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<mods:title id="9FF92EAFF821BB5D0D277E6597F7EF18">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<treatment id="6C3DA91C5110187BFF0EFF06ED5D1CBC" ID-DOI="http://doi.org/10.5281/zenodo.4688801" ID-GBIF-Taxon="159757435" ID-Zenodo-Dep="4688801" LSID="urn:lsid:plazi:treatment:6C3DA91C5110187BFF0EFF06ED5D1CBC" httpUri="http://treatment.plazi.org/id/6C3DA91C5110187BFF0EFF06ED5D1CBC" lastPageId="45" lastPageNumber="45" pageId="43" pageNumber="44">
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<heading id="BF63AF665110187DFF0EFF06EED91CF4" bold="true" box="[151,481,153,180]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<taxonomicName id="239463895110187DFF0EFF06EED91CF4" ID-CoL="ZHV" authority="Ballantyne 2009" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,481,153,180]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFF0EFF06EED91CF4" bold="true" box="[151,481,153,180]" pageId="43" pageNumber="44">
<emphasis id="D6E0C4185110187DFF0EFF06EE281CF3" bold="true" box="[151,272,153,180]" italics="true" pageId="43" pageNumber="44">Aquilonia</emphasis>
<bibRefCitation id="800565FB5110187DFE8EFF05EED91CF4" author="Ballantyne" box="[279,481,153,180]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne 2009</bibRefCitation>
</emphasis>
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</paragraph>
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<figureCitation id="7CAF048F5110187DFF0EFF22EE2D1C90" box="[151,277,189,215]" captionStart="FIGURES 64–69" captionStartId="20.[151,264,1850,1875]" captionTargetBox="[157,1431,186,1824]" captionTargetId="figure@20.[151,1435,166,1831]" captionTargetPageId="20" captionText="FIGURES 64–69. Key to genera characters 10. 64 Aquilonia costata; 65, 66 Aq. similismessoria comb. nov.; 67 Magnalata limbata; 68 Atyphella flammans; 69 Aty. aphrogeneia." figureDoi="http://doi.org/10.5281/zenodo.3508127" httpUri="https://zenodo.org/record/3508127/files/figure.png" pageId="43" pageNumber="44">Figs 64–66</figureCitation>
</paragraph>
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<paragraph id="E42B180A5110187DFF0EFE9DEC631D5D" blockId="43.[151,1241,257,374]" box="[151,859,257,282]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="C4843F245110187DFF0EFE9DEC631D5D" box="[151,859,257,282]" pageId="43" pageNumber="44">
<taxonomicName id="239463895110187DFF0EFE9DEC2C1D5D" ID-CoL="ZHV" authority="Ballantyne 2009 in Ballantyne &amp; Lambkin 2009: 28" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,788,257,282]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFF0EFE9DEFC31D5D" box="[151,251,258,282]" italics="true" pageId="43" pageNumber="44">Aquilonia</emphasis>
<bibRefCitation id="800565FB5110187DFE98FE9EEE911D5D" author="Ballantyne" box="[257,425,257,282]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">
Ballantyne
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in
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<bibRefCitation id="800565FB5110187DFE50FE9EEC2C1D5D" author="Ballantyne &amp; Lambkin" box="[457,788,257,282]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009: 28</bibRefCitation>
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, fig. 4.
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</paragraph>
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<emphasis id="D6E0C4185110187DFF0EFEBFEE1E1D7E" bold="true" box="[151,294,288,313]" pageId="43" pageNumber="44">
<typeStatus id="3B2FA6A85110187DFF0EFEBFEFF41D7F" box="[151,204,288,312]" pageId="43" pageNumber="44">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="239463895110187DFEB5FEBFED031D7E" ID-CoL="3WCQ2" authority="Lea 1921 b" authorityName="Lea" authorityYear="1921" box="[300,571,288,313]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="D6E0C4185110187DFEB5FEBFEEF11D7F" box="[300,457,288,312]" italics="true" pageId="43" pageNumber="44">Luciola costata</emphasis>
<bibRefCitation id="800565FB5110187DFE56FEBEED031D7E" author="Lea" box="[463,571,288,313]" firstAuthor="Lea" pageId="43" pageNumber="44" pagination="50 - 135" refId="ref118916" refString="Lea, A. M. (1921 b) On Australian Coleoptera of the Family Malacodermidae. Transactions of the Royal Society of South Australia, 45, 50 - 135. [Lampyridae, 65 - 68]" type="journal article" year="1921" yearSuffix="b">Lea 1921b</bibRefCitation>
</taxonomicName>
by monotypy (
<bibRefCitation id="800565FB5110187DFD4CFEBFEB191D7F" author="Ballantyne &amp; Lambkin" box="[725,1057,288,313]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="15 - 93" refId="ref114393" refString="Ballantyne, L. A. &amp; Lambkin, C. (2000) Lampyridae of Australia (Coleoptera: Lampyridae: Luciolinae: Luciolini). Memoirs of the Queensland Museum, 46 (1), 15 - 93." type="journal article" year="2000">Ballantyne &amp; Lambkin 2000: 31</bibRefCitation>
, figs 1–3, 5, 6, 8).
