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<document id="06BDA4F985D15DCD5BD7E23BFB34D2BC" ID-CLB-Dataset="28041" ID-DOI="10.11646/zootaxa.4508.3.3" ID-GBIF-Dataset="9de070ed-9c35-4131-a350-72371ee93ddf" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607216" ID-ZooBank="E53B4C53-554A-4913-8E3B-CE1C530DF004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586064605" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2018" docId="DA218790FFB72E2A248FFD677A4AFE49" docLanguage="en" docName="zootaxa.4508.3.3.pdf" docOrigin="Zootaxa 4508 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum propinquum Shavrin &amp; Smetana 2018, sp.n." docType="treatment" docVersion="8" lastPageNumber="390" masterDocId="2618FFE8FFBD2E272418FFD37818FFBC" masterDocTitle="New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China" masterLastPageNumber="402" masterPageNumber="377" pageNumber="387" updateTime="1698713048312" updateUser="ExternalLinkService">
<mods:mods id="BDF8A05AF0F45869F4E9B2C322557281" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5DBF7718FF4037922D80987FB032CDF5">New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China</mods:title>
</mods:titleInfo>
<mods:name id="EF27A7A8B4F8A6F9CB44D323695756CD" type="personal">
<mods:role id="AC6470A4524D36044B054794E48DD8EC">
<mods:roleTerm id="32DAEE957029766DCE2027EF91109E15">Author</mods:roleTerm>
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<mods:namePart id="AEE4CCB08FD10AF229C2D34C118360A4">Shavrin, Alexey V.</mods:namePart>
</mods:name>
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<mods:role id="76A0805CC4C0F5EB6E5D5B9BFA326394">
<mods:roleTerm id="3895DC9B5351CC64EFD85D1A737538B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6413177946057289B15A1610D40752C">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="036E15372F8A330AAE5D27AFCA0CA23E">text</mods:typeOfResource>
<mods:relatedItem id="E401C42493F43ADF35BA6F52812CC0C8" type="host">
<mods:titleInfo id="4EEC2C2F654D15E82915E7633AE0E58A">
<mods:title id="DE72DBE11FF3A09B9354487C6F7804FA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C2F80368F6713E6CF049D350D572F6A4">
<mods:date id="C8B4F0F637DC87981E7E0D7B4BA32007">2018</mods:date>
<mods:detail id="6400B3BE43C172131918BC0135C36759" type="pubDate">
<mods:number id="B320DB472D9909212D778944530006B3">2018-11-01</mods:number>
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<mods:detail id="97BD9AE5B70D032E873FFA71AF6B9884" type="volume">
<mods:number id="6568F387357409F7BCE816A2D2553E50">4508</mods:number>
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<mods:number id="1F2A2BD123A54BB5A45BCF495E563308">3</mods:number>
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<mods:classification id="17CD63567BE87924968D8A0975749A85">journal article</mods:classification>
<mods:identifier id="8C8B27E27964328C355ED5EF4283DA34" type="CLB-Dataset">28041</mods:identifier>
<mods:identifier id="710CFEAE9A1189307AD73D0FD9763079" type="DOI">10.11646/zootaxa.4508.3.3</mods:identifier>
<mods:identifier id="54D69CA128E58E0F33300E88B93E8A0B" type="GBIF-Dataset">9de070ed-9c35-4131-a350-72371ee93ddf</mods:identifier>
<mods:identifier id="84D3773B10E7F5C4F7132EEE60AC8C66" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D0881C17DB6AF55E05C80ADF73902446" type="Zenodo-Dep">2607216</mods:identifier>
