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<document id="E423B8A9918C986C90F32ECAA0D9817B" ID-DOI="10.5281/zenodo.4508115" ID-ISSN="2345-7600" ID-Zenodo-Dep="4508115" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612542939038" checkinUser="carolina" docAuthor="Tumpeesuwan, Chanidaporn, Naggs, Fred &amp; Panha, Somsak" docDate="2007" docId="03F44727FF9F83069C9EF9E17AADB6FF" docLanguage="en" docName="RafflesBZool.55.2.363-369.pdf" docOrigin="Raffles Bulletin of Zoology 55 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Phuphania globosa Tumpeesuwan &amp; Naggs &amp; Panha 2007, new species" docType="treatment" docVersion="4" lastPageNumber="365" masterDocId="FFCD3F5FFF9E83049F27FFF37E2BB227" masterDocTitle="A New Genus And New Species Of Dyakiid Snail (Pulmonata: Dyakiidae) From The Phu Phan Range, Northeastern Thailand" masterLastPageNumber="369" masterPageNumber="363" pageNumber="364" updateTime="1698867500997" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5CDCD1F26B0C5532651CC709A2D4F0B0">A New Genus And New Species Of Dyakiid Snail (Pulmonata: Dyakiidae) From The Phu Phan Range, Northeastern Thailand</mods:title>
</mods:titleInfo>
<mods:name id="8D7B0B947A1B78EBEB6A8AFB5A734203" type="personal">
<mods:role id="C34A2B85FA49B99551D622E7886617A9">
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<mods:namePart id="24C7D0A055A00795279DFD08E6370657">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:role id="0570054A018908575729920969752999">
<mods:roleTerm id="D97660A78CA0805CB3DBE8F3FC3F0A57">Author</mods:roleTerm>
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<mods:namePart id="283D6EAC80FD08D735D2CB0E195CBD35">Naggs, Fred</mods:namePart>
</mods:name>
<mods:name id="8E144E6F1F4AAE158D18373F20BAE4B6" type="personal">
<mods:role id="416FB4C3E8A9209544BACE81FAA5CA83">
<mods:roleTerm id="9BAC4BE633907807A62E74528C5BD80F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65B334B5DFEFB37ECBE487BCAA08528B">Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource id="15C66E03A212A42A5AD3FF8AD90D4AA1">text</mods:typeOfResource>
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<mods:title id="B01D4B1A274B0B1F9BB5D7CBE69262DF">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="6071991D1C472F8A636A6240833FC337">
<mods:date id="7FE5E7ACB27A98E506043757DACB9443">2007</mods:date>
<mods:detail id="45618834BBF1D07CF290A2FCEE4FE076" type="pubDate">
<mods:number id="853B33809649E6AA7DC53DA9B1D62776">2007-08-31</mods:number>
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<mods:number id="C4B1B50858F1E3AF5C3AA8BDA3BD153B">55</mods:number>
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<mods:identifier id="370150C95C8EE4996E1D1589C16FAB59" type="DOI">10.5281/zenodo.4508115</mods:identifier>
<mods:identifier id="DA1F2BE68CD605014223FBAB39243D09" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="183BFE14C6912DBB7D82D672DE459FB0" type="Zenodo-Dep">4508115</mods:identifier>
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<treatment id="03F44727FF9F83069C9EF9E17AADB6FF" LSID="urn:lsid:plazi:treatment:03F44727FF9F83069C9EF9E17AADB6FF" httpUri="http://treatment.plazi.org/id/03F44727FF9F83069C9EF9E17AADB6FF" lastPageId="2" lastPageNumber="365" pageId="1" pageNumber="364">
<subSubSection id="C347A5BAFF9F83059C9EF9E17A82B46D" pageId="1" pageNumber="364" type="nomenclature">
<paragraph id="8BE2F631FF9F83059C9EF9E17A82B46D" blockId="1.[814,1453,1426,1610]" pageId="1" pageNumber="364">
<taxonomicName id="4C5D8DB2FF9F83059C9EF9E17ABAB40D" ID-CoL="6VDFK" authority="Tumpeesuwan &amp; Naggs &amp; Panha, 2007" authorityName="Tumpeesuwan &amp; Naggs &amp; Panha" authorityYear="2007" box="[953,1169,1554,1578]" class="Gastropoda" family="Dyakiidae" genus="Phuphania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="364" phylum="Mollusca" rank="species" species="globosa" status="sp. nov.">
