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<document id="DFA78C4ABAA8D9893A8E9073748E302D" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A62C0631FE11FDDCFD21E184" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba frontalis CH. Bu Gao &amp; M. Coca-Abia" docType="treatment" docVersion="10" lastPageNumber="28" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="26" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8F7A246659A05E904C78150B8F21DFEC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B7BD9622898E711C123A6C860C0822A6">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:name id="45F9BD692BAF2603ACD0D3D6650A9324" type="personal">
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<mods:namePart id="984A597EE509BF75820476518C18DFD3">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="0B6527029AEE36F0C50AB61EAEF9DA61">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
<mods:nameIdentifier id="CE92750E262BD3CCC183F72F3601219B" type="email">Buzh.Gao@msn.com</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="C9FA110717C9F80D336663E68C9E5ED8">Coca-Abia, María Milagro</mods:namePart>
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<mods:nameIdentifier id="1BF87C47208B0ECBFAE961EA4C936E1F" type="email">mcoca@aragon.es</mods:nameIdentifier>
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<mods:title id="77F9CADCCBD6690A8E1885D90BD3B14F">European Journal of Taxonomy</mods:title>
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<mods:date id="54545736C64AAB0692BDCE92F6D744A0">2021</mods:date>
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<treatment id="03B56743A62C0631FE11FDDCFD21E184" ID-DOI="http://doi.org/10.5281/zenodo.4773243" ID-GBIF-Taxon="182237219" ID-Zenodo-Dep="4773243" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184" httpUri="http://treatment.plazi.org/id/03B56743A62C0631FE11FDDCFD21E184" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="C30685DEA62C0633FE11FDDCFBA8E6A1" box="[490,1098,629,656]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BA3D655A62C0633FE11FDDCFBA8E6A1" blockId="25.[490,1098,629,690]" box="[490,1098,629,656]" pageId="25" pageNumber="26">
<heading id="D0EB6139A62C0633FE11FDDCFBA8E6A1" box="[490,1098,629,656]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4C1CADD6A62C0633FE11FDDCFC7AE6A1" authority="(Fairmaire, 1886)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[490,920,629,656]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62C0633FE11FDDCFD20E6A1" bold="true" box="[490,706,629,656]" italics="true" pageId="25" pageNumber="26">Miridiba frontalis</emphasis>
(
<bibRefCitation id="EF8DABA4A62C0633FD2AFDDCFC72E6A1" author="Fairmaire L." box="[721,912,629,656]" pageId="25" pageNumber="26" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire, 1886</bibRefCitation>
)
</taxonomicName>
(valid species)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA62C0633FD23FD31FCB9E683" box="[728,859,664,690]" pageId="25" pageNumber="26" type="description">
<paragraph id="8BA3D655A62C0633FD23FD31FCB9E683" blockId="25.[490,1098,629,690]" box="[728,859,664,690]" pageId="25" pageNumber="26">
<figureCitation id="1327CAD0A62C0633FD23FD31FCB9E683" box="[728,859,664,690]" captionStart="Figs 34–36" captionStartId="25.[190,242,1916,1942]" captionTargetBox="[279,1311,1081,1866]" captionTargetId="figure-152@25.[264,1323,1078,1876]" captionTargetPageId="25" captionText="Figs 34–36. Miridiba frontalis (Fairmaire, 1886), holotype (MNHN). 34. Habitus in dorsal view. 35. Parameres in dorsal (A), ventral (B) and lateral (C) views. 36. Endophallus (everted) and temones in dorsal view. Scale bars: 34 = 5.0 mm; 35–36 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770344" httpUri="https://zenodo.org/record/4770344/files/figure.png" pageId="25" pageNumber="26">Figs 34–36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA62C0633FF46FD73FCF0E707" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BA3D655A62C0633FF46FD73FC14E6C4" blockId="25.[189,1014,730,757]" box="[189,1014,730,757]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB0CF17BA62C0633FF46FD73FC14E6C4" box="[189,1014,730,757]" pageId="25" pageNumber="26">
<treatmentCitation id="0ABDF044A62C0633FF46FD73FD50E6C4" author="Fairmaire L." box="[189,690,730,757]" page="325" pageId="25" pageNumber="26" year="1886">
<taxonomicName id="4C1CADD6A62C0633FF46FD73FD50E6C4" authority="Fairmaire, 1886: 325" authorityName="Fairmaire" authorityPageNumber="325" authorityYear="1886" box="[189,690,730,757]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62C0633FF46FD73FE51E6C5" box="[189,435,730,756]" italics="true" pageId="25" pageNumber="26">Rhizotrogus frontalis</emphasis>
<bibRefCitation id="EF8DABA4A62C0633FE40FD73FD50E6C4" author="Fairmaire L." box="[443,690,730,757]" pageId="25" pageNumber="26" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire, 1886: 325</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A62C0633FD3AFD73FD17E6C5" box="[705,757,730,756]" pageId="25" pageNumber="26">type</typeStatus>
loc.:
<collectingRegion id="49D818B7A62C0633FCCDFD73FC71E6C5" box="[822,915,730,756]" country="China" name="Yunnan" pageId="25" pageNumber="26">Yunnan</collectingRegion>
,
<collectingCountry id="F30B96C5A62C0633FC65FD73FC0BE6C4" box="[926,1001,730,757]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A62C0633FF46FCB2FCF0E707" blockId="25.[189,786,795,822]" box="[189,786,795,822]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="AB0CF17BA62C0633FF46FCB2FCF0E707" box="[189,786,795,822]" pageId="25" pageNumber="26">
<taxonomicName id="4C1CADD6A62C0633FF46FCB2FE6DE704" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[189,399,795,821]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62C0633FF46FCB2FE6DE704" box="[189,399,795,821]" italics="true" pageId="25" pageNumber="26">Miridiba frontalis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A62C0633FE57FCB2FDBEE704" author="Keith D." box="[428,604,795,822]" page="98" pageId="25" pageNumber="26" year="2005">
<bibRefCitation id="EF8DABA4A62C0633FE57FCB2FDBEE704" author="Keith D." box="[428,604,795,822]" pageId="25" pageNumber="26" pagination="93 - 102" refId="ref44569" refString="Keith D. 2005. Sur quelques Scarabaeoidea (Coleoptera) palearctiques et orientaux. Bulletin mensuel de la Societe linneenne de Lyon 74 (3): 93 - 102. Available from https: // www. persee. fr / doc / linly _ 0366 - 1326 _ 2005 _ num _ 74 _ 3 _ 13586 [accessed 19 Feb. 2021]" type="journal article" year="2005">Keith 2005: 98</bibRefCitation>
</treatmentCitation>
(combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA62C0630FF46FCF5FB82E6DD" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8BA3D655A62C0633FF46FCF5FED4E747" blockId="25.[189,310,860,886]" box="[189,310,860,886]" pageId="25" pageNumber="26">
