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<document id="BB9BD8581BE16CA9F1E54C773A96BE5F" ID-DOI="10.1649/556" ID-ISSN="1938-4394" ID-Zenodo-Dep="10103651" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630163456920" checkinUser="tatiana" docAuthor="Milléo, Julianne &amp; de Almeida, Lúcia Massutti" docDate="2003" docId="C13F8795B828FFD6FF76AC6AFC14FC03" docLanguage="en" docName="556.pdf.imf" docOrigin="The Coleopterists Bulletin 57 (3)" docSource="http://dx.doi.org/10.1649/556" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Tiphysa Mulsant 1850" docType="treatment" docVersion="3" lastPageNumber="279" masterDocId="3D06FFEDB828FFD3FFE2A823FFD9FFCD" masterDocTitle="Revision of the Genus Tiphysa Mulsant (Coleoptera, Coccinellidae, Hyperaspidinae)" masterLastPageNumber="280" masterPageNumber="274" pageNumber="274" updateTime="1699626702877" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ABEF785131CDEC808152020DC7E212C0">Revision of the Genus Tiphysa Mulsant (Coleoptera, Coccinellidae, Hyperaspidinae)</mods:title>
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<mods:namePart id="9148253A8F975AF5672DF13C48244D77">Milléo, Julianne</mods:namePart>
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<mods:namePart id="A961630DCA181A3C33B837CEF35F5C59">de Almeida, Lúcia Massutti</mods:namePart>
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<mods:title id="048F787481186137773822624FABD879">The Coleopterists Bulletin</mods:title>
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<mods:date id="034E0066E91272A8CD80C2E0EF04E8F5">2003</mods:date>
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<mods:number id="F926F7325F94A4711144599B4ADFCE5E">2003-09-30</mods:number>
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<mods:number id="3EE6178D89FBE383E8FB893BFAB6139B">57</mods:number>
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<mods:number id="E334ADBA5656B248EAAD13594C3F32D6">3</mods:number>
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<mods:identifier id="623528116332DC55373D52888EEB1BA8" type="ISSN">1938-4394</mods:identifier>
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<treatment id="C13F8795B828FFD6FF76AC6AFC14FC03" LSID="urn:lsid:plazi:treatment:C13F8795B828FFD6FF76AC6AFC14FC03" httpUri="http://treatment.plazi.org/id/C13F8795B828FFD6FF76AC6AFC14FC03" lastPageId="5" lastPageNumber="279" pageId="0" pageNumber="274">
<subSubSection id="018C6508B828FFD3FF76AC6AFC08FBB5" pageId="0" pageNumber="274" type="nomenclature">
<paragraph id="49293683B828FFD3FF76AC6AFC09FB93" blockId="0.[148,977,1097,1490]" box="[148,976,1097,1118]" pageId="0" pageNumber="274">
<heading id="126181EFB828FFD3FF76AC6AFC09FB93" box="[148,976,1097,1118]" centered="true" fontSize="9" level="2" pageId="0" pageNumber="274" reason="2">
<treatmentCitationGroup id="698611ADB828FFD3FF76AC6AFC09FB93" box="[148,976,1097,1118]" pageId="0" pageNumber="274">
<treatmentCitation id="C8371092B828FFD3FF76AC6AFE44FB93" author="Mulsant" box="[148,413,1097,1118]" page="517" pageId="0" pageNumber="274" year="1850">
<taxonomicName id="8E964D00B828FFD3FF76AC6AFE44FB93" authority="Mulsant, 1850: 517" authorityName="Mulsant" authorityPageNumber="517" authorityYear="1850" box="[148,413,1097,1118]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B828FFD3FF76AC6AFF06FB93" box="[148,223,1097,1118]" italics="true" pageId="0" pageNumber="274">Tiphysa</emphasis>
<bibRefCitation id="2D074B72B828FFD3FF04AC6AFE44FB93" author="Mulsant" box="[230,413,1097,1118]" pageId="0" pageNumber="274" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850:517</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="C8371092B828FFD3FE45AC6AFD92FB93" author="Crotch" box="[423,587,1097,1118]" page="210" pageId="0" pageNumber="274" year="1874">