<mods:identifier id="72D907C9B8B56C601DD92DEEF7819EC7" type="ZooBank">E53B4C53-554A-4913-8E3B-CE1C530DF004</mods:identifier>
</mods:mods>
<treatment id="DA218790FFB72E2A248FFD677A4AFE49" ID-DOI="http://doi.org/10.5281/zenodo.5962994" ID-GBIF-Taxon="154495880" ID-Zenodo-Dep="5962994" LSID="urn:lsid:plazi:treatment:DA218790FFB72E2A248FFD677A4AFE49" httpUri="http://treatment.plazi.org/id/DA218790FFB72E2A248FFD677A4AFE49" lastPageId="13" lastPageNumber="390" pageId="10" pageNumber="387">
<subSubSection id="1A92650DFFB72E2D248FFD677B34FD72" box="[151,812,692,718]" pageId="10" pageNumber="387" type="nomenclature">
<paragraph id="52373686FFB72E2D248FFD677B34FD72" blockId="10.[151,812,692,751]" box="[151,812,692,718]" pageId="10" pageNumber="387">
<heading id="097F81EAFFB72E2D248FFD677B34FD72" bold="true" box="[151,812,692,718]" fontSize="11" level="1" pageId="10" pageNumber="387" reason="1">
<emphasis id="60FCEA94FFB72E2D248FFD677B34FD72" bold="true" box="[151,812,692,718]" pageId="10" pageNumber="387">
<taxonomicName id="95884D05FFB72E2D248FFD677AFCFD72" ID-CoL="CZQB" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,740,692,718]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="propinquum" status="sp. nov.">
<emphasis id="60FCEA94FFB72E2D248FFD6779FCFD72" bold="true" box="[151,484,692,718]" italics="true" pageId="10" pageNumber="387">Amphichroum propinquum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="7BCF57EFFFB72E2D26E9FD677B34FD72" box="[753,812,692,718]" pageId="10" pageNumber="387" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB72E2D248FFD057999FD53" box="[151,385,726,751]" pageId="10" pageNumber="387" type="description">
<paragraph id="52373686FFB72E2D248FFD057999FD53" blockId="10.[151,812,692,751]" box="[151,385,726,751]" pageId="10" pageNumber="387">
(
<figureCitation id="CAB32A03FFB72E2D2487FD0578E4FD53" box="[159,252,726,751]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1549,1572]" captionTargetBox="[311,1291,206,1518]" captionTargetId="figure@9.[223,1363,193,1528]" captionTargetPageId="9" captionText="FIGURES 16–19. Habitus of Amphichroum (males): 16–A. propinquum sp.n. (holotype), 17–A. rotundatum sp.n. (holotype), 18–A. schuelkei sp.n. (paratype), 19–A. squamosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607228/files/figure.png" pageId="10" pageNumber="387">Figs. 16</figureCitation>
,
<figureCitation id="CAB32A03FFB72E2D2510FD047949FD53" box="[264,337,727,751]" captionStart="FIGURES 20–23" captionStartId="11.[151,264,1853,1876]" captionTargetBox="[321,1247,234,1791]" captionTargetId="figure@11.[182,1404,193,1832]" captionTargetPageId="11" captionText="FIGURES 20–23. Aedeagus of Amphichroum propinquum sp.n.(20–21: holotype) and A. rotundatum sp.n. (22–23: paratype): 20, 22–parameral view, 21, 23–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607230/files/figure.png" pageId="10" pageNumber="387">20–21</figureCitation>
,
<figureCitation id="CAB32A03FFB72E2D2545FD047962FD53" box="[349,378,727,751]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="10" pageNumber="387">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB72E2D248FFCCD7ACCFBB3" pageId="10" pageNumber="387" type="materials_examined">
<paragraph id="52373686FFB72E2D248FFCCD7BF3FC7B" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">