<heading id="D0EB6139A62C0633FF46FCF5FED4E747" bold="true" box="[189,310,860,886]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B9680A47A62C0633FF46FCF5FED4E747" bold="true" box="[189,310,860,886]" pageId="25" pageNumber="26">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A62C0630FF46FC22FB82E6DD" blockId="25.[187,1398,907,1004]" lastBlockId="26.[187,1400,265,749]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
External morphology of adult (
<figureCitation id="1327CAD0A62C0633FDB9FC22FD43E794" box="[578,673,907,933]" captionStart="Figs 34–36" captionStartId="25.[190,242,1916,1942]" captionTargetBox="[279,1311,1081,1866]" captionTargetId="figure-152@25.[264,1323,1078,1876]" captionTargetPageId="25" captionText="Figs 34–36. Miridiba frontalis (Fairmaire, 1886), holotype (MNHN). 34. Habitus in dorsal view. 35. Parameres in dorsal (A), ventral (B) and lateral (C) views. 36. Endophallus (everted) and temones in dorsal view. Scale bars: 34 = 5.0 mm; 35–36 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770344" httpUri="https://zenodo.org/record/4770344/files/figure.png" pageId="25" pageNumber="26">Fig. 34</figureCitation>
). Body size
<quantity id="4CE47BB0A62C0633FCBFFC22FC00E797" box="[836,994,907,934]" metricMagnitude="-2" metricUnit="m" metricValue="1.825" metricValueMax="1.95" metricValueMin="1.7" pageId="25" pageNumber="26" unit="mm" value="18.25" valueMax="19.5" valueMin="17.0">17–19.5 mm</quantity>
. Body surface with conspicuous pubescence on frons, basal part of elytra and pygidium. Clypeus shorter than frons, concave, with anterior edge emarginate at middle. Frontal carina well developed and sharp. Pronotum without pubescence on disc; anterior margin with short setae, without concavity at each lateral end; lateral margins smooth and glabrous, moderately sinuate at posterior half; anterior angles acute, not projected forward, elevated laterally; posterior angles obtuse. Prosternal process quadrangular-shaped. Scutellum glabrous, with punctures except lateral margins. Elytra with conspicuous pubescence on basal part, disc glabrous; epipleuron with pubescence except on apex. Foretibia with strong and sharp carina; insertion of inner spur closer third outer tooth than second one. Meso- and metatibia with transverse carina interrupted at middle, with a marked dorsal and ventral prominence; inner margin of dorsal surface with strong spines. Prepygidium regularly punctate, conspicuous short pubescence. Pygidium with punctures of irregular size and distribution; conspicuous pubescence on disc, longer toward apex; apical margin moderately widened and raised. Ventrites with conspicuous pubescence; ventrite 5 moderately depressed at posterior half, with pubescence longer on depressed part. Ventrite 6 flat, with diffuse punctures. Male genitalia. Parameres (
<figureCitation id="1327CAD0A62F0630FE47FD25FDF3E696" box="[444,529,652,679]" captionStart="Figs 34–36" captionStartId="25.[190,242,1916,1942]" captionTargetBox="[279,1311,1081,1866]" captionTargetId="figure-152@25.[264,1323,1078,1876]" captionTargetPageId="25" captionText="Figs 34–36. Miridiba frontalis (Fairmaire, 1886), holotype (MNHN). 34. Habitus in dorsal view. 35. Parameres in dorsal (A), ventral (B) and lateral (C) views. 36. Endophallus (everted) and temones in dorsal view. Scale bars: 34 = 5.0 mm; 35–36 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770344" httpUri="https://zenodo.org/record/4770344/files/figure.png" pageId="26" pageNumber="27">Fig. 35</figureCitation>
) with apices of dorsal branches moderately curveted; without pubescence or granules. Endophallus (
<figureCitation id="1327CAD0A62F0630FE0DFD06FDACE6FB" box="[502,590,687,714]" captionStart="Figs 34–36" captionStartId="25.[190,242,1916,1942]" captionTargetBox="[279,1311,1081,1866]" captionTargetId="figure-152@25.[264,1323,1078,1876]" captionTargetPageId="25" captionText="Figs 34–36. Miridiba frontalis (Fairmaire, 1886), holotype (MNHN). 34. Habitus in dorsal view. 35. Parameres in dorsal (A), ventral (B) and lateral (C) views. 36. Endophallus (everted) and temones in dorsal view. Scale bars: 34 = 5.0 mm; 35–36 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770344" httpUri="https://zenodo.org/record/4770344/files/figure.png" pageId="26" pageNumber="27">Fig. 36</figureCitation>
) with raspula of setae and spines larger toward distal end. Temones (
<figureCitation id="1327CAD0A62F0630FF3CFD7BFEFFE6DC" box="[199,285,722,749]" captionStart="Figs 34–36" captionStartId="25.[190,242,1916,1942]" captionTargetBox="[279,1311,1081,1866]" captionTargetId="figure-152@25.[264,1323,1078,1876]" captionTargetPageId="25" captionText="Figs 34–36. Miridiba frontalis (Fairmaire, 1886), holotype (MNHN). 34. Habitus in dorsal view. 35. Parameres in dorsal (A), ventral (B) and lateral (C) views. 36. Endophallus (everted) and temones in dorsal view. Scale bars: 34 = 5.0 mm; 35–36 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770344" httpUri="https://zenodo.org/record/4770344/files/figure.png" pageId="26" pageNumber="27">Fig. 36</figureCitation>
) with apophysis reduced and separated at distal end. Female unknown.
</paragraph>
</subSubSection>
<caption id="DF6386DDA62C0633FF45F8D5FCAAE3EC" ID-DOI="http://doi.org/10.5281/zenodo.4770344" ID-Zenodo-Dep="4770344" httpUri="https://zenodo.org/record/4770344/files/figure.png" pageId="25" pageNumber="26" startId="25.[190,242,1916,1942]" targetBox="[279,1311,1081,1866]" targetPageId="25">
<paragraph id="8BA3D655A62C0633FF45F8D5FCAAE3EC" blockId="25.[188,1399,1916,2013]" pageId="25" pageNumber="26">
<emphasis id="B9680A47A62C0633FF45F8D5FEB2E3A7" bold="true" box="[190,336,1916,1942]" pageId="25" pageNumber="26">Figs 34–36.</emphasis>
<taxonomicName id="4C1CADD6A62C0633FEA6F8D5FCF9E3A6" authority="(Fairmaire, 1886)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[349,795,1916,1943]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62C0633FEA6F8D5FDD7E3A7" box="[349,565,1916,1942]" italics="true" pageId="25" pageNumber="26">Miridiba frontalis</emphasis>
(
<bibRefCitation id="EF8DABA4A62C0633FDB0F8D5FCF0E3A6" author="Fairmaire L." box="[587,786,1916,1943]" pageId="25" pageNumber="26" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire, 1886</bibRefCitation>
)
</taxonomicName>
, holotype (MNHN).
<emphasis id="B9680A47A62C0633FBD8F8D5FBA2E3A7" bold="true" box="[1059,1088,1916,1942]" pageId="25" pageNumber="26">34</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A62C0633FF46F836FF38E388" bold="true" box="[189,218,1951,1977]" pageId="25" pageNumber="26">35</emphasis>
. Parameres in dorsal (A), ventral (B) and lateral (C) views.
