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<document id="45465157EF4B48EFE768109CEC7FD5D1" ID-CLB-Dataset="28041" ID-DOI="10.11646/zootaxa.4508.3.3" ID-GBIF-Dataset="9de070ed-9c35-4131-a350-72371ee93ddf" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607216" ID-ZooBank="E53B4C53-554A-4913-8E3B-CE1C530DF004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586064605" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2018" docId="DA218790FFAB2E3F248FFC0778CDFF69" docLanguage="en" docName="zootaxa.4508.3.3.pdf" docOrigin="Zootaxa 4508 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum tibetanum Shavrin &amp; Smetana 2018, sp.n." docType="treatment" docVersion="8" lastPageNumber="401" masterDocId="2618FFE8FFBD2E272418FFD37818FFBC" masterDocTitle="New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China" masterLastPageNumber="402" masterPageNumber="377" pageNumber="399" updateTime="1698713048312" updateUser="ExternalLinkService">
<mods:mods id="FAFC3AF5BB0490DE94E6AF83D994C817" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="59BD00069025CD035ED0EE24639D3E5E">New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China</mods:title>
</mods:titleInfo>
<mods:name id="7C5DD0D15FE6D035B7EC74BC0D66F1B2" type="personal">
<mods:role id="D28300DDA8EAD4C0DE606C5838226DF2">
<mods:roleTerm id="F5084B7A2237669454ADFB04C5ADD582">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53EE24B66E021E639D2DC76316B62606">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:name id="B2B229979B0FD9C581E41ED163E217CB" type="personal">
<mods:role id="A0F2A607C4035A8D06F863A001972B75">
<mods:roleTerm id="A5F5B2ED5C639B883F2CCA0B1FC6A545">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41159906382BFD0918DAF88DA8510D81">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="741E575E40558807C13E36ADD7DE6DDA">text</mods:typeOfResource>
<mods:relatedItem id="4829669B13AAEAAB6B1C73770E3FEBD8" type="host">
<mods:titleInfo id="A07D146788AB7D911D0593291CBA90AA">
<mods:title id="BA7A1CAE22899529AACB21D109D906DD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="ECAA7885105B9EDD24EBBEE9DB90D6DA">
<mods:date id="9268B4DE3B200E07D7FA542479DB3335">2018</mods:date>
<mods:detail id="94C085783C2FE8CF6D576FFA252BB61E" type="pubDate">
<mods:number id="AC7EB296D325D979DDBBEAAD4D3D8994">2018-11-01</mods:number>
</mods:detail>
<mods:detail id="4C89F275977B04C1AD5E52D3CE30540F" type="volume">
<mods:number id="1EFAB9912F488A4E58CC7C99BAFFEBD8">4508</mods:number>
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<mods:detail id="229E8F2E49E48051F0A9F54BE5AF433F" type="issue">
<mods:number id="C1C3FDB3FA686FE3A1F50AE746CC385B">3</mods:number>
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<mods:start id="EBDFAD54515DDF1671206FF36F6B4494">377</mods:start>
<mods:end id="2E9C91B5BA8FDF88226FEEE28C640B6A">402</mods:end>
</mods:extent>
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<mods:classification id="AB52B6E79051DF3AAE1CFC97240DF7F7">journal article</mods:classification>
<mods:identifier id="7B502D691CEDCB33E5173AE9C5CB2CF1" type="CLB-Dataset">28041</mods:identifier>
<mods:identifier id="FC778B086EB044184421F4015D642016" type="DOI">10.11646/zootaxa.4508.3.3</mods:identifier>
<mods:identifier id="F3D3A0FD49C16CBB3F96243BDB8CCF55" type="GBIF-Dataset">9de070ed-9c35-4131-a350-72371ee93ddf</mods:identifier>