</paragraph>
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<paragraph id="E42B180A5110187DFF0EFEA0EB541D11" blockId="43.[151,1241,257,374]" box="[151,1132,318,343]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="C4843F245110187DFF0EFEA0EB541D11" box="[151,1132,318,343]" pageId="43" pageNumber="44">
<taxonomicName id="239463895110187DFF0EFEA0EC1E1D10" ID-CoL="8MR6M" authority="Ballantyne 2009 in Ballantyne &amp; Lambkin 2009: 58" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,806,318,343]" class="Insecta" family="Lampyridae" genus="Gilvainsula" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFF0EFEA0EE351D10" box="[151,269,319,343]" italics="true" pageId="43" pageNumber="44">Gilvainsula</emphasis>
<bibRefCitation id="800565FB5110187DFE8DFEA1EE831D10" author="Ballantyne" box="[276,443,318,343]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">
Ballantyne
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</bibRefCitation>
in
<treatmentCitation id="65353E1B5110187DFE42FEA1EC1E1D10" ID-DOI="http://doi.org/10.5281/zenodo.5324364" ID-Zenodo-Dep="5324364" author="Ballantyne, L. A. &amp; Lambkin, C. L." box="[475,806,318,343]" httpUri="http://treatment.plazi.org/id/0394D665BE15FF86FF3C56342636E81A" page="58" pageId="43" pageNumber="44" year="2009">
<bibRefCitation id="800565FB5110187DFE42FEA1EC1E1D10" author="Ballantyne &amp; Lambkin" box="[475,806,318,343]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009: 58</bibRefCitation>
</treatmentCitation>
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, figs 192–203.
<emphasis id="D6E0C4185110187DFC5BFEA0EB5F1D10" bold="true" box="[962,1127,319,343]" pageId="43" pageNumber="44">New synonymy</emphasis>
.
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</paragraph>
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<emphasis id="D6E0C4185110187DFF0EFEC1EE261D31" bold="true" box="[151,286,349,374]" pageId="43" pageNumber="44">
<typeStatus id="3B2FA6A85110187DFF0EFEC1EFF41D31" box="[151,204,350,374]" pageId="43" pageNumber="44">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="239463895110187DFEB3FEC1ED041D31" authority="Olivier" authorityName="Olivier" authorityYear="1913" box="[298,572,349,374]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="messoria">
<emphasis id="D6E0C4185110187DFEB3FEC1EED51D32" box="[298,493,350,374]" italics="true" pageId="43" pageNumber="44">Atyphella messoria</emphasis>
Olivier
</taxonomicName>
designated by
<bibRefCitation id="800565FB5110187DFD4DFEC2EB181D31" author="Ballantyne &amp; Lambkin" box="[724,1056,349,374]" firstAuthor="Ballantyne" pageId="43" pageNumber="44" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009: 58</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B815110187DFF0EFE3CEE801F29" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="E42B180A5110187DFF0EFE3CEE801F29" blockId="43.[151,1437,419,2031]" pageId="43" pageNumber="44">
<emphasis id="D6E0C4185110187DFF0EFE3CEE321DF9" bold="true" box="[151,266,419,446]" pageId="43" pageNumber="44">Diagnosis</emphasis>
.
<taxonomicName id="239463895110187DFE81FE3BEEBE1DF9" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[280,390,420,446]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFE81FE3BEEBE1DF9" box="[280,390,420,446]" italics="true" pageId="43" pageNumber="44">Aquilonia</emphasis>
</taxonomicName>
is part of the
<taxonomicName id="239463895110187DFD87FE3BEDB11DF9" authorityName="Olliff" authorityYear="1890" box="[542,649,420,446]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFD87FE3BEDB11DF9" box="[542,649,420,446]" italics="true" pageId="43" pageNumber="44">Atyphella</emphasis>
</taxonomicName>
complex and has aedeagal LL visible at the sides of the ML and an aedeagal sheath which is evenly emarginated on its right side. It is distinguished from other
<taxonomicName id="239463895110187DFBE9FE57EBD01DA5" authorityName="Lacordaire" authorityYear="1857" box="[1136,1256,456,482]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="166" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
in the
<taxonomicName id="239463895110187DFAA8FE57EAA41DA5" authorityName="Olliff" authorityYear="1890" box="[1329,1436,456,482]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFAA8FE57EAA41DA5" box="[1329,1436,456,482]" italics="true" pageId="43" pageNumber="44">Atyphella</emphasis>
</taxonomicName>
‘complex’ (as defined in the key to genera), by the following: from
<emphasis id="D6E0C4185110187DFC15FE73ECC81E41" box="[908,1008,492,518]" italics="true" pageId="43" pageNumber="44">Lloydiell</emphasis>
a which has dark brown non margined elytra and a hooded basal piece of the aedeagus by the colour;
<taxonomicName id="239463895110187DFCCDFD8FECFA1E6D" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[852,962,528,554]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFCCDFD8FECFA1E6D" box="[852,962,528,554]" italics="true" pageId="43" pageNumber="44">Aquilonia</emphasis>
</taxonomicName>
males are pale brownish to orange yellow dorsally with or without a black elytral apex, and the basal piece is not hooded; from
<taxonomicName id="239463895110187DFBD9FDABEB9A1E09" authorityName="Ballantyne Pronotum" authorityYear="2009" box="[1088,1186,564,590]" class="Insecta" family="Lampyridae" genus="Convexa" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFBD9FDABEB9A1E09" box="[1088,1186,564,590]" italics="true" pageId="43" pageNumber="44">Convexa</emphasis>
</taxonomicName>
which has dark brown elytra having lateral, sutural and apical margins orange by the elytral colour; from
<taxonomicName id="239463895110187DFBC9FDC7EBF31E35" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[1104,1227,600,626]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFBC9FDC7EBF31E35" box="[1104,1227,600,626]" italics="true" pageId="43" pageNumber="44">Magnalata</emphasis>
</taxonomicName>
which has orange pronotum and black elytra with pale lateral margin, by the elytral colour; from
<taxonomicName id="239463895110187DFB9BFDE3EB551ED1" authorityName="Olliff" authorityYear="1890" box="[1026,1133,636,662]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFB9BFDE3EB551ED1" box="[1026,1133,636,662]" italics="true" pageId="43" pageNumber="44">Atyphella</emphasis>