<emphasis id="B9680A47A62C0633FC6AF836FC4CE388" bold="true" box="[913,942,1951,1977]" pageId="25" pageNumber="26">36</emphasis>
. Endophallus (everted) and temones in dorsal view. Scale bars: 34 = 5.0 mm; 35–36 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA62F0630FF46FCBDFB61E085" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BA3D655A62F0630FF46FCBDFE48E71F" blockId="26.[189,426,788,814]" box="[189,426,788,814]" pageId="26" pageNumber="27">
<heading id="D0EB6139A62F0630FF46FCBDFE48E71F" bold="true" box="[189,426,788,814]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B9680A47A62F0630FF46FCBDFE48E71F" bold="true" box="[189,426,788,814]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A62F0630FF11FCEAFEBEE76C" blockId="26.[188,1399,835,967]" box="[234,348,835,861]" pageId="26" pageNumber="27">
<emphasis id="B9680A47A62F0630FF11FCEAFEBEE76C" bold="true" box="[234,348,835,861]" pageId="26" pageNumber="27">
<typeStatus id="54A768F7A62F0630FF11FCEAFEBEE76C" box="[234,348,835,861]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA3D655A62F0630FF46FCCFFC9BE7F6" blockId="26.[188,1399,835,967]" pageId="26" pageNumber="27">
<materialsCitation id="3B74DC08A62F0630FF46FCCFFC9BE7F6" ID-GBIF-Occurrence="3114121369" collectionCode="MNHN" collectorName="Coll. Leon Fairmaire" country="China" determinerName="CH. Bu Gao &amp; M. Coca-Abia &amp; Miridiba castanea &amp; Denis Keith" location="Yunnan" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<collectingCountry id="F30B96C5A62F0630FF46FCCFFEFAE7B1" box="[189,280,870,896]" name="China" pageId="26" pageNumber="27">CHINA</collectingCountry>
•
<specimenCount id="9D1A1DDCA62F0630FEC9FCCFFEA9E7B1" box="[306,331,870,896]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
; “
<collectingRegion id="49D818B7A62F0630FE99FCCFFE27E7B1" box="[354,453,870,896]" country="China" name="Yunnan" pageId="26" pageNumber="27">Yunnan</collectingRegion>
”; “Museum Paris 1906
<collectorName id="26E9B383A62F0630FD20FCCFFC3BE7B1" box="[731,985,870,896]" pageId="26" pageNumber="27">Coll. Lèon Fairmaire</collectorName>
”; “
<taxonomicName id="4C1CADD6A62F0630FC07FCCFFA8AE7B1" authority="Fair Yunan" authorityName="Fair Yunan" box="[1020,1384,870,896]" class="Insecta" family="Scarabaeidae" genus="Rhizotrog" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FC07FCCFFB3BE7B1" box="[1020,1241,870,896]" italics="true" pageId="26" pageNumber="27">Rhizotrog frontalis</emphasis>
Fair Yunan
</taxonomicName>
”; “
<typeStatus id="54A768F7A62F0630FF3CFC20FEEBE792" box="[199,265,905,931]" pageId="26" pageNumber="27">Type</typeStatus>
”; “
<typeStatus id="54A768F7A62F0630FED4FC20FE7DE792" box="[303,415,905,931]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C1CADD6A62F0630FE50FC20FB06E792" authority="(Fairmaire, 1886) CH. Bu Gao &amp; M. Coca-Abia" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[427,1252,905,932]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FE50FC20FD60E792" box="[427,642,905,931]" italics="true" pageId="26" pageNumber="27">Miridiba frontalis</emphasis>
(
<bibRefCitation id="EF8DABA4A62F0630FD6DFC20FCBBE795" author="Fairmaire L." box="[662,857,905,932]" pageId="26" pageNumber="27" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire, 1886</bibRefCitation>
)
<determinerName id="7E7F3A91A62F0630FC96FC20FBEBE792" box="[877,1033,905,931]" pageId="26" pageNumber="27">CH. Bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A62F0630FBCCFC20FB06E792" box="[1079,1252,905,931]" pageId="26" pageNumber="27">M. Coca-Abia</determinerName>
</taxonomicName>
det. 2018”; “
<taxonomicName id="4C1CADD6A62F0630FF31FC05FD65E7F7" authority="(Wat.) Denis Keith" authorityName="Denis Keith" baseAuthorityName="Wat." box="[202,647,940,966]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B9680A47A62F0630FF31FC05FE40E7F7" box="[202,418,940,966]" italics="true" pageId="26" pageNumber="27">
<determinerName id="7E7F3A91A62F0630FF31FC05FE40E7F7" box="[202,418,940,966]" pageId="26" pageNumber="27">Miridiba castanea</determinerName>
</emphasis>
(Wat.)
<determinerName id="7E7F3A91A62F0630FE01FC05FD65E7F7" box="[506,647,940,966]" pageId="26" pageNumber="27">Denis Keith</determinerName>
</taxonomicName>
det. 2003”;
<collectionCode id="ED0D4E90A62F0630FCE3FC04FC94E7F6" box="[792,886,941,967]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A62F0630FF11FC44FE44E036" blockId="26.[189,1398,1005,1205]" box="[234,422,1005,1031]" pageId="26" pageNumber="27">
<emphasis id="B9680A47A62F0630FF11FC44FE44E036" bold="true" box="[234,422,1005,1031]" pageId="26" pageNumber="27">Other material</emphasis>
</paragraph>
<paragraph id="8BA3D655A62F0630FF46FBB9FB61E085" blockId="26.[189,1398,1005,1205]" pageId="26" pageNumber="27">
<materialsCitation id="3B74DC08A62F0630FF46FBB9FABFE01B" ID-GBIF-Occurrence="3114121323" box="[189,1373,1039,1066]" collectingDate="1979-10-06" collectionCode="SYAU" collectorName="C. - J. Yang" country="China" determinerName="C. - B. Gao" location="Guiyang" pageId="26" pageNumber="27" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Guizhou Prov.">
<collectingCountry id="F30B96C5A62F0630FF46FBB9FEFAE01B" box="[189,280,1040,1066]" name="China" pageId="26" pageNumber="27">CHINA</collectingCountry>
–
<emphasis id="B9680A47A62F0630FECEFBA6FE0BE01B" bold="true" box="[309,489,1039,1066]" pageId="26" pageNumber="27">
<collectingRegion id="49D818B7A62F0630FECEFBA6FE0BE01B" box="[309,489,1039,1066]" country="China" name="Guizhou" pageId="26" pageNumber="27">Guizhou Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="9D1A1DDCA62F0630FDFAFBB9FDA7E01B" box="[513,581,1040,1066]" pageId="26" pageNumber="27" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D1A1DDCA62F0630FDAAFBB9FD9CE01B" box="[593,638,1040,1066]" pageId="26" pageNumber="27" type="female">1 ♀</specimenCount>
;
<location id="8EC3808EA62F0630FD77FBB9FD17E01B" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184:8EC3808EA62F0630FD77FBB9FD17E01B" box="[652,757,1040,1066]" country="China" name="Guiyang" pageId="26" pageNumber="27" stateProvince="Guizhou Prov.">Guiyang</location>
;
<date id="FFA2F095A62F0630FCFBFBB9FC6FE01B" box="[768,909,1040,1066]" pageId="26" pageNumber="27" value="1979-10-06">
<collectingDate id="EFE6097DA62F0630FCFBFBB9FC6FE01B" box="[768,909,1040,1066]" pageId="26" pageNumber="27" value="1979-10-06">6 Oct. 1979</collectingDate>
</date>
;
<collectorName id="26E9B383A62F0630FC63FBB9FBF1E01B" box="[920,1043,1040,1066]" pageId="26" pageNumber="27">C.-J. Yang</collectorName>
