFB6D" blockId="120.[151,1127,219,1620]" pageId="120" pageNumber="117">
<emphasis id="B958FC5AFFEF095161C0FC9CCEDEFC6D" bold="true" box="[151,244,891,915]" pageId="120" pageNumber="117">Biology.</emphasis>
Predominantly inquilinous genus, its representatives are found within nests of birds or mammals; some species are found exclusively inside these nests where they are an active and important part of the local communities (
<bibRefCitation id="EFBD5DB9FFEF09516322FC5BCA34FC2A" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[629,1054,955,980]" pageId="120" pageNumber="117" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT 1976: 114</bibRefCitation>
). They presumably prey upon larvae of fleas and other tiny arthropods thus reducing their number within the nests. Some species, however, are occasionally collected also on carrion. At least one species (
<taxonomicName id="4C2C5BCBFFEF09516076FBFACCB8FBCA" authority="Auzat, 1923" authorityName="Auzat" authorityYear="1923" box="[289,658,1052,1076]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="B958FC5AFFEF09516076FBFACC2CFBCA" box="[289,518,1053,1076]" italics="true" pageId="120" pageNumber="117">Gnathoncus cerberus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF0951635AFBFBCCB8FBCA" author="AUZAT V." box="[525,658,1052,1076]" pageId="120" pageNumber="117" pagination="145 - 147" refId="ref112516" refString="AUZAT V. 1923: Description d'un Gnathoncus de Sardaigne (Coleopt. Hist.). Bolletino della Societa Entomologica Italiana 55: 145 - 147." type="journal article" year="1923">Auzat, 1923</bibRefCitation>
</taxonomicName>
) lives cavernicolously, probably feeding on flies and other arthropods developing in guano. Several species of
<taxonomicName id="4C2C5BCBFFEF0951621FFBDACDE2FBAA" authorityName=": THOMSON" authorityYear="1867" box="[840,968,1085,1108]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEF0951621FFBDACDE2FBAA" box="[840,968,1085,1108]" italics="true" pageId="120" pageNumber="117">Gnathoncus</emphasis>
</taxonomicName>
(e.g.
<taxonomicName id="4C2C5BCBFFEF09516556FBDACF12FB8A" baseAuthorityName="Kugelann" baseAuthorityYear="1792" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B958FC5AFFEF09516556FBDACF12FB8A" italics="true" pageId="120" pageNumber="117">Gnathoncus rotundatus</emphasis>
</taxonomicName>
) are typical synanthropes and are often collected in pigsties, dovecotes or chicken coops.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FFEF095161C0FB7CCD71FA2D" pageId="120" pageNumber="117" type="distribution">
<paragraph id="8B932048FFEF095161C0FB7CCCFDFB0D" blockId="120.[151,1127,219,1620]" pageId="120" pageNumber="117">
<emphasis id="B958FC5AFFEF095161C0FB7CCF00FB4D" bold="true" box="[151,298,1179,1203]" pageId="120" pageNumber="117">Distribution.</emphasis>
Twenty-four species and subspecies are known to occur world-wide (
<bibRefCitation id="EFBD5DB9FFEF0951654CFB7BCEFAFB2A" author="MAZUR S." pageId="120" pageNumber="117" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR 1997</bibRefCitation>
); several species were possibly distributed over the globe by human activity. However, most of the species are found in the Holarctic
<collectingRegion id="49E8EEAAFFEF095163D5FB3CCCF9FB0D" box="[642,723,1243,1267]" country="Iraq" pageId="120" pageNumber="117">Region</collectingRegion>
.
