386 lines
54 KiB
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386 lines
54 KiB
XML
<document id="06B7161C58B6D5E7EEBB6DC7C799288D" ID-DOI="10.11646/zootaxa.3956.2.2" ID-GBIF-Dataset="cc2e6574-6d74-4c04-a6fb-b5a63e67fc81" ID-ISSN="1175-5326" ID-Zenodo-Dep="237218" ID-ZooBank="80102A8C-F6C0-43C4-81A1-B33E10EE889E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461054501477" checkinUser="plazi" docAuthor="Dean, Harlan K. & Blake, James A." docDate="2015" docId="2C0487A6FFCFFF9E99EBFB1DFBF6FBA1" docLanguage="en" docName="zt03956p198.pdf" docOrigin="Zootaxa 3956 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leitoscoloplos pugettensis Pettibone 1957" docType="treatment" docVersion="9" lastPageNumber="188" masterDocId="D03DFFDEFFCBFF9B997CFF8DFFE0FFF9" masterDocTitle="The Orbiniidae (Annelida: Polychaeta) of Pacific Costa Rica" masterLastPageNumber="198" masterPageNumber="183" pageNumber="187" updateTime="1698580733129" updateUser="plazi">
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<mods:title id="B0BE0D4B68094CC79D6F627BCE03E851">The Orbiniidae (Annelida: Polychaeta) of Pacific Costa Rica</mods:title>
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<mods:date id="5BA96927F3494EC9263825E00AF56926">2015</mods:date>
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<treatment id="2C0487A6FFCFFF9E99EBFB1DFBF6FBA1" ID-DOI="http://doi.org/10.5281/zenodo.6115428" ID-GBIF-Taxon="119612717" ID-Zenodo-Dep="6115428" LSID="urn:lsid:plazi:treatment:2C0487A6FFCFFF9E99EBFB1DFBF6FBA1" httpUri="http://treatment.plazi.org/id/2C0487A6FFCFFF9E99EBFB1DFBF6FBA1" lastPageId="5" lastPageNumber="188" pageId="4" pageNumber="187">
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<subSubSection id="ECB7653BFFCFFF9F99EBFB1DFE59FB32" pageId="4" pageNumber="187" type="nomenclature">
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<paragraph id="A41236B0FFCFFF9F99EBFB1DFD54FB53" blockId="4.[151,692,1168,1228]" box="[151,692,1168,1194]" pageId="4" pageNumber="187">
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<heading id="FF5A81DCFFCFFF9F99EBFB1DFD54FB53" bold="true" box="[151,692,1168,1194]" fontSize="11" level="1" pageId="4" pageNumber="187" reason="1">
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<taxonomicName id="63AD4D33FFCFFF9F99EBFB1DFD54FB53" ID-CoL="3T3DL" authority="Pettibone, 1957" authorityName="Pettibone" authorityYear="1957" box="[151,692,1168,1194]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="pugettensis">
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<emphasis id="96D9EAA2FFCFFF9F99EBFB1DFD54FB53" bold="true" box="[151,692,1168,1194]" pageId="4" pageNumber="187">
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<emphasis id="96D9EAA2FFCFFF9F99EBFB1DFE35FB53" bold="true" box="[151,469,1168,1194]" italics="true" pageId="4" pageNumber="187">Leitoscoloplos pugettensis</emphasis>
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(
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<bibRefCitation id="C03C4B41FFCFFF9F989BFB1DFD4CFB53" author="Pettibone" box="[487,684,1168,1194]" pageId="4" pageNumber="187" refString="Pettibone, M. H. (1957) North American genera of the family Orbiniidae. Journal of the Washington Academy of Sciences, 47, 159 - 167." type="journal article" year="1957">Pettibone, 1957</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="A41236B0FFCFFF9F99EBFB3EFE59FB32" blockId="4.[151,692,1168,1228]" box="[151,441,1203,1228]" pageId="4" pageNumber="187">
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<figureCitation id="3C962A35FFCFFF9F99EBFB3EFEE1FB32" box="[151,257,1203,1228]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="4" pageNumber="187">Figures 2</figureCitation>
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(C–E), 6 (A–B)
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</paragraph>
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</subSubSection>
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<subSubSection id="ECB7653BFFCFFF9F99EBFB76FBF7F9FF" pageId="4" pageNumber="187" type="reference_group">
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<paragraph id="A41236B0FFCFFF9F99EBFB76FCACFAE8" blockId="4.[151,1436,1275,1542]" box="[151,844,1275,1297]" pageId="4" pageNumber="187">
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<taxonomicName id="63AD4D33FFCFFF9F99EBFB76FDC5FAE8" ID-CoL="79XJ7" authority="Johnson 1901: 412" authorityName="Johnson" authorityPageNumber="412" authorityYear="1901" box="[151,549,1275,1297]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="elongata">
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<emphasis id="96D9EAA2FFCFFF9F99EBFB76FEBAFAE8" box="[151,346,1275,1297]" italics="true" pageId="4" pageNumber="187">Scoloplos elongata</emphasis>
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Johnson 1901: 412
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</taxonomicName>
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–413, P1. 10, Figs. 105–110.