<emphasis id="60FCEA94FFB72E2D248FFCCD79D0FC8B" bold="true" box="[151,456,798,823]" pageId="10" pageNumber="387">Type material examined:</emphasis>
<materialsCitation id="E2E03CDBFFB72E2D25CBFCCD7B2CFC1F" ID-GBIF-Occurrence="2028139626" collectingDate="1999-07-13" collectorName="Siniaev &amp; A. Plutenko' &amp; Sammlung M. Schulke &amp; Berlin &amp; Ankauf Plutenkov" country="China" county="Moxi" determinerName="Zanetti" elevation="1700" latitude="29.65" location="Daxue Shan" longLatPrecision="1226" longitude="102.1" municipality="Gongga Shan" pageId="10" pageNumber="387" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="8D338824FFB72E2D25CBFCCD7A23FC8B" box="[467,571,798,823]" pageId="10" pageNumber="387" type="holotype">Holotype</typeStatus>
<specimenCount id="448EFD0FFFB72E2D265EFCCC7A46FC8A" box="[582,606,799,822]" pageId="10" pageNumber="387" type="male">♂</specimenCount>
: ‘
<collectingCountry id="2A9F7616FFB72E2D266AFCCD7AA6FC8B" box="[626,702,798,823]" name="China" pageId="10" pageNumber="387">China</collectingCountry>
: W-Sichuan,
<location id="5757605DFFB72E2D2743FCCC7BFFFC8B" LSID="urn:lsid:plazi:treatment:DA218790FFB72E2A248FFD677A4AFE49:5757605DFFB72E2D2743FCCC7BFFFC8B" box="[859,999,798,823]" country="China" county="Moxi" latitude="29.65" longLatPrecision="1226" longitude="102.1" municipality="Gongga Shan" name="Daxue Shan" pageId="10" pageNumber="387">Daxue Shan</location>
|
<collectingMunicipality id="B253ACFCFFB72E2D2019FCCC7CB8FC8B" box="[1025,1184,798,823]" pageId="10" pageNumber="387">Gongga Shan</collectingMunicipality>
,
<collectingCounty id="BB564E0AFFB72E2D20B7FCCD7CF7FC8B" box="[1199,1263,798,823]" pageId="10" pageNumber="387">Moxi</collectingCounty>
,
<quantity id="95709B63FFB72E2D20E5FCCC7D54FC8B" box="[1277,1356,799,823]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="387" unit="m" value="1700.0">
<elevation id="D9A5D1B5FFB72E2D20E5FCCC7D54FC8B" box="[1277,1356,799,823]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="387" unit="m" value="1700.0">1700m</elevation>
</quantity>
| 11. -
<date id="26361046FFB72E2D248FFC90790DFCE7" box="[151,277,835,859]" pageId="10" pageNumber="387" value="1999-07-13">
<collectingDate id="3672E9AEFFB72E2D248FFC90790DFCE7" box="[151,277,835,859]" pageId="10" pageNumber="387" value="1999-07-13">13.07.1999</collectingDate>
</date>
|
<geoCoordinate id="37BC5041FFB72E2D2532FC907980FCE6" box="[298,408,835,859]" direction="north" orientation="latitude" pageId="10" pageNumber="387" precision="925" value="29.65">29° 39' N</geoCoordinate>
,
<geoCoordinate id="37BC5041FFB72E2D25BCFC907A03FCE6" box="[420,539,834,859]" direction="east" orientation="longitude" pageId="10" pageNumber="387" precision="925" value="102.1">102° 06' E</geoCoordinate>
| leg. [V].
<collectorName id="FF7D5350FFB72E2D268EFC917AF2FCE7" box="[662,746,834,859]" pageId="10" pageNumber="387">Siniaev</collectorName>
&amp;
<collectorName id="FF7D5350FFB72E2D2717FC907B87FCE7" box="[783,927,834,859]" pageId="10" pageNumber="387">A. Plutenko’</collectorName>
&lt;rectangular label, printed&gt;, ‘
<taxonomicName id="95884D05FFB72E2D20ECFC917D84FCE7" authorityName="Kraatz" authorityYear="1857" box="[1268,1436,834,859]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
[handwritten] | sp. [handwritten] | det.