<mods:identifier id="C5CBBBEB36B07DE90CF8D7217799D70E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CA22C9B3D8E0A32AE91ACF7EEF32362B" type="Zenodo-Dep">2607216</mods:identifier>
<mods:identifier id="9D2B9B332561A85E28E5B0C22A0E9532" type="ZooBank">E53B4C53-554A-4913-8E3B-CE1C530DF004</mods:identifier>
</mods:mods>
<treatment id="DA218790FFAB2E3F248FFC0778CDFF69" ID-DOI="http://doi.org/10.5281/zenodo.5963006" ID-GBIF-Taxon="154495886" ID-Zenodo-Dep="5963006" LSID="urn:lsid:plazi:treatment:DA218790FFAB2E3F248FFC0778CDFF69" httpUri="http://treatment.plazi.org/id/DA218790FFAB2E3F248FFC0778CDFF69" lastPageId="24" lastPageNumber="401" pageId="22" pageNumber="399">
<subSubSection id="1A92650DFFAB2E31248FFC077B08FC52" box="[151,784,980,1006]" pageId="22" pageNumber="399" type="nomenclature">
<paragraph id="52373686FFAB2E31248FFC077B08FC52" blockId="22.[151,784,980,1039]" box="[151,784,980,1006]" pageId="22" pageNumber="399">
<heading id="097F81EAFFAB2E31248FFC077B08FC52" bold="true" box="[151,784,980,1006]" fontSize="11" level="1" pageId="22" pageNumber="399" reason="1">
<emphasis id="60FCEA94FFAB2E31248FFC077B08FC52" bold="true" box="[151,784,980,1006]" pageId="22" pageNumber="399">
<taxonomicName id="95884D05FFAB2E31248FFC077AD0FC52" ID-CoL="CZQP" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,712,980,1006]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="399" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="60FCEA94FFAB2E31248FFC0779D0FC52" bold="true" box="[151,456,980,1006]" italics="true" pageId="22" pageNumber="399">Amphichroum tibetanum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="7BCF57EFFFAB2E3126CDFC077B08FC52" box="[725,784,980,1006]" pageId="22" pageNumber="399" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFAB2E31248FFC257940FBB3" box="[151,344,1014,1039]" pageId="22" pageNumber="399" type="description">
<paragraph id="52373686FFAB2E31248FFC257940FBB3" blockId="22.[151,784,980,1039]" box="[151,344,1014,1039]" pageId="22" pageNumber="399">
(
<figureCitation id="CAB32A03FFAB2E312487FC2578E4FBB3" box="[159,252,1014,1039]" captionStart="FIGURES 32–34" captionStartId="19.[151,264,1835,1858]" captionTargetBox="[243,1357,849,1808]" captionTargetId="figure@19.[175,1411,805,1814]" captionTargetPageId="19" captionText="FIGURES 32–34. Habitus of Amphichroum (males): 32—A. subaequale sp.n. (paratype), 33—A. subelongatum sp.n. (paratype), 34—A. tibetanum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607238/files/figure.png" pageId="22" pageNumber="399">Figs. 34</figureCitation>
,
<figureCitation id="CAB32A03FFAB2E312510FC247957FBB3" box="[264,335,1015,1039]" captionStart-0="FIGURES 39–40" captionStart-1="FIGURE 41" captionStartId-0="23.[151,264,1606,1629]" captionStartId-1="24.[151,250,1057,1079]" captionTargetBox-0="[280,1350,527,1566]" captionTargetBox-1="[179,1408,267,1035]" captionTargetId-0="figure@23.[151,1436,481,1585]" captionTargetId-1="figure@24.[179,1408,267,1035]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionText-0="FIGURES 39–40. Aedeagus of Amphichroum tibetanum sp.n. (paratype): 39—parameral view, 40—lateral view. Scale bar: 0.1 mm." captionText-1="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri-0="https://zenodo.org/record/2607242/files/figure.png" httpUri-1="https://zenodo.org/record/2607244/files/figure.png" pageId="22" pageNumber="399">39–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFAB2E31248FFBED790DFACB" pageId="22" pageNumber="399" type="materials_examined">
