treatments-xml/data/DA/21/87/DA218790FFB52E2D248FFC5F7A7FFDE3.xml

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<mods:title id="C78D3C1C075194EF1BE5DA79BAEEC4C0">New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China</mods:title>
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<mods:namePart id="D7B2ED55D487EED186E757C20AD8180B">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="574B8661CF9E84CC304A981D711D60B2">Smetana, Aleš</mods:namePart>
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<mods:date id="8A4FAB9362EF10183785CEC7C2300125">2018</mods:date>
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<treatment id="DA218790FFB52E2D248FFC5F7A7FFDE3" ID-DOI="http://doi.org/10.5281/zenodo.5962992" ID-GBIF-Taxon="154495881" ID-Zenodo-Dep="5962992" LSID="urn:lsid:plazi:treatment:DA218790FFB52E2D248FFC5F7A7FFDE3" httpUri="http://treatment.plazi.org/id/DA218790FFB52E2D248FFC5F7A7FFDE3" lastPageId="10" lastPageNumber="387" pageId="8" pageNumber="385">
<subSubSection id="1A92650DFFB52E2F248FFC5F7B3BFC1A" box="[151,803,908,934]" pageId="8" pageNumber="385" type="nomenclature">
<paragraph id="52373686FFB52E2F248FFC5F7B3BFC1A" blockId="8.[151,803,908,967]" box="[151,803,908,934]" pageId="8" pageNumber="385">
<heading id="097F81EAFFB52E2F248FFC5F7B3BFC1A" bold="true" box="[151,803,908,934]" fontSize="11" level="1" pageId="8" pageNumber="385" reason="1">
<emphasis id="60FCEA94FFB52E2F248FFC5F7B3BFC1A" bold="true" box="[151,803,908,934]" pageId="8" pageNumber="385">
<taxonomicName id="95884D05FFB52E2F248FFC5F7AC3FC1A" ID-CoL="CZQ4" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,731,908,934]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="385" phylum="Arthropoda" rank="species" species="maculosum" status="sp. nov.">
<emphasis id="60FCEA94FFB52E2F248FFC5F79C3FC1A" bold="true" box="[151,475,908,934]" italics="true" pageId="8" pageNumber="385">Amphichroum maculosum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
, 
<taxonomicNameLabel id="7BCF57EFFFB52E2F26FFFC5F7B3BFC1A" box="[743,803,908,934]" pageId="8" pageNumber="385" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="1A92650DFFB52E2F248FFC7D796BFC7B" box="[151,371,942,967]" pageId="8" pageNumber="385" type="description">
<paragraph id="52373686FFB52E2F248FFC7D796BFC7B" blockId="8.[151,803,908,967]" box="[151,371,942,967]" pageId="8" pageNumber="385">
(
<figureCitation id="CAB32A03FFB52E2F2487FC7D78F6FC7B" box="[159,238,942,967]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1954,1977]" captionTargetBox="[296,1236,546,1939]" captionTargetId="figure@2.[283,1307,539,1939]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Amphichroum (males): 1–A. angustilobatum sp.n. (holotype), 2–A. assingi sp.n. (holotype), 3–A. longilobatum sp.n. (paratype), 4–A. maculosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="8" pageNumber="385">Figs. 4</figureCitation>
, 
<figureCitation id="CAB32A03FFB52E2F24E2FC7D795AFC7B" box="[250,322,942,967]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[330,1322,210,1922]" captionTargetId="figure@7.[217,1370,161,1956]" captionTargetPageId="7" captionText="FIGURES 12–15.Aedeagus of Amphichroum longilobatum sp.n.(12–13: paratype) and A. maculosum sp.n. (14–15: paratype): 12, 14–parameral view, 13, 15–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607226/files/figure.png" pageId="8" pageNumber="385">14–15</figureCitation>
