312 lines
40 KiB
XML
312 lines
40 KiB
XML
<document id="7A45F1DE8923F3C13E6FBA260ED4BEAB" ID-CLB-Dataset="302901" ID-DOI="10.11646/zootaxa.5506.1.11" ID-GBIF-Dataset="0ee6b947-49f7-49a8-adb9-7c502f9e7f81" ID-ISSN="1175-5326" ID-Zenodo-Dep="13746092" ID-ZooBank="4BD20A0F-1A6E-4389-82E0-18784A01438A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052428659" checkinUser="plazi" docAuthor="Borovec, Roman & Anderson, Robert" docDate="2024" docId="03DC87FAFFB1FF86FF17FF16D488F9D4" docLanguage="en" docName="zootaxa.5506.1.11.pdf" docOrigin="Zootaxa 5506 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5506.1.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laohajekia uniformis Borovec and Anderson 2024, n. sp." docType="treatment" docUuid="38A02A43-458B-4B41-907B-67A0D2C71B71" docUuidSource="ZooBank" docVersion="2" lastPageNumber="148" masterDocId="FFE5FF82FFB0FF85FF80FFC9D014FF85" masterDocTitle="Two new Trachyphloeini (Coleoptera; Curculionidae; Entiminae) from Laos" masterLastPageNumber="150" masterPageNumber="145" pageNumber="146" updateTime="1726053369622" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E7E1F81132C3486285AA947B9BD51B2D">Two new Trachyphloeini (Coleoptera; Curculionidae; Entiminae) from Laos</mods:title>
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<mods:name id="2D78A3344616B8994373EF4996A00AA7" type="personal">
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<mods:namePart id="26F7E7C7C63D04F4F47C5AD1EA08863A">Borovec, Roman</mods:namePart>
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<mods:affiliation id="D412BD01CC288211DC5649C6B57052C3">Sloupno 64, 503 53 Smidary, Czech Republic.</mods:affiliation>
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<mods:namePart id="5A1722C8B7E331168E669676F075EBC5">Anderson, Robert</mods:namePart>
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<mods:affiliation id="6CE8F8164BD639A03F8E67337CD9AC64">Research and Collection Division, Canadian Museum of Nature, 1740 Chemin Pink, Gatineau, PQ, J 9 J 3 N 7, Canada.</mods:affiliation>
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<mods:title id="2F5EF7FD7E06A0A82FE538A0E3F9C5B0">Zootaxa</mods:title>
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<mods:part id="AC859D4549EE941DE5A233D8ED321EBA">
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<mods:date id="F591A0C1FB7177E43461C7BD2DBF00C3">2024</mods:date>
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<mods:number id="2E4626E00F9EA36D8AC1A3D9AA0AF86C">2024-09-05</mods:number>
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<emphasis id="B901EAFEFFB1FF84FF17FF16D318FF7C" bold="true" box="[151,780,223,250]" pageId="1" pageNumber="146">
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<taxonomicName id="4C754D6FFFB1FF84FF17FF16D2D6FF7C" authority="Borovec and Anderson" authorityName="Borovec and Anderson" authorityYear="2024" box="[151,706,223,250]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="uniformis" status="sp. nov.">
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<emphasis id="B901EAFEFFB1FF84FF17FF16D18AFF7C" bold="true" box="[151,414,223,250]" italics="true" pageId="1" pageNumber="146">Laohajekia uniformis</emphasis>
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Borovec and Anderson
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<taxonomicNameLabel id="A2325785FFB1FF84FD4AFF16D318FF7C" box="[714,780,223,249]" pageId="1" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C36F6567FFB1FF84FF17FEEED157FEC7" box="[151,323,295,322]" pageId="1" pageNumber="146" type="description">
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<paragraph id="8BCA36ECFFB1FF84FF17FEEED157FEC7" blockId="1.[151,965,223,322]" box="[151,323,295,322]" pageId="1" pageNumber="146">
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(
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<figureCitation id="134E2A69FFB1FF84FF1FFEEED110FEC4" box="[159,260,295,321]" captionStart="FIGURES 1–4" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 1–4. Dorsal and lateral habitus. 1–2. Laohajekia uniformis. 3–4. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="1" pageNumber="146">Figs 1, 2</figureCitation>
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,
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<figureCitation id="134E2A69FFB1FF84FE90FEEED12FFEC7" box="[272,315,295,322]" captionStart="FIGURES 5–13" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 5–13. Structural details. 5–8. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 9–13. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="1" pageNumber="146">5–8</figureCitation>
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)
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</paragraph>
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<subSubSection id="C36F6567FFB1FF86FF17FEA2D504FD00" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146" type="materials_examined">