<determinerName id="A7EBDA42FFB72E2D265EFCB57A8DFCC3" box="[582,661,870,895]" pageId="10" pageNumber="387">Zanetti</determinerName>
2001’ &lt;rectangular label, printed&gt;, ‘
<collectorName id="FF7D5350FFB72E2D2037FCB57D2EFCC3" box="[1071,1334,870,895]" pageId="10" pageNumber="387">Sammlung M. Schülke</collectorName>
|
<collectorName id="FF7D5350FFB72E2D2151FCB57D97FCC3" box="[1353,1423,870,895]" pageId="10" pageNumber="387">Berlin</collectorName>
|
<collectorName id="FF7D5350FFB72E2D248FFC59797EFC1F" box="[151,358,906,931]" pageId="10" pageNumber="387">Ankauf Plutenkov</collectorName>
1999’ &lt;green rectangular label, printed
</materialsCitation>
&gt;, ‘
<materialsCitation id="E2E03CDBFFB72E2D274DFC587BF0FC7B" ID-GBIF-Occurrence="2028139633" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="10" pageNumber="387" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D338824FFB72E2D274DFC587BF6FC1F" box="[853,1006,907,931]" pageId="10" pageNumber="387" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="95884D05FFB72E2D201EFC597CB1FC1F" authorityName="Kraatz" authorityYear="1857" box="[1030,1193,906,931]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| propinquum
<emphasis id="60FCEA94FFB72E2D214CFC597D94FC1F" bold="true" box="[1364,1420,906,931]" pageId="10" pageNumber="387">
<taxonomicNameLabel id="7BCF57EFFFB72E2D214CFC597D94FC1F" box="[1364,1420,906,931]" pageId="10" pageNumber="387" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFB72E2D248FFC7D7908FC7B" box="[151,272,942,967]" pageId="10" pageNumber="387">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFB72E2D252CFC7C79AFFC7B" box="[308,439,943,967]" pageId="10" pageNumber="387">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt;(
<collectionCode id="3499AE43FFB72E2D27B1FC7D7BC7FC7B" box="[937,991,942,967]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="10" pageNumber="387" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="52373686FFB72E2D24DFFC007ACCFBB3" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">
<materialsCitation id="E2E03CDBFFB72E2D24DFFC007BB1FC57" ID-GBIF-Occurrence="2028139635" box="[199,937,978,1003]" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="10" pageNumber="387" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="8D338824FFB72E2D24DFFC007931FC57" box="[199,297,979,1003]" pageId="10" pageNumber="387" type="paratype">Paratype</typeStatus>
<specimenCount id="448EFD0FFFB72E2D2529FC007950FC56" box="[305,328,979,1002]" pageId="10" pageNumber="387" type="female">♀</specimenCount>
: same data as the holotype, with red rectangular label
</materialsCitation>
‘
<materialsCitation id="E2E03CDBFFB72E2D27ADFC007AD7FBB3" ID-GBIF-Occurrence="2028139624" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="10" pageNumber="387" specimenCount="1" typeStatus="paratype">
<typeStatus id="8D338824FFB72E2D27ADFC007C5EFC57" box="[949,1094,979,1003]" pageId="10" pageNumber="387" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="95884D05FFB72E2D2042FC017CE6FC57" authorityName="Kraatz" authorityYear="1857" box="[1114,1278,978,1003]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| propinquum
<emphasis id="60FCEA94FFB72E2D248FFC2578D6FBB3" bold="true" box="[151,206,1014,1039]" pageId="10" pageNumber="387">
