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<document id="F6568E2E55C9EF3063E92E7241A2ABF0" ID-DOI="10.11646/zootaxa.2225.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5320276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629632753302" checkinUser="felipe" docAuthor="Emmerich, Daniel &amp; Nieto, Carolina" docDate="2009" docId="5F198786FFCB57332FFAFBFEFE654BD9" docLanguage="en" docName="zt02225p062.pdf" docOrigin="Zootaxa 2225 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2225.1.3" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paracloeodes morellii Emmerich &amp; Nieto, 2009, sp. nov." docType="treatment" docVersion="4" lastPageNumber="62" masterDocId="A320FFFEFFC857362F6DFFEFFF854916" masterDocTitle="First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species" masterLastPageNumber="62" masterPageNumber="57" pageNumber="60" updateTime="1699087309791" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FAF4D14082B3B94A500E4D3E51D06475">First record of the genus Paracloeodes Day (Ephemeroptera: Baetidae) in Uruguay and description of two new species</mods:title>
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<mods:namePart id="A6797B691DB2CE9AD796738EFD350AD9">Emmerich, Daniel</mods:namePart>
<mods:affiliation id="7103B15AD93B570C39CB4D34322AB385">Facultad de Ciencias, Sección Entomología, PEDECIBA. Universidad de la República, Montevideo, Uruguay.</mods:affiliation>
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<mods:namePart id="3D35080057330E937CE1FD74384F4574">Nieto, Carolina</mods:namePart>
<mods:affiliation id="B6288ABC8099C0D03BBA1069DAED13B8">CONICET-INSUE-Facultad de Ciencias Naturales e IML, Miguel Lillo 205, S. M. de Tucumán (4000), Tucumán, Argentina.</mods:affiliation>
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<mods:title id="DA42252ECA2FC1EBA7EF0771080249E3">Zootaxa</mods:title>
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<mods:date id="174B7032C2600EAF0423BFEE3E3BCEF4">2009</mods:date>
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<treatment id="5F198786FFCB57332FFAFBFEFE654BD9" ID-DOI="http://doi.org/10.5281/zenodo.5333493" ID-Zenodo-Dep="5333493" LSID="urn:lsid:plazi:treatment:5F198786FFCB57332FFAFBFEFE654BD9" httpUri="http://treatment.plazi.org/id/5F198786FFCB57332FFAFBFEFE654BD9" lastPageId="5" lastPageNumber="62" pageId="3" pageNumber="60">
<subSubSection id="9FAA651BFFCB57352FFAFBFEFD874D3C" box="[151,514,1040,1067]" pageId="3" pageNumber="60" type="nomenclature">
<paragraph id="D70F3690FFCB57352FFAFBFEFD874D3C" blockId="3.[151,514,1040,1102]" box="[151,514,1040,1067]" pageId="3" pageNumber="60">
<heading id="8C4781FCFFCB57352FFAFBFEFD874D3C" bold="true" box="[151,514,1040,1067]" fontSize="11" level="1" pageId="3" pageNumber="60" reason="1">
<emphasis id="E5C4EA82FFCB57352FFAFBFEFD874D3C" bold="true" box="[151,514,1040,1067]" pageId="3" pageNumber="60">
<taxonomicName id="10B04D13FFCB57352FFAFBFEFE184D3D" ID-CoL="4CQL3" authorityName="Emmerich &amp; Nieto" authorityYear="2009" box="[151,413,1041,1067]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="morellii" status="sp. nov.">
<emphasis id="E5C4EA82FFCB57352FFAFBFEFE184D3D" bold="true" box="[151,413,1041,1067]" italics="true" pageId="3" pageNumber="60">Paracloeodes morellii</emphasis>
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<taxonomicNameLabel id="FEF757F9FFCB57352EC8FBFFFD874D3C" box="[421,514,1040,1066]" pageId="3" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="9FAA651BFFCB57352FFAFBDBFE274EB6" pageId="3" pageNumber="60" type="description">
<paragraph id="D70F3690FFCB57352FFAFBDBFEB04D58" blockId="3.[151,514,1040,1102]" box="[151,309,1076,1102]" pageId="3" pageNumber="60">
(
<figureCitation id="4F8B2A15FFCB57352FF2FBDBFEA94D58" box="[159,300,1076,1102]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Figs. 1325</figureCitation>
)
</paragraph>
<paragraph id="D70F3690FFCB57352FFAFB6FFE044CE1" blockId="3.[151,1436,1152,2030]" pageId="3" pageNumber="60">