</taxonomicName>
by the elytral colour; those
<taxonomicName id="239463895110187DFF0EFD3FEE3A1EFD" authorityName="Olliff" authorityYear="1890" box="[151,258,672,698]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFF0EFD3FEE3A1EFD" box="[151,258,672,698]" italics="true" pageId="43" pageNumber="44">Atyphella</emphasis>
</taxonomicName>
having dark brown pale margined elytra have pale margins on both lateral and sutural margins. Many
<taxonomicName id="239463895110187DFF0EFD5BEE3A1E99" authorityName="Olliff" authorityYear="1890" box="[151,258,708,734]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185110187DFF0EFD5BEE3A1E99" box="[151,258,708,734]" italics="true" pageId="43" pageNumber="44">Atyphella</emphasis>
</taxonomicName>
have both the anterior and posterior areas of the hypomeron strongly flattened, or at least the posterior area strongly adpressed. In
<taxonomicName id="239463895110187DFE57FD77EDAD1F45" authorityName="Pic" authorityYear="1929" box="[462,661,744,770]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="D6E0C4185110187DFE57FD77EDAD1F45" box="[462,661,744,770]" italics="true" pageId="43" pageNumber="44">Aquilonia costata</emphasis>
</taxonomicName>
the anterior area is not closely adpressed and the posterior area may be close or not;
<emphasis id="D6E0C4185110187DFECAFC93ED3D1F61" box="[339,517,780,806]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFE39FC92ED3D1F61" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1913" box="[416,517,781,806]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="messoria" status="comb. nov.">messoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFD97FC93EDA91F61" bold="true" box="[526,657,780,806]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFD97FC93EDA91F61" box="[526,657,780,806]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="D6E0C4185110187DFD55FC93ECFF1F61" box="[716,967,780,806]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFC80FC93ECFF1F61" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[793,967,780,806]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFC49FC93EB6B1F61" bold="true" box="[976,1107,780,806]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFC49FC93EB6B1F61" box="[976,1107,780,806]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
have only the posterior area close. Known females are macropterous and a possible larva has laterally explanate tergal margins covering the laterotergites from above.
</paragraph>
</subSubSection>
<subSubSection id="AC8E4B815110187AFF5EFCE8EC1C1838" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="E42B180A5110187DFF5EFCE8ED111FD4" blockId="43.[151,1437,419,2031]" box="[199,553,887,915]" pageId="43" pageNumber="44">
<emphasis id="D6E0C4185110187DFF5EFCE8EEEF1FD5" bold="true" box="[199,471,887,914]" pageId="43" pageNumber="44">Redescription of genus</emphasis>
.
<emphasis id="D6E0C4185110187DFE7CFCE7ED1A1FD5" bold="true" box="[485,546,888,914]" pageId="43" pageNumber="44">Male</emphasis>
.
</paragraph>
<paragraph id="E42B180A5110187DFF5EFC02EBC318F5" blockId="43.[151,1437,419,2031]" pageId="43" pageNumber="44">
Pronotum: wider than long; dorsal surface without irregularities in posterolateral areas (1) and longitudinal groove in lateral areas (2); punctation moderately dense (3); anterior margin not explanate (6); lateral margins diverging posteriorly along most of their length (C&gt;A, B) or lateral margins subparallel-sided (B=C) in
<emphasis id="D6E0C4185110187DFB63FC7BEAAD1FB9" box="[1274,1429,996,1022]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFADDFC7AEAAD1FB9" authorityName="Pic" authorityYear="1929" box="[1348,1429,997,1022]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
; width&gt; humeral width (subequal to humeral width in
<emphasis id="D6E0C4185110187DFD6FFB97ECA91865" box="[758,913,1032,1058]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFCD9FB96ECA91865" authorityName="Pic" authorityYear="1929" box="[832,913,1033,1058]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
and some
<emphasis id="D6E0C4185110187DFB90FB97EA3A1865" box="[1033,1282,1032,1058]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFBCDFB97EA3A1865" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[1108,1282,1032,1058]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFA90FB97EAB21865" bold="true" box="[1289,1418,1032,1058]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFA90FB97EAB21865" box="[1289,1418,1032,1058]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); anterolateral corners rounded obtuse; lateral margins without indentation at mid-point, and sinuosity in either horizontal or vertical plane (15, 16); without indentation in lateral margin near posterolateral corner, and irregularities at corner (17, 18); posterolateral corners rounded obtuse or approximately 90°; posterolateral corners projecting as far as median posterior margin (in
<emphasis id="D6E0C4185110187DFE6CFB07EDA918F5" box="[501,657,1176,1202]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFDD9FB06EDA918F5" authorityName="Pic" authorityYear="1929" box="[576,657,1177,1202]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
) or beyond; separated from it by scarce emarginations.
</paragraph>
<paragraph id="E42B180A5110187DFF5EFB22EA6B1959" blockId="43.[151,1437,419,2031]" pageId="43" pageNumber="44">
Hypomera: closed; median area of hypomeron not elevated in vertical direction; in the posterior area the dorsal and ventral surfaces of the pronotum may closely approach (strongly adpressed) or not in
<emphasis id="D6E0C4185110187DFBE4FB7FEA2018BD" box="[1149,1304,1248,1274]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFB5EFB7EEA2018BD" authorityName="Pic" authorityYear="1929" box="[1223,1304,1249,1274]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
; this area is not adpressed in the remaining species; pronotal width/ GHW 1.4–1.5 in
<emphasis id="D6E0C4185110187DFC53FA9BEB5D1959" box="[970,1125,1284,1310]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFB8DFA9AEB5D1959" authorityName="Pic" authorityYear="1929" box="[1044,1125,1285,1310]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
, 1.6 in other species.
