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<mods:title id="63EAC570CA1CCB2985C0509525FDA859">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<treatment id="6C3DA91C51481822FF0EFB34EE2A1D58" ID-DOI="http://doi.org/10.5281/zenodo.4688913" ID-GBIF-Taxon="159762439" ID-Zenodo-Dep="4688913" LSID="urn:lsid:plazi:treatment:6C3DA91C51481822FF0EFB34EE2A1D58" httpUri="http://treatment.plazi.org/id/6C3DA91C51481822FF0EFB34EE2A1D58" lastPageId="116" lastPageNumber="116" pageId="115" pageNumber="116">
<subSubSection id="AC8E4B8151481825FF0EFB34EEF31881" box="[151,459,1195,1222]" pageId="115" pageNumber="116" type="nomenclature">
<paragraph id="E42B180A51481825FF0EFB34EEF31881" blockId="115.[151,459,1195,1258]" box="[151,459,1195,1222]" pageId="115" pageNumber="116">
<heading id="BF63AF6651481825FF0EFB34EEF31881" bold="true" box="[151,459,1195,1222]" fontSize="11" level="1" pageId="115" pageNumber="116" reason="1">
<taxonomicName id="2394638951481825FF0EFB34EEF31881" ID-CoL="6C4W" authority="Ballantyne 2013" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[151,459,1195,1222]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851481825FF0EFB34EEF31881" bold="true" box="[151,459,1195,1222]" pageId="115" pageNumber="116">
<emphasis id="D6E0C41851481825FF0EFB34EFC21882" bold="true" box="[151,250,1195,1221]" italics="true" pageId="115" pageNumber="116">Pacifica</emphasis>
<bibRefCitation id="800565FB51481825FE9BFB33EEF31881" author="Ballantyne" box="[258,459,1196,1222]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 48" refId="ref114209" refString="Ballantyne, L., Fu, X. - H., Lambkin, C., Jeng, M. - L., Faust, L., Wijekoon, W. M. C. D., Li, D. - Q. &amp; Zhu, T. - F. (2013) Studies on South-east Asian fireflies: Abscondita, a new genus with details of life history, flashing patterns and behaviour of Abs. chinensis (L.) and Abs. terminalis (Olivier) (Coleoptera: Lampyridae: Luciolinae). Zootaxa, 3721 (1), 1 - 48. https: // doi. org / 10.11646 / zootaxa. 3721.1.1" type="journal article" year="2013">Ballantyne 2013</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="AC8E4B8151481825FF0EFB4FEE2E18AD" box="[151,278,1232,1258]" pageId="115" pageNumber="116" type="description">
<paragraph id="E42B180A51481825FF0EFB4FEE2E18AD" blockId="115.[151,459,1195,1258]" box="[151,278,1232,1258]" pageId="115" pageNumber="116">
<figureCitation id="7CAF048F51481825FF0EFB4FEE2E18AD" box="[151,278,1232,1258]" captionStart="FIGURES 22−29" captionStartId="14.[151,264,1369,1394]" captionTargetBox="[151,1435,212,1346]" captionTargetId="figure@14.[151,1435,212,1349]" captionTargetPageId="14" captionText="FIGURES 22−29. Key to genera characters 4. 22, 23, 26−28 Pacifica salomonis male; 24, 29 Pac. limbatipennis (Guadalcanal BPBM); 25 Pac. limbatifusca paratype male. 22, 24, 25 dorsal; 23 ventral; 26 aedeagus ventral; 27, 28 tergite 8 ventral and from right side showing ventral surface; upper arrows in 27, 28 indicate projections from anterior tergal arms; lower arrow in 27 indicates oblique ridge on ventral surface; 29 aedeagal sheath ventral (upper arrow indicates beginning of emargination, lower arrow indicates emargination." figureDoi="http://doi.org/10.5281/zenodo.3508108" httpUri="https://zenodo.org/record/3508108/files/figure.png" pageId="115" pageNumber="116">Figs 22−29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151481825FF0EFA8BEBC9196B" box="[151,1265,1299,1324]" pageId="115" pageNumber="116" type="reference_group">
<paragraph id="E42B180A51481825FF0EFA8BEBC9196B" blockId="115.[151,1265,1299,1355]" box="[151,1265,1299,1324]" pageId="115" pageNumber="116">
<treatmentCitationGroup id="C4843F2451481825FF0EFA8BEBC9196B" box="[151,1265,1299,1324]" pageId="115" pageNumber="116">
<taxonomicName id="2394638951481825FF0EFA8BEDF4196B" ID-CoL="6C4W" authority="Ballantyne in Ballantyne &amp; Lambkin 2013: 94" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityPageNumber="94" authorityYear="2013" box="[151,716,1299,1324]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851481825FF0EFA8BEFD3196B" box="[151,235,1300,1324]" italics="true" pageId="115" pageNumber="116">Pacifica</emphasis>
Ballantyne in
<treatmentCitation id="65353E1B51481825FE19FA8CEDF4196B" author="Ballantyne, L. A. &amp; Lambkin, C. L." box="[384,716,1299,1324]" httpUri="http://treatment.plazi.org/id/E46FF2165E5DC52CFF6FBA54FA4AE87F" page="94" pageId="115" pageNumber="116" year="2013">