</paragraph>
<paragraph id="8B932048FFEF095161C0FB1CCD71FA2D" blockId="120.[151,1127,219,1620]" pageId="120" pageNumber="117">
<emphasis id="B958FC5AFFEF095161C0FB1CCF4FFAED" bold="true" box="[151,357,1275,1299]" pageId="120" pageNumber="117">Species examined.</emphasis>
<taxonomicName id="4C2C5BCBFFEF0951603CFB1ACD0DFAEA" authority="Reichardt, 1941" authorityName="Reichardt" authorityYear="1941" box="[363,807,1275,1300]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="baeckmanni">
<emphasis id="B958FC5AFFEF0951603CFB1ACC5EFAEA" box="[363,628,1277,1300]" italics="true" pageId="120" pageNumber="117">Gnathoncus baeckmanni</emphasis>
<bibRefCitation id="EFBD5DB9FFEF0951632DFB1CCD0DFAEA" author="REICHARDT A." box="[634,807,1275,1300]" pageId="120" pageNumber="117" refId="ref120382" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">Reichardt, 1941</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516266FB1ACA48FAEA" authority="Lewis, 1907" authorityName="Lewis" authorityYear="1907" box="[817,1122,1275,1300]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="brevisternus">
<emphasis id="B958FC5AFFEF09516266FB1ACDFCFAEA" box="[817,982,1277,1300]" italics="true" pageId="120" pageNumber="117">G. brevisternus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF0951628BFB1CCA48FAEA" author="LEWIS G." box="[988,1122,1275,1300]" pageId="120" pageNumber="117" pagination="480 - 484" refId="ref117739" refString="LEWIS G. 1907: On a new species of Histeridae and notices of others. Annals and Magazine of Natural History, Series 7 20: 480 - 484." type="journal article" year="1907">Lewis, 1907</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF095161C0FAFACFB3FACA" authority="Auzat, 1917" authorityName="Auzat" authorityYear="1917" box="[151,409,1308,1332]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="buyssoni">
<emphasis id="B958FC5AFFEF095161C0FAFACF39FACA" box="[151,275,1309,1332]" italics="true" pageId="120" pageNumber="117">G. buyssoni</emphasis>
<bibRefCitation id="EFBD5DB9FFEF0951604FFAFBCFB3FACA" author="AUZAT V." box="[280,409,1308,1332]" pageId="120" pageNumber="117" pagination="184 - 185" refId="ref112459" refString="AUZAT V. 1917: Description d'un nouveau Gnathoncus de France (Col., Histeridae). Bulletin de la Societe Entomologique de France 1917: 184 - 185." type="journal article" year="1917">Auzat, 1917</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF095160F4FAFACC8FFACA" authority="Auzat, 1923" authorityName="Auzat" authorityYear="1923" box="[419,677,1308,1332]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="cerberus">
<emphasis id="B958FC5AFFEF095160F4FAFACC35FACA" box="[419,543,1309,1332]" italics="true" pageId="120" pageNumber="117">G. cerberus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF09516374FAFBCC8FFACA" author="AUZAT V." box="[547,677,1308,1332]" pageId="120" pageNumber="117" pagination="145 - 147" refId="ref112516" refString="AUZAT V. 1923: Description d'un Gnathoncus de Sardaigne (Coleopt. Hist.). Bolletino della Societa Entomologica Italiana 55: 145 - 147." type="journal article" year="1923">Auzat, 1923</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF095163F8FAFACDC3FACD" authority="(Marseul, 1862)" baseAuthorityName="Marseul" baseAuthorityYear="1862" box="[687,1001,1307,1332]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="communis">
<emphasis id="B958FC5AFFEF095163F8FAFACD13FACA" box="[687,825,1309,1332]" italics="true" pageId="120" pageNumber="117">G. communis</emphasis>
(
<bibRefCitation id="EFBD5DB9FFEF09516212FAFCCDC8FACD" author="MARSEUL S. A." box="[837,994,1307,1331]" pageId="120" pageNumber="117" pagination="677 - 758" refId="ref118029" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Serie 4 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF095162A5FAFACF94FAAD" authority="disjunctus Solskij, 1876" authorityName="Solskij" authorityYear="1876" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="disjunctus" subSpecies="disjunctus">
