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<document id="4A27319FB841EC6FD4919F9D1544D40B" ID-DOI="10.5281/zenodo.185020" ID-GBIF-Dataset="30df05c7-bf86-489b-adca-cf70db449d58" ID-ISSN="1175-5326" ID-Zenodo-Dep="185020" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459949045561" checkinUser="plazi" docAuthor="Marin, Ivan" docDate="2009" docId="03DC87A39575F632F1D42A43FB2425D5" docLanguage="en" docName="zt01971p049.pdf" docOrigin="Zootaxa 1971" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laomenes clarki Marin, 2009, sp. nov." docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFE5FFDB9579F623F1432864FFF42206" masterDocTitle="Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity" masterLastPageNumber="49" masterPageNumber="1" pageNumber="13" updateTime="1698230776601" updateUser="plazi">
<mods:mods id="D649ED6506FB902A8935D01EE17EBB4E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C436283FFC39DCF23573B844C23B5260">
<mods:title id="6BFD7F42CE58116EA788DC2D26E392E2">Crinoid-associated shrimps of the genus Laomenes A. H. Clark, 1919 (Caridea: Palaemonidae: Pontoniinae): new species and probable diversity</mods:title>
</mods:titleInfo>
<mods:name id="537417DD39032314E725F6AB8031F173" type="personal">
<mods:role id="3210D76FE6FA2D57431948E3E95DE63F">
<mods:roleTerm id="9A1F000592B1F943F53F9E98480333E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="327CA85B4F7BBEF39F788C4A85238246">Marin, Ivan</mods:namePart>
</mods:name>
<mods:typeOfResource id="F5257564381C4C8AC19520AFC0BB50FD">text</mods:typeOfResource>
<mods:relatedItem id="5835B3F0B300D0808D823F2248CB3B70" type="host">
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<mods:title id="DBC479CDC2686FE39A73E85B75BBF8A6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="321534E0152BB057C46D71C506D48225">
<mods:date id="3EAD2DC66A66CF08670DD10C78C8643B">2009</mods:date>
<mods:detail id="2E64B934CAA783FD91CD5EADB8800CA2" type="volume">
<mods:number id="32AE8FAC0B93711B53CF09EFD00E1411">1971</mods:number>
</mods:detail>
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<mods:classification id="8C3A57E1C571D2C5D36F9901D1852C7C">journal article</mods:classification>
<mods:identifier id="C2E024E633629F9179CBA90E0518C8DF" type="DOI">10.5281/zenodo.185020</mods:identifier>
<mods:identifier id="5115955A9AC84EB4DF27D1803FB15BA7" type="GBIF-Dataset">30df05c7-bf86-489b-adca-cf70db449d58</mods:identifier>
<mods:identifier id="A6EB936B34ED6F094C63807A19D316E2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F6EC17793066CAF9F8BC946DE2237E7C" type="Zenodo-Dep">185020</mods:identifier>
</mods:mods>
<treatment id="03DC87A39575F632F1D42A43FB2425D5" ID-DOI="http://doi.org/10.5281/zenodo.6213140" ID-GBIF-Taxon="119376425" ID-Zenodo-Dep="6213140" LSID="urn:lsid:plazi:treatment:03DC87A39575F632F1D42A43FB2425D5" httpUri="http://treatment.plazi.org/id/03DC87A39575F632F1D42A43FB2425D5" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13">
<subSubSection id="C36F653E9575F62FF1D42A43FEC12062" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BCA36B59575F62FF1D42A43FE312047" blockId="12.[151,453,551,612]" box="[151,453,551,577]" pageId="12" pageNumber="13">
<heading id="D08281D99575F62FF1D42A43FE312047" bold="true" box="[151,453,551,577]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="B901EAA79575F62FF1D42A43FE312047" bold="true" box="[151,453,551,577]" pageId="12" pageNumber="13">
<taxonomicName id="4C754D369575F62FF1D42A43FE942047" ID-CoL="ff7ab485-870b-43eb-8f16-2d465316db28" box="[151,352,551,577]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79575F62FF1D42A43FE942047" bold="true" box="[151,352,551,577]" italics="true" pageId="12" pageNumber="13">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9575F62FF0242A43FE312047" box="[359,453,551,577]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36B59575F62FF1D42A2EFEC12062" blockId="12.[151,453,551,612]" box="[151,309,586,612]" pageId="12" pageNumber="13">
(