<bibRefCitation id="800565FB51481825FE19FA8CEDF4196B" author="Ballantyne &amp; Lambkin" box="[384,716,1299,1324]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin 2013: 94</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
<bibRefCitation id="800565FB51481825FD4FFA8CECC1196B" author="Ballantyne &amp; Lambkin" box="[726,1017,1299,1324]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009</bibRefCitation>
figs 329–376, 397–440.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151481825FF0EFAACEDBB190C" box="[151,643,1330,1355]" pageId="115" pageNumber="116" type="type_taxon">
<paragraph id="E42B180A51481825FF0EFAACEDBB190C" blockId="115.[151,1265,1299,1355]" box="[151,643,1330,1355]" pageId="115" pageNumber="116">
<emphasis id="D6E0C41851481825FF0EFAACEE26190C" bold="true" box="[151,286,1330,1355]" pageId="115" pageNumber="116">
<typeStatus id="3B2FA6A851481825FF0EFAACEFF4190C" box="[151,204,1331,1355]" pageId="115" pageNumber="116">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="2394638951481825FEB3FAACED47190C" ID-CoL="4QNJ3" authority="(Olivier)" baseAuthorityName="Olivier" box="[298,639,1330,1355]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="salomonis">
<emphasis id="D6E0C41851481825FEB3FAACED25190C" box="[298,541,1331,1355]" italics="true" pageId="115" pageNumber="116">Pygatyphella salomonis</emphasis>
(Olivier)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B8151481822FF0EFAE7EE2A1D58" lastPageId="116" lastPageNumber="117" pageId="115" pageNumber="116" type="diagnosis">
<paragraph id="E42B180A51481822FF0EFAE7EE2A1D58" blockId="115.[151,1437,1400,2039]" lastBlockId="116.[151,1436,153,287]" lastPageId="116" lastPageNumber="117" pageId="115" pageNumber="116">
<emphasis id="D6E0C41851481825FF0EFAE7EE2A19D4" bold="true" box="[151,274,1400,1427]" pageId="115" pageNumber="116">Diagnosis.</emphasis>
Repeated with modifications from
<bibRefCitation id="800565FB51481825FD3AFAE6ECC519D4" author="Ballantyne &amp; Lambkin" box="[675,1021,1401,1427]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin (2013)</bibRefCitation>
. A genus of five species restricted to the
<collectingCountry id="9C83589A51481825FF59FA02EE4619F0" box="[192,382,1437,1463]" name="Solomon Islands" pageId="115" pageNumber="116">Solomon Islands</collectingCountry>
, with most having orange pronotum and black elytra which may be pale margined.
<bibRefCitation id="800565FB51481825FABAFA02EE52199C" author="Ballantyne &amp; Lambkin" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin (2009)</bibRefCitation>
distinguished a group of five species within
<taxonomicName id="2394638951481825FCC4FA5EEBBB199B" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[861,1155,1473,1500]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851481825FCC4FA5EECC9199C" box="[861,1009,1473,1499]" italics="true" pageId="115" pageNumber="116">Pygatyphella</emphasis>
(Ballantyne)
</taxonomicName>
as “
<taxonomicName id="2394638951481825FB2DFA5EEA70199C" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[1204,1352,1473,1499]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851481825FB2DFA5EEA70199C" box="[1204,1352,1473,1499]" italics="true" pageId="115" pageNumber="116">Pygatyphella</emphasis>
</taxonomicName>
B” and this difference was formalised in
<bibRefCitation id="800565FB51481825FD91FA7AEC6119B8" author="Ballantyne &amp; Lambkin" box="[520,857,1509,1535]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 162" refId="ref114597" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2013) Systematics and phylogenetics of Indo-Pacific Luciolinae fireflies (Coleoptera: Lampyridae) and the description of new genera. Zootaxa, 3653 (1), 1 - 162. https: // doi. org / 10.11646 / zootaxa. 3653.1.1" type="journal article" year="2013">Ballantyne &amp; Lambkin (2013)</bibRefCitation>
with the erection of a new genus
<emphasis id="D6E0C41851481825FB52FA7AEA0919B8" box="[1227,1329,1509,1535]" italics="true" pageId="115" pageNumber="116">
<taxonomicName id="2394638951481825FB52FA7AEA1419B8" authorityName="Ballantyne in Ballantyne &amp; Lambkin" authorityYear="2013" box="[1227,1324,1509,1535]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">Pacifica</taxonomicName>
.
