treatments-xml/data/03/DC/87/03DC87B0FFF2FF85FF5D3D70FE4486AD.xml
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<document id="0B5A4F710276D1F14CB173CD42DF3BC7" ID-DOI="10.11646/zootaxa.1522.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5087760" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625852368432" checkinUser="felipe" docAuthor="KNEBELSBERGER, THOMAS &amp; MILLER, MICHAEL A." docDate="2007" docId="03DC87B0FFF2FF85FF5D3D70FE4486AD" docLanguage="en" docName="zt01522p068.pdf.imf" docOrigin="Zootaxa 1522 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.1522.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Phyllodromica quadracantha Published 2007, spec. nov." docType="treatment" docUuid="79FA1D12-F1CD-4661-9D83-81F34A0705AA" docUuidSource="ZooBank" docVersion="5" lastPageNumber="19" masterDocId="FFE5FFC8FFE3FF97FFCA3F3FFF9D831F" masterDocTitle="Revision and phylogeny of the subaptera-group of Phyllodromica (Blattoptera: Blattellidae: Ectobiinae), including a parthenogenetic species and the evaluation of COI sequences for species identification (DNA barcoding)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="18" updateTime="1698998954654" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="77E87DBC4F30817C8E0513E5595D98FB">Revision and phylogeny of the subaptera-group of Phyllodromica (Blattoptera: Blattellidae: Ectobiinae), including a parthenogenetic species and the evaluation of COI sequences for species identification (DNA barcoding)</mods:title>
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<mods:namePart id="0E9D5050981044614D103F2B30C2DC1B">KNEBELSBERGER, THOMAS</mods:namePart>
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<mods:title id="6C916633F654B600DA70B77D3536D759">Zootaxa</mods:title>
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<mods:date id="EF3D544AB01D19F0F922A4FB6FC8A8BD">2007</mods:date>
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<heading id="D08281CAFFF2FF86FF5D3D70FDE28176" bold="true" box="[151,639,591,617]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="B901EAB4FFF2FF86FF5D3D70FDE28176" bold="true" box="[151,639,591,617]" pageId="17" pageNumber="18">
<taxonomicName id="4C754D25FFF2FF86FF5D3D70FE648176" ID-CoL="4H78K" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[151,505,591,617]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="quadracantha" status="sp. nov.">
<emphasis id="B901EAB4FFF2FF86FF5D3D70FE648176" bold="true" box="[151,505,591,617]" italics="true" pageId="17" pageNumber="18">Phyllodromica quadracantha</emphasis>
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,
<taxonomicNameLabel id="A23257CFFFF2FF86FDCD3D70FDE28176" box="[519,639,591,617]" pageId="17" pageNumber="18" rank="species">spec. nov.</taxonomicNameLabel>
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<paragraph id="8BCA36A6FFF2FF86FF5D3D4DFE608193" blockId="17.[151,639,591,652]" box="[151,509,626,652]" pageId="17" pageNumber="18">
(
<figureCitation id="134E2A23FFF2FF86FF553D4DFF668193" box="[159,251,626,652]" captionStart="FIGURE 8" captionStartId="35.[151,255,1618,1642]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-136@35.[264,1323,194,1578]" captionTargetPageId="35" captionText="FIGURE 8. AE Phyllodromica iberica morph #2, male. (AD) Different colour patterns of tergite 6; (E) helmet sclerite. F, G Phyllodromica iberica morph #2, female. (F) Thoracal nota; (G) abdominal tergite 5. H Right front leg of Phyllodromica iberica morph #1, male in frontal view. I Right front leg of Phyllodromica quadracantha, male in frontal view. Abbreviations: fr “frontal” part of helmet sclerite, cox coxa, fem femur, hs helmet sclerite re “rear” of helmet sclerite, tib tibia, tm tegmen. Same scale for (AD), (F, G) and (H, I). Identification: (A) Sp 266/2, (B) Sp 365/3, (C) Sp 365a/M14, (D) Sp 365a/M8, (E) Sp 267c/M1, (F, G) Sp 267c/W2, (H) Sp 510/M11, (I) Sp 203d/M1 (holotype).." figureDoi="http://doi.org/10.5281/zenodo.5087778" httpUri="https://zenodo.org/record/5087778/files/figure.png" pageId="17" pageNumber="18">Figs 8 I</figureCitation>
,
<figureCitation id="134E2A23FFF2FF86FEC33D4DFEC38193" box="[265,350,626,652]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="17" pageNumber="18">12 AI</figureCitation>