<emphasis id="E5C4EA82FFCB57352FFAFB6FFF794D8C" bold="true" box="[151,252,1152,1178]" pageId="3" pageNumber="60">Nymph.</emphasis>
Length: body:
<quantity id="10489B75FFCB57352EDAFB6EFDCE4D8D" box="[439,587,1153,1179]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="3" pageNumber="60" unit="mm" value="3.05" valueMax="3.3" valueMin="2.8">2.83.3 mm</quantity>
; cerci 1.0
<quantity id="10489B75FFCB57352DBDFB6EFCAA4D8D" box="[720,815,1153,1179]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="60" unit="mm" value="1.3">1.3 mm</quantity>
. Antennae
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, 1.5 times the head capsule. General color brownish orange. Head orange brown, with small brownish spots as in
<figureCitation id="4F8B2A15FFCB57352B15FB47FA804DD4" box="[1144,1285,1192,1218]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Figs. 1314</figureCitation>
. Frontal keel present, ocelli brownish. Antennae: scape and pedicel pale yellow, flagellum pale. Mouthparts (
<figureCitation id="4F8B2A15FFCB57352B93FB21FA0A4DFE" box="[1278,1423,1230,1256]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Figs. 1520</figureCitation>
): labrum (
<figureCitation id="4F8B2A15FFCB57352E6DFB1AFE1B4C19" box="[256,414,1269,1295]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Figs. 15 ab</figureCitation>
) dorsally with 23 subapical setae near midline, anterior margin with bifid setae. Mandibles (
<figureCitation id="4F8B2A15FFCB57352E4FFAF3FE284C20" box="[290,429,1308,1334]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Figs. 1617</figureCitation>
) without setae between prostheca and mola, incisors cleft in two sets, inner set at right angle with outer set. Left mandible (
<figureCitation id="4F8B2A15FFCB57352D0FFAADFD384C4A" box="[610,701,1346,1372]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 16</figureCitation>
) with molars with constrictions, prostheca broad with 12 denticles. Right mandible (
<figureCitation id="4F8B2A15FFCB57352E89FA86FDB84C95" box="[484,573,1385,1411]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 17</figureCitation>
) with prostheca bifid. Hypopharynx (
<figureCitation id="4F8B2A15FFCB57352B7DFA86FBEC4C95" box="[1040,1129,1385,1411]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 18</figureCitation>
) with lingua subequal to superlinguae and rounded apically. Maxillae (
<figureCitation id="4F8B2A15FFCB57352DC6FA7FFC854CBC" box="[683,768,1424,1450]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 19</figureCitation>
) with palpi two segmented, 1.5 times the length of galealacinia. Labium (
<figureCitation id="4F8B2A15FFCB57352E05FA59FE444CC6" box="[360,449,1462,1488]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 20</figureCitation>
) with segment II of palpi with rounded projection, 2 times width of segment III, segment III conical.
</paragraph>
<paragraph id="D70F3690FFCB57352FA8F9EBFDAE4FAE" blockId="3.[151,1436,1152,2030]" pageId="3" pageNumber="60">
Thorax (
<figureCitation id="4F8B2A15FFCB57352E42F9EBFE0C4F08" box="[303,393,1540,1566]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 13</figureCitation>
) orange brown. Pronotum with a pair of brownish spots in the midline. Mesonotum shadowed with brown and with small brownish spots. Metanotum brownish. Pleura reddish brown, sterna whitish. Legs (
<figureCitation id="4F8B2A15FFCB57352E28F9BEFE1F4F7D" box="[325,410,1617,1643]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 21</figureCitation>
) pale yellow. Tarsal claws 0.4 times the length of tarsi, with 2 rows of minute denticles (
<figureCitation id="4F8B2A15FFCB57352FF2F997FF7D4F84" box="[159,248,1656,1682]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 22</figureCitation>
). Hind wing pads absent. Female (
<figureCitation id="4F8B2A15FFCB57352DE2F997FD614F84" box="[655,740,1656,1682]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 14</figureCitation>
) paler than male and with a brown band in metanotum and anterior portion of fore wing pads.