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</paragraph>
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<paragraph id="A41236B0FFCFFF9F99EBFA94FCB2FAD6" blockId="4.[151,1436,1275,1542]" box="[151,850,1305,1327]" pageId="4" pageNumber="187">
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<treatmentCitationGroup id="84BD119EFFCFFF9F99EBFA94FCB2FAD6" box="[151,850,1305,1327]" pageId="4" pageNumber="187">
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<taxonomicName id="63AD4D33FFCFFF9F99EBFA94FD98FAD6" ID-CoL="6LD7J" authority="Hartman, 1944: 257" authorityName="Hartman" authorityPageNumber="257" authorityYear="1944" box="[151,632,1305,1327]" class="Polychaeta" family="Orbiniidae" genus="Haploscoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="elongata">
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<emphasis id="96D9EAA2FFCFFF9F99EBFA94FE78FAD6" box="[151,408,1305,1327]" italics="true" pageId="4" pageNumber="187">Haploscoloplos elongata</emphasis>
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:
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<treatmentCitation id="250C10A1FFCFFF9F98DAFA97FD98FAD6" author="Hartman" box="[422,632,1305,1327]" page="257" pageId="4" pageNumber="187" year="1944">
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<bibRefCitation id="C03C4B41FFCFFF9F98DAFA97FDA3FAD6" author="Hartman" box="[422,579,1305,1327]" pageId="4" pageNumber="187" refString="Hartman, O. (1944) Polychaetous annelids from California, including the descriptions of two new genera and nine new species. Allan Hancock Pacific Expeditions, 10, 239 - 310." type="journal article" year="1944">Hartman, 1944</bibRefCitation>
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: 257
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</treatmentCitation>
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</taxonomicName>
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; 1948: 30; 1955:174.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="A41236B0FFCFFF9F99EBFAB5FCA8FA72" blockId="4.[151,1436,1275,1542]" pageId="4" pageNumber="187">
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<treatmentCitationGroup id="84BD119EFFCFFF9F99EBFAB5FCA8FA72" pageId="4" pageNumber="187">
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<taxonomicName id="63AD4D33FFCFFF9F99EBFAB5FD64FAB7" ID-CoL="6LCVJ" authority="Hartman, 1957: 273" authorityName="Hartman" authorityPageNumber="273" authorityYear="1957" box="[151,644,1336,1358]" class="Polychaeta" family="Orbiniidae" genus="Haploscoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="elongatus">
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<emphasis id="96D9EAA2FFCFFF9F99EBFAB5FE43FAB7" box="[151,419,1336,1358]" italics="true" pageId="4" pageNumber="187">Haploscoloplos elongatus</emphasis>
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:
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<treatmentCitation id="250C10A1FFCFFF9F98CFFAB4FD64FAB7" author="Hartman" box="[435,644,1336,1358]" page="273" pageId="4" pageNumber="187" year="1957">
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<bibRefCitation id="C03C4B41FFCFFF9F98CFFAB4FDB1FAB7" author="Hartman" box="[435,593,1336,1358]" pageId="4" pageNumber="187" refString="Hartman, O. (1957) Orbiniidae, Apistobranchidae, Paraonidae and Longosomidae. Allan Hancock Pacific Expeditions, 15, 211 - 393." type="journal article" year="1957">Hartman, 1957</bibRefCitation>
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: 273
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</treatmentCitation>
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</taxonomicName>
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–275, pl. 26,
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<figureCitation id="3C962A35FFCFFF9F9A72FAB5FCB1FAB7" box="[782,849,1336,1358]" captionStart="FIGURE 1" captionStartId="2.[151,250,1546,1568]" captionTargetBox="[199,1393,388,1512]" captionTargetId="figure@2.[188,1400,373,1525]" captionTargetPageId="2" captionText="FIGURE 1. Leitoscoloplos multipapillatus: A, Posterior thoracic parapodium, posterior view; B, Station 29, Gulf of Nicoya specimen, thoracic setiger, anterior view; C, Station 1, Bahia Culebra specimen, posterior end and pygidium. Leitoscoloplos panamensis: D, Station 29, Gulf of Nicoya specimen, thoracic setiger, anterior view; E, Station 29, Gulf of Nicoya specimen, anterior abdominal setiger, anterior view; F, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view. Scale bars: A ‒ F = 100 µm. br = branchiae, ic = interramal cirri, neL = neuropodial lamellae, noL = notopodial lamellae, p = prostomium, per = peristomium, sp = subpodial papillae, vc = ventral cirri." httpUri="https://zenodo.org/record/237219/files/figure.png" pageId="4" pageNumber="187">Figs. 1</figureCitation>
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–11; 1960: 107–108; 1961: 265; 1963: 36; 1969: 19–22,
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<figureCitation id="3C962A35FFCFFF9F99BBFADAFEC2FA94" box="[199,290,1367,1389]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1546,1568]" captionStartId-1="3.[151,250,1500,1522]" captionStartId-2="6.[151,250,1831,1853]" captionStartId-3="7.[151,250,1824,1846]" captionStartId-4="9.[151,250,1545,1567]" captionTargetBox-0="[199,1393,388,1512]" captionTargetBox-1="[174,1411,559,1474]" captionTargetBox-2="[205,1384,507,1790]" captionTargetBox-3="[154,1428,201,1801]" captionTargetBox-4="[173,1414,311,1516]" captionTargetId-0="figure@2.[188,1400,373,1525]" captionTargetId-1="figure@3.[173,1413,555,1478]" captionTargetId-2="figure@6.[189,1397,491,1810]" captionTargetId-3="figure@7.[150,1436,193,1803]" captionTargetId-4="figure@9.