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<paragraph id="8BCA36ECFFB1FF84FF17FEA2D59DFE48" blockId="1.[151,1437,363,750]" pageId="1" pageNumber="146">
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<emphasis id="B901EAFEFFB1FF84FF17FEA2D151FE00" bold="true" box="[151,325,363,389]" pageId="1" pageNumber="146">Type material.</emphasis>
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<materialsCitation id="3B1D3CB1FFB1FF84FECCFEA2D37FFE48" collectingDate="2008-05-28" collectionCode="FMNH" collectorName="A. Newton & M. Thayer" country="Laos" elevation="820" latitude="18.352467" location="Phou Khao Khouay" longLatPrecision="1" longitude="102.81115" pageId="1" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Viangchan" typeStatus="holotype">
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<typeStatus id="54CE884EFFB1FF84FECCFEA2D1ADFE00" box="[332,441,363,389]" pageId="1" pageNumber="146" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D73FD65FFB1FF84FE43FEA5D1CEFE03" box="[451,474,364,390]" count="1" pageId="1" pageNumber="146" type="male">♂</specimenCount>
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,
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<collectingCountry id="F362767CFFB1FF84FE66FEA5D20BFE03" box="[486,543,364,390]" name="Laos" pageId="1" pageNumber="146">Laos</collectingCountry>
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,
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<collectingRegion id="49B1F80EFFB1FF84FDAAFEA2D316FE00" box="[554,770,363,389]" country="Laos" name="Viangchan" pageId="1" pageNumber="146">Vientiane Province</collectingRegion>
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,
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<location id="8EAA6037FFB1FF84FC8DFEA2D3FEFE00" LSID="urn:lsid:plazi:treatment:03DC87FAFFB1FF86FF17FF16D488F9D4:8EAA6037FFB1FF84FC8DFEA2D3FEFE00" box="[781,1002,363,389]" country="Laos" latitude="18.352467" longLatPrecision="1" longitude="102.81115" name="Phou Khao Khouay" pageId="1" pageNumber="146" stateProvince="Viangchan">Phou Khao Khouay</location>
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NBCA,
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<locationDeviation id="9A782FCEFFB1FF84FBCDFEA5D53CFE00" box="[1101,1320,363,390]" location="Ban Vangheua" pageId="1" pageNumber="146">N of Ban Vangheua</locationDeviation>
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(= Khua),
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<quantity id="4C8D9B09FFB1FF84FF17FE46D0F4FE2F" box="[151,224,399,426]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="146" unit="m" value="820.0">
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<elevation id="0058D1DFFFB1FF84FF17FE46D0F4FE2F" box="[151,224,399,426]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="146" unit="m" value="820.0">820 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE41502BFFB1FF84FF6CFE46D16AFE2F" box="[236,382,399,426]" degrees="18" direction="north" minutes="21.148" orientation="latitude" pageId="1" pageNumber="146" precision="1" value="18.352467">18°21.148 N</geoCoordinate>
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,
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<geoCoordinate id="EE41502BFFB1FF84FE09FE46D231FE2F" box="[393,549,399,426]" degrees="102" direction="east" minutes="48.669" orientation="longitude" pageId="1" pageNumber="146" precision="1" value="102.81115">102°48.669 E</geoCoordinate>
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,
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<date id="FFCB102CFFB1FF84FDB1FE46D2B6FE2C" box="[561,674,399,425]" pageId="1" pageNumber="146" value="2008-05-28">
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<collectingDate id="EF8FE9C4FFB1FF84FDB1FE46D2B6FE2C" box="[561,674,399,425]" pageId="1" pageNumber="146" value="2008-05-28">28.v.2008</collectingDate>
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</date>
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, old-growth tropical hardwood forest, FMHD2008-020; berlese, log & leaf litter,
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<collectorName id="2680533AFFB1FF84FEA5FE7AD1B4FE4B" box="[293,416,435,462]" pageId="1" pageNumber="146">A. Newton</collectorName>
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&
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<collectorName id="2680533AFFB1FF84FE44FE7AD221FE48" box="[452,565,435,461]" pageId="1" pageNumber="146">M.Thayer</collectorName>
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, ANMT site 1222 (