<emphasis id="B9292A23FF9F83059C9EF9E17ABAB40D" bold="true" box="[953,1169,1554,1578]" italics="true" pageId="1" pageNumber="364">Phuphania globosa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21A9758FF9F83059B86F9E07B09B40D" box="[1185,1314,1555,1578]" pageId="1" pageNumber="364" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="1366EAB4FF9F83059B1DF9C07A8AB46D" box="[1082,1185,1587,1610]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[138,173,2065,2086]" captionStartId-1="2.[839,874,2094,2115]" captionStartId-2="4.[114,149,1521,1542]" captionStartId-3="5.[114,149,2065,2086]" captionTargetBox-0="[137,776,791,2039]" captionTargetBox-1="[837,1476,1713,2068]" captionTargetBox-2="[115,2034,687,1441]" captionTargetBox-3="[114,1451,1193,2039]" captionTargetId-0="figure-14@2.[137,776,791,2039]" captionTargetId-1="figure-38@2.[837,1476,1713,2068]" captionTargetId-2="figure-14@4.[115,2078,687,1442]" captionTargetId-3="figure-13@5.[114,1452,1193,2040]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Fig. 2. External shell shape of Phuphania globosa, new species: A, dorsal view; B, apertural view; C, ventral view." captionText-1="Fig. 3. Live adult of Phuphania globosa, new species." captionText-2="Fig. 4. Genital system of Phuphania globosa, new species: A, genital system; B, schematic drawing of genital system; C, interior structures of amatorial organ gland (amg) and amatorial organ (am)." captionText-3="Fig. 5. Radula of Phuphania globosa, new species: A, central tooth and lateral teeth; B, left lateral teeth; C, right lateral teeth; D, left marginal teeth; E, right marginal teeth." figureDoi-0="http://doi.org/10.5281/zenodo.4508121" figureDoi-1="http://doi.org/10.5281/zenodo.4508123" figureDoi-2="http://doi.org/10.5281/zenodo.4508125" figureDoi-3="http://doi.org/10.5281/zenodo.4508127" httpUri-0="https://zenodo.org/record/4508121/files/figure.png" httpUri-1="https://zenodo.org/record/4508123/files/figure.png" httpUri-2="https://zenodo.org/record/4508125/files/figure.png" httpUri-3="https://zenodo.org/record/4508127/files/figure.png" pageId="1" pageNumber="364">Figs. 25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C347A5BAFF9F83059C09F9837BA1B5A5" pageId="1" pageNumber="364" type="materials_examined">
<paragraph id="8BE2F631FF9F83059C09F9837B29B49A" blockId="1.[814,1454,1648,1725]" pageId="1" pageNumber="364">
<emphasis id="B9292A23FF9F83059C09F9837A27B4A2" bold="true" box="[814,1036,1648,1669]" italics="true" pageId="1" pageNumber="364">Material examined. </emphasis>
<materialsCitation id="3B35FC6CFF9F83059B32F9837B29B49A" collectionCode="CUMZ" country="Thailand" latitude="17.416666" location="Ban Dong Kham Pho" longLatPrecision="1279" longitude="103.45" municipality="Waritchaphum District" pageId="1" pageNumber="364" specimenCode="CUMZ 001760" specimenCount="1" stateProvince="Sakon Nakhon" typeStatus="holotype">
<typeStatus id="54E64893FF9F83059B32F9837A5FB4A2" box="[1045,1140,1648,1669]" pageId="1" pageNumber="364" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBFB5E4AFF9F83059BA7F9837B36B4A2" box="[1152,1309,1648,1669]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="364">CUMZ 001760</specimenCode>
; type locality:
<location id="8E82A0EAFF9F83059C09F97F7DD6B486" LSID="urn:lsid:plazi:treatment:03F44727FF9F83069C9EF9E17AADB6FF:8E82A0EAFF9F83059C09F97F7DD6B486" box="[814,1021,1676,1697]" country="Thailand" latitude="17.416666" longLatPrecision="1279" longitude="103.45" municipality="Waritchaphum District" name="Ban Dong Kham Pho" pageId="1" pageNumber="364" stateProvince="Sakon Nakhon">Ban Dong Kham Pho</location>