[160,1426,301,1524]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionText-0="FIGURE 1. Leitoscoloplos multipapillatus: A, Posterior thoracic parapodium, posterior view; B, Station 29, Gulf of Nicoya specimen, thoracic setiger, anterior view; C, Station 1, Bahia Culebra specimen, posterior end and pygidium. Leitoscoloplos panamensis: D, Station 29, Gulf of Nicoya specimen, thoracic setiger, anterior view; E, Station 29, Gulf of Nicoya specimen, anterior abdominal setiger, anterior view; F, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view. Scale bars: A ‒ F = 100 µm. br = branchiae, ic = interramal cirri, neL = neuropodial lamellae, noL = notopodial lamellae, p = prostomium, per = peristomium, sp = subpodial papillae, vc = ventral cirri." captionText-1="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." captionText-2="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D & E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." captionText-3="FIGURE 4. Leitoscoloplos multipapillatus: A, Setigers 17 – 19, ventral view. Leodamas hamatus n. sp.: B, Paratype XXX Anterior region, lateral view; C, Anterior thoracic spines, setiger three, mid-neuropodial region; D, Posterior thoracic spines, setiger 15, mid-neuropodial region; E, Posterior thoracic spines, upper neuropodial region; F, Furcate notosetae, posterior abdominal region; G, Posterior emergent neuroaciculum, posterior abdominal region. Scale bars: A = 200 µm, B = 500 µm, C ‒ E, F, G = 10 µm. h = hoods, stp = stomach papillae." captionText-4="FIGURE 5. Leodamas treadwelli: A, Station 28 Oct, Gulf of Nicoya specimen, anterior and midbody, lateral view; B, Station 28 Jun, Gulf of Nicoya specimen, notopodial furcate seta. Naineris setosa: C, Playa Blanca specimen, anterior end, dorsal view; D, Playa Blanca specimen, thoracic parapodia, posterior view; E, Playa Blanca specimen, anterior abdominal parapodia, posterior view; F, Playa Blanca specimen, posterior abdominal parapodia, posterior view; G, Playa Blanca specimen, notopodial furcate seta. Scoloplos cryptospinigerus n. sp.: H, Holotype, anterior end, dorsal view; I, Holotype, posterior thoracic neuropodial spines. Scale bars: A, D ‒ E, H = 500 µm, B, G, I = 10 µm, C, F = 1000 µm. s = neuropodial spines." httpUri-0="https://zenodo.org/record/237219/files/figure.png" httpUri-1="https://zenodo.org/record/237220/files/figure.png" httpUri-2="https://zenodo.org/record/237221/files/figure.png" httpUri-3="https://zenodo.org/record/237222/files/figure.png" httpUri-4="https://zenodo.org/record/237223/files/figure.png" pageId="4" pageNumber="187">Figs. 1–5</figureCitation>
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.—
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<treatmentCitation id="250C10A1FFCFFF9F983DFADAFDF8FA94" author="Fauchald" box="[321,536,1367,1389]" page="166" pageId="4" pageNumber="187" year="1972">
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<bibRefCitation id="C03C4B41FFCFFF9F983DFADAFE09FA94" author="Fauchald" box="[321,489,1367,1389]" pageId="4" pageNumber="187" refString="Fauchald, K. (1972) Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the Eastern Pacific Ocean. Allan Hancock Monographs in Marine Biology, 7, 1 - 575." type="journal article" year="1972">Fauchald, 1972</bibRefCitation>
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: 166
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</treatmentCitation>
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||
.—
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||
<treatmentCitation id="250C10A1FFCFFF9F9B4AFADAFCCEFA94" author="Vargas" box="[566,814,1367,1389]" page="336" pageId="4" pageNumber="187" year="1985">
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<bibRefCitation id="C03C4B41FFCFFF9F9B4AFADAFD1CFA94" author="Vargas" box="[566,764,1367,1389]" pageId="4" pageNumber="187" refString="Vargas, J. A., Dean, H. K., Maurer, D. & Orellana, P. (1985) Lista preliminar de invertebrados asociadoa a los sedimentos del Golfo de Nicoya, Costa Rica. Brenesia, 24, 327 - 341." type="journal article" year="1985">
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Vargas
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<emphasis id="96D9EAA2FFCFFF9F9BFAFAD5FD5AFA94" box="[646,698,1367,1389]" italics="true" pageId="4" pageNumber="187">et al.</emphasis>
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, 1985
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</bibRefCitation>
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: 336
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</treatmentCitation>
|
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.—
|
||
<treatmentCitation id="250C10A1FFCFFF9F9A37FADAFBD6FA94" author="Maurer" box="[843,1078,1367,1389]" page="47" pageId="4" pageNumber="187" year="1988">
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<bibRefCitation id="C03C4B41FFCFFF9F9A37FADAFBF1FA94" author="Maurer" box="[843,1041,1367,1389]" pageId="4" pageNumber="187" refString="Maurer, D., Vargas, J. & Dean, H. (1988) Polychaetous Annelids from the Gulf of Nicoya, Costa Rica. Internationale Revue der Gesamten Hydrobiologie, 73 (1), 43 - 59. http: // dx. doi. org / 10.1002 / iroh. 19880730105" type="journal article" year="1988">
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Maurer
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<emphasis id="96D9EAA2FFCFFF9F9ADDFAD5FC34FA94" box="[929,980,1367,1389]" italics="true" pageId="4" pageNumber="187">et al.</emphasis>
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1988
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||
</bibRefCitation>
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: 47
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</treatmentCitation>
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.—
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||
<treatmentCitation id="250C10A1FFCFFF9F9D2FFADAFECDFA72" author="Molina-Lara" page="198" pageId="4" pageNumber="187" year="1995">
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<bibRefCitation id="C03C4B41FFCFFF9F9D2FFADAFF1DFA72" author="Molina-Lara" pageId="4" pageNumber="187" refString="Molina-Lara, O. A. & Vargas Zamora, J. A. (1995) Poliquetos (Annelida: Polychaeta) del estero d Jaltepeque, El Salvador una comparacion 1959 - 1991. Revista de Biologia Tropical, 43, 195 - 205." type="journal article" year="1995">Molina-Lara & Vargas Zamora, 1995</bibRefCitation>
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: 198
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</treatmentCitation>
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.—
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<treatmentCitation id="250C10A1FFCFFF9F9830FAF8FE19FA72" author="Dean" box="[332,505,1397,1419]" page="74" pageId="4" pageNumber="187" year="1996">
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<bibRefCitation id="C03C4B41FFCFFF9F9830FAF8FE32FA72" author="Dean" box="[332,466,1397,1419]" pageId="4" pageNumber="187" refString="Dean, H. K. (1996 a) Subtidal benthic polychaetes (Annelida) of the Gulf of Nicoya, Costa Rica. Revista Biologia Tropical, 44 (Supplement 3), 69 - 80." type="journal article" year="1996" yearSuffix="a">Dean, 1996a</bibRefCitation>
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||
: 74
|
||
</treatmentCitation>
|
||
. Not
|
||