, 
<figureCitation id="CAB32A03FFB52E2F2556FC7C7973FC7B" box="[334,363,943,967]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="8" pageNumber="385">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB52E2F248FFC257A22FB5B" pageId="8" pageNumber="385" type="materials_examined">
<paragraph id="52373686FFB52E2F248FFC257B4DFBC7" blockId="8.[151,1437,1014,2011]" pageId="8" pageNumber="385">
<emphasis id="60FCEA94FFB52E2F248FFC2579D3FBB3" bold="true" box="[151,459,1014,1039]" pageId="8" pageNumber="385">Type material examined:</emphasis>
<materialsCitation id="E2E03CDBFFB52E2F25CFFC257A83FBEB" ID-GBIF-Occurrence="2028139621" collectingDate="2009-09-06" collectorName="Diancang Shan W Dali &amp; D. W. Wrase" country="China" elevation="2970" location="Yunnan" pageId="8" pageNumber="385" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="8D338824FFB52E2F25CFFC257A27FBB3" box="[471,575,1014,1039]" pageId="8" pageNumber="385" type="holotype">Holotype</typeStatus>
<specimenCount id="448EFD0FFFB52E2F2653FC247A7BFBB2" box="[587,611,1015,1038]" pageId="8" pageNumber="385" type="male">♂</specimenCount>
: ‘ 
<collectingCountry id="2A9F7616FFB52E2F2660FC247ACEFBB3" box="[632,726,1015,1039]" name="China" pageId="8" pageNumber="385">CHINA</collectingCountry>
(
<collectingRegion id="904CF864FFB52E2F26F1FC247B5AFBB3" box="[745,834,1015,1039]" country="China" name="Yunnan" pageId="8" pageNumber="385">Yunnan</collectingRegion>
) Dali Bai Auton. Pref., | 
<collectorName id="FF7D5350FFB52E2F2060FC257D92FBB3" box="[1144,1418,1014,1039]" pageId="8" pageNumber="385">Diancang Shan W Dali</collectorName>
| 25°41'50.9-52''N / 100°06'01.7-28.3''E | ca 
<quantity id="95709B63FFB52E2F269CFBC97B04FB8E" box="[644,796,1050,1075]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="2.98" metricValueMin="2.96" pageId="8" pageNumber="385" unit="m" value="2970.0" valueMax="2980.0" valueMin="2960.0">
<elevation id="D9A5D1B5FFB52E2F269CFBC97B04FB8E" box="[644,796,1050,1075]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="2.98" metricValueMin="2.96" pageId="8" pageNumber="385" unit="m" value="2970.0" valueMax="2980.0" valueMin="2960.0">2960-2980 m</elevation>
</quantity>
(former stone pit, | in gravel /under stones) | 
<date id="26361046FFB52E2F2132FBC97D84FB8F" box="[1322,1436,1050,1075]" pageId="8" pageNumber="385" value="2009-09-06">
<collectingDate id="3672E9AEFFB52E2F2132FBC97D84FB8F" box="[1322,1436,1050,1075]" pageId="8" pageNumber="385" value="2009-09-06">6.IX.2009</collectingDate>
</date>
<collectorName id="FF7D5350FFB52E2F248FFBEC7938FBEA" box="[151,288,1087,1110]" pageId="8" pageNumber="385">D.W. Wrase</collectorName>
[31]’ &lt;rectangular label, printed
</materialsCitation>
&gt;, ‘ 
<materialsCitation id="E2E03CDBFFB52E2F26A3FBEC7B49FBC7" ID-GBIF-Occurrence="2028139617" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="8" pageNumber="385" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D338824FFB52E2F26A3FBEC7B4CFBEB" box="[699,852,1087,1111]" pageId="8" pageNumber="385" type="holotype">HOLOTYPE</typeStatus>
| 
<taxonomicName id="95884D05FFB52E2F2774FBED7C17FBEB" authorityName="Kraatz" authorityYear="1857" box="[876,1039,1086,1111]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| maculosum 
<emphasis id="60FCEA94FFB52E2F20AAFBED7CF1FBEB" bold="true" box="[1202,1257,1086,1111]" pageId="8" pageNumber="385">
