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<document id="6BD5EA9B3CC1A1B12BD413FEEA1119F3" ID-DOI="10.11646/zootaxa.2391.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5308365" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629669120741" checkinUser="felipe" docAuthor="Richards, Stephen J., Hoskin, Conrad J., Cunningham, Michael J., Mcdonald, Keith &amp; Donnellan, Stephen C." docDate="2010" docId="03FF878FFFA0200A48B3FCD1FF0CB45F" docLanguage="en" docName="zt02391p046.pdf" docOrigin="Zootaxa 2391 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2391.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hyla papua Van Kampen 1909" docType="treatment" docVersion="1" lastPageNumber="42" masterDocId="FFC6FFF7FFA920034877FFE9FFEFB023" masterDocTitle="Taxonomic re-assessment of the Australian and New Guinean green-eyed treefrogs Litoria eucnemis, L. genimaculata and L. serrata (Anura: Hylidae)" masterLastPageNumber="46" masterPageNumber="33" pageNumber="42" updateTime="1730367965011" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="C6B0CE7F893715680296032380D0E796">Taxonomic re-assessment of the Australian and New Guinean green-eyed treefrogs Litoria eucnemis, L. genimaculata and L. serrata (Anura: Hylidae)</mods:title>
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<mods:namePart id="453A6DDC2B12122FB35600C72664BC60">Richards, Stephen J.</mods:namePart>
<mods:affiliation id="535EBCB513C1BC79B14C0566103E063D">South Australian Museum, North Terrace, Adelaide, 5000, Australia. Email: Steve. Richards @ samuseum. sa. gov. au, and Conservation International, P. O. Box 1024, Atherton, Queensland 4883, Australia.</mods:affiliation>
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<mods:namePart id="723F6AEB9987395D21BF52B57D165273">Hoskin, Conrad J.</mods:namePart>
<mods:affiliation id="9A020B969E2766F67E79FAFCE0823E6C">Research School of Biology, The Australian National University, Canberra, 0200, Australia. Email: conrad. hoskin @ anu. edu. au.</mods:affiliation>
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<mods:namePart id="1D40AE5324CCA57946FC969D6E99CCDD">Cunningham, Michael J.</mods:namePart>
<mods:affiliation id="7D90CB6F7B5F24425C97C8701211C39B">School of Integrative Biology, The University of Queensland, St Lucia, 4072, Qld, Australia. &amp; Department of Zoology, University of the Free State, Private Bag X 13, Phuthaditjhaba, 9866, South Africa. Email: CunninghamMJ @ qwa. ufs. ac. za.</mods:affiliation>
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<mods:namePart id="136CE69D822EF1ECCFCDFD64D3544EC0">Mcdonald, Keith</mods:namePart>
<mods:affiliation id="59D8208E2E659B221E6A3E84775B736F">Queensland Department of Environment and Heritage, Wet Tropics District Office, P. O. Box 834, Atherton, Australia, 4883. Email: keith. mcdonald @ epa. qld. gov. au.</mods:affiliation>
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<mods:namePart id="E26B1348065AAFCC294FDF1767D93304">Donnellan, Stephen C.</mods:namePart>
<mods:affiliation id="CC046CA259D5840E09EB82A49C0612E6">South Australian Museum, North Terrace, Adelaide, 5000, Australia. Email: Steve. Richards @ samuseum. sa. gov. au, and Conservation International, P. O. Box 1024, Atherton, Queensland 4883, Australia. &amp; Australian Centre for Evolutionary Biology and Biodiversity, University of Adelaide, Adelaide 5005.</mods:affiliation>
<mods:nameIdentifier id="E5C8D9607DBCCC6CD79FF5782024E1F2" type="email">onnellan@samuseum.sa.gov.au</mods:nameIdentifier>
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<treatment id="03FF878FFFA0200A48B3FCD1FF0CB45F" LSID="urn:lsid:plazi:treatment:03FF878FFFA0200A48B3FCD1FF0CB45F" httpUri="http://treatment.plazi.org/id/03FF878FFFA0200A48B3FCD1FF0CB45F" lastPageNumber="42" pageId="9" pageNumber="42">
<subSubSection id="C34C6512FFA0200A48B3FCD1FDBFB371" box="[196,592,824,850]" pageId="9" pageNumber="42" type="nomenclature">
<paragraph id="8BE93699FFA0200A48B3FCD1FDBFB371" blockId="9.[151,1436,152,2007]" box="[196,592,824,850]" pageId="9" pageNumber="42">