<emphasis id="B958FC5AFFEF095162A5FAFACF02FAAA" italics="true" pageId="120" pageNumber="117">G. disjunctus disjunctus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF09516078FADCCF94FAAD" author="SOLSKIJ S. M." box="[303,446,1339,1363]" pageId="120" pageNumber="117" pagination="223 - 398" refId="ref121457" refString="SOLSKIJ S. M. 1876: Zhestokrylye (Coleoptera). In: FEDTSCHENKO A. P. (ed.): Puteshestvie v Turkestan, t. 2, tsh. 5, tetr. 2. Izvestiya Imperatorskago Obshchestva Lyubiteley Estestvoznaniya, Antropologii i Etnografii 21 (1): 223 - 398 (in Russian)." type="journal article" year="1876">Solskij, 1876</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF0951609FFADACD60FAAD" authority="Reitter, 1896" authorityName="Reitter" authorityYear="1896" box="[456,842,1339,1364]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="disjunctus" subSpecies="suturifer">
<emphasis id="B958FC5AFFEF0951609FFADACC9CFAAA" box="[456,694,1341,1364]" italics="true" pageId="120" pageNumber="117">G. disjunctus suturifer</emphasis>
<bibRefCitation id="EFBD5DB9FFEF095163EBFADCCD60FAAD" author="REITTER E." box="[700,842,1339,1363]" pageId="120" pageNumber="117" pagination="306 - 308" refId="ref120457" refString="REITTER E. 1896: Uebersicht der mir bekannten Gnathoncus Arten der palearktischen Fauna. Entomologische Nachrichten 22: 306 - 308." type="journal article" year="1896">Reitter, 1896</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516203FADACEE5FA8A" authority="Reichardt, 1930" authorityName="Reichardt" authorityYear="1930" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="kiritshenkoi">
<emphasis id="B958FC5AFFEF09516203FADACDD8FAAA" box="[852,1010,1341,1364]" italics="true" pageId="120" pageNumber="117">G. kiritshenkoi</emphasis>
<bibRefCitation id="EFBD5DB9FFEF095162AFFADCCEE5FA8A" author="REICHARDT A." pageId="120" pageNumber="117" pagination="285 - 304" refId="ref120298" refString="REICHARDT A. 1930: De Histeridis (Coleoptera) novis faunae palaearcticae. Annuaire du Musee Zoologique 30: 285 - 304." type="journal article" year="1930">Reichardt, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF0951618BFABACC1FFA8D" authority="(Marseul, 1862)" baseAuthorityName="Marseul" baseAuthorityYear="1862" box="[220,565,1371,1396]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="nannetensis">
<emphasis id="B958FC5AFFEF0951618BFABACF56FA8A" box="[220,380,1373,1396]" italics="true" pageId="120" pageNumber="117">G. nannetensis</emphasis>
(
<bibRefCitation id="EFBD5DB9FFEF095160DBFABCCC07FA8D" author="MARSEUL S. A." box="[396,557,1371,1395]" pageId="120" pageNumber="117" pagination="677 - 758" refId="ref118029" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Serie 4 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">Marseul, 1862</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516315FABACDADFA8D" authority="Stockmann, 1957" authorityName="Stockmann" authorityYear="1957" box="[578,903,1371,1396]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="nidorum">
<emphasis id="B958FC5AFFEF09516315FABACC94FA8A" box="[578,702,1373,1396]" italics="true" pageId="120" pageNumber="117">G. nidorum</emphasis>
<bibRefCitation id="EFBD5DB9FFEF09516390FABCCDADFA8D" author="STOCKMANN S." box="[711,903,1371,1395]" pageId="120" pageNumber="117" pagination="67 - 76" refId="ref121660" refString="STOCKMANN S. 1957: Beitrage zur Kenntnis der Koleopterenfauna Ostfennoskandiens. 5. Die Gnathoncus-Arten Ostfennoskandiens. Notulae Entomologicae 37: 67 - 76." type="journal article" year="1957">Stockmann, 1957</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF095162C4FABACEE4FA6D" authority="Reitter, 1896" authorityName="Reitter" authorityYear="1896" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B958FC5AFFEF095162C4FABACA25FA8A" box="[915,1039,1373,1396]" italics="true" pageId="120" pageNumber="117">G. potanini</emphasis>