<bibRefCitation id="C03C4B41FFCFFF9F9B4DFAF8FCA4FA72" author="Imajima" box="[561,836,1397,1419]" pageId="4" pageNumber="187" refString="Imajima, M. & Hartman, O. (1964) The polychaetous annelids of Japan. Allan Hancock Foundation Occasional Paper, 26, 1 - 452." type="journal article" year="1964">Imajima & Hartman, 1964</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCFFF9F99EBFA19FD33FA53" blockId="4.[151,1436,1275,1542]" box="[151,723,1428,1450]" pageId="4" pageNumber="187">
|
||
<treatmentCitationGroup id="84BD119EFFCFFF9F99EBFA19FD33FA53" box="[151,723,1428,1450]" pageId="4" pageNumber="187">
|
||
<taxonomicName id="63AD4D33FFCFFF9F99EBFA19FD30FA53" ID-CoL="4VSKJ" authority="Pettibone, 1957: 163" authorityName="Pettibone" authorityPageNumber="163" authorityYear="1957" box="[151,720,1428,1450]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="pugettensis" subGenus="Scoloplos">
|
||
<emphasis id="96D9EAA2FFCFFF9F99EBFA19FE0FFA53" box="[151,495,1428,1450]" italics="true" pageId="4" pageNumber="187">Scoloplos (Scoloplos) pugettensis</emphasis>
|
||
<treatmentCitation id="250C10A1FFCFFF9F988BFA19FD30FA53" author="Pettibone" box="[503,720,1428,1450]" page="163" pageId="4" pageNumber="187" year="1957">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F988BFA19FD7BFA53" author="Pettibone" box="[503,667,1428,1450]" pageId="4" pageNumber="187" refString="Pettibone, M. H. (1957) North American genera of the family Orbiniidae. Journal of the Washington Academy of Sciences, 47, 159 - 167." type="journal article" year="1957">Pettibone, 1957</bibRefCitation>
|
||
: 163
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCFFF9F99EBFA3EFABAFA1E" blockId="4.[151,1436,1275,1542]" pageId="4" pageNumber="187">
|
||
<treatmentCitationGroup id="84BD119EFFCFFF9F99EBFA3EFABAFA1E" pageId="4" pageNumber="187">
|
||
<taxonomicName id="63AD4D33FFCFFF9F99EBFA3EFDB9FA30" ID-CoL="3T3DL" authority="Blake, 1980:1" authorityName="Blake" authorityPageNumber="1" authorityYear="1980" box="[151,601,1459,1481]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="pugettensis">
|
||
<emphasis id="96D9EAA2FFCFFF9F99EBFA3EFE4DFA30" box="[151,429,1459,1481]" italics="true" pageId="4" pageNumber="187">Leitoscoloplos pugettensis</emphasis>
|
||
:
|
||
<treatmentCitation id="250C10A1FFCFFF9F98B8FA3EFDB9FA30" author="Blake" box="[452,601,1459,1481]" page="1" pageId="4" pageNumber="187" year="1980">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F98B8FA3EFDADFA30" author="Blake" box="[452,589,1459,1481]" pageId="4" pageNumber="187" refString="Blake, J. A. (1980) Larval development of Polychaeta from the northern California coast. IV. Leitoscoloplos pugettensis and Scoloplos acmeceps (Family Orbiniidae). Ophelia, 19, 1 - 18. http: // dx. doi. org / 10.1080 / 00785326.1980.10425502" type="journal article" year="1980">Blake, 1980</bibRefCitation>
|
||
:1
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
‒18,
|
||
<figureCitation id="3C962A35FFCFFF9F9BEAFA3EFD1EFA30" box="[662,766,1459,1481]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1546,1568]" captionStartId-1="3.[151,250,1500,1522]" captionStartId-2="6.[151,250,1831,1853]" captionTargetBox-0="[199,1393,388,1512]" captionTargetBox-1="[174,1411,559,1474]" captionTargetBox-2="[205,1384,507,1790]" captionTargetId-0="figure@2.[188,1400,373,1525]" captionTargetId-1="figure@3.[173,1413,555,1478]" captionTargetId-2="figure@6.[189,1397,491,1810]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionText-0="FIGURE 1. Leitoscoloplos multipapillatus: A, Posterior thoracic parapodium, posterior view; B, Station 29, Gulf of Nicoya specimen, thoracic setiger, anterior view; C, Station 1, Bahia Culebra specimen, posterior end and pygidium. Leitoscoloplos panamensis: D, Station 29, Gulf of Nicoya specimen, thoracic setiger, anterior view; E, Station 29, Gulf of Nicoya specimen, anterior abdominal setiger, anterior view; F, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view. Scale bars: A ‒ F = 100 µm. br = branchiae, ic = interramal cirri, neL = neuropodial lamellae, noL = notopodial lamellae, p = prostomium, per = peristomium, sp = subpodial papillae, vc = ventral cirri." captionText-1="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." captionText-2="FIGURE 3. Leodamas hamatus n. sp.: A, holotype, anterior end, dorsal view; B, Holotype, posterior abdominal setiger, anterior view; C, Holotype, anterior thoracic mid-neuropodial spine; D & E, Holotype, posterior thoracic mid-neuropodial spines; F, Holotype, abdominal notopodial furcate setae; G, Holotype, posterior neuropodial emergent hooked spine. Scale bars: A = 200 µm, B = 100 µm, C ‒ G = 10 µm." httpUri-0="https://zenodo.org/record/237219/files/figure.png" httpUri-1="https://zenodo.org/record/237220/files/figure.png" httpUri-2="https://zenodo.org/record/237221/files/figure.png" pageId="4" pageNumber="187">Figs. 1–3</figureCitation>
|
||
; 1996:9‒10, Fig. 1.2.—
|
||
<treatmentCitation id="250C10A1FFCFFF9F9D79FA3EFAC6FA30" author="Hobson" box="[1029,1318,1459,1481]" page="28" pageId="4" pageNumber="187" year="1981">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F9D79FA3EFAE9FA30" author="Hobson" box="[1029,1289,1459,1481]" pageId="4" pageNumber="187" refString="Hobson, K. D. & Banse, K. (1981) Sedentariate and archiannelid polychaetes of British Columbia and Washington. Canadian Bulletin of Fisheries and Aquatic Sciences, 209, 1 - 144." type="journal article" year="1981">Hobson & Banse, 1981</bibRefCitation>
|
||
:28
|
||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="250C10A1FFCFFF9F9C39FA3EFEEDFA1E" author="Mackie" page="8" pageId="4" pageNumber="187" year="1987">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F9C39FA3EFF1EFA1E" author="Mackie" pageId="4" pageNumber="187" refString="Mackie, A. S. Y. (1987) A review of species currently assigned to the genus Leitoscoloplos Day, 1977 (Polychaeta: Orbiniidae), with descriptions of species newly referred to Scoloplos Blainville, 1828. Sarsia, 72, 1 - 28." type="journal article" year="1987">Mackie, 1987</bibRefCitation>
|
||
:8
|
||
</treatmentCitation>
|
||
‒9, Fig. 8.—
|
||
<treatmentCitation id="250C10A1FFCFFF9F98F0FA5CFC9CFA1E" author="Hernandez-Alcantara" box="[396,892,1489,1511]" page="1029" pageId="4" pageNumber="187" year="1993">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F98F0FA5CFCA4FA1E" author="Hernandez-Alcantara" box="[396,836,1489,1511]" pageId="4" pageNumber="187" refString="Hernandez-Alcantara, P. & Solis-Weiss, V. (1993) New records of sedentariate polychaetous annelids from the continental shelf of the Gulf of California. Bulletin of Marine Science, 53, 1027 - 1041." type="journal article" year="1993">Hernández-Alcantara & Solis-Weiss, 1993</bibRefCitation>