<taxonomicNameLabel id="7BCF57EFFFB52E2F20AAFBED7CF1FBEB" box="[1202,1257,1086,1111]" pageId="8" pageNumber="385" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| 
<collectorName id="FF7D5350FFB52E2F211AFBED7D65FBEB" box="[1282,1405,1086,1111]" pageId="8" pageNumber="385">Shavrin A.</collectorName>
&amp; 
<collectorName id="FF7D5350FFB52E2F248FFBB07902FBC7" box="[151,282,1123,1147]" pageId="8" pageNumber="385">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="3499AE43FFB52E2F2716FBB07B50FBC7" box="[782,840,1123,1147]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="8" pageNumber="385" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="52373686FFB52E2F24DFFB547A22FB5B" blockId="8.[151,1437,1014,2011]" pageId="8" pageNumber="385">
<typeStatus id="8D338824FFB52E2F24DFFB547921FB23" box="[199,313,1159,1183]" pageId="8" pageNumber="385" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="448EFD0FFFB52E2F255BFB547976FB22" box="[323,366,1159,1183]" pageId="8" pageNumber="385" type="male">1 ♂</specimenCount>
[specimen dissected; abdomen glued under the specimen], 
<materialsCitation id="E2E03CDBFFB52E2F2014FB547B41FB7F" ID-GBIF-Occurrence="2028139636" collectionCode="CS, CSC" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="8" pageNumber="385" specimenCount="7" specimenCount-female="6" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="448EFD0FFFB52E2F2014FB547C56FB22" box="[1036,1102,1159,1183]" pageId="8" pageNumber="385" type="female">3 ♀♀</specimenCount>
: same data as the holotype (
<specimenCount id="448EFD0FFFB52E2F2188FB5478B7FB7E" pageId="8" pageNumber="385" type="male">1 ♂</specimenCount>
: 
<collectionCode id="3499AE43FFB52E2F24A5FB7878FBFB7F" box="[189,227,1195,1219]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="8" pageNumber="385">CS</collectionCode>
; 
<specimenCount id="448EFD0FFFB52E2F24E8FB78792CFB7E" box="[240,308,1195,1219]" pageId="8" pageNumber="385" type="female">3 ♀♀</specimenCount>
: 
<collectionCode id="3499AE43FFB52E2F255BFB787964FB7F" box="[323,380,1195,1219]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="8" pageNumber="385" type="Herbarium">CSC</collectionCode>
). All 
<typeStatus id="8D338824FFB52E2F25A6FB787A31FB7F" box="[446,553,1195,1219]" pageId="8" pageNumber="385" type="paratype">paratypes</typeStatus>
with red rectangular label
</materialsCitation>
‘ 
<typeStatus id="8D338824FFB52E2F277EFB787BEFFB7F" box="[870,1015,1195,1219]" pageId="8" pageNumber="385" type="paratype">PARATYPE</typeStatus>
| 
<taxonomicName id="95884D05FFB52E2F2017FB797CAAFB7F" authorityName="Kraatz" authorityYear="1857" box="[1039,1202,1194,1219]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| maculosum 
<emphasis id="60FCEA94FFB52E2F214DFB797D95FB7F" bold="true" box="[1365,1421,1194,1219]" pageId="8" pageNumber="385">
<taxonomicNameLabel id="7BCF57EFFFB52E2F214DFB797D95FB7F" box="[1365,1421,1194,1219]" pageId="8" pageNumber="385" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. des. 2018’.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB52E2D24DFFB217CADFE3B" lastPageId="10" lastPageNumber="387" pageId="8" pageNumber="385" type="description">
<paragraph id="52373686FFB52E2F24DFFB217C90FAEF" blockId="8.[151,1437,1014,2011]" pageId="8" pageNumber="385">