</paragraph>
<paragraph id="D70F3690FFCB57352FA8F92AFE2F4E6C" blockId="3.[151,1436,1152,2030]" pageId="3" pageNumber="60">
Abdomen orange brown. Color pattern: segment IIVII with medial elongate spot as in
<figureCitation id="4F8B2A15FFCB57352BBBF92AFAA94FC9" box="[1238,1324,1733,1759]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 13</figureCitation>
, segment IX brownish. Posterior margin of terga with spines as long as wide (
<figureCitation id="4F8B2A15FFCB57352CD2F903FB924E10" box="[959,1047,1772,1798]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 23</figureCitation>
). Sterna pale yellow, sternum IX brownish except at lateral sides. Gills whitish, translucent, oblong, 1.5 times the length of each tergum. Paraprocts (
<figureCitation id="4F8B2A15FFCB57352E4EF8D6FEFC4E45" box="[291,377,1849,1875]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 24</figureCitation>
) with 810 spines apically. Terminal filaments pale yellow, with crown of small spines on each segment (
<figureCitation id="4F8B2A15FFCB57352E28F88FFE184E6C" box="[325,413,1888,1914]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="3" pageNumber="60">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="D70F3690FFCB57352FA8F869FE274EB6" blockId="3.[151,1436,1152,2030]" box="[197,418,1926,1952]" pageId="3" pageNumber="60">
<emphasis id="E5C4EA82FFCB57352FA8F869FE9A4EB6" bold="true" box="[197,287,1926,1952]" pageId="3" pageNumber="60">Adults.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9FAA651BFFCB57352FA8F842FBD24EF8" pageId="3" pageNumber="60" type="etymology">
<paragraph id="D70F3690FFCB57352FA8F842FBD24EF8" blockId="3.[151,1436,1152,2030]" pageId="3" pageNumber="60">
<emphasis id="E5C4EA82FFCB57352FA8F842FED44ED1" bold="true" box="[197,337,1965,1991]" pageId="3" pageNumber="60">Etymology.</emphasis>
This species is dedicated to Dr. Enrique Morelli, entomologist of Universidad de la República (
<collectingCountry id="AFA77600FFCB57352FF2F83BFE8C4EF8" box="[159,265,2004,2030]" name="Uruguay" pageId="3" pageNumber="60">Uruguay</collectingCountry>
) who continually stimulates the study of the aquatic insects of
<collectingCountry id="AFA77600FFCB57352C87F83BFBD74EF8" box="[1002,1106,2004,2030]" name="Uruguay" pageId="3" pageNumber="60">Uruguay</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="83CF6618FFCC57322FFAF880FDF84EF8" ID-DOI="http://doi.org/10.5281/zenodo.5320282" ID-Zenodo-Dep="5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="4" pageNumber="61" startId="4.[151,269,1903,1927]" targetBox="[241,1370,191,1829]" targetPageId="4">
<paragraph id="D70F3690FFCC57322FFAF880FDF84EF8" blockId="4.[151,1436,1903,2030]" pageId="4" pageNumber="61">
<emphasis id="E5C4EA82FFCC57322FFAF880FED84E91" bold="true" box="[151,349,1903,1927]" pageId="4" pageNumber="61">FIGURES 1325.</emphasis>
<taxonomicName id="10B04D13FFCC57322E09F880FDD54E90" authorityName="Emmerich &amp; Nieto" authorityYear="2009" box="[356,592,1903,1926]" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="morellii" status="sp. nov.">
<emphasis id="E5C4EA82FFCC57322E09F880FDD54E90" box="[356,592,1903,1926]" italics="true" pageId="4" pageNumber="61">Paracloeodes morellii</emphasis>
</taxonomicName>
<emphasis id="E5C4EA82FFCC57322D3AF880FD164E91" bold="true" box="[599,659,1903,1927]" pageId="4" pageNumber="61">
<taxonomicNameLabel id="FEF757F9FFCC57322D3AF880FD164E91" box="[599,659,1903,1927]" pageId="4" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus.