<figureCitation id="134E2A309575F62FF1DC2A2EFED82062" box="[159,300,586,612]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStart-3="FIGURE 17" captionStart-4="FIGURE 18" captionStart-5="FIGURE 19" captionStartId-0="18.[151,255,1351,1375]" captionStartId-1="21.[151,255,1948,1972]" captionStartId-2="22.[151,255,1941,1965]" captionStartId-3="23.[151,255,1978,2002]" captionStartId-4="24.[151,255,1107,1131]" captionStartId-5="25.[151,255,1903,1927]" captionTargetBox-0="[177,1425,164,1310]" captionTargetBox-1="[167,1408,1321,1893]" captionTargetBox-2="[190,1417,192,1894]" captionTargetBox-3="[154,1430,189,1922]" captionTargetBox-4="[189,1391,157,1087]" captionTargetBox-5="[153,1436,194,1855]" captionTargetId-0="figure@18.[152,1440,158,1327]" captionTargetId-1="figure@21.[153,1438,1314,1904]" captionTargetId-2="figure@22.[155,1440,166,1917]" captionTargetId-3="figure@23.[152,1437,154,1968]" captionTargetId-4="figure@24.[162,1422,147,1108]" captionTargetId-5="figure@25.[153,1438,194,1857]" captionTargetPageId-0="18" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="25" captionText-0="FIGURE 13. Laomenes clarki sp. nov., male. A, distal part of carapace and rostrum. B, major pereiopod II. C, same, chela. D, minor pereiopod II. E, same, chela. F, pleopod I. G, appendix masculina. Scale 1 mm; F, G, without scale" captionText-1="FIGURE 15. Laomenes pardus sp. nov., ovigerous female. Scale 1 mm." captionText-2="FIGURE 16. Laomenes pardus sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, antennule. D, same, distolateral margin of basal antennular segment. E, distal margin of incisor process of mandible. F, maxillule. Scale 1 mm; E, without scale." captionText-3="FIGURE 17. Laomenes pardus sp. nov., ovigerous female. A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. Scale 1 mm." captionText-4="FIGURE 18. Laomenes pardus sp. nov., ovigerous female. A, pereiopod III. B, C, same, distal part of propodus and dactylus. Scale 1 mm." captionText-5="FIGURE 19. Laomenes pardus sp. nov. A, B, holotype, female, Nhatrang Bay, Vietnam. C, D, paratype, male, same locality. E, female of Laomenes cf. pardus, Manado, Indonesia. F, G, ovigerous females of Laomenes cf. pardus, Tulamben, Indonesia. H, ovigerous female of Laomenes cf. pardus, Cebu, Philippines." httpUri-0="https://zenodo.org/record/185032/files/figure.png" httpUri-1="https://zenodo.org/record/185033/files/figure.png" httpUri-2="https://zenodo.org/record/185034/files/figure.png" httpUri-3="https://zenodo.org/record/185035/files/figure.png" httpUri-4="https://zenodo.org/record/185036/files/figure.png" httpUri-5="https://zenodo.org/record/185037/files/figure.png" pageId="12" pageNumber="13">Figs. 13–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9575F62FF1D42AFEFAD2212A" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BCA36B59575F62FF1D42AFEFAD2212A" blockId="12.[151,1436,666,812]" pageId="12" pageNumber="13">
<emphasis id="B901EAA79575F62FF1D42AFEFEFF20B2" bold="true" box="[151,267,666,692]" pageId="12" pageNumber="13">Material.</emphasis>
<collectingCountry id="F36276259575F62FF0502AFEFE8D20B2" box="[275,377,666,692]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
, Nhatrang Bay: 1 ovigerous female (
<typeStatus id="54CE88179575F62FF26A2AFEFC6020B2" box="[809,916,666,692]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, pcl.
<quantity id="4C8D9B509575F62FF2902AFEFBDB20B2" box="[979,1071,666,692]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="13" unit="mm" value="5.5">5.5 mm</quantity>
) and
<specimenCount id="9D73FD3C9575F62FF5332AFEFB4B20B2" box="[1136,1215,666,692]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(
<typeStatus id="54CE88179575F62FF58C2AFEFACC20B2" box="[1231,1336,666,692]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
, pcl.
<quantity id="4C8D9B509575F62FF4342AFEFF3320DA" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="13" unit="mm" value="3.2">3.2 mm</quantity>
) (
<collectionCode id="ED64AE709575F62FF19C2AA6FEB520DA" box="[223,321,706,732]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="13">ZMMU</collectionCode>
), Nok
<collectingCountry id="F36276259575F62FF0D12AA6FE2920DA" box="[402,477,706,732]" name="Iceland" pageId="12" pageNumber="13">Island</collectingCountry>
,
<quantity id="4C8D9B509575F62FF0A92AA6FDDE20DA" box="[490,554,706,732]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="m" value="15.0">15 m</quantity>
, on crinoid
<taxonomicName id="4C754D369575F62FF3FA2AA6FC3A20DD" box="[697,974,706,732]" class="Crinoidea" family="Comasteridae" genus="Zygometra" kingdom="Animalia" order="Comatulida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="punctata">
<emphasis id="B901EAA79575F62FF3FA2AA6FCCC20DD" box="[697,824,706,731]" italics="true" pageId="12" pageNumber="13">Zygometra</emphasis>
cf.
<emphasis id="B901EAA79575F62FF22B2AA6FC3A20DD" box="[872,974,706,731]" italics="true" pageId="12" pageNumber="13">punctata</emphasis>
</taxonomicName>
,
<date id="FFCB10759575F62FF29C2AA6FBA420DA" box="[991,1104,706,732]" pageId="12" pageNumber="13" value="2006-09-04">4.ix.2006</date>
, coll. I. Marin;
<specimenCount id="9D73FD3C9575F62FF44E2AA6FAAA20DA" box="[1293,1374,706,732]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(pcl.