<bibRefCitation id="2D074B72B828FFD3FE45AC6AFD92FB93" author="Crotch" box="[423,587,1097,1118]" pageId="0" pageNumber="274" refId="ref1847" refString="Crotch, G. R. 1874. A revision of the coleopterous family Coccinellidae. London. 311 pp." type="book" year="1874">Crotch 1874:210</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C8371092B828FFD3FDB4AC6AFCDEFB93" author="Chapuis" box="[598,775,1097,1118]" page="228" pageId="0" pageNumber="274" year="1876">
<bibRefCitation id="2D074B72B828FFD3FDB4AC6AFCDEFB93" author="Chapuis" box="[598,775,1097,1118]" pageId="0" pageNumber="274" pagination="1 - 424" refId="ref1825" refString="Chapuis, F. 1876. Histoire naturelle des insectes. Genera des Coleopteres, Paris 12: 1 - 424." type="journal article" year="1876">Chapuis 1876:228</bibRefCitation>
</treatmentCitation>
, 232–233; Korschef-
</treatmentCitationGroup>
</heading>
</paragraph>
<paragraph id="49293683B828FFD3FF31AC40FC08FBB5" blockId="0.[148,977,1097,1490]" box="[211,977,1123,1144]" pageId="0" pageNumber="274">
sky 1931:208;
<bibRefCitation id="2D074B72B828FFD3FE8FAC40FD94FBB5" author="Blackwelder" box="[365,589,1123,1144]" pageId="0" pageNumber="274" pagination="343 - 550" refId="ref1781" refString="Blackwelder, R. E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bull. United States Natural History Museum Bulletin 185 (3): 343 - 550." type="journal article" year="1945">Blackwelder 1945:449</bibRefCitation>
;
<bibRefCitation id="2D074B72B828FFD3FDBFAC40FCDBFBB5" author="Gordon" box="[605,770,1123,1144]" pageId="0" pageNumber="274" pagination="1 - 46" refId="ref1952" refString="Gordon, R. D. 1987. A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology 3: 1 - 46." type="journal article" year="1987">Gordon 1987:26</bibRefCitation>
;
<bibRefCitation id="2D074B72B828FFD3FCF0AC40FC7BFBB5" author="Fursch" box="[786,930,1123,1144]" pageId="0" pageNumber="274" pagination="1 - 42" refId="ref1904" refString="Fursch, H. 1989. Newsletter for systematic research in Coccinellids. Universitat Passau 1 (1): 1 - 42." type="journal article" year="1989">Fürsch 1989:6</bibRefCitation>
, 20;
</paragraph>
</subSubSection>
<subSubSection id="018C6508B828FFD7FF31AC5DFEADFB29" lastPageId="4" lastPageNumber="278" pageId="0" pageNumber="274" type="description">
<paragraph id="49293683B828FFD3FF31AC5DFD81FB5E" blockId="0.[148,977,1097,1490]" box="[211,600,1150,1171]" pageId="0" pageNumber="274">
1990:4, 17, 62, 63;
<bibRefCitation id="2D074B72B828FFD3FE74AC5DFD8DFB5E" author="Duverger" box="[406,596,1150,1171]" pageId="0" pageNumber="274" pagination="143 - 157" refId="ref1868" refString="Duverger, C. 1989. Contribution a l'etude de Hyperaspinae 1 ere Note (Coleoptera, Coccinellidae). Bulletin de la Societe Linneenne de Bordeaux 17 (3): 143 - 157." type="journal article" year="1989">Duverger 1989:146</bibRefCitation>
.
</paragraph>
<paragraph id="49293683B828FFD3FF4EACBBFCD1FB63" blockId="0.[148,977,1097,1490]" box="[172,776,1176,1198]" pageId="0" pageNumber="274">
<emphasis id="7BE2EA91B828FFD3FF4EACBBFEEDFB60" bold="true" box="[172,308,1176,1197]" pageId="0" pageNumber="274">Type-species.</emphasis>
<taxonomicName id="8E964D00B828FFD3FEA2ACBAFDA1FB63" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[320,632,1177,1198]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="274" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="7BE2EA91B828FFD3FEA2ACBAFE3CFB63" box="[320,485,1177,1198]" italics="true" pageId="0" pageNumber="274">Tiphysa plumbea</emphasis>
<bibRefCitation id="2D074B72B828FFD3FE0FACBAFDA1FB63" author="Mulsant" box="[493,632,1177,1198]" pageId="0" pageNumber="274" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
by monotypy.