<emphasis id="60FCEA94FFB52E2F24DFFB217949FAB7" bold="true" box="[199,337,1266,1291]" pageId="8" pageNumber="385">Description</emphasis>
. Measurements (n=5): HW: 0.49–0.56; HL: 0.22–0.24; AL (
<typeStatus id="8D338824FFB52E2F2019FB217C72FAB7" box="[1025,1130,1266,1291]" pageId="8" pageNumber="385" type="holotype">holotype</typeStatus>
): 0.85; OL: 0.13–0.15; PL: 0.50–0.57; PW: 0.75–0.90; ESL: 0.80–1.05; EW: 1.00–1.24; AW: 0.85–1.10; MTbL (
<typeStatus id="8D338824FFB52E2F2089FAC57CE2FA93" box="[1169,1274,1302,1327]" pageId="8" pageNumber="385" type="holotype">holotype</typeStatus>
): 0.45; MTrL (
<typeStatus id="8D338824FFB52E2F2487FAE97910FAEF" box="[159,264,1338,1363]" pageId="8" pageNumber="385" type="holotype">holotype</typeStatus>
): 0.39 (MTrL 1–4: 0.22; MTrL 5: 0.17); AedL: 0.57; TL: 2.00 (
<typeStatus id="8D338824FFB52E2F27D6FAE97C2CFAEF" box="[974,1076,1338,1363]" pageId="8" pageNumber="385" type="holotype">holotype</typeStatus>
) –2.35.
</paragraph>
<paragraph id="52373686FFB52E2F24DFFA8D7BC2F962" blockId="8.[151,1437,1014,2011]" pageId="8" pageNumber="385">
Head and pronotum yellow-brown; posterior portion of head, median part of neck and scutellum reddishbrown; disc of pronotum with moderately wide reddish-brown spot (indistinct in some 
<typeStatus id="8D338824FFB52E2F2069FA507CC6FA27" box="[1137,1246,1411,1435]" pageId="8" pageNumber="385" type="paratype">paratypes</typeStatus>
) beginning from anterior margin, stretching and gradually narrowing toward basal margin of pronotum; antennomeres 1–4, elytra and legs yellow; maxillary palpi and abdomen reddish-brown to brown (some 
<typeStatus id="8D338824FFB52E2F2050FA187CABFA5F" box="[1096,1203,1483,1507]" pageId="8" pageNumber="385" type="paratype">paratypes</typeStatus>
with yellow-brown abdomen, with yellow apical portion and paratergites). Forebody without distinct microsculpture except for posterior part of infraorbital ridges of head with indistinct diagonal and transverse meshes, neck and scutellum with distinct isodiametric microsculpture; abdominal tergites with distinct isodiametric microsculpture. Middle and posterior portions of head with irregular, small and moderately deep punctation; pronotum with dense and irregular small punctation slightly larger as that on head, distinctly sparser on medio-posterior third and lateral portions; punctation of elytra distinctly larger and deeper than that in pronotum, markedly smaller and denser on prescutellar area; abdominal tergites with irregular and very indistinct small punctures.
</paragraph>
<paragraph id="52373686FFB52E2F24DFF93978FEF867" blockId="8.[151,1437,1014,2011]" pageId="8" pageNumber="385">
Head more than twice as wide as long, flattened in middle. Ocelli very large, indistinct due the same coloration with surface of head, located at level of posterior fourth of eyes, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palp 1.1-1.2 times as long as preceding segment, gradually narrowed apicad from base. Antenna moderately short, reaching basal margin of elytra when reclined, with antennomeres gradually widened towards apex; length × width of antennomeres (
<typeStatus id="8D338824FFB52E2F20CAF8A97D23F82F" box="[1234,1339,1914,1939]" pageId="8" pageNumber="385" type="holotype">holotype</typeStatus>
): 1: 0.11 × 0.04; 2: 0.07 × 0.04; 3: 0.07 × 0.02; 4–6: 0.06 × 0.03; 6: 0.07 × 0.04; 7–8: 0.07 × 0.05; 9–10: 0.07 × 0.06; 11: 0.13 × 0.06.