</paragraph>
<paragraph id="E42B180A5110187DFF5EFAB7EA3A1A5D" blockId="43.[151,1437,419,2031]" pageId="43" pageNumber="44">
Elytron: lateral margins either parallel-sided or slightly convex sided in
<emphasis id="D6E0C4185110187DFC75FAB7EBA31905" box="[1004,1179,1320,1346]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFBAFFAB6EBA31905" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1913" box="[1078,1179,1321,1346]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="messoria" status="comb. nov.">messoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFB3BFAB7EA1B1905" bold="true" box="[1186,1315,1320,1346]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFB3BFAB7EA1B1905" box="[1186,1315,1320,1346]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="D6E0C4185110187DFAC0FAB7EE7D1921" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFF0EFAD3EE7D1921" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[151,325,1356,1382]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFED4FAD3EEF71921" bold="true" box="[333,463,1356,1382]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFED4FAD3EEF71921" box="[333,463,1356,1382]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; apices not deflexed; punctation dense (34, 35), not linear (36), not as large as that of pronotum, nor widely and evenly spaced; epipleuron and sutural ridge extend beyond mid-point almost to apex but not extending around apex, neither thickened in apical half; two interstitial lines; elytral carina absent; in horizontal specimen viewed from below epipleuron at elytral base wide, covering humerus or almost so; viewed from above anterior margin of epipleuron arises level with or anterior to posterior margin of MS; epipleuron developed as a lateral ridge along most of length; sutural margins approximate along most of length in closed elytra.
</paragraph>
<paragraph id="E42B180A5110187DFF5EF9BBEA561BE1" blockId="43.[151,1437,419,2031]" pageId="43" pageNumber="44">
Head: gently depressed between eyes; moderately well exposed in front of pronotum, and capable of partial retraction within prothoracic cavity; eyes with moderate separation beneath at level of posterior margin of mouthpart complex; eyes above labrum moderately to widely separated; frons-vertex junction rounded, without median elevation; posterolateral eye excavation not strongly developed, not visible in resting head position; antennal sockets on head between eyes, not contiguous, always separated by &lt;ASW or subequal to ASW; clypeolabral suture present, flexible, not in front of anterior eye margin when head viewed with labrum horizontal; outer edges of labrum reach inner edges of closed mandibles or beyond. Mouthparts: functional; apical labial palpomere flattened, shaped like a broad triangle with inner edge dentate with two teeth in
<emphasis id="D6E0C4185110187DFC8AF8BFEC971B7D" box="[787,943,1824,1850]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFCC7F8BEEC971B7D" authorityName="Pic" authorityYear="1929" box="[862,943,1825,1850]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
, irregularly dentate in both
<emphasis id="D6E0C4185110187DFB75F8BFEAA41B7D" box="[1260,1436,1824,1850]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFAAEF8BEEAA41B7D" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1913" box="[1335,1436,1825,1850]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="messoria" status="comb. nov.">messoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFF0EF8DBEE201B19" bold="true" box="[151,280,1860,1886]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFF0EF8DBEE201B19" box="[151,280,1860,1886]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="D6E0C4185110187DFED6F8DBED701B19" box="[335,584,1860,1886]" italics="true" pageId="43" pageNumber="44">
Aquil.
<taxonomicName id="239463895110187DFE03F8DBED701B19" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[410,584,1860,1886]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185110187DFDD6F8DBEDE81B19" bold="true" box="[591,720,1860,1886]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="CDD379635110187DFDD6F8DBEDE81B19" box="[591,720,1860,1886]" pageId="43" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and at least half as long as apical maxillary palpomere. Antennae 11 segmented; length&gt;GHW up to twice GHW; FS1 subequal in length to, or longer than, pedicel; FS 7−9 not conspicuously shorter than rest; no FS expanded or laterally produced; all FS including FS 1 longer than wide.
</paragraph>
<paragraph id="E42B180A5110187DFF5EF82EEA261B8D" blockId="43.[151,1437,419,2031]" box="[199,1310,1968,1995]" pageId="43" pageNumber="44">Legs: with inner tarsal claw entire; without MFC; no leg segments swollen, curved or emarginated.</paragraph>
<paragraph id="E42B180A5110187AFF5EF84BEEBC1E5C" blockId="43.[151,1437,419,2031]" lastBlockId="44.[151,1437,153,1151]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
Abdomen: without cuticular remnants in association with aedeagal sheath (131, 132); no ventrites with curved posterior margins, nor extending anteriorly into emarginated posterior margin of anterior segment (106–108).
<date id="902A3ECA5117187AFAE3FF06EAA41CF4" box="[1402,1436,153,179]" pageId="44" pageNumber="45">V6</date>
with LO present, occupying almost all the ventrite surface.
<date id="902A3ECA5117187AFCA2FF22EC651C90" box="[827,861,189,215]" pageId="44" pageNumber="45">V7</date>
with LO entire, occupying more than half the surface of ventrite, and reaching to sides, and into MPP in
<emphasis id="D6E0C4185117187AFC95FF7EEC911CBC" box="[780,937,225,251]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFCC1FF7DEC911CBC" authorityName="Pic" authorityYear="1929" box="[856,937,226,251]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
only; MPP as long as wide (L=W) in
<emphasis id="D6E0C4185117187AFAC0FF7EEFD01D58" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFF0EFE99EFD01D58" authorityName="Pic" authorityYear="1929" box="[151,232,262,287]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
, short or wider than long (W&gt;L) in remainder; no PLP. T8 well sclerotised, symmetrical, visible posterior area either widest across median area (W&gt;L) or W=L; median posterior margin not emarginate; T8 not engulfing posterior margin of
<date id="902A3ECA5117187AFEECFED2EEAF1D20" box="[373,407,333,359]" pageId="44" pageNumber="45">V7</date>
nor MPP (133, 134), without well-defined median longitudinal trough; concealed anterolateral arms of T8 very short in
<emphasis id="D6E0C4185117187AFE47FEEEED421DCC" box="[478,634,369,395]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFDB0FEEDED421DCC" authorityName="Pic" authorityYear="1929" box="[553,634,370,395]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
, not as long as entire posterior portion in rest; not laterally emarginated before their origins, dorsoventral expansions absent, expanded only in horizontal plane, narrowly in
<emphasis id="D6E0C4185117187AFB63FE0AEAAD1DE8" box="[1274,1429,405,431]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFADDFE09EAAD1DE8" authorityName="Pic" authorityYear="1929" box="[1348,1429,406,431]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
, widely in
<emphasis id="D6E0C4185117187AFE90FE26EE811D94" box="[265,441,441,467]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFECDFE25EE811D94" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1913" box="[340,441,442,467]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="messoria" status="comb. nov.">messoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185117187AFE58FE26ED7A1D94" bold="true" box="[449,578,441,467]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFE58FE26ED7A1D94" box="[449,578,441,467]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="D6E0C4185117187AFDE2FE26EC4C1D94" box="[635,884,441,467]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFD5FFE26EC4C1D94" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[710,884,441,467]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185117187AFCE5FE26ECC51D94" bold="true" box="[892,1021,441,467]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFCE5FE26ECC51D94" box="[892,1021,441,467]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; without bifurcation of inner margin and ventrally directed pieces (217); lateral margins of T8 not enfolding sides of
<date id="902A3ECA5117187AFBB4FE42EB771DB0" box="[1069,1103,477,503]" pageId="44" pageNumber="45">V7</date>
(190); T7 without prolonged anterolateral corners.