<taxonomicNameLabel id="7BCF57EFFFB72E2D248FFC2578D6FBB3" box="[151,206,1014,1039]" pageId="10" pageNumber="387" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFB72E2D24FBFC257944FBB3" box="[227,348,1014,1039]" pageId="10" pageNumber="387">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFB72E2D2567FC247A1BFBB3" box="[383,515,1015,1039]" pageId="10" pageNumber="387">Smetana A.</collectorName>
des. 2018’ (
<collectionCode id="3499AE43FFB72E2D2695FC247ADFFBB3" box="[653,711,1015,1039]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="10" pageNumber="387" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB72E2A24DFFBC97D8CFE83" lastPageId="13" lastPageNumber="390" pageId="10" pageNumber="387" type="description">
<paragraph id="52373686FFB72E2D24DFFBC97BD5FBC7" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">
<emphasis id="60FCEA94FFB72E2D24DFFBC97949FB8F" bold="true" box="[199,337,1050,1075]" pageId="10" pageNumber="387">Description</emphasis>
. Measurements (n=2): HW: 0.60–0.64; HL: 0.37; AL(
<typeStatus id="8D338824FFB72E2D27D8FBC97C3FFB8F" box="[960,1063,1050,1075]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
): 1.24; OL: 0.17; PL: 0.62–0.66; PW: 0.95–1.07; ESL: 1.12–1.21; EW: 1.25–1.45; AW: 1.15–1.46; MTbL(
<typeStatus id="8D338824FFB72E2D27FDFBED7C53FBEB" box="[997,1099,1086,1111]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
): 0.65; MTrL(
<typeStatus id="8D338824FFB72E2D20E9FBED7D4EFBEB" box="[1265,1366,1086,1111]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
): 0.55 (MTrL 1–4: 0.32; MTrL 5: 0.22); AedL: 0.85; TL: 3.00 (
<typeStatus id="8D338824FFB72E2D270AFBB17B60FBC7" box="[786,888,1122,1147]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
) –3.15.
</paragraph>
<paragraph id="52373686FFB72E2D24DFFB557945FACA" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">
Body yellow-brown to reddish-brown; antennomeres 4–11 brown; mouthparts, antennomeres 1–3, legs, paratergites and apical segments of abdomen yellow (
<typeStatus id="8D338824FFB72E2D2718FB787B7AFB7F" box="[768,866,1195,1219]" pageId="10" pageNumber="387" type="paratype">paratype</typeStatus>
with yellow abdomen except for median parts of tergites 4–6).Head with irregular small and deep punctures, denser on middle portion and on infraorbital ridges and deeper between grooves; pronotum with dense punctures, smaller than those in middle part of head, with narrow impunctate longitudinal area on disc and very sparse and small punctation on basal third of pronotum (
<typeStatus id="8D338824FFB72E2D2131FAC47D88FA93" box="[1321,1424,1303,1327]" pageId="10" pageNumber="387" type="paratype">paratype</typeStatus>
); punctation of elytra very dense and deep, coarser and smaller on prescutellar area; abdominal tergites without visible punctures.
</paragraph>
<paragraph id="52373686FFB72E2D24DFFA517B7CF9BB" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">
Head 1.6–1.7 times as wide as long, slightly convex posteriorly. Distance between ocelli one and half times as long as distance betwen ocellus and posterior margin of eye. Apical segment of maxillary palp slightly longer than preceding segment. Length × width of antennomeres (
<typeStatus id="8D338824FFB72E2D26E3FA197B7CFA5F" box="[763,868,1482,1507]" pageId="10" pageNumber="387" type="holotype">holotype</typeStatus>
): 1: 0.12 × 0.05; 2: 0.13 × 0.04; 3: 0.12 × 0.03; 4: 0.10 × 0.04; 5–7: 0.11 × 0.04;8–10: 0.10 × 0.05;11: 0.14 × 0.06.