</paragraph>
<paragraph id="49293683B828FFD3FF4EAC90FC09FA1F" blockId="0.[148,977,1097,1490]" pageId="0" pageNumber="274">
<emphasis id="7BE2EA91B828FFD3FF4EAC90FE99FB05" bold="true" box="[172,320,1203,1224]" pageId="0" pageNumber="274">Redescription.</emphasis>
Body oval, convex (
<figureCitation id="D1AD2A06B828FFD3FDF7AC90FDAFFB05" box="[533,630,1203,1224]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Figs. 1, 2</figureCitation>
). Eyes finely faceted and glabrous, emarginate near antennal insertions (
<figureCitation id="D1AD2A06B828FFD3FDE7ACEDFD9DFB2E" box="[517,580,1230,1251]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Fig. 3</figureCitation>
). Antennal insertions visible. Antennae short, composed of eleven antennomeres; scape twice as long as wide; with slight lateral lobe; pedicel quadrangular; antennomeres 3–8 slender, 9–10 longer; 11 conical (
<figureCitation id="D1AD2A06B828FFD3FF79AD3EFF01FAFF" box="[155,216,1309,1330]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Fig. 4</figureCitation>
). Clypeus with apical margin arcuate and lateral margins rounded, short and projected (
<figureCitation id="D1AD2A06B828FFD3FF1BAD1BFEEAFA80" box="[249,307,1336,1357]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Fig. 3</figureCitation>
). Labrum rectangular in shape (
<figureCitation id="D1AD2A06B828FFD3FD85AD1BFD78FA80" box="[615,673,1336,1357]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Fig. 5</figureCitation>
). Mandible asymmetrical, large and robust, strongly sclerotized; with incisor teeth, apical and subapical, with pointed median molar (
<figureCitation id="D1AD2A06B828FFD3FEC8AD4EFEB1FA4F" box="[298,360,1389,1410]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Fig. 6</figureCitation>
). Maxillae sclerotized; palpomere 4 securiform with sides that diverge strongly (
<figureCitation id="D1AD2A06B828FFD3FED9ADABFEACFA50" box="[315,373,1416,1437]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="0" pageNumber="274">Fig. 7</figureCitation>
). Labium having anterior margin of ligula densely covered with small spines, having ventral median portion of ligula with several long bristles; each labial palp with basal articles shorter, and the last narrower in the distal region. Mentum
</paragraph>
<paragraph id="49293683B828FFD3FF76AE28FCAFF9EF" blockId="0.[148,886,1547,1570]" box="[148,886,1547,1570]" pageId="0" pageNumber="274">
<superScript id="BEE39BCBB828FFD3FF76AE28FF45F9DA" attach="left" box="[148,156,1547,1559]" fontSize="5" pageId="0" pageNumber="274">1</superScript>
Contribution n°. 2000, Departamento de Zoologia, Universidade Federal do Parana´.
</paragraph>
<paragraph id="49293683B828FFD3FDC0AE1CFD9BF99F" blockId="0.[546,578,1599,1618]" box="[546,578,1599,1618]" pageId="0" pageNumber="274">274</paragraph>
<caption id="1DE9660BB829FFD2FF27AD76FD3EFA4D" ID-DOI="http://doi.org/10.5281/zenodo.10103653" ID-Zenodo-Dep="10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="1" pageNumber="275" startId="1.[197,236,1365,1384]" targetBox="[176,1007,197,1344]" targetPageId="1">
<paragraph id="49293683B829FFD2FF27AD76FD3EFA4D" blockId="1.[175,1005,1365,1408]" pageId="1" pageNumber="275">
<emphasis id="7BE2EA91B829FFD2FF27AD76FEC5FAA5" bold="true" box="[197,284,1365,1384]" pageId="1" pageNumber="275">Figs. 1–8.</emphasis>
<taxonomicName id="8E964D00B829FFD2FED3AD76FD91FAA5" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[305,584,1365,1384]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="275" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="7BE2EA91B829FFD2FED3AD76FE1DFAA5" box="[305,452,1365,1384]" italics="true" pageId="1" pageNumber="275">Tiphysa plumbea</emphasis>
<bibRefCitation id="2D074B72B829FFD2FE28AD76FD91FAA5" author="Mulsant" box="[458,584,1365,1384]" pageId="1" pageNumber="275" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
.
<emphasis id="7BE2EA91B829FFD2FDB0AD76FDBAFAA5" bold="true" box="[594,611,1365,1384]" pageId="1" pageNumber="275">1)</emphasis>
Dorsal view;
<emphasis id="7BE2EA91B829FFD2FD3DAD76FD29FAA5" bold="true" box="[735,752,1365,1384]" pageId="1" pageNumber="275">2)</emphasis>
lateral view;
<emphasis id="7BE2EA91B829FFD2FC8AAD76FCA0FAA5" bold="true" box="[872,889,1365,1384]" pageId="1" pageNumber="275">3)</emphasis>
head, frontal view;
<emphasis id="7BE2EA91B829FFD2FF07AD4EFF2FFA4D" bold="true" box="[229,246,1389,1408]" pageId="1" pageNumber="275">4)</emphasis>
antenna;
<emphasis id="7BE2EA91B829FFD2FEAEAD4EFE84FA4D" bold="true" box="[332,349,1389,1408]" pageId="1" pageNumber="275">5)</emphasis>
labrum;
<emphasis id="7BE2EA91B829FFD2FE4FAD4EFE66FA4D" bold="true" box="[429,447,1389,1408]" pageId="1" pageNumber="275">6)</emphasis>
mandibles;
<emphasis id="7BE2EA91B829FFD2FDCBAD4EFDE3FA4D" bold="true" box="[553,570,1389,1408]" pageId="1" pageNumber="275">7)</emphasis>
maxilla; 8) labium.