<paragraph id="52373686FFAB2E31248FFBED7954FB5B" blockId="22.[151,1437,1086,2011]" pageId="22" pageNumber="399">
<emphasis id="60FCEA94FFAB2E31248FFBED79DDFBEB" bold="true" box="[151,453,1086,1111]" pageId="22" pageNumber="399">Type material examined:</emphasis>
<materialsCitation id="E2E03CDBFFAB2E3125D5FBED7D91FB23" ID-GBIF-Occurrence="2028139601" collectingDate="1997-06-24" collectingDateMax="1997-06-27" collectingDateMin="1997-06-24" collectorName="Jaroslav Turna &amp; Sammlung &amp; M. Schulke &amp; Berlin'" elevation="4400" latitude="29.266666" location="Yanjing-Markam" longLatPrecision="1228" longitude="98.63333" pageId="22" pageNumber="399" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="8D338824FFAB2E3125D5FBED7A2DFBEB" box="[461,565,1086,1111]" pageId="22" pageNumber="399" type="holotype">Holotype</typeStatus>
<specimenCount id="448EFD0FFFAB2E312626FBEC7A4BFBEA" box="[574,595,1087,1110]" pageId="22" pageNumber="399" type="male">♂</specimenCount>
[specimen without antennomeres 10–11]: ‘SE
<collectingRegion id="904CF864FFAB2E312076FBED7CB3FBEB" box="[1134,1195,1086,1111]" country="China" name="Xizang" pageId="22" pageNumber="399">Tibet</collectingRegion>
, “Chola Shan” pass | road
<location id="5757605DFFAB2E3124CAFBB179BCFBC7" LSID="urn:lsid:plazi:treatment:DA218790FFAB2E3F248FFC0778CDFF69:5757605DFFAB2E3124CAFBB179BCFBC7" box="[210,420,1122,1147]" latitude="29.266666" longLatPrecision="1228" longitude="98.63333" name="Yanjing-Markam" pageId="22" pageNumber="399" stateProvince="Xizang">Yanjing—Markam</location>
|
<locationDeviation id="43852FA4FFAB2E3125A6FBB17A92FBC7" box="[446,650,1122,1147]" location="Markam" pageId="22" pageNumber="399">
<quantity id="95709B63FFAB2E3125A6FBB17A10FBC7" box="[446,520,1122,1147]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="399" unit="km" value="50.0">50 km</quantity>
S
<emphasis id="60FCEA94FFAB2E312633FBB07A92FBC7" box="[555,650,1123,1147]" italics="true" pageId="22" pageNumber="399">Markam</emphasis>
</locationDeviation>
,
<quantity id="95709B63FFAB2E312683FBB07AF1FBC7" box="[667,745,1123,1147]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="22" pageNumber="399" unit="m" value="4400.0">
<elevation id="D9A5D1B5FFAB2E312683FBB07AF1FBC7" box="[667,745,1123,1147]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="22" pageNumber="399" unit="m" value="4400.0">4400m</elevation>
</quantity>
|
<geoCoordinate id="37BC5041FFAB2E31271AFBB17B78FBC7" box="[770,864,1122,1147]" direction="north" orientation="latitude" pageId="22" pageNumber="399" precision="925" value="29.266666">29°16'N</geoCoordinate>
,
<geoCoordinate id="37BC5041FFAB2E312777FBB07BD1FBC7" box="[879,969,1123,1147]" direction="east" orientation="longitude" pageId="22" pageNumber="399" precision="925" value="98.63333">98°38'E</geoCoordinate>
| mixed forest,
<date id="26361046FFAB2E31209AFBB07D3FFBC7" box="[1154,1319,1123,1147]" pageId="22" pageNumber="399" value="1997-06-24" valueMax="1997-06-27" valueMin="1997-06-24">
<collectingDate id="3672E9AEFFAB2E31209AFBB07D3FFBC7" box="[1154,1319,1123,1147]" pageId="22" pageNumber="399" value="1997-06-24" valueMax="1997-06-27" valueMin="1997-06-24">24-27.VI.1997</collectingDate>
</date>
|
<collectorName id="FF7D5350FFAB2E312158FBB178C1FB22" pageId="22" pageNumber="399">Jaroslav Turna</collectorName>
leg.’ &lt;rectangular label, printed&gt;, ‘