</paragraph>
<paragraph id="E42B180A5117187AFF5EFDBAEC111E88" blockId="44.[151,1437,153,1151]" pageId="44" pageNumber="45">
Aedeagal sheath: asymmetrical; sheath sternite evenly emarginated on its right side from the point of attachment of the tergite; anterior half of sternite broad, apically rounded; tergite without lateral arms extending anteriorly at sides of sheath sternite; tergite without projecting pieces along posterior margin of T9, anterior margin without separate transverse band but
<emphasis id="D6E0C4185117187AFE45FD0EEDEC1EEC" box="[476,724,657,683]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFDBFFD0EEDEC1EEC" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[550,724,657,683]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185117187AFD45FD0EEC651EEC" bold="true" box="[732,861,657,683]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFD45FD0EEC651EEC" box="[732,861,657,683]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
has a darker band across the anterior margin of the sheath tergite (
<bibRefCitation id="800565FB5117187AFEA2FD2AED461E88" author="Ballantyne &amp; Lambkin" box="[315,638,693,719]" firstAuthor="Ballantyne" pageId="44" pageNumber="45" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009</bibRefCitation>
figs 199, 200).
</paragraph>
<paragraph id="E42B180A5117187AFF5EFD46EEFF1FE0" blockId="44.[151,1437,153,1151]" pageId="44" pageNumber="45">
Aedeagus: L/W 3/1 or greater (&lt;3/
<quantity id="236CB5EF5117187AFDC1FD46ED471EB4" box="[600,639,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="44" pageNumber="45" unit="in" value="1.0">1 in</quantity>
<emphasis id="D6E0C4185117187AFD1EFD46EC1B1EB4" box="[647,803,729,755]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFD4BFD45EC1B1EB4" authorityName="Pic" authorityYear="1929" box="[722,803,730,755]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
), symmetrical with apices of LL visible from beneath at sides of ML, LL/ML moderate; LL of equal length, a little longer than ML, separated along inner dorsal margins but not divergent in basal half; with apices narrower than ML; preapical inner margin of LL obliquely truncate, with no obvious tooth; dorsal base of LL produced apically acute and medially entire except for
<emphasis id="D6E0C4185117187AFB18FCDAEA251F18" box="[1153,1309,837,863]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFB55FCD9EA251F18" authorityName="Pic" authorityYear="1929" box="[1228,1309,838,863]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
where it is neither produced nor emarginated; ML symmetrical; BP not strongly sclerotised, not hooded, not strongly emarginated along anterior margin.
</paragraph>
<paragraph id="E42B180A5117187AFF5EFC2EED311854" blockId="44.[151,1437,153,1151]" pageId="44" pageNumber="45">
<emphasis id="D6E0C4185117187AFF5EFC2EEE261F8C" bold="true" box="[199,286,945,971]" pageId="44" pageNumber="45">Female</emphasis>
. Macropterous and probably capable of flight. Pronotum: subparallel sided (A=C) in
<emphasis id="D6E0C4185117187AFB6EFC2EEAAD1F8C" box="[1271,1429,945,971]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFADDFC2DEAAD1F8C" authorityName="Pic" authorityYear="1929" box="[1348,1429,946,971]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
, wider across posterior area (C&gt; A, B) in
<emphasis id="D6E0C4185117187AFDC5FC4AEC6D1FA8" box="[604,853,981,1007]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFD3EFC4AEC6D1FA8" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[679,853,981,1007]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185117187AFCC5FC4AECE41FA8" bold="true" box="[860,988,981,1007]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFCC5FC4AECE41FA8" box="[860,988,981,1007]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
Pronotal width/humeral width subequal in
<emphasis id="D6E0C4185117187AFF2DFC66EE681854" box="[180,336,1017,1043]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFF66FC65EE681854" authorityName="Pic" authorityYear="1929" box="[255,336,1018,1043]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
,&gt; in remainder.