</paragraph>
</caption>
<paragraph id="49293683B829FFD2FF4DAD9EFC35F9EF" blockId="1.[175,1004,1469,1570]" pageId="1" pageNumber="275">
cordiform, with rounded apical projections; prementum truncate, with long bristles in median region between palps (
<figureCitation id="D1AD2A06B829FFD2FE1AADFBFDE2FA20" box="[504,571,1496,1517]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="1" pageNumber="275">Fig. 8</figureCitation>
). Pronotum transverse; prosternal carinae convergent and finishing together at base of prosternum; prosternal process with nearly parallel sides (
<figureCitation id="D1AD2A06B829FFD2FEDEAE2EFEA1F9EF" box="[316,376,1549,1570]" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="1" pageNumber="275">Fig. 9</figureCitation>
). Metendosternite quadrangular, with straight lateral borders and
</paragraph>
<paragraph id="49293683B82AFFD1FF76A8A3FC16FA8D" blockId="2.[148,180,128,147]" box="[148,975,128,1344]" lastBlockId="2.[148,978,1301,1344]" pageId="2" pageNumber="276">
276
<emphasis id="7BE2EA91B82AFFD1FF76AD0EFF69FA8D" bold="true" box="[148,176,1325,1344]" pageId="2" pageNumber="276">11)</emphasis>
wing;
<emphasis id="7BE2EA91B82AFFD1FF0CAD0EFED3FA8D" bold="true" box="[238,266,1325,1344]" pageId="2" pageNumber="276">12)</emphasis>
epipleuron;
<emphasis id="7BE2EA91B82AFFD1FE9AAD0EFE4DFA8D" bold="true" box="[376,404,1325,1344]" pageId="2" pageNumber="276">13)</emphasis>
anterior leg;
<emphasis id="7BE2EA91B82AFFD1FDE8AD0EFDFFFA8D" bold="true" box="[522,550,1325,1344]" pageId="2" pageNumber="276">14)</emphasis>
tarsus;
<emphasis id="7BE2EA91B82AFFD1FD89AD0EFD5EFA8D" bold="true" box="[619,647,1325,1344]" pageId="2" pageNumber="276">15)</emphasis>
abdomen male;
<emphasis id="7BE2EA91B82AFFD1FCFAAD0EFCEDFA8D" bold="true" box="[792,820,1325,1344]" pageId="2" pageNumber="276">16)</emphasis>
abdomen female.
</paragraph>
<caption id="1DE9660BB82AFFD1FF4BAD36FC0BFAE5" ID-DOI="http://doi.org/10.5281/zenodo.10103655" ID-Zenodo-Dep="10103655" box="[169,978,1301,1320]" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="2" pageNumber="276" startId="2.[169,208,1301,1320]" targetBox="[149,978,196,1279]" targetPageId="2">
<paragraph id="49293683B82AFFD1FF4BAD36FC0BFAE5" blockId="2.[148,978,1301,1344]" box="[169,978,1301,1320]" pageId="2" pageNumber="276">
<emphasis id="7BE2EA91B82AFFD1FF4BAD36FED6FAE5" bold="true" box="[169,271,1301,1320]" pageId="2" pageNumber="276">Figs. 9–16.</emphasis>
<taxonomicName id="8E964D00B82AFFD1FEC1AD36FD9FFAE5" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[291,582,1301,1320]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="276" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="7BE2EA91B82AFFD1FEC1AD36FE63FAE5" box="[291,442,1301,1320]" italics="true" pageId="2" pageNumber="276">Tiphysa plumbea</emphasis>
<bibRefCitation id="2D074B72B82AFFD1FE26AD36FD9FFAE5" author="Mulsant" box="[452,582,1301,1320]" pageId="2" pageNumber="276" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
.
<emphasis id="7BE2EA91B82AFFD1FDB1AD36FDBCFAE5" bold="true" box="[595,613,1301,1320]" pageId="2" pageNumber="276">9)</emphasis>
Prosternal process;
<emphasis id="7BE2EA91B82AFFD1FCFCAD36FCE3FAE5" bold="true" box="[798,826,1301,1320]" pageId="2" pageNumber="276">10)</emphasis>
metendosternite;
</paragraph>
</caption>
<paragraph id="49293683B82AFFD7FF76AD4DFEADFB29" blockId="2.[148,977,1390,1570]" lastBlockId="4.[148,977,1124,1571]" lastPageId="4" lastPageNumber="278" pageId="2" pageNumber="276">
very shallow V-shaped anterior border, with anterior arms nearly straight apically (
<figureCitation id="D1AD2A06B82AFFD1FC48AD4DFF77FA50" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="2" pageNumber="276">Fig. 10</figureCitation>
). Wings with reduced alar venation; first and second anal veins and Ea, Eb and P evident (
<figureCitation id="D1AD2A06B82AFFD1FF05AD80FEF6FA75" box="[231,303,1443,1464]" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="2" pageNumber="276">Fig. 11</figureCitation>
). Elytra oval with raised humeral callosity and with lateral margin not flattened (
<figureCitation id="D1AD2A06B82AFFD1FF10AD9DFE97FA1E" box="[242,334,1470,1491]" captionStart="Figs" captionStartId="1.[197,236,1365,1384]" captionTargetBox="[176,1007,197,1344]" captionTargetId="figure-45@1.[175,1007,194,1344]" captionTargetPageId="1" captionText="Figs. 1–8. Tiphysa plumbea Mulsant, 1850. 1) Dorsal view; 2) lateral view; 3) head, frontal view; 4) antenna; 5) labrum; 6) mandibles; 7) maxilla; 8) labium." figureDoi="http://doi.org/10.5281/zenodo.10103653" httpUri="https://zenodo.org/record/10103653/files/figure.png" pageId="2" pageNumber="276">Figs. 1, 2</figureCitation>