</emphasis>
It is most closely related to
<taxonomicName id="2394638951481825FEF9F996EDB01A63" authority="(Ballantyne)" authorityName="Ballantyne" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[352,648,1545,1572]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851481825FEF9F996EECC1A64" box="[352,500,1545,1571]" italics="true" pageId="115" pageNumber="116">Pygatyphella</emphasis>
(Ballantyne)
</taxonomicName>
from which it differs as follows: four species have dorsal colouration of orange pronotum (sometimes with median dark mark), and dark brown elytra which may be pale margined (most
<taxonomicName id="2394638951481825FF0EF9CEEE131A2C" baseAuthorityName="Ballantyne" baseAuthorityYear="1968" box="[151,299,1617,1643]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C41851481825FF0EF9CEEE131A2C" box="[151,299,1617,1643]" italics="true" pageId="115" pageNumber="116">Pygatyphella</emphasis>
</taxonomicName>
spp. have a pattern resembling bird droppings, with light brownish dorsum and brown markings on pronotum, base of elytra at the sides of the MS, and irregularly at or near the elytral apices); only
<taxonomicName id="2394638951481825FA99F9E9EAA41AC8" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[1280,1436,1653,1679]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="russellia">
<emphasis id="D6E0C41851481825FA99F9E9EAA41AC8" box="[1280,1436,1653,1679]" italics="true" pageId="115" pageNumber="116">Pac. russellia</emphasis>
</taxonomicName>
is completely pale dorsally; pronotum never with angulate convergence along lateral margins, or small depression present just anterior to corner; with irregularities at rounded obtuse posterolateral corners, not projecting strongly if at all beyond median posterior margin; no interstitial lines well-defined; elytral margins convex-sided; posterior area of
<date id="902A3ECA51481825FF2CF89AEFEF1B58" box="[181,215,1797,1823]" pageId="115" pageNumber="116">V7</date>
never reaching into LO (LO not medially emarginated along its posterior margin); apex of MPP rounded, or squarely or obliquely truncate; MPP with dorsal longitudinal ridge; dorsal surface of the posterior area of the MPP faced with cuticle that is not attached to the ventral surface of
<date id="902A3ECA51481825FCDBF8D2EC5C1B20" box="[834,868,1869,1895]" pageId="115" pageNumber="116">V7</date>
and ends just behind the area of muscle attachment posterior to the LO; the anterior margin of this cuticle is emarginate and continues along the ventral surface of the MPP as a ridge (e.g.
<bibRefCitation id="800565FB51481825FE1AF80AEDFE1BE8" author="Ballantyne &amp; Lambkin" box="[387,710,1941,1967]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009</bibRefCitation>
figs 342–349, especially fig. 358); ventral surface of T8 usually with a curved, slightly off–centre ridge close to posterior margin; T8 outlines in
<taxonomicName id="2394638951481825FBB6F825EA3A1B94" authorityName="Pic. Nothing" authorityYear="1923" box="[1071,1282,1977,2003]" class="Insecta" family="Lampyridae" genus="Pacifica" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="limbatipennis">
<emphasis id="D6E0C41851481825FBB6F825EA3A1B94" box="[1071,1282,1977,2003]" italics="true" pageId="115" pageNumber="116">Pac. limbatipennis</emphasis>
</taxonomicName>
,
<taxonomicName id="2394638951481825FA8BF826EAA41B94" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[1298,1436,1977,2003]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="limbatifusca">
<emphasis id="D6E0C41851481825FA8BF826EAA41B94" box="[1298,1436,1977,2003]" italics="true" pageId="115" pageNumber="116">limbatifusca</emphasis>
</taxonomicName>
and
<taxonomicName id="2394638951481825FF5EF842EE011BB0" box="[199,313,2013,2039]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="salomonis">
<emphasis id="D6E0C41851481825FF5EF842EE011BB0" box="[199,313,2013,2039]" italics="true" pageId="115" pageNumber="116">salomonis</emphasis>
</taxonomicName>
characterised (
<bibRefCitation id="800565FB51481825FE7BF842EC1E1BB0" author="Ballantyne &amp; Lambkin" box="[482,806,2013,2039]" firstAuthor="Ballantyne" pageId="115" pageNumber="116" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009</bibRefCitation>
figs 438–442); T8 with pale partly membranous ventral projections from the bases of the anterolateral prolongations; aedeagal sheath sternite posterior to the lateral tergite articulations subparallel-sided in basal 1/3 to 1/2; aedeagus L/W&lt;3; LL/ML wide; ML never asymmetrical, always with lateral teeth; anterior margin of LL never asymmetrically produced. Females are macropterous. Larvae are not associated.
</paragraph>
</subSubSection>
</treatment>
</document>
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