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<collectionCode id="ED64AE29FFB1FF84FC8CFE7AD377FE48" box="[780,867,435,461]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="146" type="Museum">FMNH</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B1D3CB1FFB1FF84FCF6FE7DD591FE48" box="[886,1413,435,462]" collectingDate="2008-05-28" collectionCode="CMNC" collectorName="A. Newton & M. Thayer" country="Laos" elevation="820" latitude="18.352467" location="Phou Khao Khouay" longLatPrecision="1" longitude="102.81115" pageId="1" pageNumber="146" specimenCount="1" specimenCount-female="1" stateProvince="Viangchan" typeStatus="paratype">
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<typeStatus id="54CE884EFFB1FF84FCF6FE7DD3C8FE4B" box="[886,988,436,462]" pageId="1" pageNumber="146" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="9D73FD65FFB1FF84FC66FE7AD404FE4B" box="[998,1040,435,462]" count="1" pageId="1" pageNumber="146" type="female">1 ♀</specimenCount>
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||
, same data as holotype (
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<collectionCode id="ED64AE29FFB1FF84FAA4FE7AD569FE48" box="[1316,1405,435,461]" pageId="1" pageNumber="146">CMNC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BCA36ECFFB1FF84FF17FE1ED343FD6B" blockId="1.[151,1437,363,750]" pageId="1" pageNumber="146">
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Body length:
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<typeStatus id="54CE884EFFB1FF84FEB0FE1ED186FE74" box="[304,402,471,497]" pageId="1" pageNumber="146" type="holotype">holotype</typeStatus>
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<quantity id="4C8D9B09FFB1FF84FE19FE1ED1EBFE77" box="[409,511,471,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" pageId="1" pageNumber="146" unit="mm" value="2.06">2.06 mm</quantity>
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,
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<typeStatus id="54CE884EFFB1FF84FD8AFE11D27EFE77" box="[522,618,472,498]" pageId="1" pageNumber="146" type="paratype">paratype</typeStatus>
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<quantity id="4C8D9B09FFB1FF84FDF1FE1ED2CCFE77" box="[625,728,471,498]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="1" pageNumber="146" unit="mm" value="2.13">2.13 mm</quantity>
|
||
. Body (
|
||
<figureCitation id="134E2A69FFB1FF84FCAEFE1ED386FE74" box="[814,914,471,497]" captionStart="FIGURES 1–4" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 1–4. Dorsal and lateral habitus. 1–2. Laohajekia uniformis. 3–4. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="1" pageNumber="146">Figs 1, 2</figureCitation>
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) dark brownish, basal parts of scapes, funicles with clubs and tarsi slightly paler, reddish brown.Appressed scales on dorsal part of body star-shaped, not completely covering integument, 2–3 across width of one interstria. Elytra with regular row of erect, narrow, long, sparse setae, slightly longer at posterior declivity than on the disc; setae apically rounded, widest at middle, at posterior declivity almost as long as the width of each interstria, distance between two setae 2–3 longer than length of one seta.Anterior margin of pronotum, head with rostrum with similar erect setae, slightly shorter than those on elytra. Antennal scapes, femora and tibiae with same appressed scales and erect setae as body, funicles with tarsi with semi-erect piliform setae, clubs densely setose with short appressed setae.
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</paragraph>
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<caption id="DF0A6664FFB1FF84FF18F9F3D4EEF9D7" ID-DOI="http://doi.org/10.5281/zenodo.13746095" ID-Zenodo-Dep="13746095" box="[152,1274,1594,1618]" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="1" pageNumber="146" startId="1.[152,270,1594,1618]" targetBox="[154,1436,771,1548]" targetPageId="1" targetType="figure">
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<paragraph id="8BCA36ECFFB1FF84FF18F9F3D4EEF9D7" blockId="1.[152,1274,1594,1618]" box="[152,1274,1594,1618]" pageId="1" pageNumber="146">
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<emphasis id="B901EAFEFFB1FF84FF18F9F3D155F9D7" bold="true" box="[152,321,1594,1618]" pageId="1" pageNumber="146">FIGURES 1–4.</emphasis>
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Dorsal and lateral habitus. 1–2.
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<taxonomicName id="4C754D6FFFB1FF84FD0AF9F3D377F9D7" authorityName="Borovec and Anderson" authorityYear="2024" box="[650,867,1594,1618]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="uniformis">
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||
<emphasis id="B901EAFEFFB1FF84FD0AF9F3D377F9D7" box="[650,867,1594,1618]" italics="true" pageId="1" pageNumber="146">Laohajekia uniformis</emphasis>
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||
</taxonomicName>
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||
. 3–4.
