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<document id="3E74240EF2FCCE8D93BB916ACB8E21A2" ID-DOI="10.11646/zootaxa.1522.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5087760" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625852368432" checkinUser="felipe" docAuthor="KNEBELSBERGER, THOMAS &amp; MILLER, MICHAEL A." docDate="2007" docId="03DC87B0FFE8FF9AFF5D3800FB96824D" docLanguage="en" docName="zt01522p068.pdf.imf" docOrigin="Zootaxa 1522 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.1522.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Phyllodromica subaptera" docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="FFE5FFC8FFE3FF97FFCA3F3FFF9D831F" masterDocTitle="Revision and phylogeny of the subaptera-group of Phyllodromica (Blattoptera: Blattellidae: Ectobiinae), including a parthenogenetic species and the evaluation of COI sequences for species identification (DNA barcoding)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="12" updateTime="1698998954654" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0B3CF9DD0E34E836C999008934D59237">Revision and phylogeny of the subaptera-group of Phyllodromica (Blattoptera: Blattellidae: Ectobiinae), including a parthenogenetic species and the evaluation of COI sequences for species identification (DNA barcoding)</mods:title>
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<mods:namePart id="7EC864EF3596567E9A0843462110136B">KNEBELSBERGER, THOMAS</mods:namePart>
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<mods:namePart id="7F40689FDC79A90B1A7039D0B7DCFF40">MILLER, MICHAEL A.</mods:namePart>
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<mods:title id="D8D5FD04BD6FC27F313E0CE4D353D916">Zootaxa</mods:title>
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<mods:date id="622B787D79E8EA4F9F6D28A0C5718E1C">2007</mods:date>
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<mods:number id="E521E4C13C398A8F99FF429F5FC39823">2007-07-09</mods:number>
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<mods:number id="0E925682C2D3817983A17A9435592A46">1522</mods:number>
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<mods:classification id="07372AD4358D168A49A6A5CDF5946B5F">journal article</mods:classification>
<mods:identifier id="E66D4D3290C6B4C58ADB3452A0388C65" type="DOI">10.11646/zootaxa.1522.1.1</mods:identifier>
<mods:identifier id="B69D7341617DB759224B148CE1C74B15" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03DC87B0FFE8FF9AFF5D3800FB96824D" ID-DOI="http://doi.org/10.5281/zenodo.5088408" ID-Zenodo-Dep="5088408" LSID="urn:lsid:plazi:treatment:03DC87B0FFE8FF9AFF5D3800FB96824D" httpUri="http://treatment.plazi.org/id/03DC87B0FFE8FF9AFF5D3800FB96824D" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="C36F652DFFE8FF9CFF5D3800FD0C8446" box="[151,657,1855,1881]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BCA36A6FFE8FF9CFF5D3800FD0C8446" blockId="11.[151,657,1855,1916]" box="[151,657,1855,1881]" pageId="11" pageNumber="12">
<heading id="D08281CAFFE8FF9CFF5D3800FD0C8446" bold="true" box="[151,657,1855,1881]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C754D25FFE8FF9CFF5D3800FD0C8446" ID-CoL="4H78M" authority="(Rambur, 1838)" baseAuthorityName="Rambur" baseAuthorityYear="1838" box="[151,657,1855,1881]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFE8FF9CFF5D3800FD0C8446" bold="true" box="[151,657,1855,1881]" pageId="11" pageNumber="12">
<emphasis id="B901EAB4FFE8FF9CFF5D3800FE5A8446" bold="true" box="[151,455,1855,1881]" italics="true" pageId="11" pageNumber="12">Phyllodromica subaptera</emphasis>
(Rambur, 1838)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F652DFFE8FF9BFF5D385DFA9A8250" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BCA36A6FFE8FF9CFF5D385DFEC58463" blockId="11.[151,657,1855,1916]" box="[151,344,1890,1916]" pageId="11" pageNumber="12">
(