<quantity id="4C8D9B509575F62FF1D42A8EFF012102" box="[151,245,746,772]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="1.5">1.5 mm</quantity>
) (
<collectionCode id="ED64AE709575F62FF04E2A8EFE9A2102" box="[269,366,746,772]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="13">ZMMU</collectionCode>
), Mun
<collectingCountry id="F36276259575F62FF0872A8EFDFA2102" box="[452,526,746,772]" name="Iceland" pageId="12" pageNumber="13">Island</collectingCountry>
,
<quantity id="4C8D9B509575F62FF35F2A8EFD7C2102" box="[540,648,746,772]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="12" pageNumber="13" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10–12 m</quantity>
, on crinoid
<emphasis id="B901EAA79575F62FF2542A8EFC352105" box="[791,961,746,771]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C754D369575F62FF2542A8EFC622105" box="[791,918,746,771]" class="Crinoidea" family="Comasteridae" genus="Zygometra" kingdom="Animalia" order="Comatulida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="genus">Zygometra</taxonomicName>
sp.
</emphasis>
,
<date id="FFCB10759575F62FF2922A8EFBA52102" box="[977,1105,746,772]" pageId="12" pageNumber="13" value="2006-11-29">29.xi.2006</date>
, coll. I. Marin;
<specimenCount id="9D73FD3C9575F62FF44E2A8EFAAA2102" box="[1293,1374,746,772]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(pcl.
<quantity id="4C8D9B509575F62FF1D42B76FF06212A" box="[151,242,786,812]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
) (
<collectionCode id="ED64AE709575F62FF04A2B76FE9E212A" box="[265,362,786,812]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="13">ZMMU</collectionCode>
), Nok
<collectingCountry id="F36276259575F62FF0FA2B76FDF7212A" box="[441,515,786,812]" name="Iceland" pageId="12" pageNumber="13">Island</collectingCountry>
,
<quantity id="4C8D9B509575F62FF34C2B76FDBA212A" box="[527,590,786,812]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="m" value="10.0">10 m</quantity>
, on crinoid
<taxonomicName id="4C754D369575F62FF39B2B76FC1C212D" box="[728,1000,786,811]" class="Crinoidea" family="Comasteridae" genus="Zygometra" kingdom="Animalia" order="Comatulida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="punctata">
<emphasis id="B901EAA79575F62FF39B2B76FC1C212D" box="[728,1000,786,811]" italics="true" pageId="12" pageNumber="13">Zygometra cf. punctata</emphasis>
</taxonomicName>
,
<date id="FFCB10759575F62FF2B42B76FB83212A" box="[1015,1143,786,812]" pageId="12" pageNumber="13" value="2006-11-25">25.xi.2006</date>
, coll. I. Marin.
</paragraph>
</subSubSection>
<caption id="DF0A663D9575F62FF1D42FC5FC0B25BE" box="[151,1023,1953,1977]" httpUri="https://zenodo.org/record/185028/files/figure.png" pageId="12" pageNumber="13" targetBox="[181,1424,895,1887]" targetPageId="12">
<paragraph id="8BCA36B59575F62FF1D42FC5FC0B25BE" blockId="12.[151,1023,1953,1977]" box="[151,1023,1953,1977]" pageId="12" pageNumber="13">
<emphasis id="B901EAA79575F62FF1D42FC5FEEE25BF" bold="true" box="[151,282,1953,1977]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
<taxonomicName id="4C754D369575F62FF0632FC6FE2425BF" box="[288,464,1954,1977]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79575F62FF0632FC6FE2425BF" box="[288,464,1954,1977]" italics="true" pageId="12" pageNumber="13">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9575F62FF0942FC5FDD925BF" box="[471,557,1953,1977]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B901EAA79575F62FF0942FC5FDD925BF" bold="true" box="[471,557,1953,1977]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
A, ovigerous female. B, male. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF0A663D9574F62EF1862FEAFEE825F4" httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="13" pageNumber="14" targetBox="[189,1406,182,1886]" targetPageId="13">
<paragraph id="8BCA36B59574F62EF1862FEAFEE825F4" blockId="13.[151,1437,1934,2034]" pageId="13" pageNumber="14">
<emphasis id="B901EAA79574F62EF1862FEAFEB625A0" bold="true" box="[197,322,1934,1958]" pageId="13" pageNumber="14">FIGURE 9</emphasis>
.
<taxonomicName id="4C754D369574F62EF0112FEBFDF025A0" box="[338,516,1935,1958]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79574F62EF0112FEBFDF025A0" box="[338,516,1935,1958]" italics="true" pageId="13" pageNumber="14">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9574F62EF34F2FEAFD9725A0" box="[524,611,1934,1958]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B901EAA79574F62EF34F2FEAFD9725A0" bold="true" box="[524,611,1934,1958]" pageId="13" pageNumber="14">sp. nov.</emphasis>
</taxonomicNameLabel>
, ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm.