,
<figureCitation id="134E2A23FFF2FF86FEA33D4DFE578193" box="[361,458,626,652]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="17" pageNumber="18">13 AH</figureCitation>
,
<figureCitation id="134E2A23FFF2FF86FE1C3D4DFE698193" box="[470,500,626,652]" captionStart="FIGURE 17" captionStartId="44.[151,255,1798,1822]" captionTargetBox="[294,1267,311,1652]" captionTargetId="figure-154@44.[294,1353,198,1785]" captionTargetPageId="44" captionText="FIGURE 17. Distribution of the bisexual species of the subaptera-group: Phyllodromica iberica morph #1, #2 and #3, and P. quadracantha; these species only occur on the Iberian Peninsula. In cases where two different symbols partly overlap, two P. iberica morphs are found at the same locality; in one case all three morphs occur together (Fig. 15, Sp 510 ca. 1° W, 40° N). P. iberica morph #3 symbols labeled with a “+” (Fig. 15, Sp 335) or an “ ” (Fig. 15, Sp 186, 270, 469 and 512) indicate the presence of morphological variations. At some localities no males have been found (“sad face” symbol) but females with spermathecae containing sperms and/or with oothecae containing male and female offspring. Both facts indicate the presence of a bisexual species of unknown specifity." figureDoi="http://doi.org/10.5281/zenodo.5087798" httpUri="https://zenodo.org/record/5087798/files/figure.png" pageId="17" pageNumber="18">17</figureCitation>
)
</paragraph>
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<taxonomicName id="4C754D25FFF2FF86FF5D3DFDFE3B81C6" box="[151,422,706,729]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFF2FF86FF5D3DFDFE3B81C6" box="[151,422,706,729]" italics="true" pageId="17" pageNumber="18">Phyllodromica subaptera</emphasis>
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morph #4. —Knebelsberger &amp; Bohn, 2003.
</paragraph>
</subSubSection>
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<materialsCitation id="3B1D3CFBFFF2FF86FF5D3C35FCE48053" collectingDate="2001-03-20" collectingDateMax="2001-03-23" collectingDateMin="2001-03-20" collectionCode="T, ZSM" collectorName="Prov. Granada, Sa." country="Spain" elevation="1100" location="Sa. de Baza" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B901EAB4FFF2FF86FF5D3C35FE94803B" bold="true" box="[151,265,778,804]" pageId="17" pageNumber="18">
<typeStatus id="54CE8804FFF2FF86FF5D3C35FE94803B" box="[151,265,778,804]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
:
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,
<collectingCountry id="F3627636FFF2FF86FE8B3C35FE1B803B" box="[321,390,778,804]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
,
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Prov. Granada,
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de Baza, ca.
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WSW Baza
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,
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,
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</date>
, leg.
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. Knebelsberger (on two slides, Sp203d/M1); deposited in
<collectionCode id="ED64AE63FFF2FF86FCFE3C0DFCE98053" box="[820,884,818,844]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="18" type="Museum">ZSM</collectionCode>
.
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<materialsCitation id="3B1D3CFBFFF2FF86FF0C3C65FDA380DB" collectingDate="2001-03-20" collectingDateMax="2001-03-23" collectingDateMin="2001-03-20" collectionCode="T, ZSM" collectorName="Prov. Granada, Sa." country="Spain" elevation="1100" location="Sa. de Baza" pageId="17" pageNumber="18" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
<emphasis id="B901EAB4FFF2FF86FF0C3C65FEDE806B" bold="true" box="[198,323,858,884]" pageId="17" pageNumber="18">
<typeStatus id="54CE8804FFF2FF86FF0C3C65FEDE806B" box="[198,323,858,884]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
:
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, same data as holotype. Sp 203d/M11 (frozen DNA; parts of body in 98 % ethanol); Sp 203d/M12 (frozen DNA; parts of body in 98 % ethanol); Sp 203d/M2 (on two slides); Sp 203d/M3 (on two slides); Sp 203d/M9 (on two slides)
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.