</paragraph>
<paragraph id="E42B180A5117187AFF5EFB82EC1C1838" blockId="44.[151,1437,153,1151]" pageId="44" pageNumber="45">
<emphasis id="D6E0C4185117187AFF5EFB82EE281870" bold="true" box="[199,272,1053,1079]" pageId="44" pageNumber="45">Larva</emphasis>
. Not associated by breeding in
<emphasis id="D6E0C4185117187AFD1BFB82EC191870" box="[642,801,1053,1079]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFD49FB81EC191870" authorityName="Pic" authorityYear="1929" box="[720,801,1054,1079]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
and association assumed only because of geographical location. Tergal margins laterally explanate with tergal margins not thickened; punctures in anterior half of tergites 2–10 no larger than remaining punctures in
<emphasis id="D6E0C4185117187AFD18FBFAEC251838" box="[641,797,1125,1151]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFD55FBF9EC251838" authorityName="Pic" authorityYear="1929" box="[716,797,1126,1151]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
<caption id="B0EB48825117187AFF0EFB31EDAB18AC" ID-Table-UUID="B0EB48825117187AFF0EFB31EDAB18AC" httpUri="http://table.plazi.org/id/B0EB48825117187AFF0EFB31EDAB18AC" pageId="44" pageNumber="45" startId="44.[151,239,1198,1222]" targetBox="[166,1418,1270,1631]" targetIsTable="true" targetPageId="44">
<paragraph id="E42B180A5117187AFF0EFB31EDAB18AC" blockId="44.[151,766,1198,1259]" pageId="44" pageNumber="45">
TABLE 5. List of species of
<taxonomicName id="239463895117187AFE4AFB31EDC61881" authority="Ballantyne 2009" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[467,766,1198,1223]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFE4AFB31ED791881" bold="true" box="[467,577,1198,1222]" italics="true" pageId="44" pageNumber="45">Aquilonia</emphasis>
<bibRefCitation id="800565FB5117187AFDDEFB30EDC61881" author="Ballantyne" box="[583,766,1198,1223]" firstAuthor="Ballantyne" pageId="44" pageNumber="45" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne 2009</bibRefCitation>
</taxonomicName>
(*Reference: 1 =
<bibRefCitation id="800565FB5117187AFEC8FB4CEDB018AC" author="Ballantyne &amp; Lambkin" box="[337,648,1235,1259]" firstAuthor="Ballantyne" pageId="44" pageNumber="45" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne &amp; Lambkin 2009</bibRefCitation>
.)
</paragraph>
</caption>
<paragraph id="E42B180A5117187AFF3FFB69EFE91A18" pageId="44" pageNumber="45">
<table id="9694EAAA5117E7A9FF3FFB69EAB21A18" box="[166,1418,1270,1631]" gridcols="7" gridrows="5" pageId="44" pageNumber="45">
<tr id="5AA41A485117E7A9FF3FFB69EAB21980" box="[166,1418,1270,1479]" gridrow="0" pageId="44" pageNumber="45">
<th id="197573345117E7A9FF3FFB69ED711980" box="[166,585,1270,1479]" gridcol="0" gridrow="0" pageId="44" pageNumber="45">Species</th>
<th id="197573345117E7A9FD10FB69ED9B1980" box="[649,675,1270,1479]" gridcol="1" gridrow="0" pageId="44" pageNumber="45">Australia</th>
<th id="197573345117E7A9FCA7FB69EC441980" box="[830,892,1270,1479]" gridcol="2" gridrow="0" pageId="44" pageNumber="45">New Guinea and Pacific islands</th>
<th id="197573345117E7A9FC2EFB69EB3C1980" box="[951,1028,1270,1479]" gridcol="3" gridrow="0" pageId="44" pageNumber="45">Location of types</th>
<th id="197573345117E7A9FB04FB69EB871980" box="[1181,1215,1270,1479]" gridcol="4" gridrow="0" pageId="44" pageNumber="45">Females associated</th>
<th id="197573345117E7A9FB64FB69EA271980" box="[1277,1311,1270,1479]" gridcol="5" gridrow="0" pageId="44" pageNumber="45">Larvae associated</th>
<th id="197573345117E7A9FAD5FB69EAB21980" box="[1356,1418,1270,1479]" gridcol="6" gridrow="0" pageId="44" pageNumber="45">References for illus- trations of species *</th>
</tr>
<tr id="5AA41A485117E7A9FF3FFA49EAB219A8" box="[166,1418,1494,1519]" gridrow="1" pageId="44" pageNumber="45" rowspan-2="1">
<th id="197573345117E7A9FF3FFA49ED7119A8" box="[166,585,1494,1519]" gridcol="0" gridrow="1" pageId="44" pageNumber="45">
<taxonomicName id="239463895117187AFF3FFA47EE4919A8" authority="(Lea 1921 b)" authorityName="Elytra" authorityYear="1921" baseAuthorityName="Lea" baseAuthorityYear="1921" box="[166,369,1494,1519]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="D6E0C4185117187AFF3FFA47EFD719A8" box="[166,239,1496,1519]" italics="true" pageId="44" pageNumber="45">costata</emphasis>
(
<bibRefCitation id="800565FB5117187AFF65FA48EE5219A8" author="Lea" box="[252,362,1494,1519]" firstAuthor="Lea" pageId="44" pageNumber="45" pagination="50 - 135" refId="ref118916" refString="Lea, A. M. (1921 b) On Australian Coleoptera of the Family Malacodermidae. Transactions of the Royal Society of South Australia, 45, 50 - 135. [Lampyridae, 65 - 68]" type="journal article" year="1921" yearSuffix="b">Lea 1921b</bibRefCitation>
)
</taxonomicName>
</th>
<td id="197573345117E7A9FD10FA49ED9B19A8" box="[649,675,1494,1519]" gridcol="1" gridrow="1" pageId="44" pageNumber="45">*</td>
<td id="197573345117E7A9FC2EFA49EB3C19A8" box="[951,1028,1494,1519]" gridcol="3" gridrow="1" pageId="44" pageNumber="45">NMVA</td>
<td id="197573345117E7A9FB04FA49EB8719A8" box="[1181,1215,1494,1519]" gridcol="4" gridrow="1" pageId="44" pageNumber="45">yes</td>
<td id="197573345117E7A9FB64FA49EA2719A8" box="[1277,1311,1494,1519]" gridcol="5" gridrow="1" pageId="44" pageNumber="45">yes</td>
<td id="197573345117E7A9FAD5FA49EAB219A8" box="[1356,1418,1494,1519]" gridcol="6" gridrow="1" pageId="44" pageNumber="45">1</td>
</tr>