<figureCitation id="134E2A23FFE8FF9CFF55385DFEB88463" box="[159,293,1890,1916]" captionStart="FIGURE 2" captionStartId="29.[151,255,1612,1636]" captionTargetBox="[270,1329,194,1565]" captionTargetId="figure-141@29.[270,1329,194,1571]" captionTargetPageId="29" captionText="FIGURE 2. Phyllodromica subaptera (female). A Thoracal nota. B Abdominal tergite 5. C Subgenital plate with two apodemal processes (a). D Dorsal complex of genitalia with additional sclerite (as): ventral view, posterior end on top. E Additional sclerite in higher enlargement (equivalent to “black frame” in D). F Ventral complex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: as additional sclerite, a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, T9 tergite 9, T10 tergite 10, v valves. Same scale for (B, C) and (D, F). Identification: Sp 203c/W4." figureDoi="http://doi.org/10.5281/zenodo.5087766" httpUri="https://zenodo.org/record/5087766/files/figure.png" pageId="11" pageNumber="12">Figs 2 A–F</figureCitation>
,
<figureCitation id="134E2A23FFE8FF9CFEFB385DFED28463" box="[305,335,1890,1916]" captionStart="FIGURE 16" captionStartId="43.[151,255,1744,1768]" captionTargetBox="[302,1270,213,1554]" captionTargetId="figure-36@43.[297,1356,100,1687]" captionTargetPageId="43" captionText="FIGURE 16. Distribution of bisexual species of the subaptera-group and the parthenogenetic species P. subaptera on the Iberian peninsula." figureDoi="http://doi.org/10.5281/zenodo.5087796" httpUri="https://zenodo.org/record/5087796/files/figure.png" pageId="11" pageNumber="12">16</figureCitation>
)
</paragraph>
<paragraph id="8BCA36A6FFE8FF9CFF5D388DFCF984D6" blockId="11.[151,868,1970,2025]" box="[151,868,1970,1993]" pageId="11" pageNumber="12">
<taxonomicName id="4C754D25FFE8FF9CFF5D388DFE7684D6" authority="Rambur, 1838" authorityName="Rambur" authorityYear="1838" box="[151,491,1970,1993]" class="Insecta" family="Blattidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFE8FF9CFF5D388DFED484D6" box="[151,329,1970,1993]" italics="true" pageId="11" pageNumber="12">Blatta subaptera</emphasis>
Rambur, 1838
</taxonomicName>
. —Faune entom.
<collectingRegion id="49B1F844FFE8FF9CFD6D388DFCBD84D6" box="[679,800,1970,1993]" country="Spain" name="Andalucia" pageId="11" pageNumber="12">Andalousie</collectingRegion>
2: 14.
</paragraph>
<paragraph id="8BCA36A6FFE8FF9CFF5D38EDFCC584F6" blockId="11.[151,868,1970,2025]" box="[151,856,2002,2025]" pageId="11" pageNumber="12">
<taxonomicName id="4C754D25FFE8FF9CFF5D38EDFE1084F6" box="[151,397,2002,2025]" class="Insecta" family="Blattidae" genus="Polyzosteria" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFE8FF9CFF5D38EDFE1084F6" box="[151,397,2002,2025]" italics="true" pageId="11" pageNumber="12">Polyzosteria subaptera</emphasis>
</taxonomicName>
.—Fischer 1853, Orth. Europ., Lipsiae: 94.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF5D3FA7FC0783B0" blockId="12.[151,1437,152,335]" box="[151,922,152,175]" pageId="12" pageNumber="13">
<taxonomicName id="4C754D25FFEFFF9BFF5D3FA7FEF583B0" box="[151,360,152,175]" class="Insecta" family="Corydiidae" genus="Aphlebia" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFF5D3FA7FEF583B0" box="[151,360,152,175]" italics="true" pageId="12" pageNumber="13">Aphlebia subaptera</emphasis>
</taxonomicName>
.—Brunner v. W. 1865, Nouv. Syst. Blatt., Wien: 73.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF5D3F87FC8583D0" blockId="12.[151,1437,152,335]" box="[151,792,184,207]" pageId="12" pageNumber="13">
<taxonomicName id="4C754D25FFEFFF9BFF5D3F87FE1683D0" box="[151,395,184,207]" class="Insecta" family="Ectobiidae" genus="Hololampra" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFF5D3F87FE1683D0" box="[151,395,184,207]" italics="true" pageId="12" pageNumber="13">Hololampra subaptera</emphasis>
</taxonomicName>
.—Kirby 1904, Syn. Cat. Orth. 1: 70.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF5D3FE7FB5783F0" blockId="12.[151,1437,152,335]" box="[151,1226,216,239]" pageId="12" pageNumber="13">
<taxonomicName id="4C754D25FFEFFF9BFF5D3FE7FDBA83F0" box="[151,551,216,239]" class="Insecta" family="Ectobiidae" genus="Hololampra" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera" subGenus="Lobolampra">
<emphasis id="B901EAB4FFEFFF9BFF5D3FE7FDBA83F0" box="[151,551,216,239]" italics="true" pageId="12" pageNumber="13">Hololampra (Lobolampra) subaptera</emphasis>
</taxonomicName>
. —Houlbert 1927, Thysan., Derm. et Orth.