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||
<taxonomicName id="4C754D6FFFB1FF84FC23F9F3D4E0F9D7" authorityName="Borovec and Anderson" authorityYear="2024" box="[931,1268,1594,1618]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="newthayorum">
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||
<emphasis id="B901EAFEFFB1FF84FC23F9F3D4E0F9D7" box="[931,1268,1594,1618]" italics="true" pageId="1" pageNumber="146">Trachyphloeosoma newthayorum</emphasis>
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</taxonomicName>
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.
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||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA36ECFFB1FF84FF47F954D529F80B" blockId="1.[151,1437,1692,2043]" pageId="1" pageNumber="146">
|
||
Rostrum (
|
||
<figureCitation id="134E2A69FFB1FF84FEB6F955D183F933" box="[310,407,1692,1719]" captionStart="FIGURES 5–13" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 5–13. Structural details. 5–8. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 9–13. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="1" pageNumber="146">Figs 5, 6</figureCitation>
|
||
) short and wide, 1.17–1.19× as wide as long, widest shortly before base, then slightly evenly tapering apicad with straight sides, apically broadly rounded; in lateral view regularly vaulted. Epifrons wide and flat, widest at base, distinctly evenly tapering apicad with straight sides, at base 1.36–1.41× as wide as at apex. Frons densely squamosa. Epistome small, U-shaped, glabrous, posteriorly faintly carinated. Antennal scrobes dorsally visible in anterior part of rostrum as narrowly reniform; in lateral view narrow, furrow-shaped, short, slightly curved, directed towards eye, separated from it by wide squamosa stripe. Eyes small, in dorsal view slightly prominent from outline of head; in lateral view distant from dorsal margin of head equal to eye diameter. Vertex flat, wide.
|
||
</paragraph>
|
||
<paragraph id="8BCA36ECFFB1FF87FF47F850D1ECFF50" blockId="1.[151,1437,1692,2043]" lastBlockId="2.[151,1437,150,357]" lastPageId="2" lastPageNumber="147" pageId="1" pageNumber="146">Antennae somewhat slender; scapes straight, evenly slightly enlarging apicad, 4.67–4.73× as long as at apex wide, at apex 0.8× as wide as club. Funicle of 7 antennomeres. Antennomeres 1 and 2 subequal in length, conical; antennomere 1 1.5–1.6× as long as wide; antennomere 2 1.7–1.8× as long as wide; antennomere 3 1.1× as wide as long; antennomere 4 isodiametric; antennomeres 5 and 6 1.1× as wide as long; antennomere 7 1.2× as wide as long; clubs 1.7–1.8× as long as wide.</paragraph>
|
||
<paragraph id="8BCA36ECFFB2FF87FF47FF16D487FEE0" blockId="2.[151,1437,150,357]" pageId="2" pageNumber="147">
|
||
Pronotum (
|
||
<figureCitation id="134E2A69FFB2FF87FEC5FF16D1BDFF7C" box="[325,425,223,249]" captionStart="FIGURES 1–4" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 1–4. Dorsal and lateral habitus. 1–2. Laohajekia uniformis. 3–4. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="2" pageNumber="147">Figs 1, 2</figureCitation>
|
||
) slender, 1.42–1.47× as wide as long, widest at midlength with slightly regularly rounded sides, equally narrowing anteriad and posteriad with anterior margin indistinctly narrower than posterior one. Disc weakly regularly domed, in lateral view regularly convex. Integument under appressed scales roughly densely punctured, punctures identical to those in elytral rows, distance between them very narrow.
|
||
</paragraph>
|
||
<caption id="DF0A6664FFB2FF87FF18F9BDD370F97D" ID-DOI="http://doi.org/10.5281/zenodo.13746097" ID-Zenodo-Dep="13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="2" pageNumber="147" startId="2.[152,270,1652,1676]" targetBox="[176,1410,392,1615]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BCA36ECFFB2FF87FF18F9BDD370F97D" blockId="2.[151,1436,1652,1784]" pageId="2" pageNumber="147">
|
||
<emphasis id="B901EAFEFFB2FF87FF18F9BDD15AF909" bold="true" box="[152,334,1652,1676]" pageId="2" pageNumber="147">FIGURES 5–13.</emphasis>
|
||
Structural details. 5–8.
|
||
<taxonomicName id="4C754D6FFFB2FF87FDBFF9BDD30DF909" authorityName="Borovec and Anderson" authorityYear="2024" box="[575,793,1652,1676]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="uniformis">
|
||
<emphasis id="B901EAFEFFB2FF87FDBFF9BDD30DF909" box="[575,793,1652,1676]" italics="true" pageId="2" pageNumber="147">Laohajekia uniformis</emphasis>
|
||
</taxonomicName>
|
||
. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 9–13.