</paragraph>
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<materialsCitation id="3B1D3CFBFFF2FF86FF0C3CEDFE448793" accessionNumber="AM600691, AM600692" collectionCode="DNATAX, ZSM, AM, T" pageId="17" pageNumber="18" specimenCode="DNATAX02867, DNATAX02868" specimenCount="3" typeStatus="paratype">
<typeStatus id="54CE8804FFF2FF86FF0C3CEDFEB180F3" box="[198,300,978,1004]" pageId="17" pageNumber="18" type="paratype">Paratype</typeStatus>
DNA and associated parts of body are stored in the
<collectionCode id="ED64AE63FFF2FF86FC563CEDFB8F80F3" box="[924,1042,978,1004]" country="Germany" httpUri="http://grbio.org/cool/cnpp-emg5" name="DNA-TAX" pageId="17" pageNumber="18">DNATAX</collectionCode>
collection of
<collectionCode id="ED64AE63FFF2FF86FB723CEDFB6980F3" box="[1208,1268,978,1004]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="17" pageNumber="18" type="Museum">ZSM</collectionCode>
under storage numbers
<specimenCode id="DBD39EDDFFF2FF86FEC13CC5FE56870B" box="[267,459,1018,1044]" collectionCode="DNATAX" country="Germany" httpUri="http://grbio.org/cool/cnpp-emg5" name="DNA-TAX" pageId="17" pageNumber="18">DNATAX02867</specimenCode>
(
<typeStatus id="54CE8804FFF2FF86FE283CC5FDD7870B" box="[482,586,1018,1044]" pageId="17" pageNumber="18" type="paratype">Paratype</typeStatus>
Sp 203d/M11) and
<specimenCode id="DBD39EDDFFF2FF86FC9C3CC5FB88870B" box="[854,1045,1018,1044]" collectionCode="DNATAX" country="Germany" httpUri="http://grbio.org/cool/cnpp-emg5" name="DNA-TAX" pageId="17" pageNumber="18">DNATAX02868</specimenCode>
(
<typeStatus id="54CE8804FFF2FF86FBE73CC5FB08870B" box="[1069,1173,1018,1044]" pageId="17" pageNumber="18" type="paratype">Paratype</typeStatus>
Sp 203d/M12). The sequences were submitted to GenBank under the accession numbers:
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<collectionCode id="ED64AE63FFF2FF86FC373B1DFBB08723" box="[1021,1069,1058,1084]" country="Australia" name="Australian Museum" pageId="17" pageNumber="18" type="Museum">AM</collectionCode>
600691
</accessionNumber>
(
<specimenCode id="DBD39EDDFFF2FF86FB533B1DFAC08723" box="[1177,1373,1058,1084]" collectionCode="DNATAX" country="Germany" httpUri="http://grbio.org/cool/cnpp-emg5" name="DNA-TAX" pageId="17" pageNumber="18">DNATAX02867</specimenCode>
) and
<accessionNumber id="9426AB45FFF2FF86FF5D3B75FE83877B" box="[151,286,1098,1124]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AM600692" pageId="17" pageNumber="18">
<collectionCode id="ED64AE63FFF2FF86FF5D3B75FF5B877B" box="[151,198,1098,1124]" country="Australia" name="Australian Museum" pageId="17" pageNumber="18" type="Museum">AM</collectionCode>
600692
</accessionNumber>
(
<specimenCode id="DBD39EDDFFF2FF86FEE73B75FE69877B" box="[301,500,1098,1124]" collectionCode="DNATAX" country="Germany" httpUri="http://grbio.org/cool/cnpp-emg5" name="DNA-TAX" pageId="17" pageNumber="18">DNATAX02868</specimenCode>
). The remaining
<specimenCount id="9D73FD2FFFF2FF86FD733B75FCA3877B" box="[697,830,1098,1124]" pageId="17" pageNumber="18" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
(Sp 203d/M2, 3, 9) are deposited in the private collection of
<collectionCode id="ED64AE63FFF2FF86FEC73B4DFEBD8793" box="[269,288,1138,1164]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="18">T</collectionCode>
. Knebelsberger
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.
</paragraph>
<paragraph id="8BCA36A6FFF2FF86FF0C3BA5FB8E87C3" blockId="17.[151,1436,778,2004]" pageId="17" pageNumber="18">
<emphasis id="B901EAB4FFF2FF86FF0C3BA5FE2087AB" bold="true" box="[198,445,1178,1204]" pageId="17" pageNumber="18">Additional material</emphasis>
. Numerous specimens from the following localities:
<collectingCountry id="F3627636FFF2FF86FBE33BA5FBF287AB" box="[1065,1135,1178,1204]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
. Sp 191, a, 195, 203, a, b, c, d, 207, a, 472, a, b, 473, a, 493, 495, 497, 498, 499, 500, 501, 502, 504, a.