<collectingCountry id="F3627636FFEFFF9BFC313FE7FBD983F0" box="[1019,1092,216,239]" name="France" pageId="12" pageNumber="13">France</collectingCountry>
2, Paris: 24.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF5D3FC7FB8A8210" blockId="12.[151,1437,152,335]" box="[151,1047,248,271]" pageId="12" pageNumber="13">
<taxonomicName id="4C754D25FFEFFF9BFF5D3FC7FEE08210" box="[151,381,248,271]" class="Insecta" family="Ectobiidae" genus="Dziriblatta" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFF5D3FC7FEE08210" box="[151,381,248,271]" italics="true" pageId="12" pageNumber="13">Dziriblatta subaptera</emphasis>
</taxonomicName>
. —Chopard 1936, Bull. Soc. Sci. nat.
<collectingCountry id="F3627636FFEFFF9BFCD23FC7FCC38210" box="[792,862,248,271]" name="Morocco" pageId="12" pageNumber="13">Maroc</collectingCountry>
16 (2): 153, 154.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF5D3E27FA9A8250" blockId="12.[151,1437,152,335]" pageId="12" pageNumber="13">
<taxonomicName id="4C754D25FFEFFF9BFF5D3E27FDD88230" baseAuthorityName="Rambur" baseAuthorityYear="1838" box="[151,581,280,303]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera" subGenus="Lobolampra">
<emphasis id="B901EAB4FFEFFF9BFF5D3E27FEA88230" box="[151,309,280,303]" italics="true" pageId="12" pageNumber="13">Phyllodromica</emphasis>
(
<emphasis id="B901EAB4FFEFFF9BFE8F3E27FE548230" box="[325,457,280,303]" italics="true" pageId="12" pageNumber="13">Lobolampra</emphasis>
)
<emphasis id="B901EAB4FFEFFF9BFE113E27FDD88230" box="[475,581,280,303]" italics="true" pageId="12" pageNumber="13">subaptera</emphasis>
</taxonomicName>
. —Princis 1971, In: Beier (Ed.): Orthopterorum Catalogus 14, s’ Gravenhage: 1113; Harz 1976, In: Harz &amp; Kaltenbach: Die Orthopteren Europas. 3, The Hague: 305, figs 904, 905.
</paragraph>
</subSubSection>
<subSubSection id="C36F652DFFEFFF9BFF5D3E45FE8C87C8" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BCA36A6FFEFFF9BFF5D3E45FBD7828B" blockId="12.[151,1436,378,884]" box="[151,1098,378,404]" pageId="12" pageNumber="13">
<materialsCitation id="3B1D3CFBFFEFFF9BFF5D3E45FBD7828B" box="[151,1098,378,404]" collectionCode="BMNH" country="Spain" location="Spain" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B901EAB4FFEFFF9BFF5D3E45FE94828B" bold="true" box="[151,265,378,404]" pageId="12" pageNumber="13">
<typeStatus id="54CE8804FFEFFF9BFF5D3E45FE94828B" box="[151,265,378,404]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D73FD2FFFEFFF9BFED23E45FEB18291" box="[280,300,378,398]" pageId="12" pageNumber="13" type="female">♀</specimenCount>
,
<collectingCountry id="F3627636FFEFFF9BFEF03E45FE1D828B" box="[314,384,378,404]" name="Spain" pageId="12" pageNumber="13">Spain</collectingCountry>
, Prov.Granada enviroment of “Grenade” (Granada);
<collectionCode id="ED64AE63FFEFFF9BFC203E45FBDB828B" box="[1002,1094,378,404]" pageId="12" pageNumber="13">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3E9DFAF2806B" blockId="12.[151,1436,378,884]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3E9DFE5A82A3" bold="true" box="[198,455,418,444]" pageId="12" pageNumber="13">Additional material.</emphasis>
Numerous specimens from the following localities:
<emphasis id="B901EAB4FFEFFF9BFB883E9DFB1782A3" bold="true" box="[1090,1162,418,444]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFB883E9DFB1782A3" box="[1090,1162,418,444]" name="Spain" pageId="12" pageNumber="13">Spain</collectingCountry>
</emphasis>