</paragraph>
</caption>
<subSubSection id="9FAA651BFFCD57332FA8FF78FCFE488C" pageId="5" pageNumber="62" type="discussion">
<paragraph id="D70F3690FFCD57332FA8FF78FCFE488C" blockId="5.[151,1436,151,719]" pageId="5" pageNumber="62">
<emphasis id="E5C4EA82FFCD57332FA8FF78FED249A7" bold="true" box="[197,343,151,177]" pageId="5" pageNumber="62">Discussion.</emphasis>
This species can be distinguished from the other species of the genus by the following combination of characters: 1) frontal keel present; 2) antennae short, 1.5 times the head capsule; 3) segment II of labial palpi with rounded projection, 2 times width of segment III (
<figureCitation id="4F8B2A15FFCD57332CBDFF0AFBAC49E9" box="[976,1065,229,255]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="5" pageNumber="62">Fig. 20</figureCitation>
); 4) tarsal claws with two rows of minute denticles (
<figureCitation id="4F8B2A15FFCD57332EE2FEE3FE6F4830" box="[399,490,268,294]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="5" pageNumber="62">Fig. 22</figureCitation>
); 5) hind wing pads absent; 6) general color pattern with head and thorax with small spots, abdomen with segment IIVII with medial elongate spot as in
<figureCitation id="4F8B2A15FFCD57332B44FEDDFB07485A" box="[1065,1154,306,332]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="5" pageNumber="62">Fig. 13</figureCitation>
;
<figureCitation id="4F8B2A15FFCD57332BFDFEDDFB25485A" box="[1168,1184,306,332]" captionStart="FIGURES 112" captionStartId="2.[151,271,1907,1931]" captionTargetBox="[240,1323,184,1842]" captionTargetId="figure-91@2.[207,1379,143,1882]" captionTargetPageId="2" captionText="FIGURES 112. Paracloeodes charrua sp. n. Nymph. 1, General view. Mouthparts (Figs. 27): 2a, labrum, left d.v., right v.v.; 2b, labrum: apical setae; 3, left mandible v.v.; 4, right mandible v.v.; 5 hypopharynx v.v.; 6, maxilla v.v.; 7, labium, left d.v., right v.v.; 8, leg I; 9, tarsal claw I; 10, posterior margin of tergum IV; 11, paraproct; 12, caudal filament detail; 12a, left cercus; 12b, terminal filament." figureDoi="http://doi.org/10.5281/zenodo.5320280" httpUri="https://zenodo.org/record/5320280/files/figure.png" pageId="5" pageNumber="62">7</figureCitation>
) posterior margin of terga with spines as long as wide (
<figureCitation id="4F8B2A15FFCD57332D50FEB6FD1C4865" box="[573,665,345,371]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="5" pageNumber="62">Fig. 23</figureCitation>
); 8) paraprocts with 810 spines apically (
<figureCitation id="4F8B2A15FFCD57332BCFFEB6FB7B4865" box="[1186,1278,345,371]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="5" pageNumber="62">Fig. 24</figureCitation>
); 9) terminal filament with crown of small spines on each segment (
<figureCitation id="4F8B2A15FFCD57332C7BFE6FFCE8488C" box="[790,877,384,410]" captionStart="FIGURES 1325" captionStartId="4.[151,269,1903,1927]" captionTargetBox="[241,1370,191,1829]" captionTargetId="figure-91@4.[200,1388,140,1879]" captionTargetPageId="4" captionText="FIGURES 1325. Paracloeodes morellii sp. n. Nymph. 13, Male general view; 14, female head and thorax. Mouthparts (Figs. 1520): 15a, labrum, left d.v., right v.v.; 15b labrum: apical setae; 16, left mandible v.v.; 17, right mandible v.v.; 18, hypopharynx v.v.; 19, maxilla v.v.; 20, labium, left d.v., right v.v.; 21, leg I; 22, tarsal claw I; 23, posterior margin of tergum IV; 24, paraproct; 25, detail of cercus." figureDoi="http://doi.org/10.5281/zenodo.5320282" httpUri="https://zenodo.org/record/5320282/files/figure.png" pageId="5" pageNumber="62">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9FAA651BFFCD57332FA8FE49FE654BD9" pageId="5" pageNumber="62" type="materials_examined">
<paragraph id="D70F3690FFCD57332FA8FE49FE654BD9" blockId="5.[151,1436,151,719]" pageId="5" pageNumber="62">
<emphasis id="E5C4EA82FFCD57332FA8FE49FEC148D6" bold="true" box="[197,324,422,448]" pageId="5" pageNumber="62">Material.</emphasis>
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<typeStatus id="080B8832FFCD57332E23FE49FE4D48D6" box="[334,456,422,448]" pageId="5" pageNumber="62" type="holotype">Holotype</typeStatus>
, male nymph:
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, km 26.5, A°
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,
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,
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,
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&amp;
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col.
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<typeStatus id="080B8832FFCD57332BC4FE22FA9A48F1" box="[1193,1311,461,487]" pageId="5" pageNumber="62" type="paratype">Paratypes</typeStatus>
: 5 nymphs same data (mouthparts, legs and paraprocts of 1 nymph mounted on slide); 1 nymph (parts on slide Depto.
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,
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,
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,
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,
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<collectingDate id="B34AE9B8FFCD57332CBDFDF5FBD24B22" box="[976,1111,538,564]" pageId="5" pageNumber="62" value="2008-02-21">21/02/2008</collectingDate>
</date>
, same collectors; 2 nymphs (1 on slide): Depto.