<collectorName id="FF7D5350FFAB2E31266FFB557AEEFB23" box="[631,758,1158,1183]" pageId="22" pageNumber="399">Sammlung</collectorName>
|
<collectorName id="FF7D5350FFAB2E312709FB547B8EFB23" box="[785,918,1158,1183]" pageId="22" pageNumber="399">M. Schülke</collectorName>
|
<collectorName id="FF7D5350FFAB2E3127AAFB557C19FB23" box="[946,1025,1158,1183]" pageId="22" pageNumber="399">Berlin’</collectorName>
&lt;green rectangular label, printed
</materialsCitation>
&gt;, ‘
<materialsCitation id="E2E03CDBFFAB2E312483FB787950FB5B" ID-GBIF-Occurrence="2028139608" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." pageId="22" pageNumber="399" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="8D338824FFAB2E312483FB78792CFB7F" box="[155,308,1195,1219]" pageId="22" pageNumber="399" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="95884D05FFAB2E312557FB7979EAFB7F" authorityName="Kraatz" authorityYear="1857" box="[335,498,1194,1219]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| tibetanum
<emphasis id="60FCEA94FFAB2E312690FB797AD8FB7F" bold="true" box="[648,704,1194,1219]" pageId="22" pageNumber="399">
<taxonomicNameLabel id="7BCF57EFFFAB2E312690FB797AD8FB7F" box="[648,704,1194,1219]" pageId="22" pageNumber="399" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFAB2E3126C3FB797B40FB7F" box="[731,856,1194,1219]" pageId="22" pageNumber="399">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFAB2E31279BFB787C12FB7F" box="[899,1034,1195,1219]" pageId="22" pageNumber="399">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="3499AE43FFAB2E31251DFB1C7927FB5B" box="[261,319,1231,1255]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="22" pageNumber="399" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="52373686FFAB2E3124DFFB20790DFACB" blockId="22.[151,1437,1086,2011]" pageId="22" pageNumber="399">
<typeStatus id="8D338824FFAB2E3124DFFB207921FAB7" box="[199,313,1267,1291]" pageId="22" pageNumber="399" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="448EFD0FFFAB2E31255CFB207975FAB7" box="[324,365,1266,1291]" pageId="22" pageNumber="399" type="male">1 ♂</specimenCount>
[specimen without antennomeres 9–11],
<materialsCitation id="E2E03CDBFFAB2E312725FB20794AFAEF" ID-GBIF-Occurrence="2028139607" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." pageId="22" pageNumber="399" specimenCount="2" specimenCount-female="2" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="448EFD0FFFAB2E312725FB207B65FAB6" box="[829,893,1267,1291]" pageId="22" pageNumber="399" type="female">2 ♀♀</specimenCount>
[one specimen without antennomeres 7–11 and other specimen without left antennomeres 3–11 and right antennomeres 8–11]: same data as the holotype, with red rectangular label
</materialsCitation>
‘
<materialsCitation id="E2E03CDBFFAB2E312545FAE87909FACB" ID-GBIF-Occurrence="2028139613" collectionCode="CS, CSC" collectorName="Shavrin A. &amp; Smetana A." pageId="22" pageNumber="399" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="8D338824FFAB2E312545FAE879F5FAEF" box="[349,493,1339,1363]" pageId="22" pageNumber="399" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="95884D05FFAB2E312619FAE97ABCFAEF" authorityName="Kraatz" authorityYear="1857" box="[513,676,1338,1363]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| tibetanum
<emphasis id="60FCEA94FFAB2E312736FAE97B7EFAEF" bold="true" box="[814,870,1338,1363]" pageId="22" pageNumber="399">