); epipleuron with strong excavation to receive apex of medium and posterior femur (
<figureCitation id="D1AD2A06B82AFFD1FED6ADFBFEA2FA20" box="[308,379,1496,1517]" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="2" pageNumber="276">Fig. 12</figureCitation>
). Femur robust and deeply excavated to receive tibia. Anterior tibia thin and strongly excavated to receive tarsus (
<figureCitation id="D1AD2A06B82AFFD1FD55ADD0FCDFF9C5" box="[695,774,1523,1544]" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="2" pageNumber="276">Fig. 13</figureCitation>
). Second tarsomere triangular in ventral view; tarsal claw appendiculate (
<figureCitation id="D1AD2A06B82AFFD1FD2CAE2EFCC7F9EF" box="[718,798,1549,1570]" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="2" pageNumber="276">Fig. 14</figureCitation>
). Abdomen with 278 incomplete post-coxal lines on the first visible sternite; seven visible segments present in the male, six in the female (
<figureCitation id="D1AD2A06B82CFFD7FE56AC5CFDF1FB59" box="[436,552,1151,1172]" captionStart="Figs" captionStartId="2.[169,208,1301,1320]" captionTargetBox="[149,978,196,1279]" captionTargetId="figure-93@2.[148,980,194,1281]" captionTargetPageId="2" captionText="Figs. 9–16. Tiphysa plumbea Mulsant, 1850. 9) Prosternal process; 10) metendosternite;" figureDoi="http://doi.org/10.5281/zenodo.10103655" httpUri="https://zenodo.org/record/10103655/files/figure.png" pageId="4" pageNumber="278">Figs. 15, 16</figureCitation>
). Male genitalia: Median lobe asymmetrical, parameres longer than median lobe. Sipho strongly sclerotized, curved. Female genitalia: Spermatheca alantoid; infundibulum present; coxites wide with slender bristles, without stylus.
</paragraph>
</subSubSection>
<caption id="1DE9660BB82BFFD0FF27ADB1FC34FA68" ID-DOI="http://doi.org/10.5281/zenodo.10103659" ID-Zenodo-Dep="10103659" box="[197,1005,1426,1445]" httpUri="https://zenodo.org/record/10103659/files/figure.png" pageId="3" pageNumber="277" startId="3.[197,236,1426,1445]" targetBox="[184,999,193,1404]" targetPageId="3">
<paragraph id="49293683B82BFFD0FF27ADB1FC34FA68" blockId="3.[197,1005,1426,1445]" box="[197,1005,1426,1445]" pageId="3" pageNumber="277">
<emphasis id="7BE2EA91B82BFFD0FF27ADB1FEE8FA68" bold="true" box="[197,305,1426,1445]" pageId="3" pageNumber="277">Figs. 17–19.</emphasis>
<taxonomicName id="8E964D00B82BFFD0FEA4ADB1FD87FA68" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[326,606,1426,1445]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="277" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="7BE2EA91B82BFFD0FEA4ADB1FE00FA68" box="[326,473,1426,1445]" italics="true" pageId="3" pageNumber="277">Tiphysa plumbea</emphasis>
<bibRefCitation id="2D074B72B82BFFD0FE02ADB1FD87FA68" author="Mulsant" box="[480,606,1426,1445]" pageId="3" pageNumber="277" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
.
<emphasis id="7BE2EA91B82BFFD0FD8AADB1FD5DFA68" bold="true" box="[616,644,1426,1445]" pageId="3" pageNumber="277">17)</emphasis>
Tegmen;
<emphasis id="7BE2EA91B82BFFD0FD3FADB1FD20FA68" bold="true" box="[733,761,1426,1445]" pageId="3" pageNumber="277">18)</emphasis>
sipho;
<emphasis id="7BE2EA91B82BFFD0FCD9ADB1FC8EFA68" bold="true" box="[827,855,1426,1445]" pageId="3" pageNumber="277">19)</emphasis>
female genitalia.
</paragraph>
</caption>
<caption id="1DE9660BB82CFFD7FF4BAC30FD5FFBEB" box="[169,646,1043,1062]" pageId="4" pageNumber="278" startId="4.[169,200,1043,1062]">
<paragraph id="49293683B82CFFD7FF4BAC30FD5FFBEB" blockId="4.[169,646,1043,1062]" box="[169,646,1043,1062]" pageId="4" pageNumber="278">
<emphasis id="7BE2EA91B82CFFD7FF4BAC30FF34FBEB" bold="true" box="[169,237,1043,1062]" pageId="4" pageNumber="278">Fig. 20.</emphasis>
<taxonomicName id="8E964D00B82CFFD7FEE1AC30FE37FBEB" authority="Crotch, 1874" authorityName="Crotch" authorityYear="1874" box="[259,494,1043,1062]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="7BE2EA91B82CFFD7FEE1AC30FEADFBEB" box="[259,372,1043,1062]" italics="true" pageId="4" pageNumber="278">Tiphysa egae</emphasis>
<bibRefCitation id="2D074B72B82CFFD7FE99AC30FE37FBEB" author="Crotch" box="[379,494,1043,1062]" pageId="4" pageNumber="278" refId="ref1847" refString="Crotch, G. R. 1874. A revision of the coleopterous family Coccinellidae. London. 311 pp." type="book" year="1874">Crotch, 1874</bibRefCitation>
</taxonomicName>
, female genitalia.