</paragraph>
</subSubSection>
<subSubSection id="C36F652DFFF2FF86FF0C3BD5FECA8633" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BCA36A6FFF2FF86FF0C3BD5FECA8633" blockId="17.[151,1436,778,2004]" pageId="17" pageNumber="18">
<emphasis id="B901EAB4FFF2FF86FF0C3BD5FECE861B" bold="true" box="[198,339,1258,1284]" pageId="17" pageNumber="18">Etymology.</emphasis>
The name of the species refers to the presence of only 4 distal tibia spines on the second pair of legs in males.
</paragraph>
</subSubSection>
<subSubSection id="C36F652DFFF2FF85FF0C3A05FE988625" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8BCA36A6FFF2FF86FF0C3A05FAF0864B" blockId="17.[151,1436,778,2004]" box="[198,1389,1338,1364]" pageId="17" pageNumber="18">
<emphasis id="B901EAB4FFF2FF86FF0C3A05FE02864B" bold="true" box="[198,415,1338,1364]" pageId="17" pageNumber="18">Description. Size.</emphasis>
Length of pronotum:
<specimenCount id="9D73FD2FFFF2FF86FD683A05FD2B8651" box="[674,694,1338,1358]" pageId="17" pageNumber="18" type="male"></specimenCount>
1.351.56 (mean 1.45) mm;
<specimenCount id="9D73FD2FFFF2FF86FBC63A05FBBD8651" box="[1036,1056,1338,1358]" pageId="17" pageNumber="18" type="female"></specimenCount>
1.711.90 (mean 1.81) mm.
</paragraph>
<paragraph id="8BCA36A6FFF2FF86FF0C3A5DFBCD8663" blockId="17.[151,1436,778,2004]" box="[198,1104,1378,1404]" pageId="17" pageNumber="18">
<emphasis id="B901EAB4FFF2FF86FF0C3A5DFE958663" bold="true" box="[198,264,1378,1404]" pageId="17" pageNumber="18">Legs.</emphasis>
Distal end of mid tibia with only 4 spines (
<figureCitation id="134E2A23FFF2FF86FCCE3A5DFCEB8663" box="[772,886,1378,1404]" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="17" pageNumber="18">Fig. 14 C</figureCitation>
), females rarely 5.
</paragraph>
<paragraph id="8BCA36A6FFF2FF86FF0C3AB5FE9F86BB" blockId="17.[151,1436,778,2004]" box="[198,258,1418,1444]" pageId="17" pageNumber="18">Male</paragraph>
<paragraph id="8BCA36A6FFF2FF86FF0C3A8DFD6584CB" blockId="17.[151,1436,778,2004]" pageId="17" pageNumber="18">
<emphasis id="B901EAB4FFF2FF86FF0C3A8DFE5186D3" bold="true" box="[198,460,1458,1484]" pageId="17" pageNumber="18">Tergites. —Tergite 6</emphasis>
. Emargination of the posterior border relatively broad and less deep than in the other species, without torus behind the transversal ridge (
<figureCitation id="134E2A23FFF2FF86FCF73AE5FC2F86EB" box="[829,946,1498,1524]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="17" pageNumber="18">Fig. 12 C</figureCitation>
).
<emphasis id="B901EAB4FFF2FF86FC023AE5FBA586EB" bold="true" box="[968,1080,1498,1524]" pageId="17" pageNumber="18">Tergite 7</emphasis>
. Median lobe long, at least as long as the distance between the bristle fields and the posterior border of the tergite, covered with numerous glandular pores (
<figureCitation id="134E2A23FFF2FF86FE913915FE4F855B" box="[347,466,1578,1604]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="17" pageNumber="18">Figs 12 D</figureCitation>
,
<figureCitation id="134E2A23FFF2FF86FE143915FD81855B" box="[478,540,1578,1604]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="17" pageNumber="18">13 A</figureCitation>
). Trough very shallow, almost in the plane of the tergite surface. Mound flat, mainly elevated behind the bristle fields. Bristle fields moderately large, in outline broadly oval to circular, immediately adjacent, rarely separated by a very narrow bristleless ridge, posteriorly appearing deepened by the elevating surface of the mound; bristle fields with indistinct borders (
<figureCitation id="134E2A23FFF2FF86FC37399DFBE985A3" box="[1021,1140,1698,1724]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="17" pageNumber="18">Fig. 13 A</figureCitation>
). The male from locality Sp 504a (
<figureCitation id="134E2A23FFF2FF86FEC239F5FE0185FB" box="[264,412,1738,1764]" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="17" pageNumber="18">Fig. 14 EG</figureCitation>
) differs slightly in the morphology of the bristle fields: the bristle fields are somewhat larger and slightly separated by a bristleless ridgelike structure (
<figureCitation id="134E2A23FFF2FF86FC4B39CDFC6D8413" box="[897,1008,1778,1804]" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="17" pageNumber="18">Fig. 14 F</figureCitation>
).