</paragraph>
</caption>
<caption id="DF0A663D9577F62DF1862DE9FBDD27C1" httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="14" pageNumber="15" targetBox="[162,1411,201,1373]" targetPageId="14">
<paragraph id="8BCA36B59577F62DF1862DE9FBDD27C1" blockId="14.[151,1436,1421,1479]" pageId="14" pageNumber="15">
<emphasis id="B901EAA79577F62DF1862DE9FEA027A3" bold="true" box="[197,340,1421,1445]" pageId="14" pageNumber="15">FIGURE 10.</emphasis>
<taxonomicName id="4C754D369577F62DF0182DE9FDF827A2" box="[347,524,1421,1444]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79577F62DF0182DE9FDF827A2" box="[347,524,1421,1444]" italics="true" pageId="14" pageNumber="15">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9577F62DF3512DE9FD9327A3" box="[530,615,1421,1445]" pageId="14" pageNumber="15" rank="species">
<emphasis id="B901EAA79577F62DF3512DE9FD9327A3" bold="true" box="[530,615,1421,1445]" pageId="14" pageNumber="15">sp. nov.</emphasis>
</taxonomicNameLabel>
, ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm.
</paragraph>
</caption>
<subSubSection id="C36F653E9577F632F1862E46FF2E27F5" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
<paragraph id="8BCA36B59577F62DF1862E46FECD2503" blockId="14.[151,1436,1570,2037]" pageId="14" pageNumber="15">
<emphasis id="B901EAA79577F62DF1862E46FEA9243A" bold="true" box="[197,349,1570,1596]" pageId="14" pageNumber="15">Description.</emphasis>
<emphasis id="B901EAA79577F62DF0292E47FDCD243A" box="[362,569,1571,1596]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="54CE88179577F62DF0292E47FE22243A" box="[362,470,1571,1596]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
female:
</emphasis>
Carapace swollen, smooth, with antennal and prominent hepatic teeth (
<figureCitation id="134E2A309577F62DF1DC2E2FFF112463" box="[159,229,1611,1637]" captionStart="FIGURE 8" captionStartId="12.[151,255,1953,1977]" captionTargetBox="[181,1424,895,1887]" captionTargetId="figure@12.[153,1438,882,1905]" captionTargetPageId="12" captionText="FIGURE 8. Laomenes clarki sp. nov. A, ovigerous female. B, male. Scale 1 mm." httpUri="https://zenodo.org/record/185028/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
A, 9A, B); hepatic tooth larger than antennal; antennal tooth sharp and slender. Rostrum long, slender, slightly compressed, slightly turned down, with 5 teeth along dorsal carina; ventral margin straight, with single tooth; proximal lateral rostral lamina well developed, with conspicuous supraocular teeth (
<figureCitation id="134E2A309577F62DF44B2EFFFAA524B3" box="[1288,1361,1691,1717]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A, B). Orbit well developed, inferior orbital angle slightly produced, rounded distally. Pterygostomial angle rounded, not projected.
</paragraph>
<paragraph id="8BCA36B59577F62DF1862F77FA6225A3" blockId="14.[151,1436,1570,2037]" pageId="14" pageNumber="15">
Abdominal somites smooth; pleura of abdominal somites I–V rounded (
<figureCitation id="134E2A309577F62DF5522F77FBA2252B" box="[1041,1110,1811,1837]" captionStart="FIGURE 8" captionStartId="12.[151,255,1953,1977]" captionTargetBox="[181,1424,895,1887]" captionTargetId="figure@12.[153,1438,882,1905]" captionTargetPageId="12" captionText="FIGURE 8. Laomenes clarki sp. nov. A, ovigerous female. B, male. Scale 1 mm." httpUri="https://zenodo.org/record/185028/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
A). Telson (
<figureCitation id="134E2A309577F62DF5A12F77FADC252B" box="[1250,1320,1811,1837]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
J) about 3 times longer than wide in proximal part, narrowing posteriorly, with 2 pairs of small dorsal spines arising close to lateral margins at 0.5 and 0.75 of telson length; 3 pairs of posterior spines contain short stout lateral, long slender intermediate and slender submedial spines slightly longer than half length of intermediate spines.
</paragraph>
<paragraph id="8BCA36B59577F62DF1862FD7FBC225F3" blockId="14.[151,1436,1570,2037]" pageId="14" pageNumber="15">
Eyes (
<figureCitation id="134E2A309577F62DF04D2FD7FEA725CB" box="[270,339,1971,1997]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
C) large and well developed, with projecting subovate cornea. Central cornea with small blunt apical papilla. Well marked accessory pigmented spot presents posterodorsally.
</paragraph>
<paragraph id="8BCA36B59576F62CF18628FCFF232354" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
Antennule (
<figureCitation id="134E2A309576F62CF01328FCFE6322B4" box="[336,407,152,178]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
D) well developed; basal segment about twice longer than wide, with distolateral angle bearing slender acute distolateral tooth and medial rounded projection (lobe) (
<figureCitation id="134E2A309576F62CF57928A4FB7622DC" box="[1082,1154,192,218]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
F); distinct submarginal ventral tooth situates near medial border of basal segment (
<figureCitation id="134E2A309576F62CF208288CFC642304" box="[843,912,232,258]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
E), with well developed stylocerite; intermediate segment slender, as long as wide, with distolateral margin produced distally; distal segment as long as wide.