</paragraph>
<caption id="06F7660EFFB42E2E248FF9DE7C0CF9FE" httpUri="https://zenodo.org/record/2607228/files/figure.png" pageId="9" pageNumber="386" startId="9.[151,264,1549,1572]" targetBox="[311,1291,206,1518]" targetPageId="9">
<paragraph id="52373686FFB42E2E248FF9DE7C0CF9FE" blockId="9.[151,1436,1549,1602]" pageId="9" pageNumber="386">
<emphasis id="60FCEA94FFB42E2E248FF9DE794DF99F" bold="true" box="[151,341,1549,1572]" pageId="9" pageNumber="386">FIGURES 16–19.</emphasis>
Habitus of 
<taxonomicName id="95884D05FFB42E2E25C8F9DE7A46F99F" authorityName="Kraatz" authorityYear="1857" box="[464,606,1549,1571]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="60FCEA94FFB42E2E25C8F9DE7A46F99F" box="[464,606,1549,1571]" italics="true" pageId="9" pageNumber="386">Amphichroum</emphasis>
</taxonomicName>
(males): 16– 
<taxonomicName id="95884D05FFB42E2E26FDF9DD7B67F998" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[741,895,1550,1572]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="propinquum" status="sp. nov.">
<emphasis id="60FCEA94FFB42E2E26FDF9DD7B67F998" box="[741,895,1550,1572]" italics="true" pageId="9" pageNumber="386">A. propinquum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB42E2E279CF9DD7BADF99F" box="[900,949,1550,1571]" pageId="9" pageNumber="386" rank="species">sp.n.</taxonomicNameLabel>
(holotype), 17– 
<taxonomicName id="95884D05FFB42E2E2043F9DD7CF6F99F" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1115,1262,1549,1571]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="rotundatum" status="sp. nov.">
<emphasis id="60FCEA94FFB42E2E2043F9DD7CF6F99F" box="[1115,1262,1549,1571]" italics="true" pageId="9" pageNumber="386">A. rotundatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB42E2E20ECF9DD7D3CF99F" box="[1268,1316,1550,1571]" pageId="9" pageNumber="386" rank="species">sp.n.</taxonomicNameLabel>
(holotype), 18– 
<taxonomicName id="95884D05FFB42E2E24A5F9FE792FF9FE" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[189,311,1580,1602]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="60FCEA94FFB42E2E24A5F9FE792FF9FE" box="[189,311,1580,1602]" italics="true" pageId="9" pageNumber="386">A. schuelkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB42E2E2525F9FE7976F9FE" box="[317,366,1581,1602]" pageId="9" pageNumber="386" rank="species">sp.n.</taxonomicNameLabel>
(paratype), 19– 
<taxonomicName id="95884D05FFB42E2E2616F9FE7AB9F9FE" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[526,673,1581,1602]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="squamosum" status="sp. nov.">
<emphasis id="60FCEA94FFB42E2E2616F9FE7AB9F9FE" box="[526,673,1581,1602]" italics="true" pageId="9" pageNumber="386">A. squamosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB42E2E26B0F9FE7AC0F9FE" box="[680,728,1581,1602]" pageId="9" pageNumber="386" rank="species">sp.n.</taxonomicNameLabel>