<tr id="5AA41A485117E7A9FF3FFA63EAB21A52" box="[166,1418,1532,1557]" gridrow="2" pageId="44" pageNumber="45" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<th id="197573345117E7A9FF3FFA63ED711A52" box="[166,585,1532,1557]" gridcol="0" gridrow="2" pageId="44" pageNumber="45">
<taxonomicName id="239463895117187AFF3FFA62EE9C1A52" authority="(Olivier 1913 a)" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1913" box="[166,420,1532,1557]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="messoria" status="comb. nov.">
<emphasis id="D6E0C4185117187AFF3FFA62EE391A53" box="[166,257,1533,1556]" italics="true" pageId="44" pageNumber="45">messoria</emphasis>
(
<bibRefCitation id="800565FB5117187AFE97FA63EEA51A52" author="Olivier" box="[270,413,1532,1557]" firstAuthor="Olivier" pageId="44" pageNumber="45" pagination="417 - 422" refId="ref120984" refString="Olivier, E. (1913 a) Lampyridae. Resultats de L'Expedition Scientifique Neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices de Dr H. R. Lorentz. Zoologie Livraison III. Nova Guinea, 9, 417 - 422." type="journal article" year="1913" yearSuffix="a">Olivier 1913a</bibRefCitation>
)
</taxonomicName>
<emphasis id="D6E0C4185117187AFE32FA62ED261A52" bold="true" box="[427,542,1533,1557]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFE32FA62ED261A52" box="[427,542,1533,1557]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="197573345117E7A9FCA7FA63EC441A52" box="[830,892,1532,1557]" gridcol="2" gridrow="2" pageId="44" pageNumber="45">*</td>
<td id="197573345117E7A9FC2EFA63EB3C1A52" box="[951,1028,1532,1557]" gridcol="3" gridrow="2" pageId="44" pageNumber="45">MNHN</td>
<td id="197573345117E7A9FAD5FA63EAB21A52" box="[1356,1418,1532,1557]" gridcol="6" gridrow="2" pageId="44" pageNumber="45">1</td>
</tr>
<tr id="5AA41A485117E7A9FF3FF9BDEAB21A7C" box="[166,1418,1570,1595]" gridrow="3" pageId="44" pageNumber="45" rowspan-1="1" rowspan-5="1">
<th id="197573345117E7A9FF3FF9BDED711A7C" box="[166,585,1570,1595]" gridcol="0" gridrow="3" pageId="44" pageNumber="45">
<taxonomicName id="239463895117187AFF3FF9BCED391A7C" authority="(Ballantyne 2009)" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[166,513,1570,1595]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similismessoria">
<emphasis id="D6E0C4185117187AFF3FF9BCEE7C1A7C" box="[166,324,1571,1595]" italics="true" pageId="44" pageNumber="45">similismessoria</emphasis>
(
<bibRefCitation id="800565FB5117187AFEC9F9BDEEC21A7C" author="Ballantyne" box="[336,506,1570,1595]" firstAuthor="Ballantyne" pageId="44" pageNumber="45" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. &amp; Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne 2009</bibRefCitation>
)
</taxonomicName>
<emphasis id="D6E0C4185117187AFD91F9BCED711A7C" bold="true" box="[520,585,1571,1595]" pageId="44" pageNumber="45">comb.</emphasis>
</th>
<td id="197573345117E7A9FCA7F9BDEC441A7C" box="[830,892,1570,1595]" gridcol="2" gridrow="3" pageId="44" pageNumber="45">*</td>
<td id="197573345117E7A9FC2EF9BDEB3C1A7C" box="[951,1028,1570,1595]" gridcol="3" gridrow="3" pageId="44" pageNumber="45">BPBM</td>
<td id="197573345117E7A9FB04F9BDEB871A7C" box="[1181,1215,1570,1595]" gridcol="4" gridrow="3" pageId="44" pageNumber="45">yes</td>
<td id="197573345117E7A9FAD5F9BDEAB21A7C" box="[1356,1418,1570,1595]" gridcol="6" gridrow="3" pageId="44" pageNumber="45">1</td>
</tr>
<tr id="5AA41A485117E7A9FF3FF9D8EAB21A18" box="[166,1418,1607,1631]" gridrow="4" pageId="44" pageNumber="45">
<th id="197573345117E7A9FF3FF9D8EAB21A18" box="[166,1418,1607,1631]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="44" pageNumber="45">nov.</th>
</tr>
</table>
</paragraph>
<subSubSection id="AC8E4B815117187BFF5EF914ED5D1CBC" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="discussion">
<paragraph id="E42B180A5117187BFF5EF914ED5D1CBC" blockId="44.[151,1437,1674,2025]" lastBlockId="45.[151,1436,153,251]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="D6E0C4185117187AFF5EF914EE031AE2" bold="true" box="[199,315,1675,1701]" pageId="44" pageNumber="45">Remarks.</emphasis>
The most recent phylogenetic analyses incorporating some molecular data indicated a close relationship between
<taxonomicName id="239463895117187AFEADF931ED351A8F" authorityName="Ho" authorityYear="2019" baseAuthorityName="Blanchard. McDermott" baseAuthorityYear="2019" box="[308,525,1710,1736]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="limbata">
<emphasis id="D6E0C4185117187AFEADF931ED351A8F" box="[308,525,1710,1736]" italics="true" pageId="44" pageNumber="45">Magnalata limbata</emphasis>
</taxonomicName>
, one species of
<taxonomicName id="239463895117187AFD5EF931EC0D1A8F" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[711,821,1710,1736]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFD5EF931EC0D1A8F" box="[711,821,1710,1736]" italics="true" pageId="44" pageNumber="45">Aquilonia</emphasis>
</taxonomicName>
, and two species of
<taxonomicName id="239463895117187AFBB8F931EB9C1A8F" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[1057,1188,1710,1736]" class="Insecta" family="Lampyridae" genus="Gilvainsula" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFBB8F931EB9C1A8F" box="[1057,1188,1710,1736]" italics="true" pageId="44" pageNumber="45">Gilvainsula</emphasis>
</taxonomicName>
, and was found in all analyses. In following these results
<taxonomicName id="239463895117187AFDABF94DED8D1AAB" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[562,693,1746,1772]" class="Insecta" family="Lampyridae" genus="Gilvainsula" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFDABF94DED8D1AAB" box="[562,693,1746,1772]" italics="true" pageId="44" pageNumber="45">Gilvainsula</emphasis>