: Sp 10a, 11a, 17, 18a, 19a, 24a, 61, b, c, 62, 63, 65, 68, 71, a, b, 72, 73, 76, 78, b, 79, c, 80 (?), 88, 89, 90, 91, 92, 93, 94, 101 (?), 105, a, 107, 108a, b, 109a, 114a, 139, 140a, b, 141, 142, 143, 148, a, b, 149 (?), 169, 172, 173a, 174a, 186b, 188a, 191a, 195, 196, 197, 198, 199, 200, 201, 202, 203, a, b, c, d, 204, 205, 205a, 206, 207, a, 208, 209, 210, 214, 229, 230, 253, 254, 256, 258, 262 (?), 263, 264, 278, 279, 280, 281, 282, 283, 284, 285, 286, 287, 288, 289, 290, 293, 320, 333, 339, 340, 342, a, 343, a, 345, 354, a, 355, a, 356, 357, 358, 359 (?), 362, 365, 377, 381, a, 383, 393, 395, 396, 397, a, b, 419, 454, 456, 469, 470, 472, a, b, 473, a, 474, a, 475, 476, 480, 481, 483, 484, 493, 494, 495, 496, 498, 503, 504, a, 505, 508. -
<emphasis id="B901EAB4FFEFFF9BFCC13D85FCEB81CB" bold="true" box="[779,886,698,724]" pageId="12" pageNumber="13">
<collectingRegion id="49B1F844FFEFFF9BFCC13D85FCEB81CB" box="[779,886,698,724]" country="Spain" name="Illes Baleares" pageId="12" pageNumber="13">Baleares</collectingRegion>
</emphasis>
: Ba 8, 15, 17, 19, 21, 23, 24, 25, 26, 27. -
<emphasis id="B901EAB4FFEFFF9BFAAE3D85FF4B81E3" bold="true" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFAAE3D85FF4B81E3" name="Portugal" pageId="12" pageNumber="13">Portugal</collectingCountry>
</emphasis>
: Po 19, 24. -
<emphasis id="B901EAB4FFEFFF9BFEBD3DDDFE5281E3" bold="true" box="[375,463,738,764]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFEBD3DDDFE5281E3" box="[375,463,738,764]" name="France" pageId="12" pageNumber="13">France</collectingCountry>
</emphasis>
: F 2, 3, 4, 5, 8, 11, 12, 13, 13a, 15, 15a, 17, 18, 19, 20, 21, 22, 25, 26a, 34, 39, 41, 42, 48, 63, 79, 91, 99, a, 100. -
<emphasis id="B901EAB4FFEFFF9BFDCC3C35FDFB803B" bold="true" box="[518,614,778,804]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFDCC3C35FDFB803B" box="[518,614,778,804]" name="France" pageId="12" pageNumber="13">Corsica</collectingCountry>
</emphasis>
: 12a, 54a. -
<emphasis id="B901EAB4FFEFFF9BFD323C35FC13803B" bold="true" box="[760,910,778,804]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFD323C35FC13803B" box="[760,910,778,804]" name="Switzerland" pageId="12" pageNumber="13">Switzerland</collectingCountry>
</emphasis>
: He 1, a, 4, a, 5, a, 6, 26, a, 27, 31. -
<emphasis id="B901EAB4FFEFFF9BFA8A3C35FAE7803B" bold="true" box="[1344,1402,778,804]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFA8A3C35FAE7803B" box="[1344,1402,778,804]" name="Italy" pageId="12" pageNumber="13">Italy</collectingCountry>
</emphasis>
: It 90, It 250. -
<emphasis id="B901EAB4FFEFFF9BFEE03C0DFEF38053" bold="true" box="[298,366,818,844]" pageId="12" pageNumber="13">
<collectingRegion id="49B1F844FFEFFF9BFEE03C0DFEF38053" box="[298,366,818,844]" country="Italy" name="Sicilia" pageId="12" pageNumber="13">Sicily</collectingRegion>
</emphasis>
: Sz 11a, 25, 26, 27, 41, 53, 54, 57, 60, 62, 64, 67, 69. -
<emphasis id="B901EAB4FFEFFF9BFBC93C0DFBFF8053" bold="true" box="[1027,1122,818,844]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFBC93C0DFBFF8053" box="[1027,1122,818,844]" name="Croatia" pageId="12" pageNumber="13">Croatia</collectingCountry>
</emphasis>
: Hr 14, 15, 20, 24, 26, 27, 28, 29. -
<emphasis id="B901EAB4FFEFFF9BFECA3C65FEF3806B" bold="true" box="[256,366,858,884]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFECA3C65FEF3806B" box="[256,366,858,884]" name="Morocco" pageId="12" pageNumber="13">Morocco</collectingCountry>
</emphasis>
: Ma 14a, 176a, 258, 259, 264, 273, 288. -
<emphasis id="B901EAB4FFEFFF9BFCA83C65FC22806B" bold="true" box="[866,959,858,884]" pageId="12" pageNumber="13">
<collectingCountry id="F3627636FFEFFF9BFCA83C65FC22806B" box="[866,959,858,884]" name="Algeria" pageId="12" pageNumber="13">Algeria</collectingCountry>
</emphasis>
: Al 14, 16. -
<emphasis id="B901EAB4FFEFFF9BFB963C65FB21806B" bold="true" box="[1116,1212,858,884]" pageId="12" pageNumber="13">
<collectingRegion id="49B1F844FFEFFF9BFB963C65FB21806B" box="[1116,1212,858,884]" country="Algeria" name="Tunesia" pageId="12" pageNumber="13">Tunesia</collectingRegion>
</emphasis>