<emphasis id="B901EAB4FFF2FF86FBCE39CDFBEE8413" bold="true" box="[1028,1139,1778,1804]" pageId="17" pageNumber="18">Tergite 8</emphasis>
. Without any sculpturing on the surface, rarely with weak depressions at the base of the anterior processes; membraneous incision between the anterior processes narrow (
<figureCitation id="134E2A23FFF2FF86FDAE387DFD4A8443" box="[612,727,1858,1884]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="17" pageNumber="18">Figs 12 E</figureCitation>
). Note: The membraneous incision between the anterior processes appears relatively broad due to an unnatural stretching of the tergite during preparation (
<figureCitation id="134E2A23FFF2FF86FADF3855FA12849B" box="[1301,1423,1898,1924]" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="17" pageNumber="18">Fig. 14 G</figureCitation>
). Cone completely missing (
<figureCitation id="134E2A23FFF2FF86FE1B38ADFDDE84B3" box="[465,579,1938,1964]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="17" pageNumber="18">Fig. 13 C</figureCitation>
); without median gap in the distribution of the bristles (
<figureCitation id="134E2A23FFF2FF86FB1B38ADFADF84B3" box="[1233,1346,1938,1964]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="17" pageNumber="18">Fig. 13 B</figureCitation>
).
<emphasis id="B901EAB4FFF2FF86FA9338ADFF4784CB" bold="true" pageId="17" pageNumber="18">Tergite 10</emphasis>
. Supraanal plate broadly rounded (
<figureCitation id="134E2A23FFF2FF86FDBC3885FD7684CB" box="[630,747,1978,2004]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="17" pageNumber="18">Fig. 12 G</figureCitation>
).
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3FA8FF2683C5" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3FA8FEA683AE" bold="true" box="[198,315,151,177]" pageId="18" pageNumber="19">Sternites.</emphasis>
—Paraproct. Medio-anterior process of the right parapoct with a bulge (
<emphasis id="B901EAB4FFF1FF85FB463FA8FB3083AE" bold="true" box="[1164,1197,151,177]" pageId="18" pageNumber="19">bu</emphasis>
) at the base (
<figureCitation id="134E2A23FFF1FF85FA823FA7FF3083C5" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="18" pageNumber="19">Fig. 14 B</figureCitation>
).
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3FD8FE35824D" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3FD8FEFB821E" bold="true" box="[198,358,231,257]" pageId="18" pageNumber="19">Genitalia. —</emphasis>
Helmet sclerite. “Frontal” part (
<emphasis id="B901EAB4FFF1FF85FD163FD8FD6E821E" bold="true" box="[732,755,231,257]" pageId="18" pageNumber="19">fr</emphasis>
) broadly rounded, in the “rear” (
<emphasis id="B901EAB4FFF1FF85FBBD3FD8FB0D821E" bold="true" box="[1143,1168,231,257]" pageId="18" pageNumber="19">re</emphasis>
) with a relatively long acuminate process (
<figureCitation id="134E2A23FFF1FF85FEB53E2FFE6C8235" box="[383,497,272,298]" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="18" pageNumber="19">Fig. 14 D</figureCitation>
) (in
<taxonomicName id="4C754D25FFF1FF85FDE13E2FFD5A8235" authority="the" authorityName="The" box="[555,711,272,298]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="12" phylum="Arthropoda" rank="species" species="iberica">
<emphasis id="B901EAB4FFF1FF85FDE13E2FFD068236" box="[555,667,272,297]" italics="true" pageId="18" pageNumber="19">P. iberica</emphasis>
the
</taxonomicName>
anterior part and the acuminate process of the helmet sclerite are reduced (
<figureCitation id="134E2A23FFF1FF85FEE53E07FE12824D" box="[303,399,312,338]" captionStart="FIGURE 8" captionStartId="35.[151,255,1618,1642]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-136@35.[264,1323,194,1578]" captionTargetPageId="35" captionText="FIGURE 8. AE Phyllodromica iberica morph #2, male. (AD) Different colour patterns of tergite 6; (E) helmet sclerite. F, G Phyllodromica iberica morph #2, female. (F) Thoracal nota; (G) abdominal tergite 5. H Right front leg of Phyllodromica iberica morph #1, male in frontal view. I Right front leg of Phyllodromica quadracantha, male in frontal view. Abbreviations: fr “frontal” part of helmet sclerite, cox coxa, fem femur, hs helmet sclerite re “rear” of helmet sclerite, tib tibia, tm tegmen. Same scale for (AD), (F, G) and (H, I). Identification: (A) Sp 266/2, (B) Sp 365/3, (C) Sp 365a/M14, (D) Sp 365a/M8, (E) Sp 267c/M1, (F, G) Sp 267c/W2, (H) Sp 510/M11, (I) Sp 203d/M1 (holotype).." figureDoi="http://doi.org/10.5281/zenodo.5087778" httpUri="https://zenodo.org/record/5087778/files/figure.png" pageId="18" pageNumber="19">Fig. 8 E</figureCitation>
)).