leg.;
<determinerName id="7E7F3A91A62F0630FBAFFBB9FB29E01B" box="[1108,1227,1040,1066]" pageId="26" pageNumber="27">C.-B. Gao</determinerName>
det.;
<collectionCode id="ED0D4E90A62F0630FAF7FBB9FABFE01B" box="[1292,1373,1040,1066]" pageId="26" pageNumber="27">SYAU</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B74DC08A62F0630FF46FB9BFC87E07D" ID-GBIF-Occurrence="3114121354" box="[189,869,1072,1101]" collectionCode="SYAU" country="China" determinerName="C. - B. Gao" location="Hainan" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Hainan Prov.">
<emphasis id="B9680A47A62F0630FF46FB9BFE83E07D" bold="true" box="[189,353,1074,1100]" pageId="26" pageNumber="27">
<collectingRegion id="49D818B7A62F0630FF46FB9BFE83E07D" box="[189,353,1074,1100]" country="China" name="Hainan" pageId="26" pageNumber="27">Hainan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="9D1A1DDCA62F0630FE8CFB9BFE41E07D" box="[375,419,1072,1100]" pageId="26" pageNumber="27" type="male">
1
<emphasis id="B9680A47A62F0630FE76FB99FE41E07D" box="[397,419,1072,1100]" italics="true" pageId="26" pageNumber="27">♂</emphasis>
</specimenCount>
;
<location id="8EC3808EA62F0630FE4AFB9BFDEBE07D" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184:8EC3808EA62F0630FE4AFB9BFDEBE07D" box="[433,521,1074,1100]" country="China" name="Hainan" pageId="26" pageNumber="27" stateProvince="Hainan Prov.">Hainan</location>
; 1952;
<determinerName id="7E7F3A91A62F0630FDA6FB9BFD36E07D" box="[605,724,1074,1100]" pageId="26" pageNumber="27">C.-B. Gao</determinerName>
det.;
<collectionCode id="ED0D4E90A62F0630FCEFFB9BFC87E07D" box="[788,869,1074,1100]" pageId="26" pageNumber="27">SYAU</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B74DC08A62F0630FC79FB9BFBB2E05E" ID-GBIF-Occurrence="3114121325" collectingDate="2005-05-08" collectionCode="SYAU" collectorName="J. - Y. Qiu" country="China" county="Xichang" determinerName="C. - B. Gao" elevation="1700" location="Yuhuangdian" municipality="Mt Lushan" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan Prov.">
<emphasis id="B9680A47A62F0630FC79FB9BFBCCE07D" bold="true" box="[898,1070,1074,1100]" pageId="26" pageNumber="27">
<collectingRegion id="49D818B7A62F0630FC79FB9BFBCCE07D" box="[898,1070,1074,1100]" country="China" name="Sichuan" pageId="26" pageNumber="27">Sichuan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="9D1A1DDCA62F0630FBBDFB9BFB96E07C" box="[1094,1140,1074,1101]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
;
<collectingCounty id="62C2AED9A62F0630FB84FB9BFB07E07D" box="[1151,1253,1074,1100]" pageId="26" pageNumber="27">Xichang</collectingCounty>
,
<collectingMunicipality id="6BC74C2FA62F0630FB0BFB9AFA90E07D" box="[1264,1394,1074,1101]" pageId="26" pageNumber="27">Mt Lushan</collectingMunicipality>
,
<location id="8EC3808EA62F0630FF46FBFCFEBEE05E" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184:8EC3808EA62F0630FF46FBFCFEBEE05E" box="[189,348,1109,1135]" country="China" county="Xichang" municipality="Mt Lushan" name="Yuhuangdian" pageId="26" pageNumber="27" stateProvince="Sichuan Prov.">Yuhuangdian</location>
;
<elevation id="00313166A62F0630FE9DFBFCFE1EE05E" box="[358,508,1109,1136]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="27" unit="m" value="1700.0">
<quantity id="4CE47BB0A62F0630FE9DFBFCFE5CE041" box="[358,446,1109,1136]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="27" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA2F095A62F0630FDF3FBFCFD75E041" box="[520,663,1109,1136]" pageId="26" pageNumber="27" value="2005-05-08">
<collectingDate id="EFE6097DA62F0630FDF3FBFCFD75E041" box="[520,663,1109,1136]" pageId="26" pageNumber="27" value="2005-05-08">8 May 2005</collectingDate>
</date>
;
<collectorName id="26E9B383A62F0630FD59FBFFFCEBE05E" box="[674,777,1109,1136]" pageId="26" pageNumber="27">J.-Y. Qiu</collectorName>
leg.;
<determinerName id="7E7F3A91A62F0630FCB2FBFCFC5DE05E" box="[841,959,1109,1135]" pageId="26" pageNumber="27">C.-B. Gao</determinerName>
det.;
<collectionCode id="ED0D4E90A62F0630FC04FBFCFBB2E05E" box="[1023,1104,1109,1135]" pageId="26" pageNumber="27">SYAU</collectionCode>
</materialsCitation>
. –
<materialsCitation id="3B74DC08A62F0630FB97FBFCFC75E0A3" ID-GBIF-Occurrence="3114121380" collectionCode="MNHN" collectorName="A. Davis &amp; Coll. Fairmaire" country="China" determinerName="Fairmaire" location="Yunnan" pageId="26" pageNumber="27" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan Prov.">
<emphasis id="B9680A47A62F0630FB97FBFCFAF4E05E" bold="true" box="[1132,1302,1109,1135]" pageId="26" pageNumber="27">
<collectingRegion id="49D818B7A62F0630FB97FBFCFAF4E05E" box="[1132,1302,1109,1135]" country="China" name="Yunnan" pageId="26" pageNumber="27">Yunnan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="9D1A1DDCA62F0630FAD7FBFCFA92E041" box="[1324,1392,1109,1136]" pageId="26" pageNumber="27" type="male">3 ♂♂</specimenCount>
;
<location id="8EC3808EA62F0630FF46FBD1FEF9E0A3" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184:8EC3808EA62F0630FF46FBD1FEF9E0A3" box="[189,283,1144,1170]" country="China" name="Yunnan" pageId="26" pageNumber="27" stateProvince="Yunnan Prov.">Yunnan</location>
;
<collectorName id="26E9B383A62F0630FED8FBD1FE68E0A3" box="[291,394,1144,1170]" pageId="26" pageNumber="27">A. Davis</collectorName>
leg.;
<determinerName id="7E7F3A91A62F0630FE30FBD1FDDFE0A3" box="[459,573,1144,1170]" pageId="26" pageNumber="27">Fairmaire</determinerName>
det.;
<collectorName id="26E9B383A62F0630FD87FBD1FCD1E0A3" box="[636,819,1144,1170]" pageId="26" pageNumber="27">Coll. Fairmaire</collectorName>
;
<collectionCode id="ED0D4E90A62F0630FCC6FBD1FC75E0A3" box="[829,919,1144,1170]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="27">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B74DC08A62F0630FC55FBD1FE0BE085" ID-GBIF-Occurrence="3114121338" collectionCode="IZCAS" collectorName="C. - W. Li" country="China" determinerName="Y. - W. Zhang" location="Kunming" pageId="26" pageNumber="27" specimenCount="18" specimenCount-female="16" specimenCount-male="2" stateProvince="Yunnan">
<specimenCount id="9D1A1DDCA62F0630FC55FBD1FC13E0A3" box="[942,1009,1144,1170]" pageId="26" pageNumber="27" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D1A1DDCA62F0630FC07FBD1FBAFE0A3" box="[1020,1101,1144,1170]" pageId="26" pageNumber="27" type="female">16 ♀♀</specimenCount>
;