: Tu 5, 8, 9, 10.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3C9AFD4180F8" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
Due to the difficulties in distinguishing the females of
<taxonomicName id="4C754D25FFEFFF9BFC8F3C9AFC4A80A1" baseAuthorityName="Rambur" baseAuthorityYear="1838" box="[837,983,933,958]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFC8F3C9AFC4A80A1" box="[837,983,933,958]" italics="true" pageId="12" pageNumber="13">P. subaptera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D25FFEFFF9BFBDA3C9AFBE280A1" box="[1040,1151,933,958]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="iberica">
<emphasis id="B901EAB4FFEFFF9BFBDA3C9AFBE280A1" box="[1040,1151,933,958]" italics="true" pageId="12" pageNumber="13">P.iberica</emphasis>
</taxonomicName>
there are some doubtful localities which are indicated by a question mark.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3CCAFE8C87C8" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3CCAFC678710" bold="true" box="[198,1018,1013,1039]" pageId="12" pageNumber="13">
Identification of the
<typeStatus id="54CE8804FFEFFF9BFE0C3CCAFDAC8710" box="[454,561,1013,1039]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
specimen as parthenogenetic female
</emphasis>
. Unfortunately, the abdomen of the
<typeStatus id="54CE8804FFEFFF9BFF5D3B22FF638728" box="[151,254,1053,1079]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
specimen is completely missing. There is, however, strong indirect evidence indicating that it is a representative of the parthenogenetic strain: the locality of the type specimen is situated within an area where only parthenogenetic animals have been found (
<figureCitation id="134E2A23FFEFFF9BFD023B52FC828798" box="[712,799,1133,1159]" captionStart="FIGURE 16" captionStartId="43.[151,255,1744,1768]" captionTargetBox="[302,1270,213,1554]" captionTargetId="figure-36@43.[297,1356,100,1687]" captionTargetPageId="43" captionText="FIGURE 16. Distribution of bisexual species of the subaptera-group and the parthenogenetic species P. subaptera on the Iberian peninsula." figureDoi="http://doi.org/10.5281/zenodo.5087796" httpUri="https://zenodo.org/record/5087796/files/figure.png" pageId="12" pageNumber="13">Fig. 16</figureCitation>
). The only bisexual species occurring in the surroundings is
<taxonomicName id="4C754D25FFEFFF9BFF263BAAFDDB87B1" authorityName="Published" authorityYear="2007" box="[236,582,1173,1198]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quadracantha">
<emphasis id="B901EAB4FFEFFF9BFF263BAAFDDB87B1" box="[236,582,1173,1198]" italics="true" pageId="12" pageNumber="13">Phyllodromica quadracantha</emphasis>
</taxonomicName>
of which the females show a completely different colour pattern on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C36F652DFFEFFF9AFF0C3BDAFB96824D" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="8BCA36A6FFEFFF9BFF0C3BDAFB9987E0" blockId="12.[151,1437,933,1999]" box="[198,1028,1252,1279]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3BDAFE0287E0" bold="true" box="[198,415,1253,1279]" pageId="12" pageNumber="13">Description. Size.</emphasis>
Length of pronotum:
<specimenCount id="9D73FD2FFFEFFF9BFD683BDBFD2B87E7" box="[674,694,1252,1272]" pageId="12" pageNumber="13" type="female">♀</specimenCount>
1,68–2,00 (mean 1,89) mm.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3A32FAA28638" blockId="12.[151,1437,933,1999]" box="[198,1343,1293,1319]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3A32FE958638" bold="true" box="[198,264,1293,1319]" pageId="12" pageNumber="13">Legs.</emphasis>
Distal end of mid tibia with 5 spines: 4 long ones and 1 shorter in ventro-posterior position.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3A0AFE4D8668" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3A0AFEDC8650" bold="true" box="[198,321,1333,1359]" pageId="12" pageNumber="13">Genitalia.</emphasis>