(holotype). Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="52373686FFB42E2E24DFF9A37B82F910" blockId="9.[151,1437,1646,2032]" pageId="9" pageNumber="386">Pronotum 1.5 times as wide as long, 1.5–1.6 times as wide as head, widest in middle, evenly narrowed both posteriad and anteriad; anterior and posterior angles widely rounded.</paragraph>
<paragraph id="52373686FFB42E2E24DFF9657A07F94F" blockId="9.[151,1437,1646,2032]" pageId="9" pageNumber="386">Elytra markedly wider than long, significantly widened apicad, reaching apical margin of abdominal tergite V, 1.6–1.8 times as long as pronotum.</paragraph>
<paragraph id="52373686FFB42E2E24DFF92D7B6BF84C" blockId="9.[151,1437,1646,2032]" pageId="9" pageNumber="386">
Male (
<figureCitation id="CAB32A03FFB42E2E2517F92D794DF8AB" box="[271,341,1790,1815]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1954,1977]" captionTargetBox="[296,1236,546,1939]" captionTargetId="figure@2.[283,1307,539,1939]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Amphichroum (males): 1–A. angustilobatum sp.n. (holotype), 2–A. assingi sp.n. (holotype), 3–A. longilobatum sp.n. (paratype), 4–A. maculosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="9" pageNumber="386">Fig. 4</figureCitation>
). Pronotum wide. Protarsomeres 1–4 very wide. Medial margin of apical part of protibia with two parallel rows of five to six short peg setae; mesotibia distinctly curved mediad, with dense row consisting of twelve to fourteen short and wide thorns. Abdominal tergite VIII with straight apical margin. Abdominal sternite VIII with slightly sinuate apical margin. Aedeagus (
<figureCitation id="CAB32A03FFB42E2E2691F8B87AF8F83F" box="[649,736,1899,1924]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[330,1322,210,1922]" captionTargetId="figure@7.[217,1370,161,1956]" captionTargetPageId="7" captionText="FIGURES 12–15.Aedeagus of Amphichroum longilobatum sp.n.(12–13: paratype) and A. maculosum sp.n. (14–15: paratype): 12, 14–parameral view, 13, 15–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607226/files/figure.png" pageId="9" pageNumber="386">Fig. 14</figureCitation>
) with median lobe markedly narrowed toward acute apex; parameres narrow, markedly exceeding apex of median lobe, with narrow setiferous lobes, with a pair of small apical and preapical setae; internal sac moderately long, with paired, very large and elongate structures in basal part and two elongated apical thorns. Aedeagus laterally as in 
<figureCitation id="CAB32A03FFB42E2E2704F8047B77F84C" box="[796,879,2007,2032]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1958,1981]" captionTargetBox="[330,1322,210,1922]" captionTargetId="figure@7.[217,1370,161,1956]" captionTargetPageId="7" captionText="FIGURES 12–15.Aedeagus of Amphichroum longilobatum sp.n.(12–13: paratype) and A. maculosum sp.n. (14–15: paratype): 12, 14–parameral view, 13, 15–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607226/files/figure.png" pageId="9" pageNumber="386">Fig. 15</figureCitation>
.