</paragraph>
<paragraph id="52373686FFB72E2D24DFF9C07CCAF9F3" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">Pronotum 1.5–1.6 times as wide as long, widest in middle, evenly narrowed both posteriad and anteriad; anterior angles widely rounded, indistinctly protruded anteriad; posterior angles widely rounded.</paragraph>
<paragraph id="52373686FFB72E2D24DFF9897913F92A" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">Elytra distinctly wider than long, reaching middle or apical part of abdominal tergite IV, 1.8 times as long as pronotum.</paragraph>
<paragraph id="52373686FFB72E2D24DFF9717AF9F906" blockId="10.[151,1436,798,1975]" box="[199,737,1698,1723]" pageId="10" pageNumber="387">Metatibia about 1.2 times as long as metatarsus.</paragraph>
<paragraph id="52373686FFB72E2D24DFF9157C5FF80B" blockId="10.[151,1436,798,1975]" pageId="10" pageNumber="387">
Male (
<figureCitation id="CAB32A03FFB72E2D2509F915797DF963" box="[273,357,1734,1759]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1549,1572]" captionTargetBox="[311,1291,206,1518]" captionTargetId="figure@9.[223,1363,193,1528]" captionTargetPageId="9" captionText="FIGURES 16–19. Habitus of Amphichroum (males): 16–A. propinquum sp.n. (holotype), 17–A. rotundatum sp.n. (holotype), 18–A. schuelkei sp.n. (paratype), 19–A. squamosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607228/files/figure.png" pageId="10" pageNumber="387">Fig. 16</figureCitation>
). Protarsomeres 1–4 wide. Medial margin of apical part of protibia with two rows of nine short peg setae; mesotibia markedly curved mediad, with dense row of twenty four short thorns. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII widely rounded. Aedeagus (
<figureCitation id="CAB32A03FFB72E2D2111F8DD7D43F89B" box="[1289,1371,1806,1831]" captionStart="FIGURES 20–23" captionStartId="11.[151,264,1853,1876]" captionTargetBox="[321,1247,234,1791]" captionTargetId="figure@11.[182,1404,193,1832]" captionTargetPageId="11" captionText="FIGURES 20–23. Aedeagus of Amphichroum propinquum sp.n.(20–21: holotype) and A. rotundatum sp.n. (22–23: paratype): 20, 22–parameral view, 21, 23–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607230/files/figure.png" pageId="10" pageNumber="387">Fig. 20</figureCitation>
) with median lobe strongly narrowed apicad, with narrow and elongate apical portion; parameres narrow, distinctly exceeding median lobe, indistinctly widened apicad, with small apical and preapical setae; internal sac with three fields of fine thorns in apical half, with two pairs of moderately large thorns in middle portion and with characteristic long, bifurcate structure in basal half. Aedeagus laterally as in
<figureCitation id="CAB32A03FFB72E2D27E8F84D7C5BF80B" box="[1008,1091,1950,1975]" captionStart="FIGURES 20–23" captionStartId="11.[151,264,1853,1876]" captionTargetBox="[321,1247,234,1791]" captionTargetId="figure@11.[182,1404,193,1832]" captionTargetPageId="11" captionText="FIGURES 20–23. Aedeagus of Amphichroum propinquum sp.n.(20–21: holotype) and A. rotundatum sp.n. (22–23: paratype): 20, 22–parameral view, 21, 23–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607230/files/figure.png" pageId="10" pageNumber="387">Fig. 21</figureCitation>
.