With an additional sclerite in the angle between the intercalary sclerite and the posterior lobe of valvifer II (
<figureCitation id="134E2A23FFEFFF9BFEF73A62FE5F8668" box="[317,450,1373,1399]" captionStart="FIGURE 2" captionStartId="29.[151,255,1612,1636]" captionTargetBox="[270,1329,194,1565]" captionTargetId="figure-141@29.[270,1329,194,1571]" captionTargetPageId="29" captionText="FIGURE 2. Phyllodromica subaptera (female). A Thoracal nota. B Abdominal tergite 5. C Subgenital plate with two apodemal processes (a). D Dorsal complex of genitalia with additional sclerite (as): ventral view, posterior end on top. E Additional sclerite in higher enlargement (equivalent to “black frame” in D). F Ventral complex of genitalia: laterosternal shelf with intersternal folds between the arms. Abbreviations: as additional sclerite, a apodemal process, bd dorsal sclerite of basivalvula, bv ventral sclerite of basivalvula, c cercus, i intersternal fold, is intercalary sclerite, l laterosternal shelf, ls laterosternite IX, pl posterior lobe of valvifer II, pp paraproct, pt paratergites, T9 tergite 9, T10 tergite 10, v valves. Same scale for (B, C) and (D, F). Identification: Sp 203c/W4." figureDoi="http://doi.org/10.5281/zenodo.5087766" httpUri="https://zenodo.org/record/5087766/files/figure.png" pageId="12" pageNumber="13">Fig. 2 D, E</figureCitation>
).
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3ABAFB8C86D8" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3ABAFEF88680" bold="true" box="[198,357,1413,1439]" pageId="12" pageNumber="13">Colouration.</emphasis>
Pro-, meso- and metanotum similarly coloured as in
<taxonomicName id="4C754D25FFEFFF9BFC2B3ABAFBCB8681" box="[993,1110,1413,1438]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="iberica">
<emphasis id="B901EAB4FFEFFF9BFC2B3ABAFBCB8681" box="[993,1110,1413,1438]" italics="true" pageId="12" pageNumber="13">P. iberica</emphasis>
</taxonomicName>
for detailed description see corresponding parts of the section “Characteristics of the
<taxonomicName id="4C754D25FFEFFF9BFCF23A92FC3086D9" baseAuthorityName="Rambur" baseAuthorityYear="1838" box="[824,941,1453,1478]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFCF23A92FC3086D9" box="[824,941,1453,1478]" italics="true" pageId="12" pageNumber="13">subaptera</emphasis>
</taxonomicName>
-group”.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C3AEAFAB48578" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C3AEAFEEC86F0" bold="true" box="[198,369,1493,1519]" pageId="12" pageNumber="13">Genetic Data.</emphasis>
1597 basepairs of the mitochondrial genome including the complete cytochrome c oxidase subunit I (COI) gene were analysed in
<specimenCount id="9D73FD2FFFEFFF9BFD953AC2FC918508" box="[607,780,1533,1559]" pageId="12" pageNumber="13" type="generic">two specimens</specimenCount>
from the
<typeStatus id="54CE8804FFEFFF9BFCB53AC2FC788508" box="[895,997,1533,1559]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
region (Sp 203e/W1, Sp 203e/W10). The sequences of
<taxonomicName id="4C754D25FFEFFF9BFEBD391AFD4E8520" authority="_ Sp" authorityName="Sp" box="[375,723,1573,1599]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFEBD391AFD388521" box="[375,677,1573,1598]" italics="true" pageId="12" pageNumber="13">Phyllodromica subaptera</emphasis>
_Sp
</taxonomicName>
203e/W1_1597bp and
<taxonomicName id="4C754D25FFEFFF9BFC39391AFAD28520" authority="_ Sp" authorityName="Sp" box="[1011,1359,1573,1599]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFC39391AFABC8521" box="[1011,1313,1573,1598]" italics="true" pageId="12" pageNumber="13">Phyllodromica subaptera</emphasis>
_Sp
</taxonomicName>