</paragraph>
</caption>
<subSubSection id="018C6508B82CFFD7FF4EACCAFF33FAFE" pageId="4" pageNumber="278" type="materials_examined">
<paragraph id="49293683B82CFFD7FF4EACCAFF33FAFE" blockId="4.[148,977,1124,1571]" pageId="4" pageNumber="278">
<emphasis id="7BE2EA91B82CFFD7FF4EACCAFE9EFB33" bold="true" box="[172,327,1257,1278]" pageId="4" pageNumber="278">Type Material.</emphasis>
<materialsCitation id="F9FE3CDEB82CFFD7FEB6ACCAFF3FFAFE" country="United Kingdom" county="Cambridge" location="University Museum of Zoology" municipality="Cambridge University" pageId="4" pageNumber="278" specimenCount="2" typeStatus="lectotype">
The
<typeStatus id="962D8821B82CFFD7FE63ACCAFE01FB33" box="[385,472,1257,1278]" pageId="4" pageNumber="278" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="962D8821B82CFFD7FDEEACCAFD84FB33" box="[524,605,1257,1278]" pageId="4" pageNumber="278" type="paratype">paratype</typeStatus>
(
<bibRefCitation id="2D074B72B82CFFD7FD8EACCAFD29FB33" author="Gordon" box="[620,752,1257,1278]" pageId="4" pageNumber="278" pagination="1 - 46" refId="ref1952" refString="Gordon, R. D. 1987. A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology 3: 1 - 46." type="journal article" year="1987">Gordon 1987</bibRefCitation>
) were studied, and are located at the
<location id="4C496058B82CFFD7FECAAD27FDB1FAD4" LSID="urn:lsid:plazi:treatment:C13F8795B828FFD6FF76AC6AFC14FC03:4C496058B82CFFD7FECAAD27FDB1FAD4" box="[296,616,1284,1305]" country="United Kingdom" county="Cambridge" municipality="Cambridge University" name="University Museum of Zoology" pageId="4" pageNumber="278">University Museum of Zoology</location>
,
<collectingMunicipality id="A94DACF9B82CFFD7FD95AD27FC8BFAD4" box="[631,850,1284,1305]" pageId="4" pageNumber="278">Cambridge University</collectingMunicipality>
,
<collectingCounty id="A0484E0FB82CFFD7FC83AD27FC14FAD4" box="[865,973,1284,1305]" pageId="4" pageNumber="278">Cambridge</collectingCounty>
,
<collectingCountry id="31817613B82CFFD7FF76AD3DFF3FFAFE" box="[148,230,1310,1331]" name="United Kingdom" pageId="4" pageNumber="278">England</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="018C6508B82CFFD6FF4EAD1AFC14FC03" lastPageId="5" lastPageNumber="279" pageId="4" pageNumber="278" type="discussion">
<paragraph id="49293683B82CFFD7FF4EAD1AFC08F9EF" blockId="4.[148,977,1124,1571]" pageId="4" pageNumber="278">
<emphasis id="7BE2EA91B82CFFD7FF4EAD1AFED7FA83" bold="true" box="[172,270,1337,1358]" pageId="4" pageNumber="278">Remarks.</emphasis>
The genus
<taxonomicName id="8E964D00B82CFFD7FE4BAD1AFE2DFA83" authorityName="Mulsant" authorityYear="1850" box="[425,500,1337,1358]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FE4BAD1AFE2DFA83" box="[425,500,1337,1358]" italics="true" pageId="4" pageNumber="278">Tiphysa</emphasis>
</taxonomicName>
was included in the tribe
<taxonomicName id="8E964D00B82CFFD7FCBFAD1AFC08FA83" authorityName="Mulsant" authorityYear="1846" box="[861,977,1337,1358]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="tribe" tribe="Hyperaspini">Hyperaspini</taxonomicName>
(
<emphasis id="7BE2EA91B82CFFD7FF7BAD77FF72FAA5" bold="true" box="[153,171,1364,1384]" italics="true" pageId="4" pageNumber="278">¼</emphasis>
<taxonomicName id="8E964D00B82CFFD7FF4FAD70FEECFAA5" authorityName="Mulsant" authorityYear="1846" box="[173,309,1363,1384]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
) by
<bibRefCitation id="2D074B72B82CFFD7FE92AD70FDCEFAA5" author="Duverger" box="[368,535,1363,1384]" pageId="4" pageNumber="278" pagination="143 - 157" refId="ref1868" refString="Duverger, C. 1989. Contribution a l'etude de Hyperaspinae 1 ere Note (Coleoptera, Coccinellidae). Bulletin de la Societe Linneenne de Bordeaux 17 (3): 143 - 157." type="journal article" year="1989">Duverger (1989)</bibRefCitation>