<bibRefCitation id="EFBD5DB9FFEF09516540FABCCEE4FA6D" author="REITTER E." pageId="120" pageNumber="117" pagination="306 - 308" refId="ref120457" refString="REITTER E. 1896: Uebersicht der mir bekannten Gnathoncus Arten der palearktischen Fauna. Entomologische Nachrichten 22: 306 - 308." type="journal article" year="1896">Reitter, 1896</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516180FA9ACC6AFA6D" authority="Kryzhanovskij, 1976" authorityName="Kryzhanovskij" authorityYear="1976" box="[215,576,1403,1428]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B958FC5AFFEF09516180FA9ACF74FA6A" box="[215,350,1405,1428]" italics="true" pageId="120" pageNumber="117">G. pygmaeus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF09516034FA9CCC6AFA6D" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[355,576,1403,1427]" pageId="120" pageNumber="117" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">Kryzhanovskij, 1976</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF0951631EFA9ACDB2FA6A" authority="(Kugelann, 1792)" baseAuthorityName="Kugelann" baseAuthorityYear="1792" box="[585,920,1403,1428]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B958FC5AFFEF0951631EFA9ACCF3FA6A" box="[585,729,1405,1428]" italics="true" pageId="120" pageNumber="117">G. rotundatus</emphasis>
(
<bibRefCitation id="EFBD5DB9FFEF095163B2FA9CCDBAFA6A" author="KUGELANN J. G." box="[741,912,1403,1428]" pageId="120" pageNumber="117" pagination="513 - 582" refId="ref117125" refString="KUGELANN J. G. 1792: Verzeichniss der in einigen Gegenden Preussens bis jetz entdeckten Kafer-Arten, nebst kurzen Nachrichten von denselben. Neustes Magazin Fur die Liebhaber der Entomologie, 1, 5: 513 - 582." type="journal article" year="1792">Kugelann, 1792</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF095162F7FA9ACF6EFA4A" authority="Bickhardt, 1920" authorityName="Bickhardt" authorityYear="1920" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="semimarginatus">
<emphasis id="B958FC5AFFEF095162F7FA9ACA4DFA6A" box="[928,1127,1405,1428]" italics="true" pageId="120" pageNumber="117">G. semimarginatus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF095161C0FA7CCF6EFA4A" author="BICKHARDT H." box="[151,324,1435,1460]" pageId="120" pageNumber="117" pagination="29 - 32" refId="ref112759" refString="BICKHARDT H. 1920: Histeridenstudien (42. Beitrag zur Kenntnis der Histeriden). Entomologische Blatter 16: 29 - 32." type="journal article" year="1920">Bickhardt, 1920</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF0951601AFA7ACD1FFA4A" authority="Olexa, 1992" authorityName="Olexa" authorityYear="1992" box="[333,821,1435,1460]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="turkmenicus" subSpecies="transcaucasianus">
<emphasis id="B958FC5AFFEF0951601AFA7ACC87FA4A" box="[333,685,1437,1460]" italics="true" pageId="120" pageNumber="117">G. turkmenicus transcaucasianus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF095163E5FA7CCD1FFA4A" author="OLEXA A." box="[690,821,1435,1460]" pageId="120" pageNumber="117" pagination="179 - 183" refId="ref119583" refString="OLEXA A. 1992: Bemerkungen zu einigen Arten der Gattung Gnathoncus Jacq. Du Val, 1858 (Insecta, Coleoptera, Histeridae) Faunistische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 18: 179 - 183." type="journal article" year="1992">Olexa, 1992</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516269FA7ACF36FA2A" authority="turkmenicus Olexa, 1992" authorityName="Olexa" authorityYear="1992" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="turkmenicus" subSpecies="turkmenicus">
<emphasis id="B958FC5AFFEF09516269FA7ACA4CFA4A" box="[830,1126,1437,1460]" italics="true" pageId="120" pageNumber="117">G. turkmenicus turkmenicus</emphasis>
<bibRefCitation id="EFBD5DB9FFEF095161C0FA5CCF36FA2A" author="OLEXA A." box="[151,284,1467,1492]" pageId="120" pageNumber="117" pagination="179 - 183" refId="ref119583" refString="OLEXA A. 1992: Bemerkungen zu einigen Arten der Gattung Gnathoncus Jacq. Du Val, 1858 (Insecta, Coleoptera, Histeridae) Faunistische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 18: 179 - 183." type="journal article" year="1992">Olexa, 1992</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF09516070FA5ACC77FA2D" authority="Normand, 1935" authorityName="Normand" authorityYear="1935" box="[295,605,1467,1492]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="species" species="wassiliefi">