<collectingRegion id="1574F872FFCD57332EEEFDAEFD934B4D" box="[387,534,577,603]" country="Uruguay" name="Cerro Largo" pageId="5" pageNumber="62">Cerro Largo</collectingRegion>
, ruta 8, km 335, A°
<location id="D26F604BFFCD57332C6BFDAEFCCA4B4D" LSID="urn:lsid:plazi:treatment:5F198786FFCB57332FFAFBFEFE654BD9:D26F604BFFCD57332C6BFDAEFCCA4B4D" box="[774,847,577,603]" country="Uruguay" county="Sarandi del Yi" latitude="-32.833332" longLatPrecision="20" longitude="-54.24861" municipality="Rio Yi" name="Otazo" pageId="5" pageNumber="62" stateProvince="Durazno">Otazo</location>
,
<geoCoordinate id="B2845057FFCD57332C36FDAEFC734B4D" box="[859,1014,577,603]" degrees="32" direction="south" minutes="50" orientation="latitude" pageId="5" pageNumber="62" precision="15" seconds="00" value="-32.833332">S 32°50´00´´</geoCoordinate>
,
<geoCoordinate id="B2845057FFCD57332B6FFDAEFB2C4B4D" box="[1026,1193,577,603]" degrees="54" direction="west" minutes="14" orientation="longitude" pageId="5" pageNumber="62" precision="15" seconds="55" value="-54.24861">W 54°14´55´´</geoCoordinate>
,
<quantity id="10489B75FFCD57332BDBFDAEFA874B4D" box="[1206,1282,577,603]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="5" pageNumber="62" unit="m" value="123.0">
<elevation id="5C9DD1A3FFCD57332BDBFDAEFA874B4D" box="[1206,1282,577,603]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="5" pageNumber="62" unit="m" value="123.0">123 m</elevation>
</quantity>
,
<date id="A30E1050FFCD57332A62FDAEFA124B4D" box="[1295,1431,577,603]" pageId="5" pageNumber="62" value="2008-02-20">
<collectingDate id="B34AE9B8FFCD57332A62FDAEFA124B4D" box="[1295,1431,577,603]" pageId="5" pageNumber="62" value="2008-02-20">20/02/2008</collectingDate>
</date>
,
<collectorName id="7A455346FFCD57332FFAFD87FF5F4B94" box="[151,218,616,642]" pageId="5" pageNumber="62">Same</collectorName>
collectors.
<collectorName id="7A455346FFCD57332E04FD87FE124B94" box="[361,407,616,642]" pageId="5" pageNumber="62">The</collectorName>
<typeStatus id="080B8832FFCD57332ECCFD87FD8E4B94" box="[417,523,616,642]" pageId="5" pageNumber="62" type="holotype">holotype</typeStatus>
and
<specimenCount id="C1B6FD19FFCD57332D25FD87FCA64B94" box="[584,803,616,642]" pageId="5" pageNumber="62" type="female" typeStatus="paratype">1 paratype female</specimenCount>
are deposited in the
<collectorName id="7A455346FFCD57332B48FD87FE9E4BBE" pageId="5" pageNumber="62">Museo de la Universidad de la República</collectorName>
,
<collectingRegion id="1574F872FFCD57332E44FD61FE414BBE" box="[297,452,654,680]" country="Uruguay" name="Montevideo" pageId="5" pageNumber="62">Montevideo</collectingRegion>
,
<collectorName id="7A455346FFCD57332EB9FD61FDC44BBE" box="[468,577,654,680]" pageId="5" pageNumber="62">Uruguay</collectorName>
(
<collectionCode id="B1A1AE55FFCD57332D39FD61FD244BBE" box="[596,673,654,680]" country="USA" lsid="urn:lsid:biocol.org:col:15341" name="Murray State University" pageId="5" pageNumber="62" type="Herbarium">MUR</collectionCode>
); remaining
<typeStatus id="080B8832FFCD57332C2EFD61FC394BBE" box="[835,956,654,680]" pageId="5" pageNumber="62" type="paratype">paratypes</typeStatus>
are housed in
<collectorName id="7A455346FFCD57332B10FD61FA124BBE" box="[1149,1431,654,680]" pageId="5" pageNumber="62">Instituto Miguel Lillo</collectorName>
,
<collectorName id="7A455346FFCD57332FFAFD5AFE8D4BD9" box="[151,264,693,719]" pageId="5" pageNumber="62">Tucumán</collectorName>
,
<collectorName id="7A455346FFCD57332E7EFD5AFE024BD9" box="[275,391,693,719]" pageId="5" pageNumber="62">Argentina</collectorName>
(
<collectionCode id="B1A1AE55FFCD57332EF5FD5AFE514BD9" box="[408,468,693,719]" country="Argentina" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="5" pageNumber="62">IML</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>