</paragraph>
<paragraph id="8BCA36B59576F62CF1862904FBAC23F4" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
Antenna (
<figureCitation id="134E2A309576F62CF07E2904FE72237C" box="[317,390,352,378]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
G) well developed, basicerite with well developed lobe, with large sharp distoventral tooth, overreaching distal margin of the segment for half of its length; carpocerite stout, about twice longer than wide; flagellum well developed; scaphocerite wide, about 3 times as long as maximal width, overreaching intermediate antennular segment, with well developed acute distolateral tooth.
</paragraph>
<paragraph id="8BCA36B59576F62CF1862A64FBEA2624" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
Epistomial horns well developed, sharp (
<figureCitation id="134E2A309576F62CF3E42A64FD19201C" box="[679,749,512,538]" captionStart="FIGURE 9" captionStartId="13.[197,300,1934,1958]" captionTargetBox="[189,1406,182,1886]" captionTargetId="figure@13.[173,1417,162,1918]" captionTargetPageId="13" captionText="FIGURE 9. Laomenes clarki sp. nov., ovigerous female. A, B, distal part of carapace and rostrum. C, eyes. D, E, antennule. F, same, distolateral margin of basal antennular segment. G, antenna. H, epistomial horns. I, thoratic sternites. Scale 1 mm." httpUri="https://zenodo.org/record/185029/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
H). Mandible robust (
<figureCitation id="134E2A309576F62CF2AE2A64FBB6201C" box="[1005,1090,512,538]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
A), without palp; incisor process broad, with 1 large and 8 small terminal teeth (
<figureCitation id="134E2A309576F62CF2442A4CFCAB2044" box="[775,863,552,578]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
B); molar process robust, with stout sharp distal teeth. Maxillule (
<figureCitation id="134E2A309576F62CF0242A34FE4A206C" box="[359,446,592,618]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
C) normal, with well developed bilobed palp; upper lacinia wide, slightly curved, flaring distally, with strong setae along distal margin; lower lacinia more slender, tapering distally, covered with simple setae. Maxilla (
<figureCitation id="134E2A309576F62CF09D2AC4FDC620BC" box="[478,562,672,698]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
D) with well developed, simple palp, pointed distally, with slender endites furnished with stiff, elongated setae; scaphognathite well developed, edging with plumose setae. Maxilliped I (
<figureCitation id="134E2A309576F62CF1DC2A94FF01210C" box="[159,245,752,778]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
E) with completely fused endites, exopod well developed with large caridean lobe bearing plumose setae; epipod divided distally. Maxilliped II with well developed exopod (
<figureCitation id="134E2A309576F62CF54E2B7CFB912134" box="[1037,1125,792,818]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
F); propodus with straight distolateral margin, situated angularly to dactylar segment covered with slender simple setae; dactylus about 3 times longer than broad, with numerous setae along distal margin; epipod square; podobranch absent. Maxilliped III (
<figureCitation id="134E2A309576F62CF0592BF4FE8321AC" box="[282,375,912,938]" captionStart="FIGURE 10" captionStartId="14.[197,300,1421,1445]" captionTargetBox="[162,1411,201,1373]" captionTargetId="figure@14.[153,1438,185,1379]" captionTargetPageId="14" captionText="FIGURE 10. Laomenes clarki sp. nov., ovigerous female. A, mandible. B, same, distal margin of incisor process. C, maxillule. D, maxilla. E, maxilliped I. F, maxilliped II. G, maxilliped III. Scale 1 mm." httpUri="https://zenodo.org/record/185030/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
G) with slender segments, exopod well developed, slightly exceeding ischiomerus; ischiomerus about 4 times longer than maximal width, tapering distally; penultimate segment slender, about 4 times as long as wide; terminal segment slender, about 5 times as long as wide, with tufts of sparse long setae along ventrolateral margins; epipod ovate, lobed; small arthrobranch present.
</paragraph>
<paragraph id="8BCA36B59576F62CF1862C54FBA526EC" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">Pereiopod I (Fig. 11A) with relatively slender chela and fingers; coxal segment with well developed curved lobe distoventrally; merus slender, about 6 times longer than wide; carpus long and slender, about 7 times longer than wide, flaring distally, with several long simple setae at carpo-propodal articulation; palm (Fig. 11B) about 2 times longer than wide, subcylindrical; dactylus and fixed finger about 4 times as long as wide, tapering distally, with cutting edges entire and tips bearing numerous setae.</paragraph>
<paragraph id="8BCA36B59576F62CF1862C9CFAB327DC" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">Pereiopod II equal in size and shape; segments robust (Fig. 11C); coxal segment with well developed curved lobe distoventrally; ischium about 3 times longer than wide; merus about 4 times as long as wide, with straight margins; carpus (Fig. 11D, E) triangular, flaring distally, with distinct notch ventrolaterally; palm cylindrical, about 4 times as long as wide, with straight margins; fingers pointed, with acute, curved tips; fixed fingers slender, with cutting edge bearing small triangular teeth; dactylus slender, about 4 times as long as wide, with cutting edge bearing small but distinct triangular teeth along in its proximal half (Fig. 11F).</paragraph>
<paragraph id="8BCA36B59576F62CF1862D8CFA6924CC" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
Pereiopod III (
<figureCitation id="134E2A309576F62CF03A2D8CFE252404" box="[377,465,1512,1538]" captionStart="FIGURE 12" captionStartId="17.[151,255,1182,1206]" captionTargetBox="[159,1428,204,1136]" captionTargetId="figure@17.[153,1438,194,1140]" captionTargetPageId="17" captionText="FIGURE 12. Laomenes clarki sp. nov., ovigerous female. A, pereiopod III. B, same, dactylus. Scale 1 mm." httpUri="https://zenodo.org/record/185031/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
A) slender, with segments unarmed; ischium about 2.5 times longer than wide; merus about 5 times as long as wide; carpus about 5 times as long as maximal width, tapering distally; propodus about 8 times as long as wide, slightly tapering distally, with straight and smooth margins, with small distoventral spine and tufts of long plumose setae distolaterally; dactylus (
<figureCitation id="134E2A309576F62CF2902E04FBD3247C" box="[979,1063,1632,1658]" captionStart="FIGURE 12" captionStartId="17.[151,255,1182,1206]" captionTargetBox="[159,1428,204,1136]" captionTargetId="figure@17.[153,1438,194,1140]" captionTargetPageId="17" captionText="FIGURE 12. Laomenes clarki sp. nov., ovigerous female. A, pereiopod III. B, same, dactylus. Scale 1 mm." httpUri="https://zenodo.org/record/185031/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
B) elongated, about 1.5 times as long as maximal width, with long, sharp and curved unguis, accessory tooth well developed, turned forward, situated at the middle of ventral margin accompanied with tufts of long slender setae. Pereiopod III–V similar.
</paragraph>
<paragraph id="8BCA36B59576F62CF1862EBCFDC1251C" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">Pleopods normal. Uropods slender, slightly exceeding telson; distolateral margin of uropodal exopod with small fixed tooth and mobile spine.</paragraph>
<paragraph id="8BCA36B59576F62CF1862F4CFB9F25E4" blockId="15.[151,1437,152,2018]" pageId="15" pageNumber="16">
<emphasis id="B901EAA79576F62CF1862F4CFE872547" box="[197,371,1832,1857]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="54CE88179576F62CF1862F4CFEC42547" box="[197,304,1832,1857]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
male
</emphasis>
(
<figureCitation id="134E2A309576F62CF0C52F4CFE3A2544" box="[390,462,1832,1858]" captionStart="FIGURE 8" captionStartId="12.[151,255,1953,1977]" captionTargetBox="[181,1424,895,1887]" captionTargetId="figure@12.[153,1438,882,1905]" captionTargetPageId="12" captionText="FIGURE 8. Laomenes clarki sp. nov. A, ovigerous female. B, male. Scale 1 mm." httpUri="https://zenodo.org/record/185028/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
B, 13A): Generally similar to
<typeStatus id="54CE88179576F62CF27B2F4CFC6A2544" box="[824,926,1832,1858]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
female. Pereiopod II slightly dissimilar in size (
<figureCitation id="134E2A309576F62CF1972F34FEDD256C" box="[212,297,1872,1898]" captionStart="FIGURE 13" captionStartId="18.[151,255,1351,1375]" captionTargetBox="[177,1425,164,1310]" captionTargetId="figure@18.[152,1440,158,1327]" captionTargetPageId="18" captionText="FIGURE 13. Laomenes clarki sp. nov., male. A, distal part of carapace and rostrum. B, major pereiopod II. C, same, chela. D, minor pereiopod II. E, same, chela. F, pleopod I. G, appendix masculina. Scale 1 mm; F, G, without scale" httpUri="https://zenodo.org/record/185032/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
B, D). Fingers of pereiopod II different; right major cheliped shares stout fingers with curved tips (
<figureCitation id="134E2A309576F62CF1DC2F1CFF032594" box="[159,247,1912,1938]" captionStart="FIGURE 13" captionStartId="18.[151,255,1351,1375]" captionTargetBox="[177,1425,164,1310]" captionTargetId="figure@18.[152,1440,158,1327]" captionTargetPageId="18" captionText="FIGURE 13. Laomenes clarki sp. nov., male. A, distal part of carapace and rostrum. B, major pereiopod II. C, same, chela. D, minor pereiopod II. E, same, chela. F, pleopod I. G, appendix masculina. Scale 1 mm; F, G, without scale" httpUri="https://zenodo.org/record/185032/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
C); left minor cheliped with fingers more slender (
<figureCitation id="134E2A309576F62CF21B2F1CFC5B2594" box="[856,943,1912,1938]" captionStart="FIGURE 13" captionStartId="18.[151,255,1351,1375]" captionTargetBox="[177,1425,164,1310]" captionTargetId="figure@18.[152,1440,158,1327]" captionTargetPageId="18" captionText="FIGURE 13. Laomenes clarki sp. nov., male. A, distal part of carapace and rostrum. B, major pereiopod II. C, same, chela. D, minor pereiopod II. E, same, chela. F, pleopod I. G, appendix masculina. Scale 1 mm; F, G, without scale" httpUri="https://zenodo.org/record/185032/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
E). Appendix interna slightly longer than appendix masculina, with numerous subterminal cincinnuli. Appendix masculina (
<figureCitation id="134E2A309576F62CF5252FC4FB4E25BC" box="[1126,1210,1952,1978]" captionStart="FIGURE 13" captionStartId="18.[151,255,1351,1375]" captionTargetBox="[177,1425,164,1310]" captionTargetId="figure@18.[152,1440,158,1327]" captionTargetPageId="18" captionText="FIGURE 13. Laomenes clarki sp. nov., male. A, distal part of carapace and rostrum. B, major pereiopod II. C, same, chela. D, minor pereiopod II. E, same, chela. F, pleopod I. G, appendix masculina. Scale 1 mm; F, G, without scale" httpUri="https://zenodo.org/record/185032/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
G) with 2 long terminal, 2 long subterminal plumose setae and 3 simple marginal setae distomedially.