|
||
<taxonomicName id="4C754D6FFFB2FF87FC9BF951D47AF935" authorityName="Borovec and Anderson" authorityYear="2024" box="[795,1134,1688,1712]" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="newthayorum">
|
||
<emphasis id="B901EAFEFFB2FF87FC9BF951D47AF935" box="[795,1134,1688,1712]" italics="true" pageId="2" pageNumber="147">Trachyphloeosoma newthayorum</emphasis>
|
||
</taxonomicName>
|
||
. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCA36ECFFB2FF87FF47F883D1A9F8E1" blockId="2.[151,1436,1866,2036]" box="[199,445,1866,1893]" pageId="2" pageNumber="147">Scutellum not visible.</paragraph>
|
||
<paragraph id="8BCA36ECFFB2FF87FF47F8A7D11BF871" blockId="2.[151,1436,1866,2036]" pageId="2" pageNumber="147">
|
||
Elytra (
|
||
<figureCitation id="134E2A69FFB2FF87FE9BF8A7D194F80D" box="[283,384,1902,1928]" captionStart="FIGURES 1–4" captionStartId="1.[152,270,1594,1618]" captionTargetBox="[154,1436,771,1548]" captionTargetId="figure-218@1.[151,1436,771,1568]" captionTargetPageId="1" captionText="FIGURES 1–4. Dorsal and lateral habitus. 1–2. Laohajekia uniformis. 3–4. Trachyphloeosoma newthayorum." figureDoi="http://doi.org/10.5281/zenodo.13746095" httpUri="https://zenodo.org/record/13746095/files/figure.png" pageId="2" pageNumber="147">Figs 1, 2</figureCitation>
|
||
) in male subglobular, 1.21× as long as wide, in female oval, 1.28× as long as wide, widest at midlength, with distinctly regularly rounded sides, apically broadly rounded; in lateral view distinctly vaulted. Base equally wide as base of pronotum. Striae distinctly roughly punctured, about twice as wide as narrow, equally flat interstriae.
|
||
</paragraph>
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FF5ED4F3FE98" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">All femora edentate. Tibiae robust, mucronate, not denticulate. Protibiae 4.27–4.31× as long as at midlength wide, with lateral margin straight and mesal margin bisinuate, apically obliquely subtruncated, fringed with short yellowish setae. Tarsi short; tarsomere 2 1.1–1.2× as wide as long; tarsomere 3 1.3–1.4× as wide as long and 1.3– 1.4× as wide as tarsomere 2; onychium 0.9–1.1× as long as tarsomere 3. Claws fused in basal part.</paragraph>
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FEEED273FE28" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
Abdominal ventrites 1.03–1.07× as long as wide, densely squamose with appressed rounded scales, with metasternal process obtuse, wider than transverse diameter of metacoxa. Ventrite 1 large, in middle twice as long as ventrite 2; ventrite 2 slightly longer than ventrite 3 or 4; ventrite
|
||
<quantity id="4C8D9B09FFB3FF86FCE2FEA7D398FE0C" box="[866,908,366,393]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="148" unit="in" value="5.0">5 in</quantity>
|
||
male subtrapezoidal, apically obtuse, in female subtriangular, apically narrowly tapering.
|
||
</paragraph>
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FE7ED57AFDB8" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
Penis (
|
||
<figureCitation id="134E2A69FFB3FF86FE92FE7ED140FE54" box="[274,340,439,465]" captionStart="FIGURES 5–13" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 5–13. Structural details. 5–8. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 9–13. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="3" pageNumber="148">Fig. 7</figureCitation>
|
||
) in dorsal view widest at apical third, from base slightly evenly enlarging to this point, then slightly tapering apicad, apically broadly rounded; in lateral view distinctly regularly curved, evenly tapering apicad, tip narrow, elongate, slightly curved outside; endophallus with one short, straight sclerite; temones long, about twice as long as body of penis. Tegmen lacking parameres, its apodeme short, only faintly longer than diameter of ring.