(
<geoCoordinate id="EE6990F6FF9F83059B2CF97F7A72B486" box="[1035,1113,1676,1697]" degrees="17" direction="north" minutes="25" orientation="latitude" pageId="1" pageNumber="364" precision="925" value="17.416666">17º25'N</geoCoordinate>
<geoCoordinate id="EE6990F6FF9F83059B47F97F7A90B486" box="[1120,1211,1676,1697]" degrees="103" direction="east" minutes="27" orientation="longitude" pageId="1" pageNumber="364" precision="925" value="103.45">103°27'E</geoCoordinate>
),
<collectingMunicipality id="6B866C4BFF9F83059BEBF97F7B80B486" box="[1228,1451,1676,1697]" pageId="1" pageNumber="364">Waritchaphum District</collectingMunicipality>
,
<collectingRegion id="499938D3FF9F83059C09F95B7A37B49A" box="[814,1052,1704,1725]" country="Thailand" name="Sakon Nakhon" pageId="1" pageNumber="364">Sakon Nakhon Province</collectingRegion>
, northeastern
<collectingCountry id="F34AB6A1FF9F83059B81F95B7AD5B49A" box="[1190,1278,1704,1725]" name="Thailand" pageId="1" pageNumber="364">Thailand</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE2F631FF9F83059C09F9137BA1B5A5" blockId="1.[814,1454,1760,1922]" pageId="1" pageNumber="364">
<materialsCitation id="3B35FC6CFF9F83059C09F9137BADB5A5" collectionCode="ZRC, NHMSU, CUMZ" latitude="16.75" location="Mukdaharn" longLatPrecision="1281" longitude="104.5" municipality="Phu Phan" pageId="1" pageNumber="364" specimenCode="CUMZ 001761, BMNH 20060750" specimenCount="6" stateProvince="Nakhon Phanom" typeStatus="paratype">
<typeStatus id="54E64893FF9F83059C09F9137DA4B4D2" box="[814,911,1760,1781]" pageId="1" pageNumber="364" type="paratype">Paratypes</typeStatus>
: nine
<typeStatus id="54E64893FF9F83059CECF9137A00B4D2" box="[971,1067,1760,1781]" pageId="1" pageNumber="364" type="paratype">paratypes</typeStatus>
:
<specimenCode id="DBFB5E4AFF9F83059B12F9137AE5B4D2" box="[1077,1230,1760,1781]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="364">CUMZ 001761</specimenCode>
; one
<typeStatus id="54E64893FF9F83059A23F9137B71B4D2" box="[1284,1370,1760,1781]" pageId="1" pageNumber="364" type="paratype">paratype</typeStatus>
:
<specimenCode id="DBFB5E4AFF9F83059A41F9137DA5B536" collectionCode="BMNH" pageId="1" pageNumber="364">BMNH 20060750</specimenCode>
and one
<typeStatus id="54E64893FF9F83059CC3F90F7A1CB536" box="[996,1079,1788,1809]" pageId="1" pageNumber="364" type="paratype">paratype</typeStatus>
in the
<collectionCode id="ED4C6EF4FF9F83059B5FF90F7A81B536" box="[1144,1194,1788,1809]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="364">ZRC</collectionCode>
. Additional
<typeStatus id="54E64893FF9F83059A3AF90F7B52B536" box="[1309,1401,1788,1809]" pageId="1" pageNumber="364" type="paratype">paratypes</typeStatus>
were collected from both the eastern and southern parts of the
<collectingMunicipality id="6B866C4BFF9F83059A75F8EB7B85B50A" box="[1362,1454,1816,1837]" pageId="1" pageNumber="364">Phu Phan</collectingMunicipality>
range through three provinces:
<collectingRegion id="499938D3FF9F83059B7FF8C77AD2B56E" box="[1112,1273,1844,1865]" country="Thailand" name="Nakhon Phanom" pageId="1" pageNumber="364">Nakhon Phanom</collectingRegion>
(east),
<location id="8E82A0EAFF9F83059A18F8C77B85B56E" LSID="urn:lsid:plazi:treatment:03F44727FF9F83069C9EF9E17AADB6FF:8E82A0EAFF9F83059A18F8C77B85B56E" box="[1343,1454,1844,1865]" latitude="16.75" longLatPrecision="1281" longitude="104.5" municipality="Phu Phan" name="Mukdaharn" pageId="1" pageNumber="364" stateProvince="Nakhon Phanom">Mukdaharn</location>
(south) and
<collectingRegion id="499938D3FF9F83059C87F8A37DC2B542" box="[928,1001,1872,1893]" country="Thailand" name="Kalasin" pageId="1" pageNumber="364">Kalasin</collectingRegion>
(south).