<emphasis id="B958FC5AFFEF09516070FA5ACF86FA2A" box="[295,428,1469,1492]" italics="true" pageId="120" pageNumber="117">G. wassiliefi</emphasis>
<bibRefCitation id="EFBD5DB9FFEF095160E4FA5CCC77FA2D" author="NORMAND H." box="[435,605,1467,1491]" pageId="120" pageNumber="117" pagination="103 - 117" refId="ref119325" refString="NORMAND H. 1935: Contribution au catalogue des Coleopteres de Tunisie (5 me fascicule). Bulletin de la Societe d'Histoire Naturelle d'Afrique du Nord 26: 103 - 117." type="journal article" year="1935">Normand, 1935</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2C5BCBFFEF0951633FFA5ACCC3FA2A" authority="Lackner, 2010" authorityName="Lackner" authorityYear="2010" box="[616,745,1469,1492]" class="Insecta" family="Histeridae" genus="Gnathoncus" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="117" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B958FC5AFFEF0951633FFA5ACCC3FA2A" box="[616,745,1469,1492]" italics="true" pageId="120" pageNumber="117">Gnathoncus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B4121FFEF095163B8FA5BCD71FA2D" box="[751,859,1468,1491]" pageId="120" pageNumber="117" rank="species">spec. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FFEF095061C0FA3CCC0FFECD" lastPageId="121" lastPageNumber="114" pageId="120" pageNumber="117" type="discussion">
<paragraph id="8B932048FFEF095061C0FA3CCC0FFECD" blockId="120.[151,1127,219,1620]" lastBlockId="121.[113,1088,219,307]" lastPageId="121" lastPageNumber="114" pageId="120" pageNumber="117">
<emphasis id="B958FC5AFFEF095161C0FA3CCF3CFA0D" bold="true" box="[151,278,1499,1523]" pageId="120" pageNumber="117">Discussion.</emphasis>
This is most likely a monophyletic taxon, well delineated by the presence of at least two synapomorphies, e.g, short hooked appendix between the fourth dorsal elytral and sutural elytral striae and double marginal epipleural stria. The longitudinally divided tenth abdominal tergite of males can be accounted among synapomorphies shared also with taxa
<taxonomicName id="4C2C5BCBFFEE09506126FF3BCF3EFF0D" authorityName=": ROSS" authorityYear="1940" box="[113,276,220,243]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEE09506126FF3BCF3EFF0D" box="[113,276,220,243]" italics="true" pageId="121" pageNumber="114">Eremosaprinus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C2C5BCBFFEE0950601BFF3BCF9FFF0D" authorityName="Marseul" authorityYear="1862" box="[332,437,220,243]" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEE0950601BFF3BCF9FFF0D" box="[332,437,220,243]" italics="true" pageId="121" pageNumber="114">Myrmetes</emphasis>
</taxonomicName>
. As mentioned above, together with
<taxonomicName id="4C2C5BCBFFEE09506260FF3BCDF0FF0D" authorityName=": ROSS" authorityYear="1940" box="[823,986,220,243]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEE09506260FF3BCDF0FF0D" box="[823,986,220,243]" italics="true" pageId="121" pageNumber="114">Eremosaprinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C5BCBFFEE0950655CFF3BCE84FEED" authorityName="Marseul" authorityYear="1862" class="Insecta" family="Histeridae" genus="Myrmetes" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEE0950655CFF3BCE84FEED" italics="true" pageId="121" pageNumber="114">Myrmetes</emphasis>
</taxonomicName>
, these three taxa constitute the most basal grade of the Palaearctic
<taxonomicName id="4C2C5BCBFFEE095062CBFF1CCA24FEED" authorityName="Blanchard" authorityYear="1845" box="[924,1038,251,275]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="114" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
(see Diagnosis of
<taxonomicName id="4C2C5BCBFFEE09506057FEFBCF89FECD" authorityName=": ROSS" authorityYear="1940" box="[256,419,284,307]" class="Insecta" family="Histeridae" genus="Eremosaprinus" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFFEE09506057FEFBCF89FECD" box="[256,419,284,307]" italics="true" pageId="121" pageNumber="114">Eremosaprinus</emphasis>
</taxonomicName>
for details).
</paragraph>
</subSubSection>
</treatment>
</document>