203e/ W10_1597bp differ in 9 positions (Appendix 5). The complete sequences are shown in Appendix 4.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF5D394AFC078418" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
DNA and associated parts of body are stored in the DNATAX collection of ZSM (Zoologische Staatssammlung München; Munich,
<collectingCountry id="F3627636FFEFFF9BFE3639A2FDF685A8" box="[508,619,1693,1719]" name="Germany" pageId="12" pageNumber="13">Germany</collectingCountry>
) under storage numbers DNATAX02861 (Sp 203e/W10) and DNATAX02862 (Sp 203e/W1). The sequences were submitted to GenBank under the accession numbers:
<accessionNumber id="9426AB45FFEFFF9BFF5D39D2FE838418" box="[151,286,1773,1799]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AM600684" pageId="12" pageNumber="13">AM600684</accessionNumber>
(DNATAX02861) and
<accessionNumber id="9426AB45FFEFFF9BFDFB39D2FD2A8418" box="[561,695,1773,1799]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AM600683" pageId="12" pageNumber="13">AM600683</accessionNumber>
(DNATAX02862).
</paragraph>
<paragraph id="8BCA36A6FFEFFF9BFF0C382AFBF384B8" blockId="12.[151,1437,933,1999]" pageId="12" pageNumber="13">
<emphasis id="B901EAB4FFEFFF9BFF0C382AFD978430" bold="true" box="[198,522,1813,1839]" pageId="12" pageNumber="13">Geographical distribution</emphasis>
(
<figureCitation id="134E2A23FFEFFF9BFDD0382AFDEC8430" box="[538,625,1813,1839]" captionStart="FIGURE 16" captionStartId="43.[151,255,1744,1768]" captionTargetBox="[302,1270,213,1554]" captionTargetId="figure-36@43.[297,1356,100,1687]" captionTargetPageId="43" captionText="FIGURE 16. Distribution of bisexual species of the subaptera-group and the parthenogenetic species P. subaptera on the Iberian peninsula." figureDoi="http://doi.org/10.5281/zenodo.5087796" httpUri="https://zenodo.org/record/5087796/files/figure.png" pageId="12" pageNumber="13">Fig. 16</figureCitation>
). The parthenogenetic species is distributed in most of the Mediterranean countries including
<collectingCountry id="F3627636FFEFFF9BFE0C3802FDB18448" box="[454,556,1853,1879]" name="Portugal" pageId="12" pageNumber="13">Portugal</collectingCountry>
,
<collectingCountry id="F3627636FFEFFF9BFDF13802FD1C8448" box="[571,641,1853,1879]" name="Spain" pageId="12" pageNumber="13">Spain</collectingCountry>
,
<collectingCountry id="F3627636FFEFFF9BFD453802FD7F8448" box="[655,738,1853,1879]" name="France" pageId="12" pageNumber="13">France</collectingCountry>
,
<collectingCountry id="F3627636FFEFFF9BFD3A3802FCBA8448" box="[752,807,1853,1879]" name="Italy" pageId="12" pageNumber="13">Italy</collectingCountry>
,
<collectingCountry id="F3627636FFEFFF9BFCFF3802FC138448" box="[821,910,1853,1879]" name="Croatia" pageId="12" pageNumber="13">Croatia</collectingCountry>
,
<collectingCountry id="F3627636FFEFFF9BFC573802FB978448" box="[925,1034,1853,1879]" name="Morocco" pageId="12" pageNumber="13">Morocco</collectingCountry>
,
<collectingCountry id="F3627636FFEFFF9BFBD23802FBEE8448" box="[1048,1139,1853,1879]" name="Algeria" pageId="12" pageNumber="13">Algeria</collectingCountry>
, Tunesia, and - quite far from the Mediterrenean Sea -
<collectingCountry id="F3627636FFEFFF9BFDCA385AFD118460" box="[512,652,1893,1919]" name="Switzerland" pageId="12" pageNumber="13">Switzerland</collectingCountry>
(in the
<collectingRegion id="49B1F844FFEFFF9BFD21385AFC138460" box="[747,910,1893,1919]" country="Switzerland" name="Valais" pageId="12" pageNumber="13">Wallis region</collectingRegion>
). It also occurs in part of the Mediterranean islands such as Baleares, Sicily,
<collectingCountry id="F3627636FFEFFF9BFDDB38B2FDF084B8" box="[529,621,1933,1959]" name="France" pageId="12" pageNumber="13">Corsica</collectingCountry>
, and the Adriatic islands Korcula and Hvar.