|
||
</paragraph>
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FD8ED504FD00" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
Spermatheca (
|
||
<figureCitation id="134E2A69FFB3FF86FEEAFD8ED1BBFDE4" box="[362,431,583,609]" captionStart="FIGURES 5–13" captionStartId="2.[152,270,1652,1676]" captionTargetBox="[176,1410,392,1615]" captionTargetId="figure-108@2.[164,1423,381,1628]" captionTargetPageId="2" captionText="FIGURES 5–13. Structural details. 5–8. Laohajekia uniformis. 5. Head and rostrum, dorsal view. 6. Head and rostrum, lateral view. 7. Penis, dorsal and lateral view. 8. Spermatheca. 9–13. Trachyphloeosoma newthayorum. 9. Head and rostrum, dorsal view. 10. Head and rostrum, lateral view. 11. Penis, dorsal and lateral view. 12. Spermatheca. 13. Sternite VIII in female. Scale bars: 0.50 mm (5, 6, 9, 10, 13), 0.25 mm (7, 11) and 0.125 mm (8, 12)." figureDoi="http://doi.org/10.5281/zenodo.13746097" httpUri="https://zenodo.org/record/13746097/files/figure.png" pageId="3" pageNumber="148">Fig. 8</figureCitation>
|
||
) with slender and regularly curved cornu; ramus isodiametric; collum wider than long, equally long as ramus but conspicuously wider, regularly rounded. Female sternite VIII not examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6567FFB3FF86FF47FD46D53EFD2C" box="[199,1322,655,681]" pageId="3" pageNumber="148" type="etymology">
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FD46D53EFD2C" blockId="3.[151,1437,151,1617]" box="[199,1322,655,681]" pageId="3" pageNumber="148">
|
||
<emphasis id="B901EAFEFFB3FF86FF47FD46D15AFD2C" bold="true" box="[199,334,655,681]" pageId="3" pageNumber="148">Etymology.</emphasis>
|
||
This species is named ‘uniformis’ in recognition of the uniformly flat elytral interstriae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6567FFB3FF86FF47FD7BD4B9FC90" pageId="3" pageNumber="148" type="biology_ecology">
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FD7BD4B9FC90" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
<emphasis id="B901EAFEFFB3FF86FF47FD7BD133FD49" bold="true" box="[199,295,690,716]" pageId="3" pageNumber="148">Biology.</emphasis>
|
||
The two known specimens were sifted from forest floor litter in a middle elevation fairly large (ca.
|
||
<quantity id="4C8D9B09FFB3FF86FA0EFD7AD0AFFD74" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="148" unit="km" value="1.0">1 km</quantity>
|
||
<superScript id="7C009BA4FFB3FF86FF3BFD1FD0D7FD61" attach="left" box="[187,195,726,740]" fontSize="6" pageId="3" pageNumber="148">2</superScript>
|
||
) patch of minimally disturbed old-growth tropical hardwood forest on level or gently rolling ground. This patch was surrounded by cultivated fields or scrubby cut-over forest, with rice paddies to the south.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6567FFB3FF86FF47FCD7D50BFB24" pageId="3" pageNumber="148" type="diagnosis">
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FCD7D23DFC04" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
<emphasis id="B901EAFEFFB3FF86FF47FCD7D1DBFCBD" bold="true" box="[199,463,798,824]" pageId="3" pageNumber="148">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C754D6FFFB3FF86FE57FCD6D2DCFCBC" authorityName="Borovec and Anderson" authorityYear="2024" box="[471,712,799,825]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="uniformis">
|
||
<emphasis id="B901EAFEFFB3FF86FE57FCD6D2DCFCBC" box="[471,712,799,825]" italics="true" pageId="3" pageNumber="148">Laohajekia uniformis</emphasis>
|
||
</taxonomicName>
|
||
is very similar to the only other known species of the genus,
|
||
<taxonomicName id="4C754D6FFFB3FF86FA06FCD6D1C0FCD8" authority="Borovec" authorityName="Borovec" authorityYear="2014" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="trachyphloeiformis">
|
||
<emphasis id="B901EAFEFFB3FF86FA06FCD6D178FCD8" italics="true" pageId="3" pageNumber="148">L. trachyphloeiformis</emphasis>
|
||
Borovec
|
||
</taxonomicName>
|
||
described from
|
||
<collectingRegion id="49B1F80EFFB3FF86FD11FC8AD395FCD8" box="[657,897,835,861]" country="Laos" name="Champasak" pageId="3" pageNumber="148">Champasak province</collectingRegion>
|
||
, also in
|
||
<collectingCountry id="F362767CFFB3FF86FC63FC8AD409FCD8" box="[995,1053,835,861]" name="Laos" pageId="3" pageNumber="148">Laos</collectingCountry>
|
||
. It is possible to separate the two species by the following characters:
|
||
</paragraph>
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FC42D22DFB94" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
<taxonomicName id="4C754D6FFFB3FF86FF47FC42D1ACFC20" authorityName="Borovec and Anderson" authorityYear="2024" box="[199,440,907,933]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="uniformis">
|
||
<emphasis id="B901EAFEFFB3FF86FF47FC42D1ACFC20" box="[199,440,907,933]" italics="true" pageId="3" pageNumber="148">Laohajekia uniformis</emphasis>
|
||
</taxonomicName>
|
||
: Antennal scrobes clearly visible in anterior part of rostrum; eyes in dorsal view slightly prominent from outline of head; erect elytral setae almost as long as width of interstria at posterior declivity; all elytral interstriae flat; rostrum 1.17–1.19× as wide as long; pronotum slender, 1.42–1.47× as wide as long; scapes slender, 4.7× as long as at apex wide.