<location id="8EC3808EA62F0630FBA1FBD1FB2FE0A3" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184:8EC3808EA62F0630FBA1FBD1FB2FE0A3" box="[1114,1229,1144,1170]" country="China" name="Kunming" pageId="26" pageNumber="27" stateProvince="Yunnan">Kunming</location>
;
<collectorName id="26E9B383A62F0630FB2CFBD1FADFE0A3" box="[1239,1341,1144,1170]" pageId="26" pageNumber="27">C.-W. Li</collectorName>
leg.;
<determinerName id="7E7F3A91A62F0630FF46FB32FEB0E085" box="[189,338,1178,1205]" pageId="26" pageNumber="27">Y.-W. Zhang</determinerName>
det.;
<collectionCode id="ED0D4E90A62F0630FE6FFB33FE0BE085" box="[404,489,1178,1204]" collectionName="China, Beijing, Chinese Academy of Sciences, Institute of Zoology" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="26" pageNumber="27">IZCAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B74DC08A62F0630FDF9FB33FB9DE085" ID-GBIF-Occurrence="3114121382" box="[514,1151,1178,1205]" collectionCode="IZCAS" country="China" determinerName="C. - B. Gao" location="Shaotong" pageId="26" pageNumber="27" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan">
<specimenCount id="9D1A1DDCA62F0630FDF9FB33FDA4E084" box="[514,582,1178,1205]" pageId="26" pageNumber="27" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D1A1DDCA62F0630FDA9FB33FD77E084" box="[594,661,1178,1205]" pageId="26" pageNumber="27" type="female">2 ♀♀</specimenCount>
;
<location id="8EC3808EA62F0630FD5FFB33FCF5E085" LSID="urn:lsid:plazi:treatment:03B56743A62C0631FE11FDDCFD21E184:8EC3808EA62F0630FD5FFB33FCF5E085" box="[676,791,1178,1204]" country="China" name="Shaotong" pageId="26" pageNumber="27" stateProvince="Yunnan">Shaotong</location>
; 1981;
<determinerName id="7E7F3A91A62F0630FC97FB33FC06E085" box="[876,996,1178,1204]" pageId="26" pageNumber="27">C.-B. Gao</determinerName>
det.;
<collectionCode id="ED0D4E90A62F0630FBDDFB33FB9DE085" box="[1062,1151,1178,1204]" collectionName="China, Beijing, Chinese Academy of Sciences, Institute of Zoology" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="26" pageNumber="27">IZCAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA62F0631FF46FB7DFAB4E17C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BA3D655A62F0630FF46FB7DFECDE0DF" blockId="26.[189,303,1236,1262]" box="[189,303,1236,1262]" pageId="26" pageNumber="27">
<heading id="D0EB6139A62F0630FF46FB7DFECDE0DF" bold="true" box="[189,303,1236,1262]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B9680A47A62F0630FF46FB7DFECDE0DF" bold="true" box="[189,303,1236,1262]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A62F0631FF46FAABFAB4E17C" blockId="26.[187,1400,1281,1967]" lastBlockId="27.[189,1398,1297,1358]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<bibRefCitation id="EF8DABA4A62F0630FF46FAABFEB6E12D" author="Keith D." box="[189,340,1281,1308]" pageId="26" pageNumber="27" pagination="93 - 102" refId="ref44569" refString="Keith D. 2005. Sur quelques Scarabaeoidea (Coleoptera) palearctiques et orientaux. Bulletin mensuel de la Societe linneenne de Lyon 74 (3): 93 - 102. Available from https: // www. persee. fr / doc / linly _ 0366 - 1326 _ 2005 _ num _ 74 _ 3 _ 13586 [accessed 19 Feb. 2021]" type="journal article" year="2005">Keith (2005)</bibRefCitation>
transferred
<taxonomicName id="4C1CADD6A62F0630FE1FFAABFD3BE12D" authorityName="Fairmaire" authorityYear="1886" box="[484,729,1282,1308]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FE1FFAABFD3BE12D" box="[484,729,1282,1308]" italics="true" pageId="26" pageNumber="27">Rhizotrogus frontalis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A62F0630FCF9FAABFC8BE12D" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[770,873,1282,1308]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A62F0630FCF9FAABFC8BE12D" box="[770,873,1282,1308]" italics="true" pageId="26" pageNumber="27">Miridiba</emphasis>
</taxonomicName>
and synonymized it with
<taxonomicName id="4C1CADD6A62F0630FB66FAABFACFE12A" box="[1181,1325,1282,1307]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B9680A47A62F0630FB66FAABFACFE12A" box="[1181,1325,1282,1307]" italics="true" pageId="26" pageNumber="27">M. castanea</emphasis>
</taxonomicName>
based on a series of male specimens labelled with the name of collector (Armand David). Indeed,
<bibRefCitation id="EF8DABA4A62F0630FAFCFA8DFEEDE153" author="Fairmaire L." pageId="26" pageNumber="27" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire (1886)</bibRefCitation>
, in the introduction of the work in which he described
<taxonomicName id="4C1CADD6A62F0630FC7EFAE1FBE4E150" authorityName="Fairmaire" authorityYear="1886" box="[901,1030,1351,1377]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FC7EFAE1FBE4E150" box="[901,1030,1351,1377]" italics="true" pageId="26" pageNumber="27">R. frontalis</emphasis>
</taxonomicName>
, thanked Armand David for the
<taxonomicName id="4C1CADD6A62F0630FF46FAC3FEDEE1B5" authorityName="Linnaeus" authorityYear="1758" box="[189,316,1386,1412]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
specimens received, which could suggest that the types of
<taxonomicName id="4C1CADD6A62F0630FBFAFAC3FB66E1B5" authorityName="Fairmaire" authorityYear="1886" box="[1025,1156,1386,1412]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FBFAFAC3FB66E1B5" box="[1025,1156,1386,1412]" italics="true" pageId="26" pageNumber="27">R. frontalis</emphasis>
</taxonomicName>
are these specimens labelled with the name of that collector. However,
<bibRefCitation id="EF8DABA4A62F0630FCDCFA25FC12E196" author="Fairmaire L." box="[807,1008,1420,1447]" pageId="26" pageNumber="27" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire (1886)</bibRefCitation>
indicated that the description of
<taxonomicName id="4C1CADD6A62F0630FF46FA19FEA3E1F8" authorityName="Fairmaire" authorityYear="1886" box="[189,321,1455,1481]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FF46FA19FEA3E1F8" box="[189,321,1455,1481]" italics="true" pageId="26" pageNumber="27">R. frontalis</emphasis>
</taxonomicName>
was based on only
<specimenCount id="9D1A1DDCA62F0630FDCFFA19FD3BE1F8" box="[564,729,1455,1482]" pageId="26" pageNumber="27" type="generic">one specimen</specimenCount>
; therefore, the series studied by
<bibRefCitation id="EF8DABA4A62F0630FB9AFA06FB1BE1FB" author="Keith D." box="[1121,1273,1455,1482]" pageId="26" pageNumber="27" pagination="93 - 102" refId="ref44569" refString="Keith D. 2005. Sur quelques Scarabaeoidea (Coleoptera) palearctiques et orientaux. Bulletin mensuel de la Societe linneenne de Lyon 74 (3): 93 - 102. Available from https: // www. persee. fr / doc / linly _ 0366 - 1326 _ 2005 _ num _ 74 _ 3 _ 13586 [accessed 19 Feb. 2021]" type="journal article" year="2005">Keith (2005)</bibRefCitation>
cannot be
<typeStatus id="54A768F7A62F0630FF46FA7BFEC4E1DD" box="[189,294,1490,1516]" pageId="26" pageNumber="27" type="syntype">syntypes</typeStatus>
. We have examined a male specimen of
<taxonomicName id="4C1CADD6A62F0630FD16FA7BFC8EE1DD" authorityName="Fairmaire" authorityYear="1886" box="[749,876,1490,1516]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FD16FA7BFC8EE1DD" box="[749,876,1490,1516]" italics="true" pageId="26" pageNumber="27">R. frontalis</emphasis>
</taxonomicName>
preserved in the collection of Léon Fairmaire (MNHN) labelled as a type and with an original label with his determination. Although
<bibRefCitation id="EF8DABA4A62F0630FB4BFA5DFA97E23E" author="Fairmaire L." box="[1200,1397,1524,1551]" pageId="26" pageNumber="27" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire (1886)</bibRefCitation>
thought that this unique specimen was a female judging from its short antennal club, it is actually a male. In fact, and contrary to other
<taxonomicName id="4C1CADD6A62F0630FD97F993FCE0E265" authorityName="Burmeister" authorityYear="1855" box="[620,770,1594,1620]" class="Insecta" family="Melolonthidae" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="tribe" tribe="Rhizotrogini">Rhizotrogini</taxonomicName>
, the species of
<taxonomicName id="4C1CADD6A62F0630FC45F993FBC7E265" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[958,1061,1594,1620]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A62F0630FC45F993FBC7E265" box="[958,1061,1594,1620]" italics="true" pageId="26" pageNumber="27">Miridiba</emphasis>
</taxonomicName>
do no show obvious sexual dimorphism in the antennal club, which is very short in both sexes. On the other hand, we agree with the transfer of
<taxonomicName id="4C1CADD6A62F0630FEC4F929FE23E2A8" authorityName="Fairmaire" authorityYear="1886" box="[319,449,1663,1689]" class="Insecta" family="Melolonthidae" genus="Rhizotrogus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FEC4F929FE23E2A8" box="[319,449,1663,1689]" italics="true" pageId="26" pageNumber="27">R. frontalis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A62F0630FE13F9D6FDACE2A8" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[488,590,1663,1689]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A62F0630FE13F9D6FDACE2A8" box="[488,590,1663,1689]" italics="true" pageId="26" pageNumber="27">Miridiba</emphasis>
</taxonomicName>
, the features of the external morphology and genitalia show that this species belongs to this genus. Moreover, the antenna 9-segmented and parameres with two dorsal and two ventral branches separated from each other, allow us include it in morphotype I “
<taxonomicName id="4C1CADD6A62F0630FB4BF96DFAA3E2EF" box="[1200,1345,1732,1758]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A62F0630FB4BF96DFAA3E2EF" box="[1200,1345,1732,1758]" italics="true" pageId="26" pageNumber="27">Trichophora</emphasis>
</taxonomicName>
”. In contrast, we do not agree with the synonymy of
<taxonomicName id="4C1CADD6A62F0630FD0FF94EFC27E330" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[756,965,1767,1793]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FD0FF94EFC27E330" box="[756,965,1767,1793]" italics="true" pageId="26" pageNumber="27">Miridiba frontalis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A62F0630FBF3F941FB75E330" box="[1032,1175,1768,1793]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B9680A47A62F0630FBF3F941FB75E330" box="[1032,1175,1768,1793]" italics="true" pageId="26" pageNumber="27">M. castanea</emphasis>
</taxonomicName>
proposed by
<bibRefCitation id="EF8DABA4A62F0630FACDF94EFEEFE315" author="Keith D." pageId="26" pageNumber="27" pagination="93 - 102" refId="ref44569" refString="Keith D. 2005. Sur quelques Scarabaeoidea (Coleoptera) palearctiques et orientaux. Bulletin mensuel de la Societe linneenne de Lyon 74 (3): 93 - 102. Available from https: // www. persee. fr / doc / linly _ 0366 - 1326 _ 2005 _ num _ 74 _ 3 _ 13586 [accessed 19 Feb. 2021]" type="journal article" year="2005">Keith (2005)</bibRefCitation>
, who did not sufficiently justify this synonymy. We have compared the type of
<taxonomicName id="4C1CADD6A62F0630FB49F8A3FAD8E315" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[1202,1338,1802,1828]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62F0630FB49F8A3FAD8E315" box="[1202,1338,1802,1828]" italics="true" pageId="26" pageNumber="27">M. frontalis</emphasis>
</taxonomicName>
with those of
<taxonomicName id="4C1CADD6A62F0630FED8F884FE52E377" box="[291,432,1837,1862]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B9680A47A62F0630FED8F884FE52E377" box="[291,432,1837,1862]" italics="true" pageId="26" pageNumber="27">M. castanea</emphasis>
</taxonomicName>
, finding them to differ by the following characters (
<tableCitation id="C69EE3EEA62F0630FBEAF885FB8FE377" box="[1041,1133,1836,1862]" captionStart="Table 2" captionStartId="27.[189,258,275,301]" captionTargetPageId="27" captionText="Table 2. Differences between holotype of M. frontalis (Fairmaire, 1886) and lectotype and paralectotype of M. castanea (Waterhouse, 1875)." httpUri="http://table.plazi.org/id/DF6386DDA62E0631FF46FEBAFD83E561" pageId="26" pageNumber="27" tableUuid="DF6386DDA62E0631FF46FEBAFD83E561">Table 2</tableCitation>
): head (pubescence on clypeus and frons, frontal carina), pronotum (anterior margin, anterior angles), the shape of the prosternal process, elytra (pubescence), foretibia (carina, inner spur), the protarsi and mesotarsi (tufts of setae), parameres (pubescence and granules at branches) and endophallus (internal structures). Accordingly,
<taxonomicName id="4C1CADD6A62E0631FF46FABBFEA4E11A" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[189,326,1297,1323]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62E0631FF46FABBFEA4E11A" box="[189,326,1297,1323]" italics="true" pageId="27" pageNumber="28">M. frontalis</emphasis>
</taxonomicName>
is established herein as valid species. The unique type specimen of
<taxonomicName id="4C1CADD6A62E0631FB9EFABBFEE3E17F" authority="(Fairmaire, 1886)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62E0631FB9EFABBFB0FE11A" box="[1125,1261,1297,1323]" italics="true" pageId="27" pageNumber="28">M. frontalis</emphasis>
(