</paragraph>
<paragraph id="52373686FFB72E2D24DFFF447CE7FF69" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="387">Female. Pronotum narrower. Protarsomeres 1–4 moderately narrow. Medial margin of protibia and mesotibia without modifications, slender. Abdominal tergite VIII and sternite VIII with rounded apical margin.</paragraph>
<paragraph id="52373686FFB72E2D24DFFF0C7CADFE3B" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="387">
<emphasis id="60FCEA94FFB72E2D24DFFF0C79AAFF44" bold="true" box="[199,434,223,248]" pageId="10" pageNumber="387">Comparative notes.</emphasis>
<taxonomicName id="95884D05FFB72E2D25A4FF337AFAFF44" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[444,738,224,248]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="maculosum" status="sp. nov.">
<emphasis id="60FCEA94FFB72E2D25A4FF337AFAFF44" box="[444,738,224,248]" italics="true" pageId="10" pageNumber="387">Amphichroum maculosum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFB72E2D26F3FF0C7B3BFF44" bold="true" box="[747,803,223,248]" pageId="10" pageNumber="387">
<taxonomicNameLabel id="7BCF57EFFFB72E2D26F3FF0C7B3BFF44" box="[747,803,223,248]" pageId="10" pageNumber="387" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from the remaining species of the genus by the presence of longitudinal reddish-brown spot on the pronotum. Based on the general shape of the apical part of the aedeagus and parameres, 
<taxonomicName id="95884D05FFB72E2D25D1FEFB7A77FEFC" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[457,623,296,320]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="maculosum" status="sp. nov.">
<emphasis id="60FCEA94FFB72E2D25D1FEFB7A77FEFC" box="[457,623,296,320]" italics="true" pageId="10" pageNumber="387">A. maculosum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFB72E2D2665FEF47AACFEFC" bold="true" box="[637,692,295,320]" pageId="10" pageNumber="387">
<taxonomicNameLabel id="7BCF57EFFFB72E2D2665FEF47AACFEFC" box="[637,692,295,320]" pageId="10" pageNumber="387" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="95884D05FFB72E2D274CFEFB7CECFEFC" authority="Coiffait, 1984" authorityName="Coiffait" authorityYear="1984" box="[852,1268,295,320]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="milkense">
<emphasis id="60FCEA94FFB72E2D274CFEFB7BC5FEFC" box="[852,989,296,320]" italics="true" pageId="10" pageNumber="387">A. milkense</emphasis>
<bibRefCitation id="36194B77FFB72E2D27F2FEF47C96FEFC" author="Coiffait, H." box="[1002,1166,295,320]" pageId="10" pageNumber="387" pagination="373 - 387" refId="ref16262" refString="Coiffait, H. (1984) Contribution a la connaissance des staphylinides de l'Himalaya (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France, New Series, 20 (4), 373 - 387." type="journal article" year="1984">Coiffait, 1984</bibRefCitation>
(
<collectingCountry id="2A9F7616FFB72E2D20BBFEF47CF3FEFC" box="[1187,1259,295,320]" name="Nepal" pageId="10" pageNumber="387">Nepal</collectingCountry>
)
</taxonomicName>
, 
<taxonomicName id="95884D05FFB72E2D211EFEFB795EFED8" authority="Cameron, 1928" authorityName="Cameron" authorityYear="1928" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="60FCEA94FFB72E2D211EFEFB7D84FEFC" box="[1286,1436,296,320]" italics="true" pageId="10" pageNumber="387">A. monticola</emphasis>
<bibRefCitation id="36194B77FFB72E2D248FFE9F795EFED8" author="Cameron, M." box="[151,326,332,356]" pageId="10" pageNumber="387" pagination="558 - 569" refId="ref16128" refString="Cameron, M. (1928) The Staphylinidae (Coleoptera) of the Third Mount Everest Expedition. The Annals and Magazine of Natural History, Series 10, 2 (12), 558 - 569. https: // doi. org / 10.1080 / 00222932808672919" type="journal article" year="1928">Cameron, 1928</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="2A9F7616FFB72E2D2540FE9F79B5FED9" box="[344,429,332,357]" name="Bhutan" pageId="10" pageNumber="387">Bhutan</collectingCountry>
, Xizang) and 