|
||
</paragraph>
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FBD2D50BFB24" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
<taxonomicName id="4C754D6FFFB3FF86FF47FBD2D235FBB0" authorityName="Borovec" authorityYear="2014" box="[199,545,1051,1077]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="trachyphloeiformis">
|
||
<emphasis id="B901EAFEFFB3FF86FF47FBD2D235FBB0" box="[199,545,1051,1077]" italics="true" pageId="3" pageNumber="148">Laohajekia trachyphloeiformis</emphasis>
|
||
</taxonomicName>
|
||
: Antennal scrobes hardly visible as short and narrow furrows in anterior part of rostrum; eyes small, in dorsal view inconspicuous; erect elytral setae about as long as half the width of interstria at posterior declivity; interstriae 2, 4 and 6 slightly elevated, all interstriae with low, scarce bumps; rostrum 1.40× as wide as long; pronotum wider, 1.51× as wide as long; scapes more robust, 3.6× as long as at apex wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6567FFB3FF86FF47FB62D488F9D4" pageId="3" pageNumber="148" type="discussion">
|
||
<paragraph id="8BCA36ECFFB3FF86FF47FB62D488F9D4" blockId="3.[151,1437,151,1617]" pageId="3" pageNumber="148">
|
||
<emphasis id="B901EAFEFFB3FF86FF47FB62D12FFB40" bold="true" box="[199,315,1195,1221]" pageId="3" pageNumber="148">Remarks.</emphasis>
|
||
The monotypic genus
|
||
<taxonomicName id="4C754D6FFFB3FF86FDBBFB62D317FB40" authority="Morimoto" authorityName="Morimoto" authorityYear="2015" box="[571,771,1195,1221]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FDBBFB62D29DFB40" box="[571,649,1195,1221]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
|
||
Morimoto
|
||
</taxonomicName>
|
||
described and known from
|
||
<collectingCountry id="F362767CFFB3FF86FBBCFB62D468FB40" box="[1084,1148,1195,1221]" name="Japan" pageId="3" pageNumber="148">Japan</collectingCountry>
|
||
following the description of
|
||
<taxonomicName id="4C754D6FFFB3FF86FF34FB06D182FB6C" authority="Borovec" authorityName="Borovec" authorityYear="2014" box="[180,406,1231,1257]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FF34FB06D126FB6C" box="[180,306,1231,1257]" italics="true" pageId="3" pageNumber="148">Laohajekia</emphasis>
|
||
Borovec
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C754D6FFFB3FF86FDDAFB06D2CCFB6C" authorityName="Borovec" authorityYear="2014" box="[602,728,1231,1257]" class="Insecta" family="Curculionidae" genus="Laohajekia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FDDAFB06D2CCFB6C" box="[602,728,1231,1257]" italics="true" pageId="3" pageNumber="148">Laohajekia</emphasis>
|
||
</taxonomicName>
|
||
. Based on its original description (
|
||
<bibRefCitation id="EFE44B1DFFB3FF86FBD6FB06D51DFB6C" author="Morimoto, K." box="[1110,1289,1230,1257]" pageId="3" pageNumber="148" refId="ref4157" refString="Morimoto, K. (2015) [new taxa] In: Morimoto, K., Nakamura, T., Kanno, K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan. Vol. 4. Touka Shobo, Fukuoka, pp. 1 - 758." type="book" year="2015">Morimoto 2015</bibRefCitation>
|
||
) it is difficult to find any distinguishing characters except the funicle of 6 antennomeres and flagellate penis in
|
||
<taxonomicName id="4C754D6FFFB3FF86FB53FB3AD535FA88" authorityName="Morimoto" authorityYear="2015" box="[1235,1313,1267,1293]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FB53FB3AD535FA88" box="[1235,1313,1267,1293]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
|
||
</taxonomicName>
|
||
. While the number of funicle antennomeres clearly distinguishes
|
||
<taxonomicName id="4C754D6FFFB3FF86FD6BFADED445FAB4" authority="Morimoto" authorityName="Morimoto" authorityYear="2015" box="[747,1105,1303,1329]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="rotundipennis">