<collectionCode id="ED4C6EF4FF9F83059B19F8A37AB3B542" box="[1086,1176,1872,1893]" pageId="1" pageNumber="364">NHMSU</collectionCode>
: 0001 (
<specimenCount id="9D5B3DB8FF9F83059BF8F8A37B7CB542" box="[1247,1367,1872,1893]" pageId="1" pageNumber="364" type="generic">3 specimens</specimenCount>
,
<geoCoordinate id="EE6990F6FF9F83059A46F8A37B85B542" box="[1377,1454,1872,1893]" degrees="16" direction="north" minutes="25" orientation="latitude" pageId="1" pageNumber="364" precision="925" value="16.416666">16º25'N</geoCoordinate>
<geoCoordinate id="EE6990F6FF9F83059C09F89E7DA1B5A5" box="[814,906,1901,1922]" degrees="103" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="364" precision="925" value="103.333336">103°20'E</geoCoordinate>
);
<collectionCode id="ED4C6EF4FF9F83059CBAF89E7DCDB5A5" box="[925,998,1901,1922]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="1" pageNumber="364">CUMZ</collectionCode>
: 001707 (
<specimenCount id="9D5B3DB8FF9F83059B6FF89E7AE9B5A5" box="[1096,1218,1901,1922]" pageId="1" pageNumber="364" type="generic">3 specimens</specimenCount>
,
<geoCoordinate id="EE6990F6FF9F83059BE9F89E7B30B5A5" box="[1230,1307,1901,1922]" degrees="16" direction="north" minutes="45" orientation="latitude" pageId="1" pageNumber="364" precision="925" value="16.75">16º45'N</geoCoordinate>
<geoCoordinate id="EE6990F6FF9F83059A04F89E7B54B5A5" box="[1315,1407,1901,1922]" degrees="104" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="364" precision="925" value="104.5">104°30'E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C347A5BAFF9F83059C09F8587AF4B5C6" pageId="1" pageNumber="364" type="etymology">
<paragraph id="8BE2F631FF9F83059C09F8587AF4B5C6" blockId="1.[814,1454,1963,2017]" pageId="1" pageNumber="364">
<emphasis id="B9292A23FF9F83059C09F8587D96B5E4" bold="true" box="[814,957,1963,1987]" italics="true" pageId="1" pageNumber="364">Etymology. </emphasis>
From the Latin
<taxonomicName id="4C5D8DB2FF9F83059B50F8587AE0B5E5" authorityName="Tumpeesuwan &amp; Naggs &amp; Panha" authorityYear="2007" box="[1143,1227,1963,1986]" class="Gastropoda" family="Dyakiidae" genus="Phuphania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="364" phylum="Mollusca" rank="species" species="globosa">
<emphasis id="B9292A23FF9F83059B50F8587AE0B5E5" box="[1143,1227,1963,1986]" italics="true" pageId="1" pageNumber="364">globosa</emphasis>
</taxonomicName>
meaning spherical with reference to the shape of the shell.
</paragraph>
</subSubSection>
<caption id="DF22A6B9FF9F83059F55F82B7F60BA64" ID-DOI="http://doi.org/10.5281/zenodo.4508119" ID-Zenodo-Dep="4508119" httpUri="https://zenodo.org/record/4508119/files/figure.png" pageId="1" pageNumber="364" startId="1.[114,151,2008,2029]" targetBox="[115,753,1079,1983]" targetPageId="1">
<paragraph id="8BE2F631FF9F83059F55F82B7F60BA64" blockId="1.[114,753,2008,2115]" pageId="1" pageNumber="364">
Fig. 1. Map of Thailand showing type locality of
<taxonomicName id="4C5D8DB2FF9F83059DA4F82B7EE8BA2D" authority="Tumpeesuwan &amp; Naggs &amp; Panha, 2007" authorityName="Tumpeesuwan &amp; Naggs &amp; Panha" authorityYear="2007" class="Gastropoda" family="Dyakiidae" genus="Phuphania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="364" phylum="Mollusca" rank="species" species="globosa" status="sp. nov.">
<emphasis id="B9292A23FF9F83059DA4F82B7EE8BA2D" italics="true" pageId="1" pageNumber="364">Phuphania globosa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21A9758FF9F83059FF2F8067F7DBA2D" box="[213,342,2037,2058]" pageId="1" pageNumber="364" rank="species">new species</taxonomicNameLabel>
, Ban Dong Kham Pho, Waritchaphum District, Sakon Nakhon Province. The location is on the east of Udon Thani Province.