|
||
:1029
|
||
</treatmentCitation>
|
||
.—de
|
||
<treatmentCitation id="250C10A1FFCFFF9F9AC0FA5CFAB6FA1E" author="Leon-Gonzalez" box="[956,1366,1489,1511]" page="172" pageId="4" pageNumber="187" year="1996">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F9AC0FA5CFACAFA1E" author="Leon-Gonzalez" box="[956,1322,1489,1511]" pageId="4" pageNumber="187" refString="de Leon-Gonzalez, J. A. & Rodriguez, J. A. (1996) Orbiniidae (Polychaeta) from soft bottom of the western coast of Baja California Peninsula, Mexico. Bulletin of Marine Science, 59, 169 - 174." type="journal article" year="1996">León-González & Rodriguez, 1996</bibRefCitation>
|
||
:172
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCFFF9F99EBFA7DFBF7F9FF" blockId="4.[151,1436,1275,1542]" box="[151,1047,1520,1542]" pageId="4" pageNumber="187">
|
||
<treatmentCitationGroup id="84BD119EFFCFFF9F99EBFA7DFBF7F9FF" box="[151,1047,1520,1542]" pageId="4" pageNumber="187">
|
||
<taxonomicName id="63AD4D33FFCFFF9F99EBFA7DFDB7F9FF" ID-CoL="854RR" authority="Vargas et al. 1985:336" authorityName="Vargas et al." authorityPageNumber="336" authorityYear="1985" box="[151,599,1520,1542]" class="Polychaeta" family="Orbiniidae" genus="Scoloplos" kingdom="Animalia" pageId="4" pageNumber="187" phylum="Annelida" rank="species" species="acmeceps">
|
||
<emphasis id="96D9EAA2FFCFFF9F99EBFA7DFE84F9FF" box="[151,356,1520,1542]" italics="true" pageId="4" pageNumber="187">Scoloplos acmeceps</emphasis>
|
||
:
|
||
<treatmentCitation id="250C10A1FFCFFF9F980FFA7CFDB7F9FF" author="Vargas" box="[371,599,1520,1542]" page="336" pageId="4" pageNumber="187" year="1985">
|
||
<bibRefCitation id="C03C4B41FFCFFF9F980FFA7CFDCCF9FF" author="Vargas" box="[371,556,1520,1542]" pageId="4" pageNumber="187" refString="Vargas, J. A., Dean, H. K., Maurer, D. & Orellana, P. (1985) Lista preliminar de invertebrados asociadoa a los sedimentos del Golfo de Nicoya, Costa Rica. Brenesia, 24, 327 - 341." type="journal article" year="1985">
|
||
Vargas
|
||
<emphasis id="96D9EAA2FFCFFF9F98C3FA7CFE12F9FF" box="[447,498,1520,1542]" italics="true" pageId="4" pageNumber="187">et al.</emphasis>
|
||
1985
|
||
</bibRefCitation>
|
||
:336
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.—Dean, 2009:146. Not
|
||
<bibRefCitation id="C03C4B41FFCFFF9F9A2AFA7DFBF3F9FF" author="Chamberlin" box="[854,1043,1520,1542]" pageId="4" pageNumber="187" refString="Chamberlin, R. V. (1919) Pacific coast Polychaeta collected by Alexander Agassiz. Bulletin of the Museum of Comparative Zoology, 63, 251 - 270." type="journal article" year="1919">Chamberlin, 1919</bibRefCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB7653BFFCFFF9F99EBF9BEFB72F8BA" pageId="4" pageNumber="187" type="materials_examined">
|
||
<paragraph id="A41236B0FFCFFF9F99EBF9BEFB72F8BA" blockId="4.[151,1437,1587,2037]" pageId="4" pageNumber="187">
|
||
<emphasis id="96D9EAA2FFCFFF9F99EBF9BEFE9AF9B5" bold="true" box="[151,378,1587,1612]" pageId="4" pageNumber="187">Material examined</emphasis>
|
||
. Gulf of Nicoya: Sta. 22, 9°48ˊ25″N, 84°52ˊ40″W,
|
||
<quantity id="63559B55FFCFFF9F9ABBF9BEFBE3F9B2" box="[967,1027,1587,1611]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="187" unit="m" value="22.0">22 m</quantity>
|
||
, muddy sand,
|
||
<date id="D0131070FFCFFF9F9DDAF9BEFAC8F9B2" box="[1190,1320,1587,1612]" pageId="4" pageNumber="187" value="1980-07-11">11 Jul 1980</date>
|
||
(2),
|
||
<date id="D0131070FFCFFF9F9C23F9BEFF32F996" pageId="4" pageNumber="187" value="1980-10-01">1 Oct 1980</date>
|
||
, 1 (
|
||
<collectionCode id="C2BCAE75FFCFFF9F9980F9DAFE86F996" box="[252,358,1623,1647]" pageId="4" pageNumber="187">MZUCR</collectionCode>
|
||
); Sta. 24, 9°49ˊ25″N, 84°41ˊ20″W,
|
||
<quantity id="63559B55FFCFFF9F9A7FF9DAFCDEF997" box="[771,830,1623,1647]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="187" unit="m" value="11.0">11 m</quantity>
|
||
, sand,
|
||
<date id="D0131070FFCFFF9F9AF5F9DBFBE7F996" box="[905,1031,1622,1647]" pageId="4" pageNumber="187" value="1980-10-01">1 Oct 1980</date>
|
||
, 4 (
|
||
<collectionCode id="C2BCAE75FFCFFF9F9D4DF9DAFB90F996" LSID="urn:lsid:biocol.org:col:33791" box="[1073,1136,1623,1647]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="187">MCZ</collectionCode>
|
||
83994),
|
||
<date id="D0131070FFCFFF9F9DA4F9DAFAB5F996" box="[1240,1365,1623,1647]" pageId="4" pageNumber="187" value="1981-06-08">8 Jun 1981</date>
|
||
(9);
|
||
<date id="D0131070FFCFFF9F9CF2F9DAFEEDF96B" pageId="4" pageNumber="187" value="1981-08-04">4 Aug 1981</date>
|
||
, 3 (
|
||
<collectionCode id="C2BCAE75FFCFFF9F9843F9F7FE4AF96B" box="[319,426,1658,1682]" pageId="4" pageNumber="187">MZUCR</collectionCode>
|
||
): Sta. 28, 9°52ˊ16″N, 84°45ˊ30"W,
|
||
<quantity id="63559B55FFCFFF9F9A2AF9F4FC75F968" box="[854,917,1657,1682]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="187" unit="m" value="26.0">26 m</quantity>
|
||
, mud,
|
||
<date id="D0131070FFCFFF9F9A96F9F7FB97F96B" box="[1002,1143,1658,1682]" pageId="4" pageNumber="187" value="1981-08-04">4 Aug 1981</date>
|
||
(2): Sta. 29, 9°54ˊ55″N, 84°45ˊ15″W,
|
||
<quantity id="63559B55FFCFFF9F984DF913FE8DF94C" box="[305,365,1694,1718]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="187" unit="m" value="18.0">18 m</quantity>
|
||
, muddy sand,
|
||
<date id="D0131070FFCFFF9F9B6FF913FD71F94F" box="[531,657,1694,1718]" pageId="4" pageNumber="187" value="1980-10-01">1 Oct 1980</date>
|
||
(1);
|
||
<date id="D0131070FFCFFF9F9BB6F913FCAFF94F" box="[714,847,1694,1718]" pageId="4" pageNumber="187" value="1982-04-01">1 Apr 1982</date>
|
||
, 1 (
|
||
<collectionCode id="C2BCAE75FFCFFF9F9A04F913FC57F94F" LSID="urn:lsid:biocol.org:col:33791" box="[888,951,1694,1718]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="187">MCZ</collectionCode>
|
||
70322), 1 (
|
||
<collectionCode id="C2BCAE75FFCFFF9F9D40F913FB9BF94F" LSID="urn:lsid:biocol.org:col:33791" box="[1084,1147,1694,1718]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="187">MCZ</collectionCode>
|
||
85626), 2 (
|
||
<collectionCode id="C2BCAE75FFCFFF9F9D83F913FADEF94F" LSID="urn:lsid:biocol.org:col:33791" box="[1279,1342,1694,1718]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="187">MCZ</collectionCode>
|
||
85695); Sta. 30, 9°54ˊ40″N, 84°45ˊ50″W,
|
||
<quantity id="63559B55FFCFFF9F9B58F94CFD82F921" box="[548,610,1729,1753]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="187" unit="m" value="18.0">18 m</quantity>
|
||
, muddy sand,
|
||
<date id="D0131070FFCFFF9F9A71F94CFC7AF920" box="[781,922,1729,1753]" pageId="4" pageNumber="187" value="1980-01-27">27 Jan 1980</date>
|
||