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3E60FBE48175" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3E60FDBF8266" bold="true" box="[198,546,351,377]" pageId="18" pageNumber="19">Colouration. —Tergites 25</emphasis>
. Dark anterior area often with short posterior extensions (
<figureCitation id="134E2A23FFF1FF85FB053E5FFADE8265" box="[1231,1347,352,378]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="18" pageNumber="19">Fig. 12 B</figureCitation>
).
<emphasis id="B901EAB4FFF1FF85FA933E60FF5382BE" bold="true" pageId="18" pageNumber="19">Tergite 6</emphasis>
. Dark area broadly reaching the posterior tergite border, behind the transversal ridge often with small lightenings medially and/or along a transversal line at some distance from the ridge (
<figureCitation id="134E2A23FFF1FF85FBBE3E8FFB7482D5" box="[1140,1257,432,458]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="18" pageNumber="19">Figs 12 C</figureCitation>
,
<figureCitation id="134E2A23FFF1FF85FB3F3E8FFAB282D5" box="[1269,1327,432,458]" captionStart="FIGURE 14" captionStartId="41.[151,255,1623,1647]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-204@41.[264,1323,194,1585]" captionTargetPageId="41" captionText="FIGURE 14 Phyllodromica quadracantha, male. A membrane glands of the right lateral region of the anterior border of tergite 6. B right paraproct with bulge on the medio-anterior process (compare Fig. 5 D). C distal end of tibia from the right mid leg (bearing 4 distal tibia spines; compare Fig. 5 E) in posterior view. D helmet sclerite, the nomenclature “fr” and “re” do not indicate the orientation of the sclerite within the animal. EG tergites 6 (E), 7 (F) and 8 (G) of specimen with slightly different tergal structures on tergite 7. Due to unnatural squeezing of the tergite 8 (G) during the mounting procedure the distance between the anterior processes appears broader than under natural conditions. Abbreviations: fr “frontal” part of helmet sclerite, bf bristle field, bu bulge, hs helmet sclerite, mg membrane glands, mp medio-anterior process of right paraproct, re “rear” of helmet sclerite, r ridge, rp right paraproct, sp spinelike process, tar tarsus, tib tibia, tr transversal trough. Same scale for (AD) and (FG). Identification: (A, EG) Sp 504a/M1, (B, C) Sp 203d/ M1(holotype). (D) Sp 203b/M7." figureDoi="http://doi.org/10.5281/zenodo.5087794" httpUri="https://zenodo.org/record/5087794/files/figure.png" pageId="18" pageNumber="19">14 E</figureCitation>
).
<emphasis id="B901EAB4FFF1FF85FA8E3E90FF3082EE" bold="true" pageId="18" pageNumber="19">Tergite 7.</emphasis>
As in tergite 6 with dark area broadly reaching the posterior border, central and posterior part darker than the remainder, bristle fields and immediate surroundings not seldom lightly coloured, trough in the middle with a whitish patch (
<figureCitation id="134E2A23FFF1FF85FE523D17FD91815D" box="[408,524,552,578]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="18" pageNumber="19">Fig. 12 D</figureCitation>
) as in the other species.
<emphasis id="B901EAB4FFF1FF85FCF83D18FC3F815E" bold="true" box="[818,930,551,577]" pageId="18" pageNumber="19">Tergite 8</emphasis>
. Dark posterior extensions medio-laterally usually not strongly pronounced, median extension often well developed (
<figureCitation id="134E2A23FFF1FF85FC313D6FFBF18175" box="[1019,1132,592,618]" captionStart="FIGURE 12" captionStartId="39.[151,255,1679,1703]" captionTargetBox="[264,1323,194,1565]" captionTargetId="figure-131@39.[264,1323,194,1781]" captionTargetPageId="39" captionText="FIGURE 12. Phyllodromica quadracantha, male (holotype). A Thoracal nota. BF Abdominal tergites 59. (B) Tergite 5; (C) tergite 6; (D) tergite 7 and (E) tergite 8 with glandular structures in the anterior part of the tergites; (F) tergite 9. G Terminalia with tergite 10 (sa), cerci and paraprocts. H Hook of left phallomere with the posterior end on the top. I Subgenital plate with remaining genital sclerites (without hook). Abbreviations: ap anterior process, bf bristle field, c cercus, ml median lobe, rp right paraproct, sa supraanal plate, tr transversal ridge, tt transversal trough. Same scale for (B, C, G) and (D, E, H). Identification: Sp 203d/M1 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5087790" httpUri="https://zenodo.org/record/5087790/files/figure.png" pageId="18" pageNumber="19">Fig. 12 E</figureCitation>
).