<taxonomicNameLabel id="7BCF57EFFFAB2E312736FAE97B7EFAEF" box="[814,870,1338,1363]" pageId="22" pageNumber="399" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFAB2E312762FAE97BEBFAEF" box="[890,1011,1338,1363]" pageId="22" pageNumber="399">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFAB2E31200EFAE87C81FAEF" box="[1046,1177,1339,1363]" pageId="22" pageNumber="399">Smetana A.</collectorName>
des. 2018’ (
<specimenCount id="448EFD0FFFAB2E31213EFAE87D49FAEE" box="[1318,1361,1339,1363]" pageId="22" pageNumber="399" type="male">1 ♂</specimenCount>
:
<collectionCode id="3499AE43FFAB2E312145FAE87D9AFAEF" box="[1373,1410,1339,1363]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="22" pageNumber="399">CS</collectionCode>
;
<specimenCount id="448EFD0FFFAB2E312196FAE878DCFACA" pageId="22" pageNumber="399" type="female">2 ♀♀</specimenCount>
:
<collectionCode id="3499AE43FFAB2E3124C8FA8C7911FACB" box="[208,265,1375,1399]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="22" pageNumber="399" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFAB2E3024DFFA517A1AF8C8" lastPageId="23" lastPageNumber="400" pageId="22" pageNumber="399" type="description">
<paragraph id="52373686FFAB2E3124DFFA517B61FA5F" blockId="22.[151,1437,1086,2011]" pageId="22" pageNumber="399">
<emphasis id="60FCEA94FFAB2E3124DFFA517949FA27" bold="true" box="[199,337,1410,1435]" pageId="22" pageNumber="399">Description</emphasis>
. Measurements (n=4): HW: 0.65–0.70; HL: 0.35–0.39; OL: 0.19–0.20; PL: 0.57–0.62; PW: 0.97– 1.04; ESL: 1.17–1.40; EW: 1.27–1.42; AW: 1.22–1.35; MTbL(
<typeStatus id="8D338824FFAB2E312746FA757BDCFA03" box="[862,964,1446,1471]" pageId="22" pageNumber="399" type="holotype">holotype</typeStatus>
): 0.72; MTrL(
<typeStatus id="8D338824FFAB2E31207EFA757CD3FA03" box="[1126,1227,1446,1471]" pageId="22" pageNumber="399" type="holotype">holotype</typeStatus>
): 0.54 (MTrL 1–4: 0.32; MTrL 5: 0.22); AedL: 0.72; TL: 3.20–3.85 (
<typeStatus id="8D338824FFAB2E3126DDFA197B35FA5F" box="[709,813,1482,1507]" pageId="22" pageNumber="399" type="holotype">holotype</typeStatus>
: 3.30).
</paragraph>
<paragraph id="52373686FFAB2E3124DFFA3D7C1DF962" blockId="22.[151,1437,1086,2011]" pageId="22" pageNumber="399">Forebody yellow-brown, with somewhat paler lateral and basal portions of pronotum; mouthparts, antennomeres 1–3 and legs yellow; apical segment of maxillary palp and antennomeres 4–11 brown; abdomen reddish-brown. Head with distinct, dense isodiametric microsculpture; microsculpture of middle part of neck as that on head but distinctly stronger; pronotum with distinct microsculpture as that on head; scutellum with distinct transverse meshes. Posterior portion of head between grooves and infraorbital ridges with several small punctures; pronotum with dense, irregular and very small punctation, distinctly sparser on medio-basal portion; punctation of elytra dense, moderately large and deep, markedly denser on prescutellar area.</paragraph>
<paragraph id="52373686FFAB2E3124DFF9397B75F89B" blockId="22.[151,1437,1086,2011]" pageId="22" pageNumber="399">
Head 1.7–1.8 times as wide as long. Length × width of antennomeres (
<typeStatus id="8D338824FFAB2E3127CEF9397C27F8BF" box="[982,1087,1770,1795]" pageId="22" pageNumber="399" type="holotype">holotype</typeStatus>
): 1: 0.15 × 0.05; 2: 0.13 × 0.05; 3: 0.12 × 0.04; 4: 0.10 × 0.05; 5–6: 0.10 × 0.06;7–9: 0.11 × 0.06.