</paragraph>
<caption id="06F7660EFFB62E2C248FF8EE7B0FF8CE" httpUri="https://zenodo.org/record/2607230/files/figure.png" pageId="11" pageNumber="388" startId="11.[151,264,1853,1876]" targetBox="[321,1247,234,1791]" targetPageId="11">
<paragraph id="52373686FFB62E2C248FF8EE7B0FF8CE" blockId="11.[151,1437,1853,1906]" pageId="11" pageNumber="388">
<emphasis id="60FCEA94FFB62E2C248FF8EE794DF8EF" bold="true" box="[151,341,1853,1876]" pageId="11" pageNumber="388">FIGURES 20–23.</emphasis>
Aedeagus of
<taxonomicName id="95884D05FFB62E2C25F9F8ED7AEAF8E8" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[481,754,1854,1876]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="388" phylum="Arthropoda" rank="species" species="propinquum" status="sp. nov.">
<emphasis id="60FCEA94FFB62E2C25F9F8ED7AEAF8E8" box="[481,754,1854,1876]" italics="true" pageId="11" pageNumber="388">Amphichroum propinquum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB62E2C26E0F8ED7B3EF8E8" box="[760,806,1854,1876]" pageId="11" pageNumber="388" rank="species">sp.n.</taxonomicNameLabel>
(20–21: holotype) and
<taxonomicName id="95884D05FFB62E2C2009F8EC7CBAF8E8" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1041,1186,1854,1876]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="388" phylum="Arthropoda" rank="species" species="rotundatum" status="sp. nov.">
<emphasis id="60FCEA94FFB62E2C2009F8EC7CBAF8E8" box="[1041,1186,1854,1876]" italics="true" pageId="11" pageNumber="388">A. rotundatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB62E2C20B1F8EC7CC1F8E8" box="[1193,1241,1855,1876]" pageId="11" pageNumber="388" rank="species">sp.n.</taxonomicNameLabel>
(22–23: paratype): 20, 22–parameral view, 21, 23–lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="06F7660EFFB12E2B248FF86C7D44F848" httpUri="https://zenodo.org/record/2607232/files/figure.png" pageId="12" pageNumber="389" startId="12.[151,264,1983,2006]" targetBox="[295,1374,198,1935]" targetPageId="12">
<paragraph id="52373686FFB12E2B248FF86C7D44F848" blockId="12.[151,1436,1983,2036]" pageId="12" pageNumber="389">
<emphasis id="60FCEA94FFB12E2B248FF86C794CF868" bold="true" box="[151,340,1983,2006]" pageId="12" pageNumber="389">FIGURES 24–28.</emphasis>
Aedeagus of
<taxonomicName id="95884D05FFB12E2B25F8F86C7ACBF869" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[480,723,1983,2005]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="60FCEA94FFB12E2B25F8F86C7ACBF869" box="[480,723,1983,2005]" italics="true" pageId="12" pageNumber="389">Amphichroum schuelkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB12E2B26C2F86C7B1FF869" box="[730,775,1983,2005]" pageId="12" pageNumber="389" rank="species">sp.n.</taxonomicNameLabel>
(24–26: paratype) and
<taxonomicName id="95884D05FFB12E2B27F7F8137C9CF86A" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1007,1156,1984,2006]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="389" phylum="Arthropoda" rank="species" species="squamosum" status="sp. nov.">
<emphasis id="60FCEA94FFB12E2B27F7F8137C9CF86A" box="[1007,1156,1984,2006]" italics="true" pageId="12" pageNumber="389">A. squamosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB12E2B2092F8137CA2F869" box="[1162,1210,1984,2005]" pageId="12" pageNumber="389" rank="species">sp.n.</taxonomicNameLabel>
(27–28: paratype): 24 (outline of internal sac shown as point line), 27–parameral view, 25–internal sac, 26, 28–lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="52373686FFB02E2A24DFFF447C5DFF69" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="390">Female. Protarsomeres 1–4 moderately narrow. Medial margin of protibia and mesotibia without modifications. Abdominal tergite VIII and sternite VIII with rounded apical margin.</paragraph>
<paragraph id="52373686FFB02E2A24DFFF0C7D8CFE83" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="390">
<emphasis id="60FCEA94FFB02E2A24DFFF0C79ABFF44" bold="true" box="[199,435,223,248]" pageId="13" pageNumber="390">Comparative notes.</emphasis>
Based on the body size, coloration, general shape of the aedeagus and the presence of similar structures within internal sac,