(1). Punta Morales, Playa Blanca, 10°04ˊN,
|
||
<geoCoordinate id="C1995077FFCFFF9F99EBF969FF19F904" box="[151,249,1764,1789]" direction="west" orientation="longitude" pageId="4" pageNumber="187" precision="925" value="-84.96667">84°58'W</geoCoordinate>
|
||
, lower intertidal, muddy sand, col: J. A. Vargas, 1986 (1); col: H.K. Dean,
|
||
<date id="D0131070FFCFFF9F9DC4F968FAB7F904" box="[1208,1367,1764,1789]" pageId="4" pageNumber="187" value="1986-12-20">20 Dec 1986</date>
|
||
(31). Guanacaste, Bahia Culebra: Sta. 5, 10º35ˊ25.2″N, 85º39ˊ31.1″W,
|
||
<quantity id="63559B55FFCFFF9F9AE8F885FC23F8E6" box="[916,963,1800,1824]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="187" unit="m" value="2.0">2 m</quantity>
|
||
,
|
||
<date id="D0131070FFCFFF9F9AAEF885FB8BF8D9" box="[978,1131,1800,1824]" pageId="4" pageNumber="187" value="2011-05-19">19 May 2011</date>
|
||
(1); Sta. 9, 10º35ˊ30.0″N, 85º39ˊ43.4″W,
|
||
<quantity id="63559B55FFCFFF9F983AF8A6FE94F8BA" box="[326,372,1835,1860]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="187" unit="m" value="5.0">5 m</quantity>
|
||
,
|
||
<date id="D0131070FFCFFF9F98FCF8A6FDF6F8BA" box="[384,534,1835,1860]" pageId="4" pageNumber="187" value="2011-05-19">19 May 2011</date>
|
||
(11). Golfo Dulce, intertidal, 1996, (1), col: H. Buttner.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB7653BFFCFFF9E99BBF8C3FCE3FE54" lastPageId="5" lastPageNumber="188" pageId="4" pageNumber="187" type="description">
|
||
<paragraph id="A41236B0FFCFFF9F99BBF8C3FB7BF80C" blockId="4.[151,1437,1587,2037]" pageId="4" pageNumber="187">
|
||
<emphasis id="96D9EAA2FFCFFF9F99BBF8C3FEB8F89E" bold="true" box="[199,344,1870,1895]" pageId="4" pageNumber="187">Description.</emphasis>
|
||
Complete specimens 12.6, 15.7, and 18.5 mm long; 0.52, 0.54 and 0.45 mm. maximum thoracic width, respectively; incomplete specimens from 0.15–1.25 mm maximum thoracic width (
|
||
<figureCitation id="3C962A35FFCFFF9F9DD8F8FFFB09F873" box="[1188,1257,1906,1931]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="4" pageNumber="187">Fig. 2</figureCitation>
|
||
C). Prostomium short, conical; eyespots lacking; paired dorsolateral nuchal organs present. Peristomium short, one-half prostomial length, subequal to setiger one. Thorax with 12–17 setigers, flattened dorso-ventrally. Branchiae from penultimate thoracic segment, occasionally in last thoracic segment of smaller individuals.
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCEFF9E99BBFF1AFCD8FEB9" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
Thoracic notopodia with triangular postsetal lobe, larger in posterior thoracic setigers; neuropodial postsetal lobe short, digitate, larger in posterior thoracic segments (
|
||
<figureCitation id="3C962A35FFCEFF9E9A27FF31FC46FF2C" box="[859,934,188,213]" captionStart="FIGURE 6" captionStartId="10.[151,250,1716,1738]" captionTargetBox="[219,1361,452,1682]" captionTargetId="figure@10.[212,1374,445,1695]" captionTargetPageId="10" captionText="FIGURE 6. Leitoscoloplos pugettensis: A, Station 9, Bahia Culebra specimen, thoracic setiger, anterior view; B, Station 9, Bahia Culebra specimen, posterior notopodial flail setae. Leodamas treadwelli: C, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view; Scoloplos cryptospinigerus n. sp.: D, Holotype, anterior end, dorsal view; E, Holotype, thoracic parapodia, posterior view; F, Paratype, abdominal parapodia, anterior view; G, Paraytpe, thoracic neurosetal spines. Scale bars: A, F = 50 µm, B = 10 µm, C, E = 100 µm, D = 500 µm, G = 20 µm." httpUri="https://zenodo.org/record/237224/files/figure.png" pageId="5" pageNumber="188">Fig. 6</figureCitation>
|
||
A). Abdominal notopodial postsetal lobes subtriangular in mid-body segments, shorter than branchiae (
|
||
<figureCitation id="3C962A35FFCEFF9E9A34FF52FC6CFF01" box="[840,908,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="5" pageNumber="188">Fig. 2</figureCitation>
|
||
D), becoming more lanceolate and subequal to branchiae posteriorly (
|
||
<figureCitation id="3C962A35FFCEFF9E98EBFE89FE3CFEE5" box="[407,476,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="5" pageNumber="188">Fig. 2</figureCitation>
|
||
E); neuropodial postsetal lobes foliose, bifurcate, medial arm twice length of lateral branch, with well-developed subpodial flange (
|
||
<figureCitation id="3C962A35FFCEFF9E9BCCFEAAFD14FEB9" box="[688,756,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1500,1522]" captionTargetBox="[174,1411,559,1474]" captionTargetId="figure@3.[173,1413,555,1478]" captionTargetPageId="3" captionText="FIGURE 2. Leitoscoloplos multipapillatus: A, Anterior end, lateral view. Leitoscoloplos panamensis: B, Station 29, Gulf of Nicoya specimen, anterior end, lateral view. Leitoscoloplos pugettensis: C, Station 22, Gulf of Nicoya specimen, anterior end, lateral view; D, Station 9, Bahia Culebra specimen, midbody abdominal setiger, anterior view; E, Station 9, Bahia Culebra specimen, posterior abdominal setiger, posterior view. Leodamas hamatus n. sp.: F, Holotype, posterior thoracic setiger, anterior view; G, Holotype, Posterior notopodial emergent hook. Scale bars: A, C = 1000 µm, B = 500 µm, D, E = 100 µm, F = 50 µm G = 10 µm." httpUri="https://zenodo.org/record/237220/files/figure.png" pageId="5" pageNumber="188">Fig. 2</figureCitation>
|
||
D, E).
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCEFF9E99BBFEC1FBD4FE71" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
Camerated capillary setae present throughout thoracic setigers and abdominal neuropodia, with an additional flail setae (
|
||
<figureCitation id="3C962A35FFCEFF9E9868FEE2FEB8FE71" box="[276,344,367,392]" captionStart="FIGURE 6" captionStartId="10.[151,250,1716,1738]" captionTargetBox="[219,1361,452,1682]" captionTargetId="figure@10.[212,1374,445,1695]" captionTargetPageId="10" captionText="FIGURE 6. Leitoscoloplos pugettensis: A, Station 9, Bahia Culebra specimen, thoracic setiger, anterior view; B, Station 9, Bahia Culebra specimen, posterior notopodial flail setae. Leodamas treadwelli: C, Station 28 Jun, Gulf of Nicoya specimen, posterior abdominal setiger, anterior view; Scoloplos cryptospinigerus n. sp.: D, Holotype, anterior end, dorsal view; E, Holotype, thoracic parapodia, posterior view; F, Paratype, abdominal parapodia, anterior view; G, Paraytpe, thoracic neurosetal spines. Scale bars: A, F = 50 µm, B = 10 µm, C, E = 100 µm, D = 500 µm, G = 20 µm." httpUri="https://zenodo.org/record/237224/files/figure.png" pageId="5" pageNumber="188">Fig. 6</figureCitation>
|
||
B) accompanying camerated capillaries in abdominal notopodia.