because of the rounded spermatheca and absence of infundibulum, but it was a mistake. We transferred here
<taxonomicName id="8E964D00B82CFFD7FC84AD4DFC68FA4E" authorityName="Mulsant" authorityYear="1850" box="[870,945,1390,1411]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FC84AD4DFC68FA4E" box="[870,945,1390,1411]" italics="true" pageId="4" pageNumber="278">Tiphysa</emphasis>
</taxonomicName>
to Brachiacanthini, together with
<taxonomicName id="8E964D00B82CFFD7FE30ADAAFD13FA53" authority="Chevrolat" authorityName="Chevrolat" authorityYear="1836" box="[466,714,1417,1438]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FE30ADAAFD86FA53" box="[466,607,1417,1438]" italics="true" pageId="4" pageNumber="278">Brachiacantha</emphasis>
Chevrolat
</taxonomicName>
<emphasis id="7BE2EA91B82CFFD7FD3BADAAFD32FA53" box="[729,747,1417,1438]" italics="true" pageId="4" pageNumber="278">in</emphasis>
Dejean, 1837,
<taxonomicName id="8E964D00B82CFFD7FC77ADAAFEFBFA75" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" class="Insecta" family="Coccinellidae" genus="Hinda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FC77ADAAFC09FA53" box="[917,976,1417,1438]" italics="true" pageId="4" pageNumber="278">Hinda</emphasis>
<bibRefCitation id="2D074B72B82CFFD7FF76AD80FEFBFA75" author="Mulsant" box="[148,290,1443,1464]" pageId="4" pageNumber="278" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8E964D00B82CFFD7FEBAAD80FDFAFA75" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[344,547,1443,1464]" class="Insecta" family="Coccinellidae" genus="Cyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FEBAAD80FE51FA75" box="[344,392,1443,1464]" italics="true" pageId="4" pageNumber="278">Cyra</emphasis>
<bibRefCitation id="2D074B72B82CFFD7FE70AD80FDFAFA75" author="Mulsant" box="[402,547,1443,1464]" pageId="4" pageNumber="278" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="8E964D00B82CFFD7FDD2AD80FDA2FA75" authorityName="Mulsant" authorityYear="1850" box="[560,635,1443,1464]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FDD2AD80FDA2FA75" box="[560,635,1443,1464]" italics="true" pageId="4" pageNumber="278">Tiphysa</emphasis>
</taxonomicName>
together with the other genera of Brachiacanthini, has integument glabrous; eyes emarginate near antennal insertions; eleven antennomeres; 4° maxillary palpomere securiform; 2° tarsomere triangular; six abdominal segments visible on female and seven in male; spermatheca alantoid and infundibulum present. It differs from
<taxonomicName id="8E964D00B82CFFD7FDE0AE2DFD56F9EE" authorityName="Chevrolat" authorityYear="1836" box="[514,655,1550,1571]" class="Insecta" family="Coccinellidae" genus="Brachiacantha" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82CFFD7FDE0AE2DFD56F9EE" box="[514,655,1550,1571]" italics="true" pageId="4" pageNumber="278">Brachiacantha</emphasis>
</taxonomicName>
by presence of a sharp spine on
</paragraph>
<caption id="1DE9660BB82DFFD6FF27AAEBFC34FD16" ID-DOI="http://doi.org/10.5281/zenodo.10103661" ID-Zenodo-Dep="10103661" box="[197,1005,712,731]" httpUri="https://zenodo.org/record/10103661/files/figure.png" pageId="5" pageNumber="279" startId="5.[197,236,712,731]" targetBox="[175,1007,199,690]" targetPageId="5">
<paragraph id="49293683B82DFFD6FF27AAEBFC34FD16" blockId="5.[175,1005,712,755]" box="[197,1005,712,731]" pageId="5" pageNumber="279">
<emphasis id="7BE2EA91B82DFFD6FF27AAEBFEE8FD16" bold="true" box="[197,305,712,731]" pageId="5" pageNumber="279">Figs. 21–22.</emphasis>
Dorsal habitus.