<taxonomicName id="4C564D1AFFA0200A48B3FCD1FDA4B371" authority="Van Kampen, 1909" authorityName="Van Kampen" authorityYear="1909" box="[196,587,824,850]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="papua">
<emphasis id="B922EA8BFFA0200A48B3FCD1FEBCB372" box="[196,339,824,849]" italics="true" pageId="9" pageNumber="42">Hyla papua</emphasis>
<bibRefCitation id="EFC74B68FFA0200A492BFCD1FDA4B371" author="van Kampen, P. N." box="[348,587,824,850]" pageId="9" pageNumber="42" pagination="31 - 49" refId="ref6625" refString="van Kampen, P. N. (1909) Die Amphibienfauna von Neu-Guinea, nach der Ausbeute der Neiderlandischen Sud-Neu- Guinea-Expeditionen von 1904 - 1905 und 1907. Nova Guinea, 9, 31 - 49." type="journal article" year="1909">Van Kampen, 1909</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6512FFA0200A4A2EFCD1FD42B354" pageId="9" pageNumber="42" type="materials_examined">
<paragraph id="8BE93699FFA0200A4A2EFCD1FD42B354" blockId="9.[151,1436,152,2007]" pageId="9" pageNumber="42">
<materialsCitation id="3B3E3CC4FFA0200A4A2EFCD1FD47B354" pageId="9" pageNumber="42">
<typeStatus id="54ED883BFFA0200A4A2EFCD1FD20B371" box="[601,719,824,850]" pageId="9" pageNumber="42" type="holotype">Holotype</typeStatus>
: ZMA 5679, according to
<bibRefCitation id="EFC74B68FFA0200A4C61FCD1FB0AB371" author="van Tuijl, L." box="[1046,1253,824,850]" pageId="9" pageNumber="42" pagination="125 - 141" refId="ref6663" refString="van Tuijl, L. (1995) Revised catalogue of the type specimens of recent amphibians and reptiles in the Zoologisch Museum, University of Amsterdam, The Netherlands. Bulletin of the Zoologisch Museum of the University of Amsterdam, 14, 125 - 141." type="journal article" year="1995">Van Tuijl (1995)</bibRefCitation>
. Type locality:
<location id="8E896042FFA0200A48E0FCB4FEA7B354" LSID="urn:lsid:plazi:treatment:03FF878FFFA0200A48B3FCD1FF0CB45F:8E896042FFA0200A48E0FCB4FEA7B354" box="[151,328,861,887]" country="Indonesia" municipality="The Lorentz River" name="Lorentz River" pageId="9" pageNumber="42" stateProvince="Papua Barat">Lorentz River</location>
,
<collectingRegion id="4992F87BFFA0200A4921FCB4FDF2B354" box="[342,541,861,887]" country="Indonesia" name="Papua Barat" pageId="9" pageNumber="42">Papua Province</collectingRegion>
,
<collectingCountry id="F3417609FFA0200A4A5CFCB4FD47B354" box="[555,680,861,887]" name="Indonesia" pageId="9" pageNumber="42">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C6512FFA0200A4AC1FCB4FF0CB45F" pageId="9" pageNumber="42" type="discussion">
<paragraph id="8BE93699FFA0200A4AC1FCB4FF0CB45F" blockId="9.[151,1436,152,2007]" pageId="9" pageNumber="42">
This name is currently synonymised with
<taxonomicName id="4C564D1AFFA0200A4CBFFCB4FA7BB355" authorityName="Horst" authorityYear="1883" box="[1224,1428,861,886]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A4CBFFCB4FA7BB355" box="[1224,1428,861,886]" italics="true" pageId="9" pageNumber="42">L. genimaculata</emphasis>
</taxonomicName>
. However re-examination of the
<typeStatus id="54ED883BFFA0200A4A63FC6BFD91B3BF" box="[532,638,898,924]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
, and collection of fresh material in the vicinity of the type locality, indicate that this species is distinct from
<taxonomicName id="4C564D1AFFA0200A4AF5FC41FCACB3E2" authorityName="Horst" authorityYear="1883" box="[642,835,936,961]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A4AF5FC41FCACB3E2" box="[642,835,936,961]" italics="true" pageId="9" pageNumber="42">L. genimaculata</emphasis>
</taxonomicName>
and that the name
<taxonomicName id="4C564D1AFFA0200A4C5EFC41FB7BB3E2" authorityName="Van Kampen" authorityYear="1909" box="[1065,1172,936,961]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="papua">
<emphasis id="B922EA8BFFA0200A4C5EFC41FB7BB3E2" box="[1065,1172,936,961]" italics="true" pageId="9" pageNumber="42">L. papua</emphasis>
</taxonomicName>
should be resurrected from its synonymy.