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCEFF9E99BBFE19FCE3FE54" blockId="5.[151,1436,151,1112]" box="[199,771,404,429]" pageId="5" pageNumber="188">Pygidium with long paired dorso-lateral anal cirri.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB7653BFFCEFF9E99BBFE3AFCBEFCA4" pageId="5" pageNumber="188" type="discussion">
|
||
<paragraph id="A41236B0FFCEFF9E99BBFE3AFAFAFDEE" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
<emphasis id="96D9EAA2FFCEFF9E99BBFE3AFEDBFE29" bold="true" box="[199,315,439,464]" pageId="5" pageNumber="188">Remarks.</emphasis>
|
||
<taxonomicName id="63AD4D33FFCEFF9E983AFE35FD94FE36" box="[326,628,440,464]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="pugettensis">
|
||
<emphasis id="96D9EAA2FFCEFF9E983AFE35FD94FE36" box="[326,628,440,464]" italics="true" pageId="5" pageNumber="188">Leitoscoloplos pugettensis</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="C03C4B41FFCEFF9E9A2EFE3AFBF1FE29" author="Pettibone" box="[850,1041,439,464]" pageId="5" pageNumber="188" refString="Pettibone, M. H. (1957) North American genera of the family Orbiniidae. Journal of the Washington Academy of Sciences, 47, 159 - 167." type="journal article" year="1957">Pettibone (1957)</bibRefCitation>
|
||
from Puget Sound in Washington State from mud and gravelly sand. She synonymized
|
||
<taxonomicName id="63AD4D33FFCEFF9E9B82FE51FB0CFE0D" authority="Johnson, 1901" authorityName="Johnson" authorityYear="1901" box="[766,1260,476,501]" class="Polychaeta" family="Orbiniidae" genus="Haploscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="elongatus">
|
||
<emphasis id="96D9EAA2FFCEFF9E9B82FE51FBC7FE0D" box="[766,1063,476,500]" italics="true" pageId="5" pageNumber="188">Haploscoloplos elongatus</emphasis>
|
||
(Johnson, 1901)
|
||
</taxonomicName>
|
||
, also described from Puget Sound, and later reported from British
|
||
<collectingCountry id="DCBA7620FFCEFF9E9BAAFE72FCA7FDE1" box="[726,839,511,536]" name="United States of America" pageId="5" pageNumber="188">Columbia</collectingCountry>
|
||
,
|
||
<collectingCountry id="DCBA7620FFCEFF9E9A29FE72FC4CFDE1" box="[853,940,511,536]" name="Canada" pageId="5" pageNumber="188">Canada</collectingCountry>
|
||
, California and Pacific
|
||
<collectingCountry id="DCBA7620FFCEFF9E9DC6FD8DFAF6FDEE" box="[1210,1302,512,535]" name="Panama" pageId="5" pageNumber="188">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCEFF9E99BBFDA9FA71FD7C" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
The variability in thoracic length of this species may be correlated with the body size of Costa Rican specimens.
|
||
<bibRefCitation id="C03C4B41FFCEFF9E9862FDCAFE28FD99" author="Mackie" box="[286,456,583,608]" pageId="5" pageNumber="188" refString="Mackie, A. S. Y. (1987) A review of species currently assigned to the genus Leitoscoloplos Day, 1977 (Polychaeta: Orbiniidae), with descriptions of species newly referred to Scoloplos Blainville, 1828. Sarsia, 72, 1 - 28." type="journal article" year="1987">Mackie (1987)</bibRefCitation>
|
||
reported 19–20 thoracic setigers in specimens from Washington State and
|
||
<quantity id="63559B55FFCEFF9E9C49FDC5FA7CFD99" box="[1333,1436,583,608]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.588" metricValueMin="4.826" pageId="5" pageNumber="188" unit="in" value="20.5" valueMax="22.0" valueMin="19.0">19–22 in</quantity>
|
||
specimens from British
|
||
<collectingCountry id="DCBA7620FFCEFF9E98D4FDE1FDF6FD7C" box="[424,534,620,645]" name="United States of America" pageId="5" pageNumber="188">Columbia</collectingCountry>
|
||
while
|
||
<bibRefCitation id="C03C4B41FFCEFF9E9B19FDE1FD19FD7C" author="Blake" box="[613,761,620,645]" pageId="5" pageNumber="188" refString="Blake, J. A. (1996) Family Orbiniidae Hartman, 1942. In: Blake, J. A., Hilbig, B. & Scott, P. H. (Eds.), Taxonomic Atlas of the benthic fauna of the Santa Maria Basin and the Western Santa Barbara Channel, Volume 6. The Annelida, Part 3. Santa Barbara Museum of Natural History, Santa Barbara, California, pp. 1 - 26." type="book chapter" year="1996">Blake (1996)</bibRefCitation>
|
||
reported thorax lengths of
|
||
<quantity id="63559B55FFCEFF9E9D53FDE1FB74FD7C" box="[1071,1172,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="5.08" metricValueMin="3.556" pageId="5" pageNumber="188" unit="in" value="17.0" valueMax="20.0" valueMin="14.0">14–20 in</quantity>
|
||
California specimens.
|
||
</paragraph>
|
||
<paragraph id="A41236B0FFCEFF9E99BBFD02FCBEFCA4" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
<bibRefCitation id="C03C4B41FFCEFF9E99BBFD02FE8FFD51" author="Mackie" box="[199,367,655,680]" pageId="5" pageNumber="188" refString="Mackie, A. S. Y. (1987) A review of species currently assigned to the genus Leitoscoloplos Day, 1977 (Polychaeta: Orbiniidae), with descriptions of species newly referred to Scoloplos Blainville, 1828. Sarsia, 72, 1 - 28." type="journal article" year="1987">Mackie (1987)</bibRefCitation>
|
||
examined
|
||
<typeStatus id="7B168812FFCEFF9E988DFD1DFDBCFD51" box="[497,604,656,680]" pageId="5" pageNumber="188" type="paratype">paratypes</typeStatus>
|
||
of this species from Puget Sound and found branchiae first occurred as small papillae on setigers
|
||
<quantity id="63559B55FFCEFF9E98A8FD39FDDDFD34" box="[468,573,692,717]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="5" pageNumber="188" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
|
||
specimens with 19–22 thoracic setigers.