<emphasis id="7BE2EA91B82DFFD6FE2CAAEBFE33FD16" bold="true" box="[462,490,712,731]" pageId="5" pageNumber="279">21)</emphasis>
<taxonomicName id="8E964D00B82DFFD6FE13AAEBFCD1FD16" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[497,776,712,731]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="plumbea">
<emphasis id="7BE2EA91B82DFFD6FE13AAEBFD5DFD16" box="[497,644,712,731]" italics="true" pageId="5" pageNumber="279">Tiphysa plumbea</emphasis>
<bibRefCitation id="2D074B72B82DFFD6FD68AAEBFCD1FD16" author="Mulsant" box="[650,776,712,731]" pageId="5" pageNumber="279" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
;
<emphasis id="7BE2EA91B82DFFD6FCF0AAEBFCF6FD16" bold="true" box="[786,815,712,731]" pageId="5" pageNumber="279">22)</emphasis>
<taxonomicName id="8E964D00B82DFFD6FCD4AAEBFC33FD16" authority="Crotch" authorityName="Crotch" authorityYear="1874" box="[822,1002,712,731]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="species" species="egae">
<emphasis id="7BE2EA91B82DFFD6FCD4AAEBFC7EFD16" box="[822,935,712,731]" italics="true" pageId="5" pageNumber="279">Tiphysa egae</emphasis>
Crotch
</taxonomicName>
,
</paragraph>
</caption>
<paragraph id="49293683B82DFFD6FF4DAAC3FE9BFD3E" blockId="5.[175,1005,712,755]" box="[175,322,736,755]" pageId="5" pageNumber="279">
1874 (
<typeStatus id="962D8821B82DFFD6FF05AAC3FEE0FD3E" box="[231,313,736,755]" pageId="5" pageNumber="279" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="49293683B82DFFD6FF4DAB17FC14FC03" blockId="5.[175,1004,820,974]" pageId="5" pageNumber="279">
anterior tibia; and from
<taxonomicName id="8E964D00B82DFFD6FE73AB17FE15FC84" authorityName="Mulsant" authorityYear="1850" box="[401,460,820,841]" class="Insecta" family="Coccinellidae" genus="Hinda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82DFFD6FE73AB17FE15FC84" box="[401,460,820,841]" italics="true" pageId="5" pageNumber="279">Hinda</emphasis>
</taxonomicName>
by the strongly serrated anterior border of tibia (Almeida and Milléo 1999) and from
<taxonomicName id="8E964D00B82DFFD6FE21AB6CFE2AFCA9" authorityName="Mulsant" authorityYear="1850" box="[451,499,847,868]" class="Insecta" family="Coccinellidae" genus="Cyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82DFFD6FE21AB6CFE2AFCA9" box="[451,499,847,868]" italics="true" pageId="5" pageNumber="279">Cyra</emphasis>
</taxonomicName>
by the thin unornamented tibia without ornamentation (
<bibRefCitation id="2D074B72B82DFFD6FF06AB4AFE47FCB3" author="Milleo" box="[228,414,873,894]" pageId="5" pageNumber="279" pagination="391 - 405" refId="ref2002" refString="Milleo, J., L. M. Almeida, and I. M. M. Lima. 1997. Contribuicao ao estudo de Brachiacanthadini (Coleoptera, Coccinellidae, Hyperaspinae). Revista Brasileira de Zoologia 14 (2): 391 - 405." type="journal article" year="1997">
Milléo
<emphasis id="7BE2EA91B82DFFD6FECCAB4AFEBBFCB3" box="[302,354,873,894]" italics="true" pageId="5" pageNumber="279">et al.</emphasis>
1997
</bibRefCitation>
).
<taxonomicName id="8E964D00B82DFFD6FE50AB4AFE24FCB3" authorityName="Mulsant" authorityYear="1850" box="[434,509,873,894]" class="Insecta" family="Coccinellidae" genus="Tiphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82DFFD6FE50AB4AFE24FCB3" box="[434,509,873,894]" italics="true" pageId="5" pageNumber="279">Tiphysa</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="8E964D00B82DFFD6FD65AB4AFCADFCB3" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[647,884,873,894]" class="Insecta" family="Coccinellidae" genus="Thalassa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82DFFD6FD65AB4AFD04FCB3" box="[647,733,873,894]" italics="true" pageId="5" pageNumber="279">Thalassa</emphasis>
<bibRefCitation id="2D074B72B82DFFD6FD05AB4AFCADFCB3" author="Mulsant" box="[743,884,873,894]" pageId="5" pageNumber="279" pagination="1 - 1104" refId="ref2050" refString="Mulsant, M. E. 1850. Species de Coleopteres trimeres securipalpes. Annales Sciences Physique et Naturelle, Lyon 2 (2): 1 - 1104." type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
of the tribe
<taxonomicName id="8E964D00B82DFFD6FF4DABA7FEE1FC54" authorityName="Mulsant" authorityYear="1846" box="[175,312,900,921]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
, in both color and size, but differs from
<taxonomicName id="8E964D00B82DFFD6FD2AABA7FCC7FC54" authorityName="Mulsant" authorityYear="1850" box="[712,798,900,921]" class="Insecta" family="Coccinellidae" genus="Thalassa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="7BE2EA91B82DFFD6FD2AABA7FCC7FC54" box="[712,798,900,921]" italics="true" pageId="5" pageNumber="279">Thalassa</emphasis>
</taxonomicName>
in the more rounded body; the absence of emargination of the eyes; shape of anterior tibia; male and female with six abdominal segments; spermatheca globular and absence of infundibulum.
</paragraph>
</subSubSection>
</treatment>
</document>
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