<collectingMunicipality id="6B8DACE3FFA0200A49F2FC24FDB0B3C4" box="[389,607,973,999]" pageId="9" pageNumber="42">The Lorentz River</collectingMunicipality>
population is clearly distinguished from
<taxonomicName id="4C564D1AFFA0200A4C3EFC24FAE7B3C5" authorityName="Horst" authorityYear="1883" box="[1097,1288,973,999]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="genimaculata">
<emphasis id="B922EA8BFFA0200A4C3EFC24FBB6B3C5" box="[1097,1113,973,998]" italics="true" pageId="9" pageNumber="42">L</emphasis>
.
<emphasis id="B922EA8BFFA0200A4C1FFC24FAE7B3C5" box="[1128,1288,973,998]" italics="true" pageId="9" pageNumber="42">genimaculata</emphasis>
</taxonomicName>
by its much smaller size, adult males 30.133.6 mm SVL (n = 8,
<typeStatus id="54ED883BFFA0200A4B7EFC1BFC80B42F" box="[777,879,1010,1036]" pageId="9" pageNumber="42" type="holotype">holotype</typeStatus>
= 30.5 mm). A re-description of this species is in preparation but, here, we refrain from resurrecting the name because it is currently preoccupied by
<taxonomicName id="4C564D1AFFA0200A48E0FBD4FEC9B475" authorityName="Stejneger" authorityYear="1916" box="[151,294,1085,1110]" class="Amphibia" family="Pelodryadidae" genus="Nyctimystes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B922EA8BFFA0200A48E0FBD4FEC9B475" box="[151,294,1085,1110]" italics="true" pageId="9" pageNumber="42">Nyctimystes</emphasis>
</taxonomicName>
[=
<taxonomicName id="4C564D1AFFA0200A493EFBD4FE73B475" box="[329,412,1085,1110]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B922EA8BFFA0200A493EFBD4FE73B475" box="[329,412,1085,1110]" italics="true" pageId="9" pageNumber="42">Litoria</emphasis>
</taxonomicName>
]
<taxonomicName id="4C564D1AFFA0200A49D9FBD4FD7AB474" authority="(Boulenger)" baseAuthorityName="Boulenger" box="[430,661,1085,1111]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="species" species="papua">
<emphasis id="B922EA8BFFA0200A49D9FBD4FE17B475" box="[430,504,1085,1110]" italics="true" pageId="9" pageNumber="42">papua</emphasis>
(Boulenger)
</taxonomicName>
, due to the synonymy of
<taxonomicName id="4C564D1AFFA0200A4BBFFBD4FBB9B475" authorityName="Stejneger" authorityYear="1916" box="[968,1110,1085,1110]" class="Amphibia" family="Pelodryadidae" genus="Nyctimystes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B922EA8BFFA0200A4BBFFBD4FBB9B475" box="[968,1110,1085,1110]" italics="true" pageId="9" pageNumber="42">Nyctimystes</emphasis>
</taxonomicName>
within
<taxonomicName id="4C564D1AFFA0200A4CC3FBD4FF31B45F" authority="(Frost et al. 2006)" baseAuthorityName="Frost" baseAuthorityYear="2006" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="9" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B922EA8BFFA0200A4CC3FBD4FAE8B475" box="[1204,1287,1085,1110]" italics="true" pageId="9" pageNumber="42">Litoria</emphasis>
(
<bibRefCitation id="EFC74B68FFA0200A4D60FBD4FF38B45F" author="Frost &amp; Grant, T. &amp; Faivovich, J. &amp; Bain, R. &amp; Haas, A. &amp; Haddad, C. F. B. &amp; de SA, R. O. &amp; Channing, A. &amp; Wilkinson, M. &amp; Donnellan, S. C. &amp; Raxworthy, C. &amp; Campbell, J. A. &amp; Blotto, B. L. &amp; Moler, P. &amp; Drewes, R. &amp; Nussbaum, R. A. &amp; Lynch, J. D. &amp; Green, D. &amp; Wheeler, W." pageId="9" pageNumber="42" refId="ref5389" refString="Frost, D. R., Grant, T., Faivovich, J., Bain, R., Haas, A., Haddad, C. F. B., de SA, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R., Nussbaum, R. A., Lynch, J. D., Green, D. &amp; Wheeler, W. (2006) The amphibian tree of life. Bulletin of the American Museum of Natural History, 297, 370 pages." type="book" year="2006">
Frost
<emphasis id="B922EA8BFFA0200A4D28FBD4FA7BB475" box="[1375,1428,1085,1110]" italics="true" pageId="9" pageNumber="42">et al</emphasis>
. 2006
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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