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF5D3D47FF70818D" blockId="18.[151,1436,151,1458]" box="[151,237,632,658]" pageId="18" pageNumber="19">Female</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3DA0FED981FD" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3DA0FE9A81A6" bold="true" box="[198,263,671,697]" pageId="18" pageNumber="19">Legs.</emphasis>
Distal end of mid tibia usually with only 4 long spines, rarely with a fifth short spine in ventro-posterior position.
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3DD0FCA6802D" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3DD0FEDC8016" bold="true" box="[198,321,751,777]" pageId="18" pageNumber="19">Genitalia.</emphasis>
No additional sclerotization in the angle between intercalary sclerite and the posterior lobe of valvifere II, the latter very broad, egg-shaped (
<figureCitation id="134E2A23FFF1FF85FD733C27FCB3802D" box="[697,814,792,818]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="18" pageNumber="19">Fig. 13 G</figureCitation>
).
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3C00FD9A80E5" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3C00FEF88046" bold="true" box="[198,357,831,857]" pageId="18" pageNumber="19">Colouration.</emphasis>
Disk of pronotum regularly mottled with variously shaped dark spots; anterior border never with a dark crescent (
<figureCitation id="134E2A23FFF1FF85FE503C57FD8E809D" box="[410,531,872,898]" captionStart="FIGURE 13" captionStartId="40.[151,255,1605,1629]" captionTargetBox="[269,1318,199,1564]" captionTargetId="figure-181@40.[264,1323,194,1567]" captionTargetPageId="40" captionText="FIGURE 13 Phyllodromica quadracantha. AC male, SEM pictures of glandular structures of tergites 7 and 8 in dorsal view. (A) Tergite 7; (B) tergite 8, bristles are present also in the median tergite region (white arrow heads); (C) anteriormedian region of tergite 8, no conelike process present. DH female. (D) Thoracal nota; (E) Tergite 5; (F) Subgenital plate; (G) dorsal complex of genitalia in ventral view, posterior end on top, without additional sclerite between pl and is (compare Fig. 2 D, E); (H) ventral komplex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, tr transversal ridge, T9 tergite 9, T10 tergite 10, v valves. Same scale for (D, E). Identification: (AD) Sp 499/M1, (EI) Sp 203d/W1." figureDoi="http://doi.org/10.5281/zenodo.5087792" httpUri="https://zenodo.org/record/5087792/files/figure.png" pageId="18" pageNumber="19">Fig. 13 D</figureCitation>
). Transparent margin of the pronotum not well set off, especially anteriorly and in the latero-posterior region with numerous small dots. Meso- and metanotum with dark anterior part ending shortly behind the transversal ridge, from there with 6 to 7 bar-shaped posterior projections, remaining surface quite regularly mottled.
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3B38FA058785" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3B38FEED873E" bold="true" box="[198,368,1031,1057]" pageId="18" pageNumber="19">Genetic Data.</emphasis>
1597 basepairs of the mitochondrial genome including the complete cytochrome c oxidase subunit I (COI) gene were analysed. in
<specimenCount id="9D73FD2FFFF1FF85FDA33B0FFC198755" box="[617,900,1072,1098]" pageId="18" pageNumber="19" type="generic" typeStatus="paratype">two paratype specimens</specimenCount>
(Sp203d/M11, Sp203d/M12). The sequences of
<taxonomicName id="4C754D25FFF1FF85FF723B67FD8E876E" authorityName="Published" authorityYear="2007" box="[184,531,1112,1137]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadracantha">
<emphasis id="B901EAB4FFF1FF85FF723B67FD8E876E" box="[184,531,1112,1137]" italics="true" pageId="18" pageNumber="19">Phyllodromica quadracantha</emphasis>
</taxonomicName>
_
<typeStatus id="54CE8804FFF1FF85FDE83B67FD15876D" box="[546,648,1112,1138]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
Sp203d/M11_1585bp and
<taxonomicName id="4C754D25FFF1FF85FC063B67FABB876E" authorityName="Published" authorityYear="2007" box="[972,1318,1112,1137]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadracantha">
<emphasis id="B901EAB4FFF1FF85FC063B67FABB876E" box="[972,1318,1112,1137]" italics="true" pageId="18" pageNumber="19">Phyllodromica quadracantha</emphasis>
</taxonomicName>
_
<typeStatus id="54CE8804FFF1FF85FAFC3B67FA01876D" box="[1334,1436,1112,1138]" pageId="18" pageNumber="19" type="paratype">Paratype</typeStatus>
Sp203d/M12_1585bp differ in 19 positions (Appendix 5). The complete sequences are shown in Appendix 4.