</paragraph>
</caption>
<subSubSection id="C347A5BAFF9F83059C09F7FA7D5FBA67" pageId="1" pageNumber="364" type="diagnosis">
<paragraph id="8BE2F631FF9F83059C09F7FA7D5FBA67" blockId="1.[814,1454,2057,2112]" pageId="1" pageNumber="364">
<emphasis id="B9292A23FF9F83059C09F7FA7D91BA06" bold="true" box="[814,954,2057,2081]" italics="true" pageId="1" pageNumber="364">Diagnosis. </emphasis>
Diagnosis of species as described for genus above.
</paragraph>
</subSubSection>
<subSubSection id="C347A5BAFF9C83069FADFF327AADB6FF" pageId="2" pageNumber="365" type="description">
<paragraph id="8BE2F631FF9C83069FADFF327FBEB05E" blockId="2.[138,777,193,633]" pageId="2" pageNumber="365">
<emphasis id="B9292A23FF9C83069FADFF327C1CB2FE" bold="true" box="[138,567,193,217]" italics="true" pageId="2" pageNumber="365">Description of key features of species. </emphasis>
Shell: medium size for
<taxonomicName id="4C5D8DB2FF9C83069F8AFF117F30B2DE" authorityName="Gude &amp; Woodward" authorityYear="1921" box="[173,283,226,249]" class="Gastropoda" family="Dyakiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="365" phylum="Mollusca" rank="family">Dyakiidae</taxonomicName>
, dextral, semiglobose, and thin with 5-6 slightly convex whorls. Aperture large ovate and distinctly oblique, lip neither expanded nor reflected. Reproductive anatomy: the amatorial organ is a thick-walled stoutly cylindrical sac, distally fused with a gland consisting of an internal cluster of numerous small ducts, closely connected apically to form a large cap covering the amatorial organ. Penis cylindrical almost the same size and length as free oviduct. Epiphallus short, equal to penis length, straight. Vas deferens small, one and half times penial length, entering from free oviduct and passing close to distal end of epiphallus. Gametolytic sac inflated, gametolytic duct enters near proximal part of amatorial organ (
<figureCitation id="1366EAB4FF9C83069E63FD917FA3B05E" box="[324,392,610,633]" captionStart="Fig" captionStartId="4.[114,149,1521,1542]" captionTargetBox="[115,2034,687,1441]" captionTargetId="figure-14@4.[115,2078,687,1442]" captionTargetPageId="4" captionText="Fig. 4. Genital system of Phuphania globosa, new species: A, genital system; B, schematic drawing of genital system; C, interior structures of amatorial organ gland (amg) and amatorial organ (am)." figureDoi="http://doi.org/10.5281/zenodo.4508125" httpUri="https://zenodo.org/record/4508125/files/figure.png" pageId="2" pageNumber="365">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BE2F631FF9C83069FADFD527D23B0FE" blockId="2.[138,776,673,729]" pageId="2" pageNumber="365">
<emphasis id="B9292A23FF9C83069FADFD527FB4B09E" bold="true" box="[138,415,673,697]" italics="true" pageId="2" pageNumber="365">
Description of
<typeStatus id="54E64893FF9C83069E0EFD527FA3B09E" box="[297,392,673,697]" pageId="2" pageNumber="365" type="holotype">holotype</typeStatus>
.