|
||
<bibRefCitation id="C03C4B41FFCEFF9E9D4DFD39FB1DFD35" author="Pettibone" box="[1073,1277,692,717]" pageId="5" pageNumber="188" refString="Pettibone, M. H. (1957) North American genera of the family Orbiniidae. Journal of the Washington Academy of Sciences, 47, 159 - 167." type="journal article" year="1957">Pettibone (1957)</bibRefCitation>
|
||
reported the branchiae from setigers
|
||
<quantity id="63559B55FFCEFF9E98D0FD5AFDF1FD09" box="[428,529,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="5" pageNumber="188" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">15–18 in</quantity>
|
||
specimens with 19–22 setigers. Blake’s (1996) California specimens were of a greater size range, with branchiae from setigers 14–20 with the onset of branchiae from the one or two most posterior thoracic segments. Costa Rican specimens agree with Blake’s California material in that the branchiae are almost always present from the last two thoracic segments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB7653BFFCEFF9E99BBFCE4FD51FBF6" pageId="5" pageNumber="188" type="reference_group">
|
||
<paragraph id="A41236B0FFCEFF9E99BBFCE4FD51FBF6" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
<taxonomicName id="63AD4D33FFCEFF9E99BBFCE4FE82FC86" ID-CoL="3T3DL" box="[199,354,872,896]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="pugettensis">
|
||
<emphasis id="96D9EAA2FFCEFF9E99BBFCE4FF36FC79" box="[199,214,873,896]" italics="true" pageId="5" pageNumber="188">L</emphasis>
|
||
.
|
||
<emphasis id="96D9EAA2FFCEFF9E9998FCE5FE82FC86" box="[228,354,872,895]" italics="true" pageId="5" pageNumber="188">pugettensis</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="63AD4D33FFCEFF9E9884FCE4FC5AFC79" ID-CoL="3T3CX" authority="Hartmann-Schroder, 1965" authorityName="Hartmann-Schroder" authorityYear="1965" box="[504,954,871,896]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="chilensis">
|
||
<emphasis id="96D9EAA2FFCEFF9E9884FCE4FD98FC79" box="[504,632,872,896]" italics="true" pageId="5" pageNumber="188">L. chilensis</emphasis>
|
||
(
|
||
<bibRefCitation id="C03C4B41FFCEFF9E9BF4FCEAFC52FC79" author="Hartmann-Schroder" box="[648,946,871,896]" pageId="5" pageNumber="188" refString="Hartmann-Schroder, G. (1965) Zur kenntnis des sublitoals der chilenischen kuste unter besonder berucksichtigung der polychaeten und ostracoden. Teil II. Die polychaeten des sublitorals. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 62, 59 - 305." type="journal article" year="1965">Hartmann-Schröder, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
which is redescribed by
|
||
<bibRefCitation id="C03C4B41FFCEFF9E9DAAFCEAFA9CFC79" author="Mackie" box="[1238,1404,871,896]" pageId="5" pageNumber="188" refString="Mackie, A. S. Y. (1987) A review of species currently assigned to the genus Leitoscoloplos Day, 1977 (Polychaeta: Orbiniidae), with descriptions of species newly referred to Scoloplos Blainville, 1828. Sarsia, 72, 1 - 28." type="journal article" year="1987">Mackie (1987)</bibRefCitation>
|
||
as having a maximum of 15–16 thoracic setigers, thoracic branchiae, and morphologically similar abdominal notopodial and neuropodial lamellae. It differs from
|
||
<taxonomicName id="63AD4D33FFCEFF9E9A7BFC3CFC6BFC31" ID-CoL="3T3CX" box="[775,907,944,968]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="chilensis">
|
||
<emphasis id="96D9EAA2FFCEFF9E9A7BFC3CFC6BFC31" box="[775,907,944,968]" italics="true" pageId="5" pageNumber="188">L. chilensis</emphasis>
|
||
</taxonomicName>
|
||
in the more triangular neuropodial postsetal lobes in the thorax and the shape of the branchiae which are medially expanded in
|
||
<emphasis id="96D9EAA2FFCEFF9E9D2BFC58FB00FC15" box="[1111,1248,980,1004]" italics="true" pageId="5" pageNumber="188">
|
||
<taxonomicName id="63AD4D33FFCEFF9E9D2BFC58FB3CFC15" ID-CoL="3T3CX" box="[1111,1244,980,1004]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="chilensis">L. chilensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Flail setae have not been previously reported for
|
||
<emphasis id="96D9EAA2FFCEFF9E9B71FC74FD51FBF6" box="[525,689,1016,1040]" italics="true" pageId="5" pageNumber="188">
|
||
<taxonomicName id="63AD4D33FFCEFF9E9B71FC74FD4DFBF6" ID-CoL="3T3DL" box="[525,685,1016,1040]" class="Polychaeta" family="Orbiniidae" genus="Leitoscoloplos" kingdom="Animalia" pageId="5" pageNumber="188" phylum="Annelida" rank="species" species="pugettensis">L. pugettensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB7653BFFCEFF9E99BBFB91FBF6FBA1" pageId="5" pageNumber="188" type="distribution">
|
||
<paragraph id="A41236B0FFCEFF9E99BBFB91FBF6FBA1" blockId="5.[151,1436,151,1112]" pageId="5" pageNumber="188">
|
||
<emphasis id="96D9EAA2FFCEFF9E99BBFB91FE80FBCC" bold="true" box="[199,352,1052,1077]" pageId="5" pageNumber="188">Distribution.</emphasis>
|
||
Eastern Pacific Ocean from Alaska to
|
||
<collectingCountry id="DCBA7620FFCEFF9E9A6BFB91FC74FBCC" box="[791,916,1052,1077]" name="Costa Rica" pageId="5" pageNumber="188">Costa Rica</collectingCountry>
|
||
, intertidal to
|
||
<specimenCount id="B2ABFD39FFCEFF9E9D54FB91FB96FBCD" box="[1064,1142,1052,1076]" pageId="5" pageNumber="188" type="generic">
|
||
<quantity id="63559B55FFCEFF9E9D54FB91FB91FBCD" box="[1064,1137,1052,1076]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="188" unit="m" value="220.0">220 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Found in muddy to sandy sediments from
|
||
<quantity id="63559B55FFCEFF9E9832FBCDFE55FBAE" box="[334,437,1088,1112]" metricMagnitude="1" metricUnit="m" metricValue="1.65" metricValueMax="2.2" metricValueMin="1.1" pageId="5" pageNumber="188" unit="m" value="16.5" valueMax="22.0" valueMin="11.0">
|
||
11‒
|
||
<specimenCount id="B2ABFD39FFCEFF9E980AFBCDFE55FBAE" box="[374,437,1088,1112]" pageId="5" pageNumber="188" type="generic">22 m</specimenCount>
|
||
</quantity>
|
||
. in the Gulf of Nicoya and 1.5‒
|
||
<quantity id="63559B55FFCEFF9E9A61FBB2FCA7FBAE" box="[797,839,1087,1112]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="188" unit="m" value="5.0">5 m</quantity>
|
||
in Bahia Culebra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |