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<document id="57F582D004700A19987DD6840E4F7683" ID-DOI="10.5281/zenodo.156671" ID-GBIF-Dataset="ad8505f5-c05a-4567-9f98-7618610ae888" ID-ISSN="1175­5326" ID-Zenodo-Dep="156671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459595750768" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="03C287EB1A30FF8DFECEB432A08E70EA" docLanguage="en" docName="zt00165.pdf" docOrigin="Zootaxa 165" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Mayatyphlus Gusarov" docType="treatment" docVersion="9" lastPageNumber="3" masterDocId="FFFBFF931A31FF8FFFC6B774A1097114" masterDocTitle="Mayatyphlus carltoni Gusarov, a new genus and species of leptotyphline staphylinid beetle from Belize (Coleoptera: Staphylinidae: Leptotyphlinae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1698202588245" updateUser="plazi">
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<mods:title id="543469BA0B231D5D5DC4209D15E2A278">Mayatyphlus carltoni Gusarov, a new genus and species of leptotyphline staphylinid beetle from Belize (Coleoptera: Staphylinidae: Leptotyphlinae)</mods:title>
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<treatment id="03C287EB1A30FF8DFECEB432A08E70EA" ID-DOI="http://doi.org/10.5281/zenodo.6277097" ID-GBIF-Taxon="119331234" ID-Zenodo-Dep="6277097" LSID="urn:lsid:plazi:treatment:03C287EB1A30FF8DFECEB432A08E70EA" httpUri="http://treatment.plazi.org/id/03C287EB1A30FF8DFECEB432A08E70EA" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection id="C37165761A30FF8EFECEB432A2007274" box="[264,777,838,864]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BD436FD1A30FF8EFECEB432A2007274" blockId="1.[264,777,838,864]" box="[264,777,838,864]" pageId="1" pageNumber="2">
<heading id="D09C81911A30FF8EFECEB432A2007274" bold="true" box="[264,777,838,864]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B91FEAEF1A30FF8EFECEB432A2007274" bold="true" box="[264,777,838,864]" pageId="1" pageNumber="2">
<taxonomicName id="4C6B4D7E1A30FF8EFECEB432A31E7274" ID-CoL="8XMMP" authority="Gusarov" authorityName="Gusarov" box="[264,535,838,864]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A30FF8EFECEB432A0AD7274" bold="true" box="[264,420,838,864]" italics="true" pageId="1" pageNumber="2">Mayatyphlus</emphasis>
Gusarov
</taxonomicName>
, gen. n. (
<figureCitation id="13502A781A30FF8EFD40B432A2097274" box="[646,768,838,864]" captionStart-0="FIGURES 1 ­ 12" captionStart-1="FIGURES 13 ­ 18" captionStartId-0="2.[264,383,1668,1692]" captionStartId-1="3.[264,383,1768,1792]" captionTargetBox-0="[321,1264,618,1634]" captionTargetBox-1="[352,1236,931,1733]" captionTargetId-0="figure@2.[312,1275,609,1641]" captionTargetId-1="figure@3.[345,1242,924,1742]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." captionText-1="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri-0="https://zenodo.org/record/156672/files/figure.png" httpUri-1="https://zenodo.org/record/156673/files/figure.png" pageId="1" pageNumber="2">Figs. 1­18</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37165761A30FF8EFECEB4E5A5A375D7" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BD436FD1A30FF8EFECEB4E5A5A375D7" blockId="1.[264,1324,913,1899]" pageId="1" pageNumber="2">
<emphasis id="B91FEAEF1A30FF8EFECEB4E5A08172BF" bold="true" box="[264,392,913,939]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
Based on the presence of procoxal fissure, dilated second and third articles of maxillar palpus and the absence of deep transverse basal furrows of abdominal sterna,
<taxonomicName id="4C6B4D7E1A30FF8EFECEB496A0A972EF" box="[264,416,994,1019]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A30FF8EFECEB496A0A972EF" box="[264,416,994,1019]" italics="true" pageId="1" pageNumber="2">Mayatyphlus</emphasis>
</taxonomicName>
is assigned to the tribe
<taxonomicName id="4C6B4D7E1A30FF8EFD7CB495A2FF72EF" authority="Coiffait, 1963" authorityName="Coiffait" authorityYear="1963" box="[698,1014,993,1019]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Neotyphlini">
Neotyphlini
<bibRefCitation id="EFFA4B0C1A30FF8EFC88B495A2FF72EF" author="Coiffait" box="[846,1014,993,1019]" pageId="1" pageNumber="6" refString="Coiffait, H. (1963) Les Leptotyphlitae (Col. Staphylinidae) du Chili. In: Delamare Deboutteville, Cl. &amp; Rapoport, E. (Eds.), Biologie de l'Amerique Australe; Vol. II. Paris: Editions du CNRS, pp. 353 - 369." type="book chapter" year="1963">Coiffait, 1963</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C6B4D7E1A30FF8EFBCCB496A5AB72EF" box="[1034,1186,994,1019]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A30FF8EFBCCB496A5AB72EF" box="[1034,1186,994,1019]" italics="true" pageId="1" pageNumber="2">Mayatyphlus</emphasis>
</taxonomicName>
can be distinguished from the other genera of that tribe by the combination of the following characters: labrum with straight anterior margin and single medial tubercle; mandibles with single subapical tooth; prostheca present; last segment of maxillar palpus short; gular sutures almost contiguous anteriorly; frontal swelling above antennal insertions interrupted in the middle; short prosternal process; and short and narrow parameres.
</paragraph>
</subSubSection>
<subSubSection id="C37165761A30FF8EFEFEB3A6A2CC742F" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BD436FD1A30FF8EFEFEB3A6A2CC742F" blockId="1.[264,1324,913,1899]" pageId="1" pageNumber="2">
<taxonomicName id="4C6B4D7E1A30FF8EFEFEB3A6A0D975FF" ID-CoL="8XMMP" box="[312,464,1234,1259]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A30FF8EFEFEB3A6A0D975FF" box="[312,464,1234,1259]" italics="true" pageId="1" pageNumber="2">Mayatyphlus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6B4D7E1A30FF8EFDBFB3A6A56875FF" ID-CoL="8XKP7" authority="Coiffait &amp; Decou, 1972" authorityName="Coiffait &amp; Decou" authorityYear="1972" box="[633,1121,1233,1259]" class="Insecta" family="Staphylinidae" genus="Cubanotyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A30FF8EFDBFB3A6A22275FF" box="[633,811,1234,1259]" italics="true" pageId="1" pageNumber="2">Cubanotyphlus</emphasis>
<bibRefCitation id="EFFA4B0C1A30FF8EFCFEB3A5A56875FF" author="Coiffait" box="[824,1121,1233,1259]" pageId="1" pageNumber="6" refString="Coiffait, H. &amp; Decou, V. (1972) Recherches sur les Coleopteres hypoges de Cuba. I. Staphylinidae- Leptotyphlinae: Cubanotyphlus jimenezi n. g., n. sp. Revue d'Ecologie et de Biologie du Sol, 9 (1), 131 - 138." type="journal article" year="1972">Coiffait &amp; Decou, 1972</bibRefCitation>
</taxonomicName>
, a genus known from
<collectingCountry id="F37C766D1A30FF8EFE8EB38DA0807407" box="[328,393,1273,1299]" name="Cuba" pageId="1" pageNumber="2">Cuba</collectingCountry>
, Florida and
<collectingCountry id="F37C766D1A30FF8EFDE2B38DA3AF7407" box="[548,678,1273,1299]" name="Guatemala" pageId="1" pageNumber="2">Guatemala</collectingCountry>
, in having labrum with straight anterior margin; antennal article 3 without setae; and short and narrow parameres.
</paragraph>
</subSubSection>
<subSubSection id="C37165761A30FF8DFEFEB23DA2947091" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="8BD436FD1A30FF8EFEFEB23DA5CF7477" blockId="1.[264,1324,913,1899]" box="[312,1222,1353,1379]" pageId="1" pageNumber="2">
<emphasis id="B91FEAEF1A30FF8EFEFEB23DA0D97477" bold="true" box="[312,464,1353,1379]" pageId="1" pageNumber="2">Description.</emphasis>
Length
<quantity id="4C939B181A30FF8EFDFFB23DA3CD7477" box="[569,708,1353,1379]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="2" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7­0.8 mm</quantity>
. Body brownish yellow, poorly pigmented.
</paragraph>
<paragraph id="8BD436FD1A30FF8EFEFEB205A06E77DF" blockId="1.[264,1324,913,1899]" pageId="1" pageNumber="2">
Head with slightly convex sides (
<figureCitation id="13502A781A30FF8EFD08B205A235749F" box="[718,828,1393,1419]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Figs. 1­2</figureCitation>
). Labrum (
<figureCitation id="13502A781A30FF8EFC16B205A510749F" box="[976,1049,1393,1419]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) with straight anterior margin (in exactly dorsal view), with small medial tubercle (visible in oblique dorsal view:
<figureCitation id="13502A781A30FF8EFECEB2B5A04574CF" box="[264,332,1473,1499]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). Mandibles (
<figureCitation id="13502A781A30FF8EFE3EB2B5A36B74CF" box="[504,610,1473,1499]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Figs. 5­6</figureCitation>
) with single subapical tooth and developed prostheca. Maxillar palpus with dilated articles 2 and 3, article 3 with broad base, last article short and narrow (
<figureCitation id="13502A781A30FF8EFEADB165A0BB773F" box="[363,434,1553,1579]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 8</figureCitation>
). Antenna with articles 3­10 transverse, article 3 without setae, articles 4­8 with normal setae arranged in one belt, articles 9­11 with normal setae arranged in two belts and with clavate setae in subapical portions (
<figureCitation id="13502A781A30FF8EFC89B115A29E776F" box="[847,919,1633,1659]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
). Gular sutures almost contiguous anteriorly (
<figureCitation id="13502A781A30FF8EFE4BB1FDA0DE77B7" box="[397,471,1673,1699]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
). Frontal swelling above antennal insertions interrupted in the middle (
<figureCitation id="13502A781A30FF8EFED7B1C5A05F77DF" box="[273,342,1713,1739]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BD436FD1A30FF8EFEFEB1ADA0547657" blockId="1.[264,1324,913,1899]" pageId="1" pageNumber="2">
Procoxal fissure present as anterolateral notch in procoxal cavity (
<figureCitation id="13502A781A30FF8EFB8DB1ADA59D77E7" box="[1099,1172,1753,1779]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
). Prosternal process short (
<figureCitation id="13502A781A30FF8EFE75B075A0F2760F" box="[435,507,1793,1819]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
). Meso­ and metathorax as in
<figureCitation id="13502A781A30FF8EFCAAB075A2BC760F" box="[876,949,1793,1819]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
. Tarsi with three articles (
<figureCitation id="13502A781A30FF8EFB36B075A0597657" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="1" pageNumber="2">Figs. 10­12</figureCitation>
).
</paragraph>
<paragraph id="8BD436FD1A30FF8EFEFEB025A29F767F" blockId="1.[264,1324,913,1899]" box="[312,918,1873,1899]" pageId="1" pageNumber="2">Abdominal sterna without transverse basal furrows.</paragraph>
<paragraph id="8BD436FD1A33FF8DFEFEB668A2947091" blockId="2.[264,1323,284,510]" pageId="2" pageNumber="3">
Male protarsus with adhesive setae (
<figureCitation id="13502A781A33FF8DFD2EB668A2497022" box="[744,832,284,310]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
). Aedeagus, when retracted in abdomen, with basal orifice facing right. Parameres short and narrow (
<figureCitation id="13502A781A33FF8DFC17B630A552704A" box="[977,1115,324,350]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="2" pageNumber="3">Figs. 13­15</figureCitation>
).
<emphasis id="B91FEAEF1A33FF8DFEFEB61FA0D37091" bold="true" box="[312,474,363,389]" pageId="2" pageNumber="3">
<typeStatus id="54D0885F1A33FF8DFEFEB61FA07C7091" box="[312,373,363,389]" pageId="2" pageNumber="3">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6B4D7E1A33FF8DFE2EB618A2467092" authority="Gusarov" authorityName="Gusarov" box="[488,847,364,390]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="carltoni" status="sp. nov.">
<emphasis id="B91FEAEF1A33FF8DFE2EB618A3ED7091" box="[488,740,364,389]" italics="true" pageId="2" pageNumber="3">Mayatyphlus carltoni</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="B91FEAEF1A33FF8DFC9DB61FA2947091" bold="true" box="[859,925,363,389]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A22C57941A33FF8DFC9DB61FA2947091" box="[859,925,363,389]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37165761A33FF8DFEFEB6E7A08E70EA" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BD436FD1A33FF8DFEFEB6E7A08E70EA" blockId="2.[264,1323,284,510]" pageId="2" pageNumber="3">
<emphasis id="B91FEAEF1A33FF8DFEFEB6E7A0CC70B9" bold="true" box="[312,453,403,429]" pageId="2" pageNumber="3">Etymology.</emphasis>
The name
<taxonomicName id="4C6B4D7E1A33FF8DFD9BB6E0A3FC70B9" box="[605,757,404,429]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A33FF8DFD9BB6E0A3FC70B9" box="[605,757,404,429]" italics="true" pageId="2" pageNumber="3">Mayatyphlus</emphasis>
</taxonomicName>
is derived from the word &quot;Maya&quot; (indigenous people of Yucatan and Central
<collectingCountry id="F37C766D1A33FF8DFDB7B6C8A3D170C2" box="[625,728,444,470]" name="United States of America" pageId="2" pageNumber="3">America</collectingCountry>
) and the Greek adjective ó (blind). Gender: masculine.
</paragraph>
</subSubSection>
</treatment>
</document>