<bibRefCitation id="EF8DABA4A62E0631FB06FAB8FF18E17F" author="Fairmaire L." pageId="27" pageNumber="28" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire, 1886</bibRefCitation>
)
</taxonomicName>
found in coll. Léon Fairmaire (MNHN) is a
<typeStatus id="54A768F7A62E0631FCF3FA9AFC8CE17C" box="[776,878,1331,1357]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
by monotypy, and is labelled as
<typeStatus id="54A768F7A62E0631FB12FA9AFAB0E17C" box="[1257,1362,1331,1357]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF6386DDA62E0631FF46FEBAFD83E561" ID-Table-UUID="DF6386DDA62E0631FF46FEBAFD83E561" httpUri="http://table.plazi.org/id/DF6386DDA62E0631FF46FEBAFD83E561" pageId="27" pageNumber="28" startId="27.[189,258,275,301]" targetBox="[200,1383,387,1221]" targetIsTable="true" targetPageId="27">
<paragraph id="8BA3D655A62E0631FF46FEBAFD83E561" blockId="27.[188,1398,275,337]" pageId="27" pageNumber="28">
<emphasis id="B9680A47A62E0631FF46FEBAFEFCE51C" bold="true" box="[189,286,275,301]" pageId="27" pageNumber="28">Table 2.</emphasis>
Differences between holotype of
<taxonomicName id="4C1CADD6A62E0631FD58FEBDFBE2E51F" authority="(Fairmaire, 1886)" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[675,1024,275,302]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62E0631FD58FEBDFCC9E51C" box="[675,811,275,301]" italics="true" pageId="27" pageNumber="28">M. frontalis</emphasis>
(
<bibRefCitation id="EF8DABA4A62E0631FCC0FEBAFC1AE51F" author="Fairmaire L." box="[827,1016,275,302]" pageId="27" pageNumber="28" pagination="303 - 356" refId="ref43851" refString="Fairmaire L. 1886. Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe entomologique de France 6 (6): 303 - 356. Available from https: // www. biodiversitylibrary. org / page / 34026721 # page / 309 / mode / 1 up [accessed 18 Feb. 2021]." type="journal article" year="1886">Fairmaire, 1886</bibRefCitation>
)
</taxonomicName>
and lectotype and paralectotype of
<taxonomicName id="4C1CADD6A62E0631FF26FE9EFDB8E560" authority="(Waterhouse, 1875)" authorityName="M. Coca-Abia" baseAuthorityName="Watrerhouse" baseAuthorityYear="1875" box="[221,602,310,337]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B9680A47A62E0631FF26FE9EFE8EE561" box="[221,364,311,336]" italics="true" pageId="27" pageNumber="28">M. castanea</emphasis>
(
<bibRefCitation id="EF8DABA4A62E0631FE80FE9FFDB0E560" author="Waterhouse C. O." box="[379,594,310,337]" pageId="27" pageNumber="28" pagination="71 - 116" refId="ref45987" refString="Waterhouse C. O. 1875. VI. On the Lamellicorn Coleoptera of Japan. Transactions of the Royal Entomological Society of London 23 (1): 71 - 116. https: // doi. org / 10.1111 / j. 1365 - 2311.1875. tb 01901. x" type="journal article" year="1875">Waterhouse, 1875</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA3D655A62E0631FF33FE2AFB96E0F5" pageId="27" pageNumber="28">
<table id="F91C24F5A62EF9D5FF33FE2AFA85E0F4" box="[200,1383,387,1221]" gridcols="3" gridrows="14" pageId="27" pageNumber="28">
<tr id="352CD417A62EF9D5FF33FE2AFA85E5AA" box="[200,1383,387,411]" gridrow="0" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FE2AFDDFE5AA" box="[200,573,387,411]" gridcol="0" gridrow="0" pageId="27" pageNumber="28">
<emphasis id="B9680A47A62E0631FF33FE2AFEDFE5AA" bold="true" box="[200,317,387,411]" pageId="27" pageNumber="28">Character</emphasis>
</th>
<th id="76FDBD6BA62EF9D5FDAEFE2AFC47E5AA" box="[597,933,387,411]" gridcol="1" gridrow="0" pageId="27" pageNumber="28">
<taxonomicName id="4C1CADD6A62E0631FDAEFE2DFD35E5AA" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[597,727,387,411]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B9680A47A62E0631FDAEFE2DFD35E5AA" bold="true" box="[597,727,387,411]" italics="true" pageId="27" pageNumber="28">M. frontalis</emphasis>
</taxonomicName>
</th>
<th id="76FDBD6BA62EF9D5FC46FE2AFA85E5AA" box="[957,1383,387,411]" gridcol="2" gridrow="0" pageId="27" pageNumber="28">
<taxonomicName id="4C1CADD6A62E0631FC46FE2DFBA0E5AA" authorityName="M. Coca-Abia" baseAuthorityName="Watrerhouse" baseAuthorityYear="1875" box="[957,1090,388,411]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B9680A47A62E0631FC46FE2DFBA0E5AA" bold="true" box="[957,1090,388,411]" italics="true" pageId="27" pageNumber="28">M. castanea</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="352CD417A62EF9D5FF33FE69FA85E5E9" box="[200,1383,448,472]" gridrow="1" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FE69FDDFE5E9" box="[200,573,448,472]" gridcol="0" gridrow="1" pageId="27" pageNumber="28">Clypeus length</th>
<td id="76FDBD6BA62EF9D5FDAEFE69FC47E5E9" box="[597,933,448,472]" gridcol="1" gridrow="1" pageId="27" pageNumber="28">shorter than the frons</td>
<td id="76FDBD6BA62EF9D5FC46FE69FA85E5E9" box="[957,1383,448,472]" gridcol="2" gridrow="1" pageId="27" pageNumber="28">as long as the frons</td>
</tr>
<tr id="352CD417A62EF9D5FF33FE55FA85E625" box="[200,1383,508,532]" gridrow="2" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FE55FDDFE625" box="[200,573,508,532]" gridcol="0" gridrow="2" pageId="27" pageNumber="28">Clypeus shape</th>
<td id="76FDBD6BA62EF9D5FDAEFE55FC47E625" box="[597,933,508,532]" gridcol="1" gridrow="2" pageId="27" pageNumber="28">concave</td>
<td id="76FDBD6BA62EF9D5FC46FE55FA85E625" box="[957,1383,508,532]" gridcol="2" gridrow="2" pageId="27" pageNumber="28">flat</td>
</tr>
<tr id="352CD417A62EF9D5FF33FD86FA85E656" box="[200,1383,559,615]" gridrow="3" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FD86FDDFE656" box="[200,573,559,615]" gridcol="0" gridrow="3" pageId="27" pageNumber="28">Frontal pubescence</th>
<td id="76FDBD6BA62EF9D5FDAEFD86FC47E656" box="[597,933,559,615]" gridcol="1" gridrow="3" pageId="27" pageNumber="28">present and long</td>
<td id="76FDBD6BA62EF9D5FC46FD86FA85E656" box="[957,1383,559,615]" gridcol="2" gridrow="3" pageId="27" pageNumber="28">absent or, at most, very short hardly noticeable</td>
</tr>
<tr id="352CD417A62EF9D5FF33FD28FA85E6A8" box="[200,1383,641,665]" gridrow="4" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FD28FDDFE6A8" box="[200,573,641,665]" gridcol="0" gridrow="4" pageId="27" pageNumber="28">Frontal carina</th>
<td id="76FDBD6BA62EF9D5FDAEFD28FC47E6A8" box="[597,933,641,665]" gridcol="1" gridrow="4" pageId="27" pageNumber="28">high, sharp.</td>
<td id="76FDBD6BA62EF9D5FC46FD28FA85E6A8" box="[957,1383,641,665]" gridcol="2" gridrow="4" pageId="27" pageNumber="28">low, not sharp</td>
</tr>