</paragraph>
<paragraph id="52373686FFAB2E3124DFF8E0798EF8D3" blockId="22.[151,1437,1086,2011]" pageId="22" pageNumber="399">Pronotum 1.6–1.7 times as wide as long, 1.4 times as wide as head, widest in middle, evenly narrowed both posteriad and anteriad.</paragraph>
<paragraph id="52373686FFAB2E3124DFF8A97C81F82E" blockId="22.[151,1437,1086,2011]" box="[199,1177,1914,1939]" pageId="22" pageNumber="399">Elytra slightly wider than long and slightly widened apicad, twice as long as pronotum.</paragraph>
<paragraph id="52373686FFAB2E3124DFF84D7A84F80A" blockId="22.[151,1437,1086,2011]" box="[199,668,1950,1975]" pageId="22" pageNumber="399">Metatibia 1.3 times as long as metatarsus.</paragraph>
<paragraph id="52373686FFAB2E3024DFF8117B07FED8" blockId="22.[151,1437,1086,2011]" lastBlockId="23.[151,1436,151,429]" lastPageId="23" lastPageNumber="400" pageId="22" pageNumber="399">
Male (
<figureCitation id="CAB32A03FFAB2E312508F811797CF867" box="[272,356,1986,2011]" captionStart="FIGURES 32–34" captionStartId="19.[151,264,1835,1858]" captionTargetBox="[243,1357,849,1808]" captionTargetId="figure@19.[175,1411,805,1814]" captionTargetPageId="19" captionText="FIGURES 32–34. Habitus of Amphichroum (males): 32—A. subaequale sp.n. (paratype), 33—A. subelongatum sp.n. (paratype), 34—A. tibetanum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607238/files/figure.png" pageId="22" pageNumber="399">Fig. 34</figureCitation>
). Protarsomeres 1–4 wide. Medial margin of apical part of protibia with two parallel rows of ten short peg setae; mesotibia moderately strongly curved mediad, with dense row of twenty short and wide thorns, slightly elongated toward apex. Abdominal tergite VIII and sternite VIII with straight apical margin. Aedeagus (
<figureCitation id="CAB32A03FFAA2E302487FF0C78E9FF44" box="[159,241,223,248]" captionStart="FIGURES 39–40" captionStartId="23.[151,264,1606,1629]" captionTargetBox="[280,1350,527,1566]" captionTargetId="figure@23.[151,1436,481,1585]" captionTargetPageId="23" captionText="FIGURES 39–40. Aedeagus of Amphichroum tibetanum sp.n. (paratype): 39—parameral view, 40—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607242/files/figure.png" pageId="23" pageNumber="400">Fig. 39</figureCitation>
) with median lobe gradually narrowed toward rounded apex; parameres narrow, distinctly exceeding apex of median lobe, with two pairs of apical and preapical short setae; internal sac long, with extensive field of densely set fine thorns in apical and middle portion and with wide field of strongly sclerotized large thorns and oval structure in basal portion.Aedeagus laterally as in
<figureCitation id="CAB32A03FFAA2E3026D0FE9F7B03FED8" box="[712,795,332,357]" captionStart="FIGURES 39–40" captionStartId="23.[151,264,1606,1629]" captionTargetBox="[280,1350,527,1566]" captionTargetId="figure@23.[151,1436,481,1585]" captionTargetPageId="23" captionText="FIGURES 39–40. Aedeagus of Amphichroum tibetanum sp.n. (paratype): 39—parameral view, 40—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607242/files/figure.png" pageId="23" pageNumber="400">Fig. 40</figureCitation>
.
</paragraph>
<paragraph id="52373686FFAA2E3024DFFEBC7AA5FE11" blockId="23.[151,1436,151,429]" pageId="23" pageNumber="400">Female. Protarsomeres 1–4 moderately narrow. Pro- and mesotibiae without modifications. Apical margin of abdominal tergite VIII and sternite VIII rounded.</paragraph>
<caption id="06F7660EFFAA2E30248FF99578F3F9C7" httpUri="https://zenodo.org/record/2607242/files/figure.png" pageId="23" pageNumber="400" startId="23.[151,264,1606,1629]" targetBox="[280,1350,527,1566]" targetPageId="23">
<paragraph id="52373686FFAA2E30248FF99578F3F9C7" blockId="23.[151,1436,1606,1659]" pageId="23" pageNumber="400">
<emphasis id="60FCEA94FFAA2E30248FF995794EF9E0" bold="true" box="[151,342,1606,1629]" pageId="23" pageNumber="400">FIGURES 39–40.</emphasis>
Aedeagus of
<taxonomicName id="95884D05FFAA2E3025FFF9957AFBF9E0" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[487,739,1606,1628]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="400" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="60FCEA94FFAA2E3025FFF9957AFBF9E0" box="[487,739,1606,1628]" italics="true" pageId="23" pageNumber="400">Amphichroum tibetanum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFAA2E3026F4F9947B07F9E0" bold="true" box="[748,799,1607,1628]" pageId="23" pageNumber="400">
<taxonomicNameLabel id="7BCF57EFFFAA2E3026F4F9947B07F9E0" box="[748,799,1607,1628]" pageId="23" pageNumber="400" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
(paratype): 39—parameral view, 40—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="52373686FFAA2E3024DFF97B7A1AF8C8" blockId="23.[151,1436,1704,2017]" pageId="23" pageNumber="400">
<emphasis id="60FCEA94FFAA2E3024DFF97B79A8F97D" bold="true" box="[199,432,1704,1729]" pageId="23" pageNumber="400">Comparative notes.</emphasis>
Based on the body size, shape of long parameres significantly exceeding median lobe and on the presence of fields of large thorns in the internal sac,
<taxonomicName id="95884D05FFAA2E30272FF91E7BDFF958" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[823,967,1740,1764]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="400" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="60FCEA94FFAA2E30272FF91E7BDFF958" box="[823,967,1740,1764]" italics="true" pageId="23" pageNumber="400">A. tibetanum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFAA2E3027D7F91F7C1FF959" bold="true" box="[975,1031,1740,1765]" pageId="23" pageNumber="400">
<taxonomicNameLabel id="7BCF57EFFFAA2E3027D7F91F7C1FF959" box="[975,1031,1740,1765]" pageId="23" pageNumber="400" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="95884D05FFAA2E302084F91E7D13F958" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1180,1291,1741,1764]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="400" phylum="Arthropoda" rank="species" species="assingi" status="sp. nov.">
<emphasis id="60FCEA94FFAA2E302084F91E7D13F958" box="[1180,1291,1741,1764]" italics="true" pageId="23" pageNumber="400">A. assingi</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFAA2E30210CF91F7D53F959" bold="true" box="[1300,1355,1740,1765]" pageId="23" pageNumber="400">
<taxonomicNameLabel id="7BCF57EFFFAA2E30210CF91F7D53F959" box="[1300,1355,1740,1765]" pageId="23" pageNumber="400" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="95884D05FFAA2E30219CF91E7901F8B4" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="400" phylum="Arthropoda" rank="species" species="rotundatum" status="sp. nov.">
<emphasis id="60FCEA94FFAA2E30219CF91E7901F8B4" italics="true" pageId="23" pageNumber="400">A. rotundatum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFAA2E30253AF9237941F8B5" bold="true" box="[290,345,1776,1801]" pageId="23" pageNumber="400">
<taxonomicNameLabel id="7BCF57EFFFAA2E30253AF9237941F8B5" box="[290,345,1776,1801]" pageId="23" pageNumber="400" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