</paragraph>
<paragraph id="8BCA36A6FFEFFF9AFF0C388AFE3683C5" blockId="12.[151,1437,933,1999]" lastBlockId="13.[151,1436,152,338]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<taxonomicName id="4C754D25FFEFFF9BFF0C388AFEC484D1" box="[198,345,1973,1998]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEFFF9BFF0C388AFEC484D1" box="[198,345,1973,1998]" italics="true" pageId="12" pageNumber="13">P. subaptera</emphasis>
</taxonomicName>
is also reported from
<collectingCountry id="F3627636FFEFFF9BFDA8388AFCCE84D0" box="[610,851,1973,1999]" name="Bosnia and Herzegovina" pageId="12" pageNumber="13">Bosnia-Herzegovina</collectingCountry>
and
<collectingCountry id="F3627636FFEFFF9BFC44388AFB8D84D0" box="[910,1040,1973,1999]" name="Macedonia" pageId="12" pageNumber="13">Macedonia</collectingCountry>
(Us &amp; Matjevev 1967),
<collectingCountry id="F3627636FFEFFF9BFAFD388AFA0184D0" box="[1335,1436,1973,1999]" name="Bulgaria" pageId="12" pageNumber="13">Bulgaria</collectingCountry>
(Drenski 1939, Buresch &amp; Peschev 1957) and
<collectingCountry id="F3627636FFEEFF9AFD7D3FA7FC9783AD" box="[695,778,152,178]" name="Greece" pageId="13" pageNumber="14">Greece</collectingCountry>
(Burr et al. 1923, Werner 1927), but we did not see representatives from there.
</paragraph>
<paragraph id="8BCA36A6FFEEFF9AFF0C3FD8FB96824D" blockId="13.[151,1436,152,338]" pageId="13" pageNumber="14">
<emphasis id="B901EAB4FFEEFF9AFF0C3FD8FD41821E" bold="true" box="[198,732,231,257]" pageId="13" pageNumber="14">
Other species of the
<taxonomicName id="4C754D25FFEEFF9AFE0B3FD8FDA5821E" box="[449,568,231,257]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="subaptera">
<emphasis id="B901EAB4FFEEFF9AFE0B3FD8FDA5821E" bold="true" box="[449,568,231,257]" italics="true" pageId="13" pageNumber="14">subaptera</emphasis>
</taxonomicName>
-group found
</emphasis>
. Bisexual females of unknown specificity: Po 19, 24; Sp 88, 89, 90, 91, 94, 108, 333, 393;
<taxonomicName id="4C754D25FFEEFF9AFE353E2FFDED8236" box="[511,624,272,297]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="iberica">
<emphasis id="B901EAB4FFEEFF9AFE353E2FFDED8236" box="[511,624,272,297]" italics="true" pageId="13" pageNumber="14">P. iberica</emphasis>
</taxonomicName>
: morph #1: Sp 419; morph #2: Sp 365; morph #3: Sp148, 186, 381, 395;
<taxonomicName id="4C754D25FFEEFF9AFF183E07FE5F824D" authority=": Sp" authorityName="Sp" box="[210,450,312,338]" class="Insecta" family="Ectobiidae" genus="Phyllodromica" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quadracantha">
<emphasis id="B901EAB4FFEEFF9AFF183E07FE0E824E" box="[210,403,312,337]" italics="true" pageId="13" pageNumber="14">P. quadracantha</emphasis>
: Sp
</taxonomicName>
191, 195, 203, 207, 472, 473, 493, 495, 498, 504.
</paragraph>
</subSubSection>
</treatment>
</document>
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