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3B98FAA587F5" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3B98FD9387DE" bold="true" box="[198,526,1191,1217]" pageId="18" pageNumber="19">Geographical distribution</emphasis>
(
<figureCitation id="134E2A23FFF1FF85FDEA3B97FDE487DD" box="[544,633,1192,1218]" captionStart="FIGURE 17" captionStartId="44.[151,255,1798,1822]" captionTargetBox="[294,1267,311,1652]" captionTargetId="figure-154@44.[294,1353,198,1785]" captionTargetPageId="44" captionText="FIGURE 17. Distribution of the bisexual species of the subaptera-group: Phyllodromica iberica morph #1, #2 and #3, and P. quadracantha; these species only occur on the Iberian Peninsula. In cases where two different symbols partly overlap, two P. iberica morphs are found at the same locality; in one case all three morphs occur together (Fig. 15, Sp 510 ca. 1° W, 40° N). P. iberica morph #3 symbols labeled with a “+” (Fig. 15, Sp 335) or an “ ” (Fig. 15, Sp 186, 270, 469 and 512) indicate the presence of morphological variations. At some localities no males have been found (“sad face” symbol) but females with spermathecae containing sperms and/or with oothecae containing male and female offspring. Both facts indicate the presence of a bisexual species of unknown specifity." figureDoi="http://doi.org/10.5281/zenodo.5087798" httpUri="https://zenodo.org/record/5087798/files/figure.png" pageId="18" pageNumber="19">Fig. 17</figureCitation>
). Relatively small distribution area in the southeast of
<collectingCountry id="F3627636FFF1FF85FADB3B97FAC987DD" box="[1297,1364,1192,1218]" name="Spain" pageId="18" pageNumber="19">Spain</collectingCountry>
separated from other bisexual species. One isolated population (Sp 504) has been found further northeast.
</paragraph>
<paragraph id="8BCA36A6FFF1FF85FF0C3BC8FE988625" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3BC8FD72860E" bold="true" box="[198,751,1271,1297]" pageId="18" pageNumber="19">
Other species of the
<taxonomicName id="4C754D25FFF1FF85FE003BC8FDDC860E" box="[458,577,1271,1297]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFF1FF85FE003BC8FDDC860E" bold="true" box="[458,577,1271,1297]" italics="true" pageId="18" pageNumber="19">subaptera</emphasis>
</taxonomicName>
-group found.
</emphasis>
<taxonomicName id="4C754D25FFF1FF85FD3D3BC7FC26860D" authority=": Sp" authorityName="Sp" box="[759,955,1272,1298]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFF1FF85FD3D3BC7FC16860E" box="[759,907,1272,1297]" italics="true" pageId="18" pageNumber="19">P. subaptera</emphasis>
: Sp
</taxonomicName>
191, 195, 203, 207, 472, 473, 493, 495, 498, 504.
</paragraph>
</subSubSection>
<subSubSection id="C36F652DFFF1FF85FF0C3A78FE4486AD" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BCA36A6FFF1FF85FF0C3A78FE4486AD" blockId="18.[151,1436,151,1458]" pageId="18" pageNumber="19">
<emphasis id="B901EAB4FFF1FF85FF0C3A78FEA2867E" bold="true" box="[198,319,1351,1377]" pageId="18" pageNumber="19">Remarks.</emphasis>
At the locality Sp504 only
<specimenCount id="9D73FD2FFFF1FF85FD423A77FD6E867D" box="[648,755,1352,1378]" pageId="18" pageNumber="19" type="male">one male</specimenCount>
(and some females) has been found. It shows remarkable differences in the structures of tergite 7, which might indicate a taxonomic separation. More material is necessary to clarify the situation.
</paragraph>
</subSubSection>
</treatment>
</document>