</paragraph>
<paragraph id="8BCA36B59569F633F1D42FD4FD4325F5" blockId="16.[151,1436,1968,2035]" pageId="16" pageNumber="17">
<emphasis id="B901EAA79569F633F1D42FD4FEC025CC" bold="true" box="[151,308,1968,1994]" pageId="16" pageNumber="17">FIGURE 11.</emphasis>
<taxonomicName id="4C754D369569F633F0782FD5FE0925CC" box="[315,509,1969,1994]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79569F633F0782FD5FE0925CC" box="[315,509,1969,1994]" italics="true" pageId="16" pageNumber="17">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9569F633F3462FD4FD9625CC" box="[517,610,1968,1994]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B901EAA79569F633F3462FD4FD9625CC" bold="true" box="[517,610,1968,1994]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
, ovigerous female. A, pereiopod I. B, same, chela. C, pereiopod II. D, E, same, carpus. F, same, fingers. Scale
<quantity id="4C8D9B509569F633F3282FBDFD4625F5" box="[619,690,2009,2035]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<caption id="DF0A663D9568F632F1D42CFAFAE126B3" box="[151,1301,1182,1206]" httpUri="https://zenodo.org/record/185031/files/figure.png" pageId="17" pageNumber="18" targetBox="[159,1428,204,1136]" targetPageId="17">
<paragraph id="8BCA36B59568F632F1D42CFAFAE126B3" blockId="17.[151,1301,1182,1206]" box="[151,1301,1182,1206]" pageId="17" pageNumber="18">
<emphasis id="B901EAA79568F632F1D42CFAFED226B0" bold="true" box="[151,294,1182,1206]" pageId="17" pageNumber="18">FIGURE 12.</emphasis>
<taxonomicName id="4C754D369568F632F06E2CFAFE2A26B3" box="[301,478,1182,1205]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79568F632F06E2CFAFE2A26B3" box="[301,478,1182,1205]" italics="true" pageId="17" pageNumber="18">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9568F632F0A72CFAFDCE26B0" box="[484,570,1182,1206]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B901EAA79568F632F0A72CFAFDCE26B0" bold="true" box="[484,570,1182,1206]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
, ovigerous female. A, pereiopod III. B, same, dactylus. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8BCA36B59568F632F1862D74FF2E27F5" blockId="17.[151,1437,1296,2003]" pageId="17" pageNumber="18">
<emphasis id="B901EAA79568F632F1862D74FEA0272C" bold="true" box="[197,340,1296,1322]" pageId="17" pageNumber="18">Coloration.</emphasis>
General coloration of the body in females is dark red corresponding to the coloration of the crinoid host (Fig. 14). Distal part of carapace, rostrum, eyestalks, antennule, antenna, telson and uropods lighter red than general body. Carapace with 2 light red longitudinal bands laterally and 2 wide transverse bands dorsally (Fig 14A, B). Several wide red bands transversing each segment of abdomen reaching the beginning of pleura, but never edging ventral margin of pleura (Fig. 14C). Males are completely red (Fig. 14D).
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9568F632F1862E64FD142535" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BCA36B59568F632F1862E64FDB324E4" blockId="17.[151,1437,1296,2003]" pageId="17" pageNumber="18">
<emphasis id="B901EAA79568F632F1862E64FECA241C" bold="true" box="[197,318,1536,1562]" pageId="17" pageNumber="18">Remarks.</emphasis>
<taxonomicName id="4C754D369568F632F00B2E65FDF9241C" box="[328,525,1537,1562]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="clarki" status="sp. nov.">
<emphasis id="B901EAA79568F632F00B2E65FDF9241C" box="[328,525,1537,1562]" italics="true" pageId="17" pageNumber="18">Laomenes clarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9568F632F3552E64FD82241C" box="[534,630,1536,1562]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B901EAA79568F632F3552E64FD82241C" bold="true" box="[534,630,1536,1562]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easy distinguished from the species of the “
<taxonomicName id="4C754D369568F632F5A62E65FA7B241C" box="[1253,1423,1537,1562]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B901EAA79568F632F5A62E65FA7B241C" box="[1253,1423,1537,1562]" italics="true" pageId="17" pageNumber="18">L.amboinensis</emphasis>
</taxonomicName>
” group and other species of the genus by unusually long distoventral tooth on basicerite of antenna and elongated dactylus of pereiopod III. The coloration of this species clearly differs from
<taxonomicName id="4C754D369568F632F51A2E35FAFE246C" box="[1113,1290,1617,1642]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B901EAA79568F632F51A2E35FAFE246C" box="[1113,1290,1617,1642]" italics="true" pageId="17" pageNumber="18">L. amboinensis</emphasis>
</taxonomicName>