</emphasis>
Shell height from apex to aperture
<quantity id="4CA55BD4FF9C83069FADFD317ED5B0FE" box="[138,254,706,729]" metricMagnitude="-2" metricUnit="m" metricValue="3.408" pageId="2" pageNumber="365" unit="mm" value="34.08">34.08 mm</quantity>
, diameter
<quantity id="4CA55BD4FF9C83069E51FD317FC0B0FE" box="[374,491,706,729]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="2" pageNumber="365" unit="mm" value="28.7">28.70 mm</quantity>
, with 5¾ convex whorls,
</paragraph>
<paragraph id="8BE2F631FF9C83069C63FF327A90B3FF" blockId="2.[836,1475,193,472]" pageId="2" pageNumber="365">
dextral, semiglobose, thin, body whorl large consisting of almost ¾ of the shell width, rounded, not deflected or distinctly angulated at periphery, fulvous above, embryonic whorls smooth, subsequent whorls with fine radial growth lines. Aperture large ovate, distinctly oblique, lip neither expanded nor reflected. Umbilicus narrow (
<figureCitation id="1366EAB4FF9C83069A3DFE927B76B35F" box="[1306,1373,353,376]" captionStart="Fig" captionStartId="2.[138,173,2065,2086]" captionTargetBox="[137,776,791,2039]" captionTargetId="figure-14@2.[137,776,791,2039]" captionTargetPageId="2" captionText="Fig. 2. External shell shape of Phuphania globosa, new species: A, dorsal view; B, apertural view; C, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4508121" httpUri="https://zenodo.org/record/4508121/files/figure.png" pageId="2" pageNumber="365">Fig. 2</figureCitation>
). Ground colour of body grey, marked with black spots. Foot fringe distinct, about
<quantity id="4CA55BD4FF9C83069CC6FE527A36B39F" box="[993,1053,417,440]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="365" unit="mm" value="5.0">5 mm</quantity>
wide, foot sole undivided, caudal gland without overhanging lobe (
<figureCitation id="1366EAB4FF9C83069B4DFE327A86B3FF" box="[1130,1197,449,472]" captionStart="Fig" captionStartId="2.[839,874,2094,2115]" captionTargetBox="[837,1476,1713,2068]" captionTargetId="figure-38@2.[837,1476,1713,2068]" captionTargetPageId="2" captionText="Fig. 3. Live adult of Phuphania globosa, new species." figureDoi="http://doi.org/10.5281/zenodo.4508123" httpUri="https://zenodo.org/record/4508123/files/figure.png" pageId="2" pageNumber="365">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BE2F631FF9C83069C63FDF27A15B63F" blockId="2.[836,1475,513,1048]" pageId="2" pageNumber="365">
<emphasis id="B9292A23FF9C83069C63FDF27DD4B03E" bold="true" box="[836,1023,513,537]" italics="true" pageId="2" pageNumber="365">Genital system. </emphasis>
Vas deferens marginally longer than penis and free oviduct, entering epiphallus subapically. Epiphallus almost equal in length to that of penis and to free oviduct, curved at junction with penis. Penis somewhat clavate, proximally wider than epiphallus, distal quarter narrows to width of epiphallus. Penial retractor muscle inserts near proximal end of epiphallus. Amatorial organ a stoutly cylindrical sac, inner surface of its prepuce with circular folds, muscular papilla of amatorial organ terminates in a minute pore, without thorn. The amatorial organ gland forms an ovate cap on the amatorial organ and consists of numerous ducts, leading from secretory cells lining the distal walls, the papilla forms a distal cap. Free oviduct rather long; a short wide duct proximal to the base of the amotorial organ prepuce expands into a large, elongate gametolytic sac, which possesses an apical ligament that attaches to the prostate gland (
<figureCitation id="1366EAB4FF9C83069CCAFBF27A1AB63F" box="[1005,1073,1025,1048]" captionStart="Fig" captionStartId="4.[114,149,1521,1542]" captionTargetBox="[115,2034,687,1441]" captionTargetId="figure-14@4.[115,2078,687,1442]" captionTargetPageId="4" captionText="Fig. 4. Genital system of Phuphania globosa, new species: A, genital system; B, schematic drawing of genital system; C, interior structures of amatorial organ gland (amg) and amatorial organ (am)." figureDoi="http://doi.org/10.5281/zenodo.4508125" httpUri="https://zenodo.org/record/4508125/files/figure.png" pageId="2" pageNumber="365">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BE2F631FF9C83069C63FBB27AADB6FF" blockId="2.[836,1475,1089,1240]" pageId="2" pageNumber="365">
<emphasis id="B9292A23FF9C83069C63FBB27D9EB67E" bold="true" box="[836,949,1089,1113]" italics="true" pageId="2" pageNumber="365">Radula. </emphasis>
Central tooth indistinct tricuspid, narrow and elongate, the first and the second admedians tricuspid, narrow and elongate. Lateral teeth bicuspid with tiny ectocone, narrow and elongate. Marginal teeth narrow and elongate without lateral cusps (
<figureCitation id="1366EAB4FF9C83069B11FB327A53B6FF" box="[1078,1144,1217,1240]" captionStart="Fig" captionStartId="5.[114,149,2065,2086]" captionTargetBox="[114,1451,1193,2039]" captionTargetId="figure-13@5.[114,1452,1193,2040]" captionTargetPageId="5" captionText="Fig. 5. Radula of Phuphania globosa, new species: A, central tooth and lateral teeth; B, left lateral teeth; C, right lateral teeth; D, left marginal teeth; E, right marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.4508127" httpUri="https://zenodo.org/record/4508127/files/figure.png" pageId="2" pageNumber="365">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>