<taxonomicName id="95884D05FFB02E2A2626FED77AFDFEA0" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[574,741,260,284]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="390" phylum="Arthropoda" rank="species" species="propinquum" status="sp. nov.">
<emphasis id="60FCEA94FFB02E2A2626FED77AFDFEA0" box="[574,741,260,284]" italics="true" pageId="13" pageNumber="390">A. propinquum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFB02E2A26F4FED77B3CFEA1" bold="true" box="[748,804,260,285]" pageId="13" pageNumber="390">
<taxonomicNameLabel id="7BCF57EFFFB02E2A26F4FED77B3CFEA1" box="[748,804,260,285]" pageId="13" pageNumber="390" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="95884D05FFB02E2A27A9FED77C7DFEA0" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[945,1125,260,284]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="390" phylum="Arthropoda" rank="species" species="longilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFB02E2A27A9FED77C7DFEA0" box="[945,1125,260,284]" italics="true" pageId="13" pageNumber="390">A. longilobatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB02E2A2074FED77CBCFEA1" box="[1132,1188,260,285]" pageId="13" pageNumber="390" rank="species">
<emphasis id="60FCEA94FFB02E2A2074FED77CBCFEA1" bold="true" box="[1132,1188,260,285]" pageId="13" pageNumber="390">sp.n.</emphasis>
</taxonomicNameLabel>
, from which it differs by shorter apical antennomeres, elytra and apical portion of the median lobe, and by the details of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB02E2A24DFFE9F7D3FFE11" pageId="13" pageNumber="390" type="distribution">
<paragraph id="52373686FFB02E2A24DFFE9F79AAFE34" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="390">
<emphasis id="60FCEA94FFB02E2A24DFFE9F7978FED9" bold="true" box="[199,352,332,357]" pageId="13" pageNumber="390">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="8D338824FFB02E2A27A4FE9E7BF4FED9" box="[956,1004,333,357]" pageId="13" pageNumber="390">type</typeStatus>
locality in the Gongga Shan range in
<collectingRegion id="904CF864FFB02E2A248FFEBC78ECFE34" box="[151,244,367,392]" country="China" name="Sichuan" pageId="13" pageNumber="390">Sichuan</collectingRegion>
,
<collectingCountry id="2A9F7616FFB02E2A24E7FEBC795AFE34" box="[255,322,367,392]" name="China" pageId="13" pageNumber="390">China</collectingCountry>
(
<figureCitation id="CAB32A03FFB02E2A2549FEBC79BDFE34" box="[337,421,367,392]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="13" pageNumber="390">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="52373686FFB02E2A24DFFE477D3FFE11" blockId="13.[151,1436,151,501]" box="[199,1319,404,429]" pageId="13" pageNumber="390">
<emphasis id="60FCEA94FFB02E2A24DFFE477952FE11" bold="true" box="[199,330,404,429]" pageId="13" pageNumber="390">Bionomics.</emphasis>
Specimens were collected at elevation
<quantity id="95709B63FFB02E2A271CFE477B41FE10" box="[772,857,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="390" unit="m" value="1700.0">1700 m</quantity>
a.s.l. Collection details are not available.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB02E2A24DFFE647A4AFE49" pageId="13" pageNumber="390" type="etymology">
<paragraph id="52373686FFB02E2A24DFFE647A4AFE49" blockId="13.[151,1436,151,501]" pageId="13" pageNumber="390">
<emphasis id="60FCEA94FFB02E2A24DFFE647956FE6C" bold="true" box="[199,334,439,464]" pageId="13" pageNumber="390">Etymology.</emphasis>
The specific epithet is a Latin adjective
<emphasis id="60FCEA94FFB02E2A2703FE6B7B82FE73" box="[795,922,440,463]" italics="true" pageId="13" pageNumber="390">propinquus</emphasis>
, -
<emphasis id="60FCEA94FFB02E2A27AAFE6A7BD8FE6C" box="[946,960,441,464]" italics="true" pageId="13" pageNumber="390">a</emphasis>
, -
<emphasis id="60FCEA94FFB02E2A27C0FE6A7BE2FE6C" box="[984,1018,441,464]" italics="true" pageId="13" pageNumber="390">um</emphasis>
(related).It refers to the similarity of this species with some of its congeners.
</paragraph>
</subSubSection>
</treatment>
</document>
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