by the presence of 2 wide dorsal transverse bands on carapace (vs. 4 thin bands in
<taxonomicName id="4C754D369568F632F2B42E1DFB5D2494" box="[1015,1193,1657,1682]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B901EAA79568F632F2B42E1DFB5D2494" box="[1015,1193,1657,1682]" italics="true" pageId="17" pageNumber="18">L. amboinensis</emphasis>
</taxonomicName>
) and the absence of lighter edging on the lateral and ventral margins of the abdominal pleura. For additional diagnostic diagnosis see remarks under
<taxonomicName id="4C754D369568F632F0302EADFE1624E4" box="[371,482,1737,1762]" class="Malacostraca" family="Palaemonidae" genus="Laomenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="B901EAA79568F632F0302EADFE1624E4" box="[371,482,1737,1762]" italics="true" pageId="17" pageNumber="18">L. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257DC9568F632F0A92EACFDB324E4" box="[490,583,1736,1762]" pageId="17" pageNumber="18" rank="species">
<emphasis id="B901EAA79568F632F0A92EACFDB324E4" bold="true" box="[490,583,1736,1762]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BCA36B59568F632F1862E94FD142535" blockId="17.[151,1437,1296,2003]" pageId="17" pageNumber="18">
<emphasis id="B901EAA79568F632F1862E94FEF3250C" bold="true" box="[197,263,1776,1802]" pageId="17" pageNumber="18">Host.</emphasis>
The specimens were collected from a zygometrid crinoid identified by author as
<taxonomicName id="4C754D369568F632F5912E95FDA22535" authority="A.H. Clark, 1912" authorityName="A.H. Clark" authorityYear="1912" class="Crinoidea" family="Comasteridae" genus="Zygometra" kingdom="Animalia" order="Comatulida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="punctata">
<emphasis id="B901EAA79568F632F5912E95FF312534" italics="true" pageId="17" pageNumber="18">Zygometra punctata</emphasis>
A.H. Clark, 1912 (Zygometridae)
</taxonomicName>
(
<figureCitation id="134E2A309568F632F3262F7DFD482535" box="[613,700,1817,1843]" captionStart="FIGURE 34" captionStartId="42.[151,255,1893,1917]" captionTargetBox="[151,1436,182,1846]" captionTargetId="figure@42.[151,1436,182,1846]" captionTargetPageId="42" captionText="FIGURE 34. Crinoid hosts of the representatives of the genus Laomenes A. H. Clark. A, Himerometra robustipinna (Carpenter, 1881) (Himerometridae). B, Amphimetra tesselata (Müller, 1841) (Himerometridae). C, Oxycomanthus benetti (Müller, 1841) (Comasteridae). D, F, Comantheria (Comanthus) briareus (Bell, 1882) (Comasteridae). E, Zygometra punctata A. H. Clark, 1912 (Zygometridae)." httpUri="https://zenodo.org/record/185052/files/figure.png" pageId="17" pageNumber="18">Fig. 34</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9568F632F1862F24FC6825AD" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BCA36B59568F632F1862F24FC6825AD" blockId="17.[151,1437,1296,2003]" pageId="17" pageNumber="18">
<emphasis id="B901EAA79568F632F1862F24FEA6255C" bold="true" box="[197,338,1856,1882]" pageId="17" pageNumber="18">Etymology.</emphasis>
The species is named in honour of American scientist Dr. Austin H. Clark (
<date id="FFCB10759568F632F5AC2F25FA8D255D" box="[1263,1401,1857,1883]" pageId="17" pageNumber="18" value="1880" valueMax="1954">1880–1954</date>
), a specialist in taxonomy of Echinoderms, who suggested generic name for the genus as well as described crinoid host of the species —
<taxonomicName id="4C754D369568F632F0952FF5FC6C25AD" authority="A.H. Clark, 1912" authorityName="A.H. Clark" authorityYear="1912" box="[470,920,1937,1963]" class="Crinoidea" family="Comasteridae" genus="Zygometra" kingdom="Animalia" order="Comatulida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="punctata">
<emphasis id="B901EAA79568F632F0952FF5FD3625AC" box="[470,706,1937,1962]" italics="true" pageId="17" pageNumber="18">Zygometra punctata</emphasis>
A.H. Clark, 1912
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F653E9568F632F1862FDCFB2425D5" box="[197,1232,1976,2003]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BCA36B59568F632F1862FDCFB2425D5" blockId="17.[151,1437,1296,2003]" box="[197,1232,1976,2003]" pageId="17" pageNumber="18">
<emphasis id="B901EAA79568F632F1862FDCFE9225D4" bold="true" box="[197,358,1976,2002]" pageId="17" pageNumber="18">Distribution.</emphasis>
The species is known only from the
<typeStatus id="54CE88179568F632F2552FDDFCBC25D5" box="[790,840,1977,2003]" pageId="17" pageNumber="18">type</typeStatus>
locality, Nhatrang Bay,
<collectingCountry id="F36276259568F632F5262FDDFB3825D5" box="[1125,1228,1977,2003]" name="Vietnam" pageId="17" pageNumber="18">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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