Based on the shape of apex of the median lobe, it’s most similar to
<taxonomicName id="95884D05FFAA2E302069F9237D0DF8B4" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1137,1301,1776,1800]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="400" phylum="Arthropoda" rank="species" species="rotundatum" status="sp. nov.">
<emphasis id="60FCEA94FFAA2E302069F9237D0DF8B4" box="[1137,1301,1776,1800]" italics="true" pageId="23" pageNumber="400">A. rotundatum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFAA2E302105F9237D4DF8B5" bold="true" box="[1309,1365,1776,1801]" pageId="23" pageNumber="400">
<taxonomicNameLabel id="7BCF57EFFFAA2E302105F9237D4DF8B5" box="[1309,1365,1776,1801]" pageId="23" pageNumber="400" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it differs by the paler coloration of the body, and sparser and smaller punctation of the head. From
<taxonomicName id="95884D05FFAA2E302135F8C67D84F890" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1325,1436,1813,1836]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="400" phylum="Arthropoda" rank="species" species="assingi" status="sp. nov.">
<emphasis id="60FCEA94FFAA2E302135F8C67D84F890" box="[1325,1436,1813,1836]" italics="true" pageId="23" pageNumber="400">A. assingi</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFAA2E30248FF8EB78D6F8ED" bold="true" box="[151,206,1848,1873]" pageId="23" pageNumber="400">
<taxonomicNameLabel id="7BCF57EFFFAA2E30248FF8EB78D6F8ED" box="[151,206,1848,1873]" pageId="23" pageNumber="400" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
it differs by the shape of anterior angles of the pronotum not protruded anteriad. From both species it differs by the details of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFAA2E3024DFF8537D09F85D" pageId="23" pageNumber="400" type="distribution">
<paragraph id="52373686FFAA2E3024DFF85379DDF801" blockId="23.[151,1436,1704,2017]" pageId="23" pageNumber="400">
<emphasis id="60FCEA94FFAA2E3024DFF8537978F825" bold="true" box="[199,352,1920,1945]" pageId="23" pageNumber="400">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="8D338824FFAA2E3027E8F8527C38F825" box="[1008,1056,1921,1945]" pageId="23" pageNumber="400">type</typeStatus>
locality (
<figureCitation id="CAB32A03FFAA2E302080F8537CE8F824" box="[1176,1264,1920,1945]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="23" pageNumber="400">Fig. 41</figureCitation>
) in the Chola Mountains in
<collectingRegion id="904CF864FFAA2E30252AF8777977F801" box="[306,367,1956,1981]" country="China" name="Xizang" pageId="23" pageNumber="400">Tibet</collectingRegion>
,
<collectingCountry id="2A9F7616FFAA2E302563F87779DAF801" box="[379,450,1956,1981]" name="China" pageId="23" pageNumber="400">China</collectingCountry>
.
</paragraph>
<paragraph id="52373686FFAA2E3024DFF81B7D09F85D" blockId="23.[151,1436,1704,2017]" box="[199,1297,1992,2017]" pageId="23" pageNumber="400">
<emphasis id="60FCEA94FFAA2E3024DFF81B7952F85D" bold="true" box="[199,330,1992,2017]" pageId="23" pageNumber="400">Bionomics.</emphasis>
All specimens were taken by sifting mixed forest floor litter at elevation
<quantity id="95709B63FFAA2E302098F81B7CCDF85C" box="[1152,1237,1992,2016]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="23" pageNumber="400" unit="m" value="4400.0">4400 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFA52E3F24DFFF4478CDFF69" pageId="24" pageNumber="401" type="etymology">
<paragraph id="52373686FFA52E3F24DFFF4478CDFF69" blockId="24.[151,1436,151,213]" pageId="24" pageNumber="401">
<emphasis id="60FCEA94FFA52E3F24DFFF447956FF0C" bold="true" box="[199,334,151,176]" pageId="24" pageNumber="401">Etymology.</emphasis>
The specific epithet is the latinized adjective derived from the area the specimens were collected from.
</paragraph>
</subSubSection>
</treatment>
</document>
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