</taxonomicName>
is synonymised with
<taxonomicName id="239463895117187AFC2BF94DEB181AAB" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[946,1056,1746,1772]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFC2BF94DEB181AAB" box="[946,1056,1746,1772]" italics="true" pageId="44" pageNumber="45">Aquilonia</emphasis>
</taxonomicName>
, while
<taxonomicName id="239463895117187AFBE1F94DEBCB1AAB" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[1144,1267,1746,1772]" class="Insecta" family="Lampyridae" genus="Magnalata" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFBE1F94DEBCB1AAB" box="[1144,1267,1746,1772]" italics="true" pageId="44" pageNumber="45">Magnalata</emphasis>
</taxonomicName>
is retained for one species. It was consequently necessary to redescribe
<taxonomicName id="239463895117187AFCB3F969ECA01B57" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[810,920,1782,1808]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFCB3F969ECA01B57" box="[810,920,1782,1808]" italics="true" pageId="44" pageNumber="45">Aquilonia</emphasis>
</taxonomicName>
. Although the three species included now in
<taxonomicName id="239463895117187AFF0EF885EE3D1B73" authorityName="Ballantyne 2009 in Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,261,1818,1844]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0C4185117187AFF0EF885EE3D1B73" box="[151,261,1818,1844]" italics="true" pageId="44" pageNumber="45">Aquilonia</emphasis>
</taxonomicName>
were grouped by phylogenetic analyses (
<bibRefCitation id="800565FB5117187AFD43F885ECA61B72" author="Jusoh" box="[730,926,1818,1845]" etAl="et al." firstAuthor="Jusoh" pageId="44" pageNumber="45" pagination="1 - 71" refId="ref118381" refString="Jusoh, W. F. A., Ballantyne, L., Lambkin, C. L., Hashim, N. R. &amp; Wahlberg, N. (2018) The firefly genus Pteroptyx Olivier revisited (Coleoptera: Lampyridae: Luciolinae). Zootaxa, 4456 (1), 1 - 71. https: // doi. org / 10.11646 / zootaxa. 4456.1.1" type="journal article" year="2018">
Jusoh
<emphasis id="D6E0C4185117187AFCBAF884EC651B73" box="[803,861,1818,1844]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2018
</bibRefCitation>
) there is little obvious commonality between them if colour is considered (none of these species had molecular data determined and the analysis was based on morphology only). One species is pale brown dorsally, one orange with black elytral apices, one pale brown with brown elytral apices. Their commonality lies however in their occurrence in a fairly restricted area of the Australopacific region.
<taxonomicName id="239463895117187AFECFF835ED251B83" authorityName="Elytra" baseAuthorityName="Lea" baseAuthorityYear="1921" box="[342,541,1962,1988]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="D6E0C4185117187AFECFF835ED251B83" box="[342,541,1962,1988]" italics="true" pageId="44" pageNumber="45">Aquilonia costata</emphasis>
</taxonomicName>
is known only from northern
<collectingCountry id="9C83589A5117187AFCF5F835ECEB1B83" box="[876,979,1962,1988]" name="Australia" pageId="44" pageNumber="45">Australia</collectingCountry>
with one possible record from islands in the Torres Strait.
<taxonomicName id="239463895117187AFECEF851ED081BAF" authorityName="Ho" authorityYear="2019" baseAuthorityName="Olivier" baseAuthorityYear="1913" box="[343,560,1998,2024]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="messoria" status="comb. nov.">
<emphasis id="D6E0C4185117187AFECEF851ED081BAF" box="[343,560,1998,2024]" italics="true" pageId="44" pageNumber="45">Aquilonia messoria</emphasis>
</taxonomicName>
<emphasis id="D6E0C4185117187AFDAEF850ED8E1BAE" bold="true" box="[567,694,1999,2025]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFDAEF850ED8E1BAE" box="[567,694,1999,2025]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="D6E0C4185117187AFD72F851ECDB1BAF" box="[747,995,1998,2024]" italics="true" pageId="44" pageNumber="45">
Aquil.
<taxonomicName id="239463895117187AFCACF851ECDB1BAF" authorityName="Ho" authorityYear="2019" baseAuthorityName="Ballantyne" baseAuthorityYear="2009" box="[821,995,1998,2024]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similismessoria" status="comb. nov.">similismessoria</taxonomicName>
</emphasis>
<emphasis id="D6E0C4185117187AFC73F850EB521BAE" bold="true" box="[1002,1130,1999,2025]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="CDD379635117187AFC73F850EB521BAE" box="[1002,1130,1999,2025]" pageId="44" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
are both known from a few records along the eastern coast of the island of New
<collectingCountry id="9C83589A5116187BFD7CFF06EC0E1CF4" box="[741,822,153,179]" name="Guinea-Bissau" pageId="45" pageNumber="46">Guinea</collectingCountry>
near Buna Bay and also off shore islands. The generic description above indicates considerable morphological differences between
<emphasis id="D6E0C4185116187BFB99FF22EBA61C90" box="[1024,1182,189,215]" italics="true" pageId="45" pageNumber="46">
Aquil.
<taxonomicName id="239463895116187BFBD4FF21EBA61C90" authorityName="Elytra" baseAuthorityName="Lea" baseAuthorityYear="1921" box="[1101,1182,190,215]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="costata">costata</taxonomicName>
</emphasis>
and the other species, especially in the outline of the pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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