<tr id="352CD417A62EF9D5FF33FD14FA85E6E7" box="[200,1383,701,726]" gridrow="5" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FD14FDDFE6E7" box="[200,573,701,726]" gridcol="0" gridrow="5" pageId="27" pageNumber="28">Pronotal anterior edge</th>
<td id="76FDBD6BA62EF9D5FDAEFD14FC47E6E7" box="[597,933,701,726]" gridcol="1" gridrow="5" pageId="27" pageNumber="28">gently wavy behind eyes</td>
<td id="76FDBD6BA62EF9D5FC46FD14FA85E6E7" box="[957,1383,701,726]" gridcol="2" gridrow="5" pageId="27" pageNumber="28">without wavy behind eyes</td>
</tr>
<tr id="352CD417A62EF9D5FF33FD53FA85E723" box="[200,1383,762,786]" gridrow="6" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FD53FDDFE723" box="[200,573,762,786]" gridcol="0" gridrow="6" pageId="27" pageNumber="28">Anterior angles</th>
<td id="76FDBD6BA62EF9D5FDAEFD53FC47E723" box="[597,933,762,786]" gridcol="1" gridrow="6" pageId="27" pageNumber="28">acute</td>
<td id="76FDBD6BA62EF9D5FC46FD53FA85E723" box="[957,1383,762,786]" gridcol="2" gridrow="6" pageId="27" pageNumber="28">obtuse</td>
</tr>
<tr id="352CD417A62EF9D5FF33FC9FFA85E77F" box="[200,1383,822,846]" gridrow="7" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FC9FFDDFE77F" box="[200,573,822,846]" gridcol="0" gridrow="7" pageId="27" pageNumber="28">Prosternal process</th>
<td id="76FDBD6BA62EF9D5FDAEFC9FFC47E77F" box="[597,933,822,846]" gridcol="1" gridrow="7" pageId="27" pageNumber="28">quadrangular shaped</td>
<td id="76FDBD6BA62EF9D5FC46FC9FFA85E77F" box="[957,1383,822,846]" gridcol="2" gridrow="7" pageId="27" pageNumber="28">rounded with a bulge on ventral face</td>
</tr>
<tr id="352CD417A62EF9D5FF33FCC0FA85E790" box="[200,1383,873,929]" gridrow="8" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FCC0FDDFE790" box="[200,573,873,929]" gridcol="0" gridrow="8" pageId="27" pageNumber="28">Elytral pubescence</th>
<td id="76FDBD6BA62EF9D5FDAEFCC0FC47E790" box="[597,933,873,929]" gridcol="1" gridrow="8" pageId="27" pageNumber="28">Long on basal part</td>
<td id="76FDBD6BA62EF9D5FC46FCC0FA85E790" box="[957,1383,873,929]" gridcol="2" gridrow="8" pageId="27" pageNumber="28">absent or, at most, very short hardly noticeable</td>
</tr>
<tr id="352CD417A62EF9D5FF33FC12FA85E7E2" box="[200,1383,955,979]" gridrow="9" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FC12FDDFE7E2" box="[200,573,955,979]" gridcol="0" gridrow="9" pageId="27" pageNumber="28">Carina of the protibia</th>
<td id="76FDBD6BA62EF9D5FDAEFC12FC47E7E2" box="[597,933,955,979]" gridcol="1" gridrow="9" pageId="27" pageNumber="28">strongly elevated. Sharp</td>
<td id="76FDBD6BA62EF9D5FC46FC12FA85E7E2" box="[957,1383,955,979]" gridcol="2" gridrow="9" pageId="27" pageNumber="28">gently elevated</td>
</tr>
<tr id="352CD417A62EF9D5FF33FC5FFA85E03E" box="[200,1383,1014,1039]" gridrow="10" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FC5FFDDFE03E" box="[200,573,1014,1039]" gridcol="0" gridrow="10" pageId="27" pageNumber="28">Protibia, insertion of the inner spur</th>
<td id="76FDBD6BA62EF9D5FDAEFC5FFC47E03E" box="[597,933,1014,1039]" gridcol="1" gridrow="10" pageId="27" pageNumber="28">closer than 3th than 2nd tooth</td>
<td id="76FDBD6BA62EF9D5FC46FC5FFA85E03E" box="[957,1383,1014,1039]" gridcol="2" gridrow="10" pageId="27" pageNumber="28">equidistant between the 2nd and 3th tooth</td>
</tr>
<tr id="352CD417A62EF9D5FF33FB9DFA85E07D" box="[200,1383,1076,1100]" gridrow="11" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FB9DFDDFE07D" box="[200,573,1076,1100]" gridcol="0" gridrow="11" pageId="27" pageNumber="28">Pro- and metatarsi</th>
<td id="76FDBD6BA62EF9D5FDAEFB9DFC47E07D" box="[597,933,1076,1100]" gridcol="1" gridrow="11" pageId="27" pageNumber="28">conspicuous tufts of setae</td>
<td id="76FDBD6BA62EF9D5FC46FB9DFA85E07D" box="[957,1383,1076,1100]" gridcol="2" gridrow="11" pageId="27" pageNumber="28">inconspicuous tufts of setae</td>
</tr>
<tr id="352CD417A62EF9D5FF33FBD9FA85E0B9" box="[200,1383,1136,1160]" gridrow="12" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FBD9FDDFE0B9" box="[200,573,1136,1160]" gridcol="0" gridrow="12" pageId="27" pageNumber="28">Branches of the parameres</th>
<td id="76FDBD6BA62EF9D5FDAEFBD9FC47E0B9" box="[597,933,1136,1160]" gridcol="1" gridrow="12" pageId="27" pageNumber="28">without pubescence or granules</td>
<td id="76FDBD6BA62EF9D5FC46FBD9FA85E0B9" box="[957,1383,1136,1160]" gridcol="2" gridrow="12" pageId="27" pageNumber="28">with pubescence or granules</td>
</tr>
<tr id="352CD417A62EF9D5FF33FB04FA85E0F4" box="[200,1383,1197,1221]" gridrow="13" pageId="27" pageNumber="28">
<th id="76FDBD6BA62EF9D5FF33FB04FDDFE0F4" box="[200,573,1197,1221]" gridcol="0" gridrow="13" pageId="27" pageNumber="28">Endophallum</th>
<td id="76FDBD6BA62EF9D5FDAEFB04FC47E0F4" box="[597,933,1197,1221]" gridcol="1" gridrow="13" pageId="27" pageNumber="28">with soft spurs</td>
<td id="76FDBD6BA62EF9D5FC46FB04FA85E0F4" box="[957,1383,1197,1221]" gridcol="2" gridrow="13" pageId="27" pageNumber="28">with strong spurs</td>
</tr>
</table>
</paragraph>
<subSubSection id="C30685DEA62E0631FF46FAC4FD21E184" box="[189,707,1389,1461]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BA3D655A62E0631FF46FAC4FEB4E1B6" blockId="27.[189,342,1389,1415]" box="[189,342,1389,1415]" pageId="27" pageNumber="28">
<heading id="D0EB6139A62E0631FF46FAC4FEB4E1B6" bold="true" box="[189,342,1389,1415]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B9680A47A62E0631FF46FAC4FEB4E1B6" bold="true" box="[189,342,1389,1415]" pageId="27" pageNumber="28">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A62E0631FF46FA32FD21E184" blockId="27.[189,707,1435,1461]" box="[189,707,1435,1461]" pageId="27" pageNumber="28">
<collectingCountry id="F30B96C5A62E0631FF46FA32FEE1E184" box="[189,259,1435,1461]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
(
<collectingRegion id="49D818B7A62E0631FEE8FA32FE8EE184" box="[275,364,1435,1461]" country="China" name="Hainan" pageId="27" pageNumber="28">Hainan</collectingRegion>
,
<collectingRegion id="49D818B7A62E0631FE83FA32FE02E184" box="[376,480,1435,1461]" country="China" name="Guizhou" pageId="27" pageNumber="28">Guizhou</collectingRegion>
,
<collectingRegion id="49D818B7A62E0631FE10FA32FDAEE184" box="[491,588,1435,1461]" country="China" name="Sichuan" pageId="27" pageNumber="28">Sichuan</collectingRegion>
,
<collectingRegion id="49D818B7A62E0631FDACFA32FD54E184" box="[599,694,1435,1461]" country="China" name="Yunnan" pageId="27" pageNumber="28">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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