<taxonomicName id="95884D05FFB72E2D2648FE9F7C13FED9" authority="Coiffait, 1978" authorityName="Coiffait" authorityYear="1978" box="[592,1035,332,357]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="387" phylum="Arthropoda" rank="species" species="reticulatum">
<emphasis id="60FCEA94FFB72E2D2648FE9F7AEAFED8" box="[592,754,332,356]" italics="true" pageId="10" pageNumber="387">A. reticulatum</emphasis>
<bibRefCitation id="36194B77FFB72E2D26E4FE9F7B83FED8" author="Coiffait, H." box="[764,923,332,357]" pageId="10" pageNumber="387" pagination="109 - 150" refId="ref16216" refString="Coiffait, H. (1978) Ergebnisse der Bhutan-Expedition 1972 des Naturhistorischen Museums in Basel. Coleoptera: Fam. Staphylinidae Subfam. Paederinae, Euaesthetinae, Piestinae, Osoriinae et Omalinae [sic]. Entomologica Basiliensia, 3, 109 - 150." type="journal article" year="1978">Coiffait, 1978</bibRefCitation>
(
<collectingCountry id="2A9F7616FFB72E2D27B4FE9F7C1AFED9" box="[940,1026,332,357]" name="Bhutan" pageId="10" pageNumber="387">Bhutan</collectingCountry>
)
</taxonomicName>
, but it differs by the smaller body, paler forebody, wider basal portion of the aedeagus and different structures of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB72E2D24DFFE4779F1FDA4" pageId="10" pageNumber="387" type="distribution">
<paragraph id="52373686FFB72E2D24DFFE4779B5FE6C" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="387">
<emphasis id="60FCEA94FFB72E2D24DFFE477978FE11" bold="true" box="[199,352,404,429]" pageId="10" pageNumber="387">Distribution.</emphasis>
The new species is at present known only from the 
<typeStatus id="8D338824FFB72E2D27A9FE467BF9FE11" box="[945,993,405,429]" pageId="10" pageNumber="387">type</typeStatus>
locality in the Diancang Shan range in 
<collectingRegion id="904CF864FFB72E2D248FFE6B78E8FE73" box="[151,240,440,463]" country="China" name="Yunnan" pageId="10" pageNumber="387">Yunnan</collectingRegion>
, 
<collectingCountry id="2A9F7616FFB72E2D24E2FE647925FE6C" box="[250,317,439,464]" name="China" pageId="10" pageNumber="387">China</collectingCountry>
(
<figureCitation id="CAB32A03FFB72E2D2555FE6479B8FE6C" box="[333,416,439,464]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="10" pageNumber="387">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="52373686FFB72E2D24DFFE0F79F1FDA4" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="387">
<emphasis id="60FCEA94FFB72E2D24DFFE0F7952FE49" bold="true" box="[199,330,476,501]" pageId="10" pageNumber="387">Bionomics.</emphasis>
Specimens were collected at elevations from 
<quantity id="95709B63FFB72E2D2745FE0F7C0DFE48" box="[861,1045,476,501]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="2.98" metricValueMin="2.96" pageId="10" pageNumber="387" unit="m" value="2970.0" valueMax="2980.0" valueMin="2960.0">
<date id="26361046FFB72E2D2745FE0F7BEEFE48" box="[861,1014,476,501]" pageId="10" pageNumber="387" value="2960" valueMax="2980">2960 to 2980</date>
m
</quantity>
a.s.l. from gravel and from under stones near a former stone pit.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB72E2D24DFFDF77A7FFDE3" pageId="10" pageNumber="387" type="etymology">
<paragraph id="52373686FFB72E2D24DFFDF77A7FFDE3" blockId="10.[151,1436,151,608]" pageId="10" pageNumber="387">
<emphasis id="60FCEA94FFB72E2D24DFFDF77956FD81" bold="true" box="[199,334,548,573]" pageId="10" pageNumber="387">Etymology.</emphasis>
The specific epithet is the Latin adjective 
<emphasis id="60FCEA94FFB72E2D2729FDF77BBFFD80" box="[817,935,548,572]" italics="true" pageId="10" pageNumber="387">maculosus</emphasis>
, - 
<emphasis id="60FCEA94FFB72E2D27D8FDF67BD6FD80" box="[960,974,549,572]" italics="true" pageId="10" pageNumber="387">a</emphasis>
, - 
<emphasis id="60FCEA94FFB72E2D27FEFDF67C10FD80" box="[998,1032,549,572]" italics="true" pageId="10" pageNumber="387">um</emphasis>
(with spot or spots). It refers to the presence of the spot on disc of pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>