|
||
<emphasis id="B901EAFEFFB3FF86FD6BFADED3CEFAB4" box="[747,986,1303,1329]" italics="true" pageId="3" pageNumber="148">Tokara rotundipennis</emphasis>
|
||
Morimoto
|
||
</taxonomicName>
|
||
from both
|
||
<taxonomicName id="4C754D6FFFB3FF86FB48FADED588FAB4" box="[1224,1436,1303,1329]" pageId="3" pageNumber="148">
|
||
<emphasis id="B901EAFEFFB3FF86FB48FADED552FAB4" box="[1224,1350,1303,1329]" italics="true" pageId="3" pageNumber="148">Laohajekia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F362767CFFB3FF86FF57FAF2D104FAD0" box="[215,272,1339,1365]" name="Laos" pageId="3" pageNumber="148">Laos</collectingCountry>
|
||
, it is difficult to use this as a basis for distinguishing the genus. The number of funicle antennomeres often varies in Palaearctic
|
||
<taxonomicName id="4C754D6FFFB3FF86FE5AFA96D29CFAFC" baseAuthorityName="Borovec and Anderson" baseAuthorityYear="2022" box="[474,648,1375,1401]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="tribe" tribe="Trachyphloeini">Trachyphloeini</taxonomicName>
|
||
. For example, the nominal genus
|
||
<taxonomicName id="4C754D6FFFB3FF86FBA9FA96D526FAFC" authority="Germar" authorityName="Germar" authorityYear="1817" box="[1065,1330,1375,1401]" class="Insecta" family="Curculionidae" genus="Trachyphloeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FBA9FA96D4DAFAFC" box="[1065,1230,1375,1401]" italics="true" pageId="3" pageNumber="148">Trachyphloeus</emphasis>
|
||
Germar
|
||
</taxonomicName>
|
||
contains species with funicles having 5, 6 or 7 antennomeres. Also, the existence of a flagellum varies within species of a genus. For example, species of
|
||
<taxonomicName id="4C754D6FFFB3FF86FD83FA6ED305FA44" authority="Schoenherr" authorityName="Schoenherr" authorityYear="1845" box="[515,785,1447,1473]" class="Insecta" family="Curculionidae" genus="Porophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FD83FA6ED29CFA44" box="[515,648,1447,1473]" italics="true" pageId="3" pageNumber="148">Porophorus</emphasis>
|
||
Schoenherr
|
||
</taxonomicName>
|
||
(Tanyrhynchini Schoenherr) or
|
||
<taxonomicName id="4C754D6FFFB3FF86FB01FA6ED589FA44" authority="Schoenherr" authorityName="Schoenherr" authorityYear="1847" box="[1153,1437,1447,1473]" class="Insecta" family="Curculionidae" genus="Glyptosomus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FB01FA6ED507FA44" box="[1153,1299,1447,1473]" italics="true" pageId="3" pageNumber="148">Glyptosomus</emphasis>
|
||
Schoenherr
|
||
</taxonomicName>
|
||
(Embrithini Marshall) either contain a long and conspicuous flagellum or the flagellum is completely absent. In all other characters, including male and female genitalia, which are well-illustrated in the description of
|
||
<taxonomicName id="4C754D6FFFB3FF86FA9CFA26D57EF98C" authorityName="Morimoto" authorityYear="2015" box="[1308,1386,1519,1545]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FA9CFA26D57EF98C" box="[1308,1386,1519,1545]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
|
||
</taxonomicName>
|
||
, the characters of the two genera are very similar. It is apparent both genera are very close, but without the possibility of examining material of
|
||
<taxonomicName id="4C754D6FFFB3FF86FE15F9FED1F7F9D4" authorityName="Morimoto" authorityYear="2015" box="[405,483,1591,1617]" class="Insecta" family="Curculionidae" genus="Tokara" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EAFEFFB3FF86FE15F9FED1F7F9D4" box="[405,483,1591,1617]" italics="true" pageId="3" pageNumber="148">Tokara</emphasis>
|
||
</taxonomicName>
|
||
, it is not possible to synonymize (distinguish) the two genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |