796 lines
120 KiB
XML
796 lines
120 KiB
XML
<document id="BBDEB3C74333C35C96DE2B0A23429159" ID-CLB-Dataset="22100" ID-DOI="10.5852/ejt.2017.321" ID-GBIF-Dataset="caeead37-b6ce-4f03-96d2-ae8c8beb8573" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589562339121" checkinUser="carolina" docAuthor="Maxwell, Erin E. & Stumpf, Sebastian" docDate="2017" docId="03B22572FF9DFF92FDD2F8BEBD97C8A5" docLanguage="en" docName="ejt-2017-321.pdf.imf" docOrigin="European Journal of Taxonomy 321 (321)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Saurorhynchus hauffi Maxwell & Stumpf 2017, sp. nov." docType="treatment" docUuid="CF1D94D7-EE54-4E13-B059-97C2536D27FA" docUuidSource="ZooBank" docVersion="10" lastPageNumber="20" masterDocId="FF8B5D0AFF91FF81FFB1FFF9BF47CD01" masterDocTitle="Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany" masterLastPageNumber="29" masterPageNumber="1" pageNumber="13" updateTime="1698839856834" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C3723BAC9A44E3E2B5AECCD2E294401A">Revision of Saurorhynchus (Actinopterygii: Saurichthyidae) from the Early Jurassic of England and Germany</mods:title>
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<mods:namePart id="4BD61F1DA6F14F8ED6156ADECEE16715">Maxwell, Erin E.</mods:namePart>
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<mods:affiliation id="FDC0067A6CA07FB9B23B135CF051B608">Staatliches Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany. Institut für Geographie und Geologie, Ernst-Moritz-Arndt-Universität Greifswald, Friedrich-Ludwig-Jahn Strasse 17 a, 17489 Greifswald, Germany.</mods:affiliation>
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<mods:nameIdentifier id="4B45E333DF38BFBB9A5236DDED27A9E2" type="email">erin.maxwell@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="7833930D06C8521F54C8235CFD3622CE">Stumpf, Sebastian</mods:namePart>
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<mods:affiliation id="04B1F96BD6A25961F1B6DDABAC393CD3">Email: sebastian. stumpf @ stud. uni-greifswald. de & urn: lsid: zoobank. org: author: 8 DADF 851 - CFA 5 - 469 E- 9628 - AD 020803 B 424</mods:affiliation>
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<mods:nameIdentifier id="1215130B90B23CDA7C69F4EB97E974E1" type="email">sebastian.stumpf@stud.uni-greifswald.de</mods:nameIdentifier>
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<mods:title id="752C510B75ECDFB87A060E3EB734EAB4">European Journal of Taxonomy</mods:title>
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<mods:date id="FA7971FEF8E6707A5DAB393377076784">2017</mods:date>
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<treatment id="03B22572FF9DFF92FDD2F8BEBD97C8A5" ID-DOI="http://doi.org/10.5281/zenodo.3848072" ID-GBIF-Taxon="164159994" ID-Zenodo-Dep="3848072" LSID="urn:lsid:zoobank.org:act:CF1D94D7-EE54-4E13-B059-97C2536D27FA" httpUri="http://treatment.plazi.org/id/03B22572FF9DFF92FDD2F8BEBD97C8A5" lastPageId="19" lastPageNumber="20" pageId="12" pageNumber="13">
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<subSubSection id="C301C7EFFF9DFF8DFDD2F8BEBC96CA63" box="[611,977,1863,1890]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BA49464FF9DFF8DFDD2F8BEBC96CA63" blockId="12.[368,1221,1863,1962]" box="[611,977,1863,1890]" pageId="12" pageNumber="13">
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<heading id="D0EC2308FF9DFF8DFDD2F8BEBC96CA63" box="[611,977,1863,1890]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
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<taxonomicName id="4C1BEFE7FF9DFF8DFDD2F8BEBC28CA63" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[611,879,1863,1890]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
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<emphasis id="B96F4876FF9DFF8DFDD2F8BEBC28CA63" bold="true" box="[611,879,1863,1890]" italics="true" pageId="12" pageNumber="13">Saurorhynchus hauffi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25CF50DFF9DFF8DFCC7F8B1BC96CA63" box="[886,977,1864,1890]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C301C7EFFF9DFF8DFEC1F892BB41CAA8" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BA49464FF9DFF8DFEC1F892BB82CA87" blockId="12.[368,1221,1863,1962]" box="[368,1221,1899,1926]" pageId="12" pageNumber="13">
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<uri id="FF8A9866FF9DFF8DFEC1F892BB82CA87" box="[368,1221,1899,1926]" pageId="12" pageNumber="13">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBDAEB1FF9DFF8DFD20F892BB82CA87" box="[657,1221,1899,1926]" pageId="12" pageNumber="13">CF1D94D7-EE54-4E13-B059-97C2536D27FA</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BA49464FF9DFF8DFD9FF876BB41CAA8" blockId="12.[368,1221,1863,1962]" box="[558,1030,1935,1962]" pageId="12" pageNumber="13">
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<figureCitation id="132088E1FF9DFF8DFD9FF876BD35CAA8" box="[558,626,1935,1961]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="12" pageNumber="13">Figs 1</figureCitation>
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D–E, 2C, 4C, F, 5C–D, 6C, 7B–D
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</paragraph>
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</subSubSection>
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<subSubSection id="C301C7EFFF9DFF8CFF0CF828BEC4CCD0" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="8BA49464FF9DFF8DFF0CF828BC2CCAEA" blockId="12.[189,875,2000,2027]" box="[189,875,2000,2027]" pageId="12" pageNumber="13">
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<treatmentCitationGroup id="AB0BB34AFF9DFF8DFF0CF828BC21CAEA" box="[189,870,2000,2027]" pageId="12" pageNumber="13">
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<treatmentCitation id="0ABAB275FF9DFF8DFF0CF828BC5CCAEA" author="Quenstedt" box="[189,795,2000,2027]" page="29" pageId="12" pageNumber="13" year="1856">
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<taxonomicName id="4C1BEFE7FF9DFF8DFF0CF828BD3BCAEA" authority="Quenstedt, 1856" authorityName="Quenstedt" authorityYear="1856" box="[189,636,2000,2027]" class="Actinopterygii" family="Aspidorhynchidae" genus="Belonostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aspidorhychiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="acutus">
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||
<emphasis id="B96F4876FF9DFF8DFF0CF828BEF1CAEA" box="[189,438,2001,2027]" italics="true" pageId="12" pageNumber="13">Belonostomus acutus</emphasis>
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<bibRefCitation id="EF8AE995FF9DFF8DFE0CF828BD3BCAEA" author="Quenstedt F. A." box="[445,636,2000,2027]" pageId="12" pageNumber="13" refId="ref14382" refString="Quenstedt F. A. 1856 - 1858. Der Jura. H. Laupp, Tubingen." type="book" year="1856">Quenstedt, 1856</bibRefCitation>
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</taxonomicName>
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||
–1858: pl. 29
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||
</treatmentCitation>
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||
, fig. 8
|
||
</treatmentCitationGroup>
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.
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||
</paragraph>
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||
<paragraph id="8BA49464FF9CFF8CFF0CFEF2BC33CC27" blockId="13.[189,884,267,294]" box="[189,884,267,294]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB0BB34AFF9CFF8CFF0CFEF2BC33CC27" box="[189,884,267,294]" pageId="13" pageNumber="14">
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<treatmentCitation id="0ABAB275FF9CFF8CFF0CFEF2BC41CC24" author="Woodward A. S." box="[189,774,267,294]" page="18" pageId="13" pageNumber="14" year="1895">
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<taxonomicName id="4C1BEFE7FF9CFF8CFF0CFEF2BC41CC24" authority="Woodward, 1895: 18" authorityName="Woodward" authorityPageNumber="18" authorityYear="1895" box="[189,774,267,294]" class="Actinopterygii" family="Saurichthyidae" genus="Belonorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="brevirostris">
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<emphasis id="B96F4876FF9CFF8CFF0CFEF2BD4ECC24" box="[189,521,267,293]" italics="true" pageId="13" pageNumber="14">Belonorhynchus brevirostris</emphasis>
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||
<bibRefCitation id="EF8AE995FF9CFF8CFDA1FEF2BC41CC24" author="Woodward A. S." box="[528,774,267,294]" pageId="13" pageNumber="14" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward, 1895: 18</bibRefCitation>
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||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(partim).
|
||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BA49464FF9CFF8CFF0CFEB4BC6ACC66" blockId="13.[189,1399,333,465]" box="[189,813,333,359]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB0BB34AFF9CFF8CFF0CFEB4BC6ACC66" box="[189,813,333,359]" pageId="13" pageNumber="14">
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<taxonomicName id="4C1BEFE7FF9CFF8CFF0CFEB4BD4ECC66" authorityName="Woodward" authorityYear="1895" box="[189,521,333,359]" class="Actinopterygii" family="Saurichthyidae" genus="Belonorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="brevirostris">
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||
<emphasis id="B96F4876FF9CFF8CFF0CFEB4BD4ECC66" box="[189,521,333,359]" italics="true" pageId="13" pageNumber="14">Belonorhynchus brevirostris</emphasis>
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</taxonomicName>
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||
–
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||
<treatmentCitation id="0ABAB275FF9CFF8CFDAEFEB4BDA6CC66" box="[543,737,333,359]" pageId="13" pageNumber="14">
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<bibRefCitation id="EF8AE995FF9CFF8CFDAEFEB4BDA6CC66" author="Woodward A. S." box="[543,737,333,359]" pageId="13" pageNumber="14" pagination="455 - 472" refId="ref15290" refString="Woodward A. S. 1899. On the fossil fishes of the Upper Lias of Whitby. Part IV. Proceedings of the Yorkshire Geological and Polytechnic Society 13: 455 - 472." type="journal article" year="1899">Woodward 1899</bibRefCitation>
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||
</treatmentCitation>
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||
: fig. 1
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9CFF8CFF0CFE89BA30CC8A" blockId="13.[189,1399,333,465]" box="[189,1399,368,395]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="AB0BB34AFF9CFF8CFF0CFE89BA30CC8A" box="[189,1399,368,395]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFF0CFE89BEBDCC8B" authorityName="Stensio" authorityYear="1925" box="[189,506,368,394]" class="Actinopterygii" family="Saurichthyidae" genus="Acidorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="brevirostris">
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<emphasis id="B96F4876FF9CFF8CFF0CFE89BEBDCC8B" box="[189,506,368,394]" italics="true" pageId="13" pageNumber="14">Acidorhynchus brevirostris</emphasis>
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||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ABAB275FF9CFF8CFDA6FE89BDDACC8B" box="[535,669,368,394]" pageId="13" pageNumber="14">
|
||
<bibRefCitation id="EF8AE995FF9CFF8CFDA6FE89BDDACC8B" author="Hauff B." box="[535,669,368,394]" pageId="13" pageNumber="14" pagination="214 - 248" refId="ref13522" refString="Hauff B. 1938. Uber Acidorhynchus aus den Posidonienschiefern von Holzmaden. Palaontologische Zeitschrift 20: 214 - 248. https: // doi. org / 10.1007 / BF 03041918" type="journal article" year="1938">Hauff 1938</bibRefCitation>
|
||
</treatmentCitation>
|
||
partim: pl. 22, figs b,c. —
|
||
<bibRefCitation id="EF8AE995FF9CFF8CFC66FE89BB2ACC8A" author="Thies D." box="[983,1133,368,395]" pageId="13" pageNumber="14" pagination="183 - 203" refId="ref14987" refString="Thies D. 1985. Funde von Acidorhynchus brevirostris (Woodward 1895) aus dem Posidonienschiefer (Unter-Toarcium) NW-Deutschlands. Palaeontographica Abt. A 187 (4 - 6): 183 - 203." type="journal article" year="1985">
|
||
<treatmentCitation id="0ABAB275FF9CFF8CFC66FE89BB23CC8A" box="[983,1124,368,395]" pageId="13" pageNumber="14">Thies (1985</treatmentCitation>
|
||
)
|
||
</bibRefCitation>
|
||
partim: pl. 1, figs 1, 5.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9CFF8CFF0CFE6ABEC4CCD0" blockId="13.[189,1399,333,465]" pageId="13" pageNumber="14">
|
||
<treatmentCitationGroup id="AB0BB34AFF9CFF8CFF0CFE6ABEC4CCD0" pageId="13" pageNumber="14">
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<taxonomicName id="4C1BEFE7FF9CFF8CFF0CFE6ABEBCCCAC" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[189,507,403,429]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9CFF8CFF0CFE6ABEBCCCAC" box="[189,507,403,429]" italics="true" pageId="13" pageNumber="14">Saurorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ABAB275FF9CFF8CFDA9FE6ABC4FCCAC" box="[536,776,403,429]" pageId="13" pageNumber="14">
|
||
<bibRefCitation id="EF8AE995FF9CFF8CFDA9FE6ABC4FCCAC" author="Hauff B. & Hauff R. B." box="[536,776,403,429]" pageId="13" pageNumber="14" refId="ref13554" refString="Hauff B. & Hauff R. B. 1981. Das Holzmadenbuch. Urweltmuseum Hauff, Holzmaden." type="book" year="1981">Hauff & Hauff 1981</bibRefCitation>
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||
</treatmentCitation>
|
||
: fig. 76. —
|
||
<treatmentCitation id="0ABAB275FF9CFF8CFC27FE6ABBC4CCAC" box="[918,1155,403,429]" pageId="13" pageNumber="14">
|
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<bibRefCitation id="EF8AE995FF9CFF8CFC27FE6ABBC4CCAC" author="Urlichs M. & Wild R. & Ziegler B." box="[918,1155,403,429]" pageId="13" pageNumber="14" pagination="1 - 95" refId="ref15023" refString="Urlichs M., Wild R. & Ziegler B. 1994. Der Posidonien-Schiefer und seine Fossilien. Stuttgarter Beitrage zur Naturkunde C 36: 1 - 95." type="journal article" year="1994">
|
||
Urlichs
|
||
<emphasis id="B96F4876FF9CFF8CFC42FE6DBB69CCAC" box="[1011,1070,403,429]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||
(1994)
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
: fig. 79. —
|
||
<treatmentCitation id="0ABAB275FF9CFF8CFAA0FE6ABE4BCCD0" pageId="13" pageNumber="14">
|
||
<bibRefCitation id="EF8AE995FF9CFF8CFAA0FE6ABE4BCCD0" author="Bottcher R." pageId="13" pageNumber="14" pagination="83 - 96" refId="ref12970" refString="Bottcher R. 1998. Leben und Tod im Meer des Posidonienschiefers. In: Heizmann E. P. J. (ed.) Erdgeschichte mitteleuropaischer Regionen (2): vom Schwarzwald zum Ries: 83 - 96. Verlag Dr. Friedrich Pfeil, Munich." type="book chapter" year="1998">Böttcher (1998)</bibRefCitation>
|
||
</treatmentCitation>
|
||
: fig. 7.15.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9CFF8CFF0CFE01BE8ECF62" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8BA49464FF9CFF8CFF0CFE01BE03CF13" blockId="13.[189,324,504,530]" box="[189,324,504,530]" pageId="13" pageNumber="14">
|
||
<heading id="D0EC2308FF9CFF8CFF0CFE01BE03CF13" bold="true" box="[189,324,504,530]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B96F4876FF9CFF8CFF0CFE01BE03CF13" bold="true" box="[189,324,504,530]" pageId="13" pageNumber="14">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9CFF8CFF0CFDDFBE8ECF62" blockId="13.[189,1398,550,611]" pageId="13" pageNumber="14">Named in honour of Bernhard Hauff, whose doctoral dissertation described the saurichthyid fishes from the Holzmaden region.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9CFF8EFF0CFD72BC27CCF1" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BA49464FF9CFF8CFF0CFD72BECBCFA4" blockId="13.[189,396,651,677]" box="[189,396,651,677]" pageId="13" pageNumber="14">
|
||
<heading id="D0EC2308FF9CFF8CFF0CFD72BECBCFA4" bold="true" box="[189,396,651,677]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B96F4876FF9CFF8CFF0CFD72BECBCFA4" bold="true" box="[189,396,651,677]" pageId="13" pageNumber="14">Material studied</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B739E39FF9CFF8CFF5BFD43BD04CE3C" ID-GBIF-Occurrence="2620280303" collectionCode="SMNS" country="Germany" location="GERMANY" pageId="13" pageNumber="14" specimenCode="SMNS 55057" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="8BA49464FF9CFF8CFF5BFD43BE1BCFD5" blockId="13.[189,1399,698,829]" box="[234,348,698,724]" pageId="13" pageNumber="14">
|
||
<emphasis id="B96F4876FF9CFF8CFF5BFD43BE1BCFD5" bold="true" box="[234,348,698,724]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54A02AC6FF9CFF8CFF5BFD43BE1BCFD5" box="[234,348,698,724]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9CFF8CFF0CFD24BD04CE3C" blockId="13.[189,1399,698,829]" pageId="13" pageNumber="14">
|
||
<collectingCountry id="F30CD4F4FF9CFF8CFF0CFD24BE14CFF6" box="[189,339,733,759]" name="Germany" pageId="13" pageNumber="14">GERMANY</collectingCountry>
|
||
: Articulated skeleton lacking median and pelvic fins (
|
||
<collectionCode id="ED0A0CA1FF9CFF8CFC0EFD24BB56CFF6" box="[959,1041,733,759]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" pageId="13" pageNumber="14">SMNS</collectionCode>
|
||
55057,
|
||
<figureCitation id="132088E1FF9CFF8CFBDEFD24BB8DCFF6" box="[1135,1226,733,759]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="13" pageNumber="14">
|
||
Fig. 1
|
||
<collectionCode id="ED0A0CA1FF9CFF8CFB1EFD24BB8DCFF6" box="[1199,1226,733,759]" pageId="13" pageNumber="14">D</collectionCode>
|
||
</figureCitation>
|
||
). Although the skull of this specimen is not perfectly preserved, it was selected as the
|
||
<typeStatus id="54A02AC6FF9CFF8CFBB4FCF9BB2BCE1B" box="[1029,1132,768,794]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
based on the presence of extensive postcranial material.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<caption id="DF64C4ECFF9CFF8CFF0CF907BEA2CAED" ID-DOI="http://doi.org/10.5281/zenodo.3829293" ID-Zenodo-Dep="3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1790,1816]" targetBox="[189,1399,869,1754]" targetPageId="13">
|
||
<paragraph id="8BA49464FF9CFF8CFF0CF907BEA2CAED" blockId="13.[189,1399,1790,2028]" pageId="13" pageNumber="14">
|
||
<emphasis id="B96F4876FF9CFF8CFF0CF907BE4FCA18" bold="true" box="[189,264,1790,1817]" pageId="13" pageNumber="14">Fig. 5.</emphasis>
|
||
Characters distinguishing
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFD88F907BCB2CA18" authority="(Agassiz, 1844)" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[569,1013,1790,1817]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9CFF8CFD88F907BC7DCA19" box="[569,826,1790,1816]" italics="true" pageId="13" pageNumber="14">Saurorhynchus acutus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF9CFF8CFCF6F907BCAACA18" author="Agassiz L." box="[839,1005,1790,1817]" pageId="13" pageNumber="14" refId="ref12560" refString="Agassiz L. 1844. Recherches sur les Poissons Fossiles. II. Contenant l'Histoire de l'Ordre des Ganoides. Vol. 2, Part 2. Petitpierre, Neuchatel." type="book" year="1844">Agassiz, 1844</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFB98F906BBC0CA19" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1065,1159,1790,1817]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9CFF8CFB98F906BBC0CA19" box="[1065,1159,1790,1817]" italics="true" pageId="13" pageNumber="14">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9CFF8CFB3AF906BBA4CA18" box="[1163,1251,1791,1817]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96F4876FF9CFF8CFB56F907BBBBCA19" bold="true" box="[1255,1276,1790,1816]" pageId="13" pageNumber="14">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFAB9F906BA37CA19" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[1288,1392,1791,1817]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9CFF8CFAB9F906BA37CA19" box="[1288,1392,1791,1817]" italics="true" pageId="13" pageNumber="14">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
, narial region (SMNS 56923).
|
||
<emphasis id="B96F4876FF9CFF8CFDAFF8DBBD75CA3D" bold="true" box="[542,562,1826,1852]" pageId="13" pageNumber="14">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFDF0F8DBBDEBCA3D" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[577,684,1826,1852]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9CFF8CFDF0F8DBBDEBCA3D" box="[577,684,1826,1852]" italics="true" pageId="13" pageNumber="14">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
, lateral extrascapular-dermopterotic contact (SMNS 57039).
|
||
<emphasis id="B96F4876FF9CFF8CFF0CF8BCBF95CA5E" bold="true" box="[189,210,1861,1887]" pageId="13" pageNumber="14">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFF6FF8BCBE7BCA5E" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[222,316,1861,1887]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9CFF8CFF6FF8BCBE7BCA5E" box="[222,316,1861,1887]" italics="true" pageId="13" pageNumber="14">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9CFF8CFEF1F8BCBEDCCA5E" box="[320,411,1861,1887]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, narial region (SMNS 51888).
|
||
<emphasis id="B96F4876FF9CFF8CFD49F8BCBC4ACA5E" bold="true" box="[760,781,1861,1887]" pageId="13" pageNumber="14">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9CFF8CFCA8F8BCBC30CA5E" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[793,887,1861,1887]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9CFF8CFCA8F8BCBC30CA5E" box="[793,887,1861,1887]" italics="true" pageId="13" pageNumber="14">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9CFF8CFCCAF8BCBC92CA5E" box="[891,981,1861,1887]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA49464FF9FFF8FFF5BFEF2BE8CCC24" blockId="14.[189,1399,267,539]" box="[234,459,267,293]" pageId="14" pageNumber="15">
|
||
<emphasis id="B96F4876FF9FFF8FFF5BFEF2BE8CCC24" bold="true" box="[234,459,267,293]" pageId="14" pageNumber="15">Referred material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9FFF8FFF0CFED7BEE7CF1A" blockId="14.[189,1399,267,539]" pageId="14" pageNumber="15">
|
||
<collectingCountry id="F30CD4F4FF9FFF8FFF0CFED7BE14CC49" box="[189,339,302,328]" name="Germany" pageId="14" pageNumber="15">GERMANY</collectingCountry>
|
||
: all referred material consists of isolated skulls, unless otherwise noted.
|
||
<materialsCitation id="3B739E39FF9FFF8FFB00FED7BED2CC6D" ID-GBIF-Occurrence="3763234302" collectionCode="SMNS" location="Bad Boll" pageId="14" pageNumber="15" specimenCode="SMNS 96878" specimenCount="1">
|
||
<location id="8EC4C2BFFF9FFF8FFB00FED7BA59CC49" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFB00FED7BA59CC49" box="[1201,1310,302,328]" name="Bad Boll" pageId="14" pageNumber="15">Bad Boll</location>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFAFDFEB9BA30CC4F" attach="left" box="[1356,1399,320,334]" fontSize="6" pageId="14" pageNumber="15">10–12</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFF74FEA8BE34CC6D" box="[197,371,337,364]" pageId="14" pageNumber="15">SMNS 96878</specimenCode>
|
||
/1);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFE11FEA8BCFACC6D" ID-GBIF-Occurrence="3763234321" box="[416,957,337,369]" collectionCode="C" location="Dotternhausen" pageId="14" pageNumber="15" specimenCode="SMNS 51007" specimenCount="1">
|
||
<location id="8EC4C2BFFF9FFF8FFE11FEA8BD09CC6A" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFE11FEA8BD09CC6A" box="[416,590,337,363]" name="Dotternhausen" pageId="14" pageNumber="15">Dotternhausen</location>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFDCFFE9ABDC0CC70" attach="left" box="[638,647,355,369]" fontSize="6" pageId="14" pageNumber="15">4</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFD24FEA8BC79CC6D" box="[661,830,337,364]" pageId="14" pageNumber="15">SMNS 51007</specimenCode>
|
||
,
|
||
<figureCitation id="132088E1FF9FFF8FFCFEFEA8BCE8CC6D" box="[847,943,337,364]" captionStart="Fig" captionStartId="14.[189,232,1828,1854]" captionTargetBox="[198,1392,811,1788]" captionTargetId="figure@14.[195,1392,808,1788]" captionTargetPageId="14" captionText="Fig. 6. Posteriormost laniary dentition. A. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV P 4878), note the lingually curved crowns. B. Saurorhynchus anningae sp. nov. (NHMUK PV P 27569), note the incisivlücke associated with the premaxillary laniary tooth and the absence of lateral crypts associated with the two flanking mandibular laniaries. Photos © The Trustees of the Natural History Museum, London. C. S. hauffi sp. nov. (SMNS 51007), mid-rostral dentition illustrating the relationship between the incisivlücken, laniaries, and flanking smaller teeth. Scale bars: 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3829295" httpUri="https://zenodo.org/record/3829295/files/figure.png" pageId="14" pageNumber="15">
|
||
Fig. 6
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFC28FEA8BCE8CC6D" box="[921,943,337,364]" pageId="14" pageNumber="15">C</collectionCode>
|
||
</figureCitation>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFC78FEA8BA31CC6D" ID-GBIF-Occurrence="3763234306" box="[969,1398,337,369]" collectionCode="SMNS" location="Dotternhausen" municipality="Dotternhausen" pageId="14" pageNumber="15" specimenCode="SMNS 58394" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFC78FEA8BB30CC6A" box="[969,1143,337,363]" pageId="14" pageNumber="15">Dotternhausen</collectingMunicipality>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFB16FE9ABBF7CC70" attach="left" box="[1191,1200,355,369]" fontSize="6" pageId="14" pageNumber="15">6</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFB0FFEA8BA2DCC6D" box="[1214,1386,337,364]" pageId="14" pageNumber="15">SMNS 58394</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFF0CFE8DBC11CC8E" ID-GBIF-Occurrence="3763234313" box="[189,854,372,399]" collectionCode="NLMH" location="Subzone" municipality="Haverlahwiese" pageId="14" pageNumber="15" specimenCode="NLMH 70598" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFF0CFE8DBE29CC8F" box="[189,366,372,398]" pageId="14" pageNumber="15">Haverlahwiese</collectingMunicipality>
|
||
,
|
||
<taxonomicName id="4C1BEFE7FF9FFF8FFECEFE8DBDC6CC8F" authority="Subzone" authorityName="Subzone" box="[383,641,372,398]" class="Cephalopoda" family="Hildoceratidae" genus="Eleganticeras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="elegantulum">
|
||
<emphasis id="B96F4876FF9FFF8FFECEFE8DBD57CC8F" box="[383,528,372,398]" italics="true" pageId="14" pageNumber="15">elegantulum</emphasis>
|
||
<location id="8EC4C2BFFF9FFF8FFDADFE8DBDC6CC8F" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFDADFE8DBDC6CC8F" box="[540,641,372,398]" municipality="Haverlahwiese" name="Subzone" pageId="14" pageNumber="15">Subzone</location>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFD24FE8CBC0FCC8E" box="[661,840,372,399]" pageId="14" pageNumber="15">NLMH 70598</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFCD0FE8DBBE0CC8E" ID-GBIF-Occurrence="3763234305" box="[865,1191,372,404]" collectionCode="MHH" location="Holzmaden" pageId="14" pageNumber="15" specimenCode="MHH 2" specimenCount="1">
|
||
<location id="8EC4C2BFFF9FFF8FFCD0FE8DBCACCC8F" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFCD0FE8DBCACCC8F" box="[865,1003,372,398]" name="Holzmaden" pageId="14" pageNumber="15">Holzmaden</location>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFBADFE7FBB62CC95" attach="left" box="[1052,1061,390,404]" fontSize="6" pageId="14" pageNumber="15">3</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFB85FE8CBBD0CC8E" box="[1076,1175,373,399]" pageId="14" pageNumber="15">MHH 2</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFB02FE8DBE3FCCB3" ID-GBIF-Occurrence="3763234323" collectionCode="SMNS" location="Holzmaden" municipality="Holzmaden" pageId="14" pageNumber="15" specimenCode="SMNS 55302" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFB02FE8DBA7ACC8F" box="[1203,1341,372,398]" pageId="14" pageNumber="15">Holzmaden</collectingMunicipality>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFADFFE7FBA30CC95" attach="left" box="[1390,1399,390,404]" fontSize="6" pageId="14" pageNumber="15">4</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFF74FE61BE2BCCB3" box="[197,364,407,434]" pageId="14" pageNumber="15">SMNS 55302</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFECEFE61BC9DCCB3" ID-GBIF-Occurrence="3763234303" box="[383,986,408,434]" collectionCode="GG, B" location="Subzone" municipality="Grimmen" pageId="14" pageNumber="15" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFECEFE61BEB4CCB3" box="[383,499,408,434]" pageId="14" pageNumber="15">Grimmen</collectingMunicipality>
|
||
,
|
||
<taxonomicName id="4C1BEFE7FF9FFF8FFE4FFE60BD9FCCB3" authority="Subzone" authorityName="Subzone" box="[510,728,408,434]" class="Cephalopoda" family="Hildoceratidae" genus="Cleviceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="exaratum">
|
||
<emphasis id="B96F4876FF9FFF8FFE4FFE60BD2ACCB3" box="[510,621,409,434]" italics="true" pageId="14" pageNumber="15">exaratum</emphasis>
|
||
<location id="8EC4C2BFFF9FFF8FFDC2FE61BD9FCCB3" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFDC2FE61BD9FCCB3" box="[627,728,408,434]" municipality="Grimmen" name="Subzone" pageId="14" pageNumber="15">Subzone</location>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFD56FE61BC54CCB3" box="[743,787,408,434]" pageId="14" pageNumber="15">GG</collectionCode>
|
||
20001,
|
||
<figureCitation id="132088E1FF9FFF8FFCC0FE61BC8CCCB3" box="[881,971,408,434]" captionStart="Fig" captionStartId="9.[189,232,1718,1744]" captionTargetBox="[195,1396,311,1622]" captionTargetId="graphics@9.[317,1359,731,1036]" captionTargetPageId="9" captionText="Fig. 3. Dorsal skull roof. A. Saurorhynchus anningae sp. nov. (NHMUK PV P 964); note that this skull has been dorsoventrally compressed. B. Saurorhynchus hauffi sp. nov., three-dimensionally preserved skull GG 20001. Fine dashed lines indicate sensory canals, large dashes indicate broken or incomplete areas; grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l.ex = lateral extrascapular; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; p = parietal; pm = rostropremaxilla; so.c = supraorbital sensory canal. Scale bars = 10 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829288" httpUri="https://zenodo.org/record/3829288/files/figure.png" pageId="14" pageNumber="15">
|
||
Fig. 3
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFC04FE61BC8CCCB3" box="[949,971,408,434]" pageId="14" pageNumber="15">B</collectionCode>
|
||
</figureCitation>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFC50FE61BA31CCB3" ID-GBIF-Occurrence="3763234317" box="[993,1398,407,434]" collectionCode="D" location="Ohmden" pageId="14" pageNumber="15" specimenCode="SMNS 53980" specimenCount="1">
|
||
<location id="8EC4C2BFFF9FFF8FFC50FE61BB01CCB3" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFC50FE61BB01CCB3" box="[993,1094,408,434]" name="Ohmden" pageId="14" pageNumber="15">Ohmden</location>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFBE4FE61BBBECCB3" box="[1109,1273,407,434]" pageId="14" pageNumber="15">SMNS 53980</specimenCode>
|
||
,
|
||
<figureCitation id="132088E1FF9FFF8FFABDFE61BA2FCCB3" box="[1292,1384,407,434]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="14" pageNumber="15">
|
||
Fig. 5
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFAE1FE6EBA2FCCB3" box="[1360,1384,407,434]" pageId="14" pageNumber="15">D</collectionCode>
|
||
</figureCitation>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFF0CFE42BD52CCD7" ID-GBIF-Occurrence="3763234304" box="[189,533,443,475]" collectionCode="SMNS" location="Ohmden" municipality="Ohmden" pageId="14" pageNumber="15" specimenCode="SMNS 50075" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFF0CFE42BE62CCD4" box="[189,293,443,469]" pageId="14" pageNumber="15">Ohmden</collectingMunicipality>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFEFEFE34BE1FCCDA" attach="left" box="[335,344,461,475]" fontSize="6" pageId="14" pageNumber="15">3</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFED2FE42BD4FCCD7" box="[355,520,443,470]" pageId="14" pageNumber="15">SMNS 50075</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFDAAFE42BB44CCD7" ID-GBIF-Occurrence="3763234316" box="[539,1027,443,475]" collectionCode="C" location="Ohmden" pageId="14" pageNumber="15" specimenCode="SMNS 51888" specimenCount="4" specimenCount-female="4">
|
||
<location id="8EC4C2BFFF9FFF8FFDAAFE42BDC4CCD4" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFDAAFE42BDC4CCD4" box="[539,643,443,469]" name="Ohmden" pageId="14" pageNumber="15">Ohmden</location>
|
||
, εII
|
||
<subScript id="179F9621FF9FFF8FFD1DFE34BDF2CCDA" attach="left" box="[684,693,461,475]" fontSize="6" pageId="14" pageNumber="15">4</subScript>
|
||
(
|
||
<specimenCode id="DBBD3C1FFF9FFF8FFD71FE42BC24CCD7" box="[704,867,443,470]" pageId="14" pageNumber="15">SMNS 51888</specimenCode>
|
||
,
|
||
<figureCitation id="132088E1FF9FFF8FFCDFFE42BC82CCD4" box="[878,965,443,469]" captionStart="Fig" captionStartId="10.[189,232,1683,1709]" captionTargetBox="[189,1399,933,1642]" captionTargetId="figure@10.[189,1399,929,1642]" captionTargetPageId="10" captionText="Fig. 4. Orbitotemporal region, illustrating variation in the braincase of Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov. (based on NHMUK PV P 36227). B, E. Saurorhynchus brevirostris (Woodward, 1895) (photo pertains to NHMUK PV P 4878, mirrored). C, F. Saurorhynchus hauffi sp. nov. (photo is of SMNS 51888, mirrored). D. Saurorhynchus acutus (Agassiz, 1844) (based on SMNS 87737). Abbreviations: asc = ascending process of the parasphenoid; bs = basisphenoid; dpt = dermopterotic; dsp = dermosphenotic; f.ic+ep = foramen for the internal carotid artery and efferent pseudobranchial artery; f.oma = f.om = foramen for the great ophthalmic artery; my = posterior myodome; n = neomorph; nc = neurocranium; orb = orbit; psr = parasphenoid rostrum; rot = foramen for the lateral otic ramus; spi = ventral opening of the spiracular canal; tf.c = trigeminofacial chamber. Scale bars: E–F = 2 mm. E. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829291" httpUri="https://zenodo.org/record/3829291/files/figure.png" pageId="14" pageNumber="15">
|
||
<location id="8EC4C2BFFF9FFF8FFCDFFE42BCE6CCD4" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFCDFFE42BCE6CCD4" box="[878,929,443,469]" name="Figs" pageId="14" pageNumber="15">Figs</location>
|
||
<specimenCount id="9D1D5FEDFF9FFF8FFC17FE42BC82CCD4" box="[934,965,443,469]" pageId="14" pageNumber="15" type="female">4F</specimenCount>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="132088E1FF9FFF8FFC7EFE42BCB3CCD7" box="[975,1012,443,470]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="14" pageNumber="15">
|
||
5
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFC6FFE42BCB3CCD7" box="[990,1012,443,470]" pageId="14" pageNumber="15">C</collectionCode>
|
||
</figureCitation>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFBB8FE42BE38CCF9" ID-GBIF-Occurrence="3763234319" collectionCode="GZG" location="Subzone" municipality="Schandelah" pageId="14" pageNumber="15" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFBB8FE42BBD4CCD4" box="[1033,1171,443,469]" pageId="14" pageNumber="15">Schandelah</collectingMunicipality>
|
||
,
|
||
<taxonomicName id="4C1BEFE7FF9FFF8FFB2CFE45BA31CCD4" authority="Subzone" authorityName="Subzone" box="[1181,1398,443,469]" class="Cephalopoda" family="Hildoceratidae" genus="Cleviceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="exaratum">
|
||
<emphasis id="B96F4876FF9FFF8FFB2CFE45BA4BCCD4" box="[1181,1292,444,469]" italics="true" pageId="14" pageNumber="15">exaratum</emphasis>
|
||
<location id="8EC4C2BFFF9FFF8FFAA0FE42BA31CCD4" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFAA0FE42BA31CCD4" box="[1297,1398,443,469]" municipality="Schandelah" name="Subzone" pageId="14" pageNumber="15">Subzone</location>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFF74FE27BE40CCF9" box="[197,263,478,504]" pageId="14" pageNumber="15">GZG</collectionCode>
|
||
.V.27931);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B739E39FF9FFF8FFE38FE27BEE7CF1A" ID-GBIF-Occurrence="3763234308" collectionCode="NMB" location="serpentinum Zone" municipality="Schandelah" pageId="14" pageNumber="15" specimenCount="1">
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFE38FE27BD54CCF9" box="[393,531,478,504]" pageId="14" pageNumber="15">Schandelah</collectingMunicipality>
|
||
,
|
||
<location id="8EC4C2BFFF9FFF8FFD92FE26BDB9CCF8" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFD92FE26BDB9CCF8" box="[547,766,479,505]" municipality="Schandelah" name="serpentinum Zone" pageId="14" pageNumber="15">
|
||
<emphasis id="B96F4876FF9FFF8FFD92FE26BDF3CCF9" box="[547,692,479,504]" italics="true" pageId="14" pageNumber="15">serpentinum</emphasis>
|
||
Zone
|
||
</location>
|
||
, articulated skeleton, caudal region and median fins absent (
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFEA6FDF8BE1BCF1A" box="[279,348,513,539]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="14" pageNumber="15">NMB</collectionCode>
|
||
373).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9FFF8FFF0CFDBABCECCF5C" blockId="14.[189,939,578,605]" box="[189,939,578,605]" pageId="14" pageNumber="15">
|
||
<materialsCitation id="3B739E39FF9FFF8FFF0CFDBABCE1CF5C" ID-GBIF-Occurrence="3763234314" box="[189,934,578,605]" collectionCode="NHMUK, OR" country="United Kingdom" location="Toarcian" municipality="Whitby" pageId="14" pageNumber="15" specimenCount="1">
|
||
<collectingCountry id="F30CD4F4FF9FFF8FFF0CFDBABE84CF5C" box="[189,451,579,605]" name="United Kingdom" pageId="14" pageNumber="15">UNITED KINGDOM</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="6BC00E1EFF9FFF8FFE7FFDBABD60CF5C" box="[462,551,579,605]" pageId="14" pageNumber="15">Whitby</collectingMunicipality>
|
||
,
|
||
<location id="8EC4C2BFFF9FFF8FFD83FDBABDDFCF5C" LSID="urn:lsid:plazi:treatment:03B22572FF9DFF92FDD2F8BEBD97C8A5:8EC4C2BFFF9FFF8FFD83FDBABDDFCF5C" box="[562,664,579,605]" country="United Kingdom" municipality="Whitby" name="Toarcian" pageId="14" pageNumber="15">Toarcian</location>
|
||
(
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFD19FDBABC5ECF5C" box="[680,793,579,605]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="15" type="Museum">NHMUK</collectionCode>
|
||
<collectionCode id="ED0A0CA1FF9FFF8FFC91FDBABC0ECF5C" box="[800,841,579,605]" pageId="14" pageNumber="15">OR</collectionCode>
|
||
39153)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF64C4ECFF9FFF8FFF0CF8DDBB3FCAEE" ID-DOI="http://doi.org/10.5281/zenodo.3829295" ID-Zenodo-Dep="3829295" httpUri="https://zenodo.org/record/3829295/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1828,1854]" targetBox="[198,1392,811,1788]" targetPageId="14">
|
||
<paragraph id="8BA49464FF9FFF8FFF0CF8DDBB3FCAEE" blockId="14.[189,1399,1828,2031]" pageId="14" pageNumber="15">
|
||
<emphasis id="B96F4876FF9FFF8FFF0CF8DDBE4ECA3F" bold="true" box="[189,265,1828,1854]" pageId="14" pageNumber="15">Fig. 6.</emphasis>
|
||
Posteriormost laniary dentition.
|
||
<emphasis id="B96F4876FF9FFF8FFD39F8DDBDDACA3F" bold="true" box="[648,669,1828,1854]" pageId="14" pageNumber="15">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9FFF8FFD1AF8DDBB8CCA3E" authority="(Woodward, 1895)" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[683,1227,1828,1855]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9FFF8FFD1AF8DDBCAECA3F" box="[683,1001,1828,1854]" italics="true" pageId="14" pageNumber="15">Saurorhynchus brevirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF9FFF8FFC46F8DDBB84CA3E" author="Woodward A. S." box="[1015,1219,1828,1855]" pageId="14" pageNumber="15" refId="ref15257" refString="Woodward A. S. 1895. Catalogue of the Fossil Fishes in the British Museum (Natural History). Part III. British Museum (Natural History), London." type="book" year="1895">Woodward, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(NHMUK PV P 4878), note the lingually curved crowns.
|
||
<emphasis id="B96F4876FF9FFF8FFD01F8B1BD83CA63" bold="true" box="[688,708,1864,1890]" pageId="14" pageNumber="15">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9FFF8FFD60F8BEBCB2CA60" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[721,1013,1863,1889]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9FFF8FFD60F8BEBCB2CA60" box="[721,1013,1863,1889]" italics="true" pageId="14" pageNumber="15">Saurorhynchus anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9FFF8FFC4DF8B1BB12CA63" box="[1020,1109,1864,1890]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NHMUK PV P 27569), note the incisivlücke associated with the premaxillary laniary tooth and the absence of lateral crypts associated with the two flanking mandibular laniaries. Photos © The Trustees of the Natural History Museum, London.
|
||
<emphasis id="B96F4876FF9FFF8FFE2BF848BEE8CACA" bold="true" box="[410,431,1969,1995]" pageId="14" pageNumber="15">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF9FFF8FFE0CF84BBD5ACACA" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[445,541,1969,1996]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9FFF8FFE0CF84BBD5ACACA" box="[445,541,1969,1996]" italics="true" pageId="14" pageNumber="15">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9FFF8FFD92F84BBD3ACACD" box="[547,637,1970,1996]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(SMNS 51007), mid-rostral dentition illustrating the relationship between the incisivlücken, laniaries, and flanking smaller teeth. Scale bars: 1 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFEF2BED3CC24" blockId="15.[189,404,267,293]" box="[189,404,267,293]" pageId="15" pageNumber="16">
|
||
<heading id="D0EC2308FF9EFF8EFF0CFEF2BED3CC24" bold="true" box="[189,404,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFEF2BED3CC24" bold="true" box="[189,404,267,293]" pageId="15" pageNumber="16">Stratum typicum</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFEC5BEEDCC7D" blockId="15.[189,1398,316,380]" pageId="15" pageNumber="16">
|
||
<collectingCountry id="F30CD4F4FF9EFF8EFF0CFEC5BE14CC57" box="[189,339,316,342]" name="Germany" pageId="15" pageNumber="16">GERMANY</collectingCountry>
|
||
: Posidonienschiefer Formation, horizon εII4 (uppermost
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFC40FEC5BBC3CC57" box="[1009,1156,316,342]" class="Cephalopoda" family="Dactylioceratidae" genus="Dactylioceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="semicelatum">
|
||
<emphasis id="B96F4876FF9EFF8EFC40FEC5BBC3CC57" box="[1009,1156,316,342]" italics="true" pageId="15" pageNumber="16">semicelatum</emphasis>
|
||
</taxonomicName>
|
||
Subzone-lowermost
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFF0CFE9BBEDACC7D" authority="Subzone" authorityName="Subzone" box="[189,413,353,380]" class="Cephalopoda" family="Hildoceratidae" genus="Cleviceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="exaratum">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFE9BBE6BCC7A" box="[189,300,354,379]" italics="true" pageId="15" pageNumber="16">exaratum</emphasis>
|
||
Subzone
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFE5CBE2ECCBE" blockId="15.[189,361,421,447]" box="[189,361,421,447]" pageId="15" pageNumber="16">
|
||
<heading id="D0EC2308FF9EFF8EFF0CFE5CBE2ECCBE" bold="true" box="[189,361,421,447]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFE5CBE2ECCBE" bold="true" box="[189,361,421,447]" pageId="15" pageNumber="16">Locus typicus</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFE2FBC27CCF1" blockId="15.[189,864,470,496]" box="[189,864,470,496]" pageId="15" pageNumber="16">
|
||
<collectingCountry id="F30CD4F4FF9EFF8EFF0CFE2FBE14CCF1" box="[189,339,470,496]" name="Germany" pageId="15" pageNumber="16">GERMANY</collectingCountry>
|
||
: Zell unter Aichelberg,
|
||
<collectingRegion id="49DF5A86FF9EFF8EFDD6FE2FBC1BCCF1" box="[615,860,470,496]" country="Germany" name="Baden-Wuerttemberg" pageId="15" pageNumber="16">Baden-Württemberg</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9EFF8EFF0CFDE0BA63C996" pageId="15" pageNumber="16" type="diagnosis">
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFDE0BE8ECF32" blockId="15.[189,457,537,563]" box="[189,457,537,563]" pageId="15" pageNumber="16">
|
||
<heading id="D0EC2308FF9EFF8EFF0CFDE0BE8ECF32" bold="true" box="[189,457,537,563]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFDE0BE8ECF32" bold="true" box="[189,457,537,563]" pageId="15" pageNumber="16">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFDB3BA63C996" blockId="15.[189,1399,586,1175]" pageId="15" pageNumber="16">
|
||
This species can be differentiated from all other
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFCB1FDB3BCF7CF65" authorityName="Reis" authorityYear="1892" box="[768,944,586,612]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF9EFF8EFCB1FDB3BCF7CF65" box="[768,944,586,612]" italics="true" pageId="15" pageNumber="16">Saurorhynchus</emphasis>
|
||
</taxonomicName>
|
||
species by the following combination of characters: short, deep skull; posterior dorsal skull roof deflected dorsally relative to the long axis of the skull (parallel in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFE6BFD6FBD00CFAE" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[474,583,662,688]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9EFF8EFE6BFD6FBD00CFAE" box="[474,583,662,688]" italics="true" pageId="15" pageNumber="16">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFD35FD6FBC57CFAE" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[644,784,662,688]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFD35FD6FBC57CFAE" box="[644,784,662,688]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFCABFD6FBC3CCFB1" box="[794,891,662,688]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); postorbital segment approximately equal in length to the depth of the lower jaw (approximately twice as long in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFBBCFD42BBDFCFD4" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1037,1176,699,725]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFBBCFD42BBDFCFD4" box="[1037,1176,699,725]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFB10FD42BA47CFD4" box="[1185,1280,699,725]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); overbite slight or absent (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFDB3FD18BDEDCFFB" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[514,682,736,763]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9EFF8EFDB3FD18BDEDCFFB" box="[514,682,736,763]" italics="true" pageId="15" pageNumber="16">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); maxilla not ventrally deflected under the orbit (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFF0CFCFFBE6DCE21" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[189,298,774,800]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFCFFBE6DCE21" box="[189,298,774,800]" italics="true" pageId="15" pageNumber="16">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
); coarse, reticular ornamentation on the dermal bones of skull roof (ornamentation reduced in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFF6AFCD5BE01CE44" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[219,326,812,838]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9EFF8EFF6AFCD5BE01CE44" box="[219,326,812,838]" italics="true" pageId="15" pageNumber="16">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
); parasphenoid edentulous ventral to the orbit (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFC54FCD5BB28CE44" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[997,1135,812,838]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFC54FCD5BB28CE44" box="[997,1135,812,838]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFBC6FCD5BB91CE47" box="[1143,1238,812,838]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); foramen for the internal carotid and efferent pseudobranchial arteries centered ventral to the anterior basisphenoid and oriented laterally (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFD8AFC8EBD80CE91" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[571,711,887,913]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFD8AFC8EBD80CE91" box="[571,711,887,913]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFD7EFC8EBC6CCE90" box="[719,811,887,913]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFCD4FC8EBB4CCE91" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[869,1035,886,913]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9EFF8EFCD4FC8EBB4CCE91" box="[869,1035,886,913]" italics="true" pageId="15" pageNumber="16">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); foramen for the ophthalmica magna artery situated between two ridges on the anterior surface of the basisphenoid (anterior surface of basisphenoid flat and foramen oriented anterolaterally in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFCCCFC3BBB66CEDA" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[893,1057,961,988]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9EFF8EFCCCFC3BBB66CEDA" box="[893,1057,961,988]" italics="true" pageId="15" pageNumber="16">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); posterior edge of mandible strongly sinusoidal (straight or only weakly sinusoidal in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFCEBFC1EBCBAC900" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[858,1021,999,1025]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9EFF8EFCEBFC1EBCBAC900" box="[858,1021,999,1025]" italics="true" pageId="15" pageNumber="16">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFB84FC1EBBF9C900" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1077,1214,999,1025]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFB84FC1EBBF9C900" box="[1077,1214,999,1025]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFB74FC1EBA64C900" box="[1221,1315,999,1025]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); angle between the posterior and ventral edges of the mandible less than 80 degrees (more than 90 degrees in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFF6EFBCBBE2AC94D" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[223,365,1074,1100]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFF6EFBCBBE2AC94D" box="[223,365,1074,1100]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFEC9FBCBBE9DC94D" box="[376,474,1074,1100]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); mandibular sensory canal positioned along the dorsal half of the posterior mandible; anterior narial opening ovate (elongate in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFC94FBA1BCC8C970" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[805,911,1112,1138]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9EFF8EFC94FBA1BCC8C970" box="[805,911,1112,1138]" italics="true" pageId="15" pageNumber="16">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
); subnarial laniary teeth absent (unlike in
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFF0CFB84BE26C996" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[189,353,1149,1175]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFB84BE26C996" box="[189,353,1149,1175]" italics="true" pageId="15" pageNumber="16">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); lateral supracleithrum bearing prominent anterior process pierced by a foramen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF9EFF92FF0CFB39BCB5C9EB" lastPageId="19" lastPageNumber="20" pageId="15" pageNumber="16" type="description">
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFB39BE09C9DB" blockId="15.[189,334,1216,1242]" box="[189,334,1216,1242]" pageId="15" pageNumber="16">
|
||
<heading id="D0EC2308FF9EFF8EFF0CFB39BE09C9DB" bold="true" box="[189,334,1216,1242]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B96F4876FF9EFF8EFF0CFB39BE09C9DB" bold="true" box="[189,334,1216,1242]" pageId="15" pageNumber="16">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFB08BA30C87D" blockId="15.[189,1399,1265,1405]" pageId="15" pageNumber="16">
|
||
As with
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFEAAFB0BBEE4C80A" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[283,419,1266,1292]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFEAAFB0BBEE4C80A" box="[283,419,1266,1292]" italics="true" pageId="15" pageNumber="16">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFE19FB0BBD47C80D" box="[424,512,1266,1292]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, material now attributed to
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFC88FB08BB76C80A" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[825,1073,1265,1291]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFC88FB08BB76C80A" box="[825,1073,1265,1291]" italics="true" pageId="15" pageNumber="16">Saurorhynchus hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFB87FB0BBBC9C80D" box="[1078,1166,1266,1292]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been previously described (
|
||
<bibRefCitation id="EF8AE995FF9EFF8EFE8DFAEEBE80C830" author="Hauff B." box="[316,455,1303,1329]" pageId="15" pageNumber="16" pagination="214 - 248" refId="ref13522" refString="Hauff B. 1938. Uber Acidorhynchus aus den Posidonienschiefern von Holzmaden. Palaontologische Zeitschrift 20: 214 - 248. https: // doi. org / 10.1007 / BF 03041918" type="journal article" year="1938">Hauff 1938</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8AE995FF9EFF8EFE63FAEEBD10C833" author="Thies D." box="[466,599,1303,1330]" pageId="15" pageNumber="16" pagination="183 - 203" refId="ref14987" refString="Thies D. 1985. Funde von Acidorhynchus brevirostris (Woodward 1895) aus dem Posidonienschiefer (Unter-Toarcium) NW-Deutschlands. Palaeontographica Abt. A 187 (4 - 6): 183 - 203." type="journal article" year="1985">Thies 1985</bibRefCitation>
|
||
). However, as much of the material on which the initial descriptions were based is referable to
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFE48FAC4BD22C857" baseAuthorityName="Agassiz" baseAuthorityYear="1844" box="[505,613,1341,1367]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="acutus">
|
||
<emphasis id="B96F4876FF9EFF8EFE48FAC4BD22C857" box="[505,613,1341,1367]" italics="true" pageId="15" pageNumber="16">S. acutus</emphasis>
|
||
</taxonomicName>
|
||
, a redescription is necessary.
|
||
<taxonomicName id="4C1BEFE7FF9EFF8EFC7BFAC4BB6BC857" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[970,1068,1340,1367]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF9EFF8EFC7BFAC4BB6BC857" box="[970,1068,1340,1367]" italics="true" pageId="15" pageNumber="16">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF9EFF8EFB84FAC4BBD5C856" box="[1077,1170,1341,1367]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
exhibits lower jaw lengths of up to
|
||
<quantity id="4CE33981FF9EFF8EFEC7FA9BBE92C87D" box="[374,469,1378,1405]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="15" pageNumber="16" unit="mm" value="115.0">115 mm</quantity>
|
||
(Suppl. Table 1); this would correspond to a fish approximately
|
||
<quantity id="4CE33981FF9EFF8EFB0DFA9BBA44C87D" box="[1212,1283,1378,1404]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="cm" value="40.0">40 cm</quantity>
|
||
in length.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CFA5CBB97CA10" blockId="15.[189,1399,1445,1810]" pageId="15" pageNumber="16">The rostropremaxilla is the dominant bone in the rostrum, dorsoventrally compressed at the anterior tip and becoming strongly laterally compressed posteriorly. The posterior rostropremaxilla bears a ventral process that forms an overlapping suture with the maxilla ventral to the external narial openings, as well as a dorsal process that approaches the anterior narial opening. Between these two processes, an anterior ventral process of the nasaloantorbital bearing the sensory canal is enclosed. The rostropremaxilla is ornamented with longitudinal grooves. Incisivlücken are lacking at the anteriormost tip. The rostropremaxilla bears approximately 30 laniary teeth. The anterior teeth are very small, and become progressively larger towards the midpoint of the jaw. The sensory pit line runs along the lateral surface of the premaxilla immediately dorsal to tooth row. The sensory pits become increasingly obscured by incisivlücken anteriorly. The presence or absence of interrostrals could not be confirmed.</paragraph>
|
||
<paragraph id="8BA49464FF9EFF8EFF0CF8C2BCE0CAEA" blockId="15.[189,1399,1851,2027]" pageId="15" pageNumber="16">The maxilla extends anteriorly as far as the first laniary tooth. The suborbital bar is dorsoventrally compressed and bears denticles on its occlusal surface. Posterior to the orbit and anterior to the deepest point of the posterior lamina, a small concave embayment is present along the ventral edge of the maxilla. The postorbital lamina is ornamented with pits, the more posterior of which are elongated in such a way that they appear directed towards the suborbital bar.</paragraph>
|
||
<paragraph id="8BA49464FF81FF91FF0CFEF2BBC6CF84" blockId="16.[189,1399,267,646]" pageId="16" pageNumber="17">
|
||
The nasaloantorbital is triangular in lateral view. The posterior end encloses two external narial openings: a small, round posterior opening and a large, oval to reniform anterior opening (
|
||
<figureCitation id="132088E1FF81FF91FBC0FED7BB89CC48" box="[1137,1230,302,329]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="16" pageNumber="17">Fig. 5C</figureCitation>
|
||
). Between the two narial openings runs the supraorbital sensory canal. This canal splits into two branches ventrally; the junction is contained within the nasaloantorbital. Because the lacrimal is often damaged or missing, the ventral edge of the nasaloantorbital is often clearly exposed and has a complex morphology dictated by the orientation of the canals. There is no evidence for separate ossifications of the nasal and antorbital. The nasaloantorbital articulates with the frontal dorsally and the premaxilla ventrally. Ornamentation can be quite strong in the around the narial openings, but becomes reduced anteriorly. Between the posterior narial opening and the orbit is an area of strong dermal ornamentation associated with an element posterior to the nasaloantorbital and excluding it from contact with the anterior orbital margin. This element appears to be anamesic and is tentatively identified as a supraorbital.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF81FF91FF0CFD54BA30CFEB" blockId="16.[189,1399,685,747]" pageId="16" pageNumber="17">
|
||
The orbital series is heavily ossified. The frontal is not excluded from the dorsal edge of the orbit (
|
||
<figureCitation id="132088E1FF81FF91FF77FD29BE62CFEB" box="[198,293,720,746]" captionStart="Fig" captionStartId="9.[189,232,1718,1744]" captionTargetBox="[195,1396,311,1622]" captionTargetId="graphics@9.[317,1359,731,1036]" captionTargetPageId="9" captionText="Fig. 3. Dorsal skull roof. A. Saurorhynchus anningae sp. nov. (NHMUK PV P 964); note that this skull has been dorsoventrally compressed. B. Saurorhynchus hauffi sp. nov., three-dimensionally preserved skull GG 20001. Fine dashed lines indicate sensory canals, large dashes indicate broken or incomplete areas; grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l.ex = lateral extrascapular; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; p = parietal; pm = rostropremaxilla; so.c = supraorbital sensory canal. Scale bars = 10 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829288" httpUri="https://zenodo.org/record/3829288/files/figure.png" pageId="16" pageNumber="17">Fig. 3B</figureCitation>
|
||
), in contrast to previous reconstructions (
|
||
<bibRefCitation id="EF8AE995FF81FF91FCAEFD29BCE9CFEB" author="Hauff B." box="[799,942,720,746]" pageId="16" pageNumber="17" pagination="214 - 248" refId="ref13522" refString="Hauff B. 1938. Uber Acidorhynchus aus den Posidonienschiefern von Holzmaden. Palaontologische Zeitschrift 20: 214 - 248. https: // doi. org / 10.1007 / BF 03041918" type="journal article" year="1938">Hauff 1938</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8AE995FF81FF91FC0CFD29BB0ECFEA" author="Thies D." box="[957,1097,720,747]" pageId="16" pageNumber="17" pagination="183 - 203" refId="ref14987" refString="Thies D. 1985. Funde von Acidorhynchus brevirostris (Woodward 1895) aus dem Posidonienschiefer (Unter-Toarcium) NW-Deutschlands. Palaeontographica Abt. A 187 (4 - 6): 183 - 203." type="journal article" year="1985">Thies 1985</bibRefCitation>
|
||
). The lacrimal is weakly
|
||
</paragraph>
|
||
<caption id="DF64C4ECFF81FF91FF0CF920BD13CAEB" ID-DOI="http://doi.org/10.5281/zenodo.3829297" ID-Zenodo-Dep="3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1753,1779]" targetBox="[189,1399,836,1707]" targetPageId="16">
|
||
<paragraph id="8BA49464FF81FF91FF0CF920BD13CAEB" blockId="16.[189,1399,1753,2027]" pageId="16" pageNumber="17">
|
||
<emphasis id="B96F4876FF81FF91FF0CF920BE4BCBF5" bold="true" box="[189,268,1753,1780]" pageId="16" pageNumber="17">Fig. 7.</emphasis>
|
||
Postcranium, Early Jurassic saurichthyids.
|
||
<emphasis id="B96F4876FF81FF91FCA7F920BC6CCBF2" bold="true" box="[790,811,1753,1779]" pageId="16" pageNumber="17">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF81FF91FC8DF923BB25CBF5" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[828,1122,1754,1780]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF81FF91FC8DF923BB25CBF5" box="[828,1122,1754,1780]" italics="true" pageId="16" pageNumber="17">Saurorhynchus anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF81FF91FBDDF923BB8CCBF5" box="[1132,1227,1754,1780]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790).
|
||
<emphasis id="B96F4876FF81FF91FB82F904BB2DCA16" bold="true" box="[1075,1130,1789,1815]" pageId="16" pageNumber="17">B–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C1BEFE7FF81FF91FBCAF904BA35CA16" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[1147,1394,1789,1815]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF81FF91FBCAF904BA35CA16" box="[1147,1394,1789,1815]" italics="true" pageId="16" pageNumber="17">Saurorhynchus hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF81FF91FF0CF8D8BE5DCA3A" box="[189,282,1825,1851]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, SMNS 55057.
|
||
<emphasis id="B96F4876FF81FF91FE64F8D9BEAECA3B" bold="true" box="[469,489,1824,1850]" pageId="16" pageNumber="17">B</emphasis>
|
||
. Lepidotrichia of the anal fin.
|
||
<emphasis id="B96F4876FF81FF91FCFDF8D9BC26CA3B" bold="true" box="[844,865,1824,1850]" pageId="16" pageNumber="17">C</emphasis>
|
||
. Relationship between the neural and haemal arches and the axonosts.
|
||
<emphasis id="B96F4876FF81FF91FE5AF8BABD47CA5C" bold="true" box="[491,512,1859,1885]" pageId="16" pageNumber="17">D</emphasis>
|
||
. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA49464FF80FF90FF0CFEF2BB8CCC48" blockId="17.[189,1398,267,330]" pageId="17" pageNumber="18">ossified and anteroposteriorly elongate, articulating with the maxilla ventrally, premaxilla anteriorly, and nasaloantorbital and supraorbital dorsally. It transmits the infraorbital sensory canal.</paragraph>
|
||
<paragraph id="8BA49464FF80FF90FF0CFE8BBCB4CF1C" blockId="17.[189,1399,370,542]" pageId="17" pageNumber="18">The dermosphenotic is variable in size, and forms the posterodorsal edge of the orbit. It carries the infraorbital canal. Ventral to the dermosphenotic, an elongate infraorbital forms the majority of the posterior orbit and carries the sensory canal. Infraorbitals along the ventral edge of the orbit are extremely weakly ossified (just a thin tube of bone surrounding the infraorbital canal, not in direct articulation with more anterior or posterior elements), and as such are rarely preserved.</paragraph>
|
||
<paragraph id="8BA49464FF80FF90FF0CFDBCBEB1CE80" blockId="17.[189,1399,581,898]" pageId="17" pageNumber="18">
|
||
Posterior to the dermosphenotic and fused with the lateral dermopterotic, an elongate, anamesic element is sometimes preserved. As with the dermosphenotic, it is somewhat variable in terms of size. It is often missing, and even when present the sutural contact can be very difficult to see. This element is most similar to the “supraspiracular plate” of
|
||
<taxonomicName id="4C1BEFE7FF80FF90FD23FD4ABB79CFCD" authority="Stensio, 1932" authorityName="Stensio" authorityYear="1932" box="[658,1086,690,716]" class="Actinopterygii" family="Birgeriidae" genus="Birgeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Palaeonisciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="groenlandica">
|
||
<emphasis id="B96F4876FF80FF90FD23FD4ABCD0CFCD" box="[658,919,690,716]" italics="true" pageId="17" pageNumber="18">Birgeria groenlandica</emphasis>
|
||
Stensiö,
|
||
<quantity id="4CE33981FF80FF90FBB2FD4BBB79CFCD" box="[1027,1086,690,716]" metricMagnitude="1" metricUnit="m" metricValue="4.907279999999999" pageId="17" pageNumber="18" unit="in" value="1932.0">1932</quantity>
|
||
</taxonomicName>
|
||
in shape, position, and the absence of a sensory canal (
|
||
<bibRefCitation id="EF8AE995FF80FF90FDB8FD2FBDEECFF1" author="Nielsen E." box="[521,681,726,752]" pageId="17" pageNumber="18" pagination="1 - 309" refId="ref14241" refString="Nielsen E. 1949. Studies on Triassic Fishes II. Palaeozoologica Groenlandica 3: 1 - 309." type="journal article" year="1949">Nielsen 1949</bibRefCitation>
|
||
). The ventral dermosphenotic in
|
||
<taxonomicName id="4C1BEFE7FF80FF90FB85FD2EBA30CFF1" authority="Aldinger, 1931" authorityName="Aldinger" authorityYear="1931" box="[1076,1399,726,752]" class="Actinopterygii" family="Birgeriidae" genus="Birgeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Palaeonisciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="stensioei">
|
||
<emphasis id="B96F4876FF80FF90FB85FD2EBBFACFF1" box="[1076,1213,727,752]" italics="true" pageId="17" pageNumber="18">B. stensioei</emphasis>
|
||
Aldinger, 1931
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF80FF90FF74FD02BD57CE14" author="Romano C. & Brinkmann W." box="[197,528,762,789]" pageId="17" pageNumber="18" pagination="17 - 31" refId="ref14762" refString="Romano C. & Brinkmann W. 2009. Reappraisal of the lower actinopterygian Birgeria stensioei Aldinger, 1931 (Osteichthyes; Birgeriidae) from the Middle Triassic of Monte San Giorgio (Switzerland) and Besano (Italy). Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen 252 (1): 17 - 31. https: // doi. org / 10.1127 / 0077 - 7749 / 2009 / 0252 - 0017" type="journal article" year="2009">Romano & Brinkmann 2009</bibRefCitation>
|
||
) was later homologized with the “supraspiracular plate” of
|
||
<taxonomicName id="4C1BEFE7FF80FF90FB07FD02BA28CE14" authorityName="Stensio" authorityYear="1932" box="[1206,1391,763,789]" class="Actinopterygii" family="Birgeriidae" genus="Birgeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Palaeonisciformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="groenlandica">
|
||
<emphasis id="B96F4876FF80FF90FB07FD02BA28CE14" box="[1206,1391,763,789]" italics="true" pageId="17" pageNumber="18">B. groenlandica</emphasis>
|
||
</taxonomicName>
|
||
, however the ventral dermosphenotic should contact the infraorbital and carry the infraorbital canal (
|
||
<bibRefCitation id="EF8AE995FF80FF90FF74FCBABE1ECE5C" author="Poplin C. M." box="[197,345,835,861]" pageId="17" pageNumber="18" pagination="165 - 178" refId="ref14309" refString="Poplin C. M. 2004. The dermosphenotic in early actinopterygians, a nomenclatural problem. In: Arratia G. & Tintori A. (eds) Mesozoic fishes 3, Systematics, Paleoenvironments and Biodiversity: 165 - 178. Verlag Dr Friedrich Pfeil, Munich." type="book chapter" year="2004">Poplin 2004</bibRefCitation>
|
||
), which the elongate element in
|
||
<taxonomicName id="4C1BEFE7FF80FF90FD55FCBABC01CE5C" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[740,838,835,861]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF80FF90FD55FCBABC01CE5C" box="[740,838,835,861]" italics="true" pageId="17" pageNumber="18">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF80FF90FCFEFCBDBCEBCE5F" box="[847,940,836,862]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
does not. We tentatively consider this element to be a neomorph.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF80FF90FF0CFC53BA33C978" blockId="17.[189,1399,938,1145]" pageId="17" pageNumber="18">
|
||
The dermopterotic is a broad dorsally convex sheet of bone making up the posterior skull roof (
|
||
<figureCitation id="132088E1FF80FF90FABCFC53BA2FCEC5" box="[1293,1384,938,964]" captionStart="Fig" captionStartId="9.[189,232,1718,1744]" captionTargetBox="[195,1396,311,1622]" captionTargetId="graphics@9.[317,1359,731,1036]" captionTargetPageId="9" captionText="Fig. 3. Dorsal skull roof. A. Saurorhynchus anningae sp. nov. (NHMUK PV P 964); note that this skull has been dorsoventrally compressed. B. Saurorhynchus hauffi sp. nov., three-dimensionally preserved skull GG 20001. Fine dashed lines indicate sensory canals, large dashes indicate broken or incomplete areas; grey lines indicate bones that were present but where the exact location of the sutural contact is unclear, and question marks indicate areas of uncertainty. Abbreviations: dpt = dermopterotic; dsp = dermophenotic; f = frontal; io = infraorbital; io.c = infraorbital sensory canal; l.ex = lateral extrascapular; mio.c = medial branch of the infraorbital sensory canal; n = neomorph; na-ao = nasaloantorbital; p = parietal; pm = rostropremaxilla; so.c = supraorbital sensory canal. Scale bars = 10 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829288" httpUri="https://zenodo.org/record/3829288/files/figure.png" pageId="17" pageNumber="18">Fig. 3B</figureCitation>
|
||
). A descending process forms an interdigitating suture with the ascending process of the parasphenoid, and creates a notch into which slots the posterior end of the neomorph. Ornamentation generally consists of pits. There is an elongate pore medial to the hyomandibular contact indicating that the medial process of the infraorbital sensory canal was present. Anterior to this canal are the parietals. These form a large, oval ossification. Ornamentation is reticular. No evidence of sensory canals or pores could be detected.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF80FF90FF0CFB5BBC1AC805" blockId="17.[189,1399,1185,1284]" pageId="17" pageNumber="18">
|
||
An elongate lateral extrascapular is sutured to the lateral dermopterotic, dorsal to the hyomandibula (
|
||
<figureCitation id="132088E1FF80FF90FF77FB3CBE64C9E1" box="[198,291,1221,1248]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="17" pageNumber="18">Fig. 5D</figureCitation>
|
||
). It transmits the posterior infraorbital canal.A medial extrascapular is also present, and together they form a v-shaped articulation with the dermopterotic.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF80FF90FF0CFAD5BCDCC88E" blockId="17.[189,1399,1324,1423]" pageId="17" pageNumber="18">
|
||
A dermohyal was present, as the dorsal portion of the hyomandibular bears the pitted ornamentation characteristic of dermal bones (
|
||
<figureCitation id="132088E1FF80FF90FDFEFAA9BDF1C86A" box="[591,694,1360,1387]" captionStart="Fig" captionStartId="13.[189,232,1790,1816]" captionTargetBox="[189,1399,869,1754]" captionTargetId="figure@13.[189,1399,869,1754]" captionTargetPageId="13" captionText="Fig. 5. Characters distinguishing Saurorhynchus acutus (Agassiz, 1844) and S. hauffi sp. nov. A. S. acutus, narial region (SMNS 56923). B. S. acutus, lateral extrascapular-dermopterotic contact (SMNS 57039). C. S. hauffi sp. nov., narial region (SMNS 51888). D. S. hauffi sp. nov., lateral extrascapular-dermopterotic contact (SMNS 53980, mirrored). Abbreviations: dpt = dermopterotic; dh = dermohyal; f = frontal; l.ex = lateral extrascapular; m = maxilla; na-ao = nasaloantorbital; orb = orbit; pm = rostropremaxilla; pop = preopercle; pop* = damaged preopercle; ps = parasphenoid; so = supraorbital. Scale bars: A, C–D = 2 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3829293" httpUri="https://zenodo.org/record/3829293/files/figure.png" pageId="17" pageNumber="18">Fig. 5D</figureCitation>
|
||
). A prominent posteriorly oriented joint surface on the ventrolateral dermopterotic articulates with the hyomandibula.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF80FF90FF0CFA4EBCAFCBF2" blockId="17.[189,1399,1463,1780]" pageId="17" pageNumber="18">
|
||
The palate is not exposed in any of the skulls attributable to this species, however the parasphenoid is often well-exposed in lateral view. The parasphenoid rostrum is curved and edentulous ventral to the orbit; the posterior stem is straight and dorsally displaced. The ascending process of the parasphenoid projects dorsolaterally, and is ornamented with fine striations. The foramen for the ophthalmic artery is closer to the anterior than posterior edge of the ventral ascending process (
|
||
<figureCitation id="132088E1FF80FF90FBC0F9B1BB95CB63" box="[1137,1234,1608,1634]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="17" pageNumber="18">Fig. 1D</figureCitation>
|
||
). There is no expanded tooth plate ventral to the ascending process, and the entire parasphenoid posterior to the anterior margin of the orbit is edentulous. The posterior stem of the parasphenoid is broad. Anteriorly, a central ridge and two lateral ridges are present on the ventral surface of the posterior stem; posteriorly the central ridge diminishes such that it is absent at the posterior end.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF80FF93FF0CF8E2BBACCC48" blockId="17.[189,1399,1819,2027]" lastBlockId="18.[189,1399,267,330]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
Posterior to the orbit, the large foramen for the internal carotid and efferent pseudobranchial artery is contained within the basisphenoid (
|
||
<figureCitation id="132088E1FF80FF90FD3DF8B9BC57CA5B" box="[652,784,1856,1882]" captionStart="Fig" captionStartId="10.[189,232,1683,1709]" captionTargetBox="[189,1399,933,1642]" captionTargetId="figure@10.[189,1399,929,1642]" captionTargetPageId="10" captionText="Fig. 4. Orbitotemporal region, illustrating variation in the braincase of Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov. (based on NHMUK PV P 36227). B, E. Saurorhynchus brevirostris (Woodward, 1895) (photo pertains to NHMUK PV P 4878, mirrored). C, F. Saurorhynchus hauffi sp. nov. (photo is of SMNS 51888, mirrored). D. Saurorhynchus acutus (Agassiz, 1844) (based on SMNS 87737). Abbreviations: asc = ascending process of the parasphenoid; bs = basisphenoid; dpt = dermopterotic; dsp = dermosphenotic; f.ic+ep = foramen for the internal carotid artery and efferent pseudobranchial artery; f.oma = f.om = foramen for the great ophthalmic artery; my = posterior myodome; n = neomorph; nc = neurocranium; orb = orbit; psr = parasphenoid rostrum; rot = foramen for the lateral otic ramus; spi = ventral opening of the spiracular canal; tf.c = trigeminofacial chamber. Scale bars: E–F = 2 mm. E. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829291" httpUri="https://zenodo.org/record/3829291/files/figure.png" pageId="17" pageNumber="18">Fig. 4C, F</figureCitation>
|
||
). In one specimen (SMNS 55302) this foramen is subdivided into anterior and posterior components by a pillar of bone. On the anterior surface of the basisphenoid, the large foramen for the ophthalmica magna artery is recessed in a groove laterally bordered by cristae. Posterior to the basisphenoid, the space between the ascending process and basisphenoid has been interpreted as the posterior myodome. The posterior myodome appears to be roofed by an independent ossification dividing the posterior myodome from the trigeminofacialis chamber. Dorsal to this ossification, the foramen for the ramus oticus lateralis emerges from the neurocranium.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF83FF93FF0CFE88BBCACFAA" blockId="18.[189,1399,369,684]" pageId="18" pageNumber="19">
|
||
The preopercle is relatively narrow, with the dorsal ramus being narrower than the posterior ramus. It carries the preopercular sensory canal. Unlike previous reconstructions (
|
||
<bibRefCitation id="EF8AE995FF83FF93FB95FE6CBBF5CCB1" author="Hauff B." box="[1060,1202,405,432]" pageId="18" pageNumber="19" pagination="214 - 248" refId="ref13522" refString="Hauff B. 1938. Uber Acidorhynchus aus den Posidonienschiefern von Holzmaden. Palaontologische Zeitschrift 20: 214 - 248. https: // doi. org / 10.1007 / BF 03041918" type="journal article" year="1938">Hauff 1938</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8AE995FF83FF93FB70FE6CBA35CCAE" author="Gardiner B. G." box="[1217,1394,405,431]" pageId="18" pageNumber="19" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8AE995FF83FF93FF0CFE40BE03CCD5" author="Thies D." box="[189,324,441,468]" pageId="18" pageNumber="19" pagination="183 - 203" refId="ref14987" refString="Thies D. 1985. Funde von Acidorhynchus brevirostris (Woodward 1895) aus dem Posidonienschiefer (Unter-Toarcium) NW-Deutschlands. Palaeontographica Abt. A 187 (4 - 6): 183 - 203." type="journal article" year="1985">Thies 1985</bibRefCitation>
|
||
), it is clear that the preopercular sensory canal runs along the length of the dorsal ramus as well as the posterior ramus, as in
|
||
<taxonomicName id="4C1BEFE7FF83FF93FD35FE24BC48CCF6" authorityName="Reis" authorityYear="1892" box="[644,783,477,503]" class="Actinopterygii" family="Saurichthyidae" genus="Saurichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF83FF93FD35FE24BC48CCF6" box="[644,783,477,503]" italics="true" pageId="18" pageNumber="19">Saurichthys</emphasis>
|
||
</taxonomicName>
|
||
. The dorsal edge of the preopercle bears two tiny processes, the anterior of which is directed dorsoanteriorly and the posterior of which is directed anteromedially. These presumably articulate with the hyomandibula. The suture between the posterior ramus of the preopercle and the maxilla is easy to differentiate based on the change in the direction of ornamentation at the suture. The quadratojugal is rectangular in shape and meets the maxilla anteriorly and the preopercle dorsally and posteriorly. It contacts the articular ventromedially.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF83FF93FF0CFD2ABB13C930" blockId="18.[189,1399,723,1073]" pageId="18" pageNumber="19">The lateral lower jaw consists of four elements: a tiny articular, a small supraangular, an expanded angular, and the dentary. The articular is in contact with the supraangular anteriorly. The dorsal portion of the angular is very thin and often eroded, making it appear that the sensory canal is located near the dorsal edge of this element. However, that is not the case. The mandibular sensory canal is contained entirely within the angular, and runs closer to the occlusal jaw margin than the ventral edge. It gives rise to descending canals. The posterior edge of the angular is strongly sinusoidal, and projects posterior to the jaw joint. The angular is also strongly ornamented with pitting and some reticulation; there are a series of “growth lines” running parallel to the ventral edge. The dentary itself has a woody texture, with the grain oriented parallel to the long axis of the mandible. The ventral part of the angular-dentary suture begins as at a posteroventrally angle dorsally, but becomes deflected ventrally.</paragraph>
|
||
<paragraph id="8BA49464FF83FF93FF0CFBA0BCC5C84A" blockId="18.[189,1399,1113,1356]" pageId="18" pageNumber="19">
|
||
The laniary teeth are simple cones bearing an acutely pointed acrodin cap, and in general appear to be entirely unornamented and lack any differentiation between tooth base and “collar ganoine” (
|
||
<figureCitation id="132088E1FF83FF93FABAFB84BA2EC999" box="[1291,1385,1149,1176]" captionStart="Fig" captionStartId="14.[189,232,1828,1854]" captionTargetBox="[198,1392,811,1788]" captionTargetId="figure@14.[195,1392,808,1788]" captionTargetPageId="14" captionText="Fig. 6. Posteriormost laniary dentition. A. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV P 4878), note the lingually curved crowns. B. Saurorhynchus anningae sp. nov. (NHMUK PV P 27569), note the incisivlücke associated with the premaxillary laniary tooth and the absence of lateral crypts associated with the two flanking mandibular laniaries. Photos © The Trustees of the Natural History Museum, London. C. S. hauffi sp. nov. (SMNS 51007), mid-rostral dentition illustrating the relationship between the incisivlücken, laniaries, and flanking smaller teeth. Scale bars: 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3829295" httpUri="https://zenodo.org/record/3829295/files/figure.png" pageId="18" pageNumber="19">Fig. 6C</figureCitation>
|
||
). However, a single specimen (SMNS 51007) does show extremely faint corrugation of the collar region. We consider that its small maximum tooth size is possibly a reason for restricted expression of corrugation in the collar region (the specimens of
|
||
<taxonomicName id="4C1BEFE7FF83FF93FDCFFB10BC4EC802" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[638,777,1257,1283]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF83FF93FDCFFB10BC4EC802" box="[638,777,1257,1283]" italics="true" pageId="18" pageNumber="19">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF83FF93FCA0FB13BC2AC805" box="[785,877,1258,1284]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
that clearly show a corrugated collar region are among the largest specimens, far larger than the largest
|
||
<taxonomicName id="4C1BEFE7FF83FF93FC34FAF4BCA0C826" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[901,999,1293,1319]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF83FF93FC34FAF4BCA0C826" box="[901,999,1293,1319]" italics="true" pageId="18" pageNumber="19">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF83FF93FC41FAF4BB17C826" box="[1008,1104,1293,1319]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Some variability in the degree of expression must also be assumed (
|
||
<bibRefCitation id="EF8AE995FF83FF93FD74FAC8BC34C84D" author="Gardiner B. G." box="[709,883,1329,1356]" pageId="18" pageNumber="19" pagination="241 - 384" refId="ref13301" refString="Gardiner B. G. 1960. A revision of certain actinopterygian and coelacanth fishes, chiefly from the Lower Lias. Bulletin of the British Museum (Natural History): Geology 4 (7): 241 - 384." type="journal article" year="1960">Gardiner 1960</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF83FF93FF5BFA8ABECDC88C" blockId="18.[189,1399,1395,1673]" box="[234,394,1395,1421]" pageId="18" pageNumber="19">
|
||
<emphasis id="B96F4876FF83FF93FF5BFA8ABECDC88C" bold="true" box="[234,394,1395,1421]" pageId="18" pageNumber="19">Postcranium</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF83FF93FF0CFA61BA21CB88" blockId="18.[189,1399,1395,1673]" pageId="18" pageNumber="19">
|
||
PAIRED FINS. The supracleithra are seldom preserved. They appear to be thin and relatively straight dorsally, ventrally bearing a prominent anterior process that is pierced by a foramen on its medial surface. There is no equivalent to this ventral anterior process of the supracleithrum within
|
||
<taxonomicName id="4C1BEFE7FF83FF93FB37FA26BA56C8F8" authorityName="Reis" authorityYear="1892" box="[1158,1297,1503,1529]" class="Actinopterygii" family="Saurichthyidae" genus="Saurichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF83FF93FB37FA26BA56C8F8" box="[1158,1297,1503,1529]" italics="true" pageId="18" pageNumber="19">Saurichthys</emphasis>
|
||
</taxonomicName>
|
||
, but this morphology is shared with
|
||
<taxonomicName id="4C1BEFE7FF83FF93FE4CF9FABDE6CB1C" baseAuthorityName="Woodward" baseAuthorityYear="1895" box="[509,673,1539,1565]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B96F4876FF83FF93FE4CF9FABDE6CB1C" box="[509,673,1539,1565]" italics="true" pageId="18" pageNumber="19">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
. The cleithra are triradiate, with a robust posterior ramus. The paired fins are even more rarely preserved. In NMB 373, the pectoral fins are triangular in shape, with the longest and most robust lepidotrichia located close to the leading edge of the fin. A precise count of lepidotrichia is not possible. In no specimens referred to
|
||
<taxonomicName id="4C1BEFE7FF83FF93FCC6F996BC90CB88" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[887,983,1647,1673]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="hauffi" status="sp. nov.">
|
||
<emphasis id="B96F4876FF83FF93FCC6F996BC90CB88" box="[887,983,1647,1673]" italics="true" pageId="18" pageNumber="19">S. hauffi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF83FF93FC6EF996BB7ECB88" box="[991,1081,1647,1673]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are pelvic fins preserved.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF83FF92FF0CF948BE9CCC93" blockId="18.[189,1399,1713,2027]" lastBlockId="19.[189,1399,267,402]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
AXIAL SKELETON. The following description is based on SMNS 55057. Sixty-three abdominal and 117 caudal neural arch-like elements are preserved anterior to the caudal fin, for a total of 180 (90 vertebral segments). As indicated by the low abdominal neural arch count, the abdominal-caudal transition is displaced anteriorly relative to the anal fin (
|
||
<figureCitation id="132088E1FF83FF93FD74F8E4BC64CA36" box="[709,803,1821,1847]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[274,1313,297,1817]" captionTargetId="figure@5.[274,1313,297,1817]" captionTargetPageId="5" captionText="Fig. 1. Holotype specimens of Early Jurassic saurichthyids. A. Saurorhynchus acutus (Agassiz, 1844) (NHMUK PV P 4268). B. Saurorhynchus brevirostris (Woodward, 1895) (NHMUK PV OR 40726). C. Saurorhynchus anningae sp. nov. (NHMUK PV P 3791). D–E. Saurorhynchus hauffi sp. nov. (SMNS 55057). Scale bars: A, C = 20 mm; E = 50 mm. Photos A–C © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829284" httpUri="https://zenodo.org/record/3829284/files/figure.png" pageId="18" pageNumber="19">Fig. 1D</figureCitation>
|
||
). The neural arch-like elements consist of a short, robust base, a moderately long prezygapophysis and a long, tapering neural spine. The medial surface of alternating neural arches is pierced by a foramen for the intersegmental vessel (
|
||
<bibRefCitation id="EF8AE995FF83FF93FB30F89CBA75CA81" author="Wu F. - X. & Sun Y. - L. & Hao W. - C. & Jiang D. - Y. & Sun Z. - Y." box="[1153,1330,1893,1920]" pageId="18" pageNumber="19" pagination="249 - 267" refId="ref15442" refString="Wu F. - X., Sun Y. - L., Hao W. - C., Jiang D. - Y. & Sun Z. - Y. 2015. A new species of Saurichthys (Actinopterygii; Saurichthyiformes) from the Middle Triassic of southwestern China, with remarks on pattern of the axial skeleton of saurichthyid fishes. Neues Jahrbuch fur Geologie und Palaontologie Abhandlungen 275 (3): 249 - 267. https: // doi. org / 10.1127 / njgpa / 2015 / 0462" type="journal article" year="2015">
|
||
Wu
|
||
<emphasis id="B96F4876FF83FF93FB05F89FBBA8CA7E" box="[1204,1263,1893,1919]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). The neural spines become longer more posteriorly in the column, and become forked immediately anterior to the insertion for the dorsal fin. Posterior to the dorsal fin, the neural spines are not forked for a short distance, but become bifurcated again and posteriorly inclined towards the caudal fin. The haemal spines are short, robust hooks anteriorly, lying dorsal to the gastric mass. Some of them are also bifurcated in this region. They lengthen at the end of the abdominal cavity, and become bifurcated or even trifurcated dorsal to the anal fin (
|
||
<figureCitation id="132088E1FF82FF92FE0AFEADBD50CC6F" box="[443,535,340,366]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="19" pageNumber="20">Fig. 7C</figureCitation>
|
||
). Posterior to the anal fin, the haemal spines become shorter, more robust, and posteriorly inclined.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF82FF92FF0CFE42BEA0CEF3" blockId="19.[189,1399,442,1011]" pageId="19" pageNumber="20">
|
||
MEDIAN FINS. The median fins are not complete. Eight elongate dorsal axonosts are preserved in the
|
||
<typeStatus id="54A02AC6FF82FF92FF0CFE27BE60CCF9" box="[189,295,478,504]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
, and no dorsal lepidotrichia. The anteriormost axonost is not differentiated relative to more posterior axonosts. Twelve anal axonosts are preserved; again the anteriormost is not differentiated (
|
||
<figureCitation id="132088E1FF82FF92FF77FDDFBE64CF40" box="[198,291,550,577]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="19" pageNumber="20">Fig. 7B</figureCitation>
|
||
). The two forks of the neural and haemal spines insert on either side of an axonost. There is a gap between the axonosts and the lepidotrichia, and a single roundish, extremely weakly ossified baseost is preserved posteriorly. Four basal fulcra are present. The preserved lepidotrichia are unsegmented and are not distally bifurcated. Fringing fulcra are present, with an arrangement similar to that described by
|
||
<bibRefCitation id="EF8AE995FF82FF92FF52FD4EBED7CFD3" author="Stensio E." box="[227,400,695,722]" pageId="19" pageNumber="20" pagination="1 - 126" refId="ref14959" refString="Stensio E. 1925. Triassic fishes from Spitzbergen, Part II. Kungliga Svenska Vetenskapsakademiens Handlingar Tredje Serien 2 (1): 1 - 126." type="journal article" year="1925">Stensiö (1925)</bibRefCitation>
|
||
for the pelvic fins of
|
||
<taxonomicName id="4C1BEFE7FF82FF92FD22FD41BC4BCFD0" baseAuthorityName="Stensio" baseAuthorityYear="1925" box="[659,780,696,722]" class="Actinopterygii" family="Saurichthyidae" genus="Saurichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="ornatus">
|
||
<emphasis id="B96F4876FF82FF92FD22FD41BC4BCFD0" box="[659,780,696,722]" italics="true" pageId="19" pageNumber="20">S. ornatus</emphasis>
|
||
</taxonomicName>
|
||
, but differing in that the fringing fulcrum and distal lepidotrichium appear to be fused, especially rostrally. When fused, a suture is present between the fulcra and underlying lepidotrichia. The posterior anal fin consists of finer, soft rays. A heavily ossified element associated with more poorly ossified endochondral fragments is also preserved in this region. It is bilaterally symmetrical, forked anteriorly with each of the two rami bifurcating again. The posterior end bears three small projections, with the medial projection being slightly longer than the other two. This element is tentatively interpreted as the axonost plate (
|
||
<figureCitation id="132088E1FF82FF92FC3DFC69BCADCEAB" box="[908,1002,912,938]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="19" pageNumber="20">Fig. 7B</figureCitation>
|
||
), based on comparisons in shape and position with the same element in
|
||
<taxonomicName id="4C1BEFE7FF82FF92FD13FC4CBC88CECF" authority="(Schwarz 1970)" baseAuthorityName="Schwarz" baseAuthorityYear="1970" box="[674,975,948,974]" class="Actinopterygii" family="Birgeriidae" genus="Birgeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Palaeonisciformes" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96F4876FF82FF92FD13FC4CBC43CECF" box="[674,772,949,974]" italics="true" pageId="19" pageNumber="20">Birgeria</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8AE995FF82FF92FCA9FC4DBC80CECF" author="Schwarz W." box="[792,967,948,974]" pageId="19" pageNumber="20" pagination="1 - 93" refId="ref14934" refString="Schwarz W. 1970. Die Triasfauna der Tessiner Kalkalpen XX. Birgeria stensioi Aldinger. Schweizerische Palaontologische Abhandlungen 89: 1 - 93." type="journal article" year="1970">Schwarz 1970</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Caudal radials are preserved, but lepidotrichia are missing.
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF82FF92FF0CFBE2BCB5C9EB" blockId="19.[189,1399,1051,1258]" pageId="19" pageNumber="20">
|
||
SQUAMATION. The only scale row present anterior to the median fins is the mid-dorsal scale row. Scales in this row are elongate, needle-like, and unornamented (
|
||
<figureCitation id="132088E1FF82FF92FCCBFBC6BC9DC958" box="[890,986,1087,1113]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="19" pageNumber="20">Fig. 7C</figureCitation>
|
||
). Posterior to the median fins, no scales are present, unlike in
|
||
<taxonomicName id="4C1BEFE7FF82FF92FDB6FB9ABDD6C97C" authority="Maxwell & Stumpf, 2017" authorityName="Maxwell & Stumpf" authorityYear="2017" box="[519,657,1123,1149]" class="Actinopterygii" family="Saurichthyidae" genus="Saurorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurichthyiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="anningae" status="sp. nov.">
|
||
<emphasis id="B96F4876FF82FF92FDB6FB9ABDD6C97C" box="[519,657,1123,1149]" italics="true" pageId="19" pageNumber="20">S. anningae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25CF50DFF82FF92FD29FB9ABDB4C97C" box="[664,755,1123,1149]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in which, at minimum, the mid-ventral row is ossified. Immediately anterior to the caudal fin, the caudal peduncle is encased in robust mid-dorsal and midventral scales, which grade into basal fulcra posteriorly (
|
||
<figureCitation id="132088E1FF82FF92FCE3FB52BCF7C9C7" box="[850,944,1195,1222]" captionStart="Fig" captionStartId="16.[189,232,1753,1779]" captionTargetBox="[189,1399,836,1707]" captionTargetId="figure@16.[189,1399,832,1707]" captionTargetPageId="16" captionText="Fig. 7. Postcranium, Early Jurassic saurichthyids. A. Saurorhynchus anningae sp. nov., neural arches and squamation in the posterior abdominal region (NHMUK PV P 3790). B–D. Saurorhynchus hauffi sp. nov., SMNS 55057. B. Lepidotrichia of the anal fin. C. Relationship between the neural and haemal arches and the axonosts. D. Caudal peduncle. Abbreviations: af.ax = axonosts of the anal fin; ax.p = axonost plate; bf = basal fulcra; cf.r = caudal fin radials; df.ax = axonosts of the dorsal fin; ff = fringing fulcra; hs = haemal spine; lep = lepidotrichia; mds = mid-dorsal scale row; mvs = mid-ventral scale row; na = neural arch; ns = neural spine. Scale bars: A = 1 mm, B–D = 5 mm. A. Photo © The Trustees of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.3829297" httpUri="https://zenodo.org/record/3829297/files/figure.png" pageId="19" pageNumber="20">Fig. 7D</figureCitation>
|
||
). The scales in the caudal peduncle are more robust than the abdominal scales, but are also smooth in texture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C301C7EFFF82FF92FF0CFAE8BD97C8A5" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="8BA49464FF82FF92FF0CFAE8BE08C82A" blockId="19.[189,335,1297,1323]" box="[189,335,1297,1323]" pageId="19" pageNumber="20">
|
||
<heading id="D0EC2308FF82FF92FF0CFAE8BE08C82A" bold="true" box="[189,335,1297,1323]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B96F4876FF82FF92FF0CFAE8BE08C82A" bold="true" box="[189,335,1297,1323]" pageId="19" pageNumber="20">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA49464FF82FF92FF0CFAB8BD97C8A5" blockId="19.[189,1399,1345,1444]" pageId="19" pageNumber="20">
|
||
Early Jurassic, Toarcian,
|
||
<taxonomicName id="4C1BEFE7FF82FF92FE46FABBBDAFC85D" authority="Zone" authorityName="Zone" box="[503,744,1346,1372]" class="Cephalopoda" family="Dactylioceratidae" genus="Dactylioceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="tenuicostatum">
|
||
<emphasis id="B96F4876FF82FF92FE46FABBBDDBC85A" box="[503,668,1346,1371]" italics="true" pageId="19" pageNumber="20">tenuicostatum</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1BEFE7FF82FF92FD4BFAB8BB47C85A" authority="Subzone" authorityName="Subzone" box="[762,1024,1345,1371]" class="Cephalopoda" family="Dactylioceratidae" genus="Dactylioceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="semicelatum">
|
||
<emphasis id="B96F4876FF82FF92FD4BFAB8BCCAC85A" box="[762,909,1345,1371]" italics="true" pageId="19" pageNumber="20">semicelatum</emphasis>
|
||
Subzone
|
||
</taxonomicName>
|
||
to
|
||
<emphasis id="B96F4876FF82FF92FB83FABBBB84C85A" box="[1074,1219,1346,1371]" italics="true" pageId="19" pageNumber="20">serpentinum</emphasis>
|
||
Zone, Baden-Württemberg,
|
||
<collectingCountry id="F30CD4F4FF82FF92FEDCFA9CBE99C87E" box="[365,478,1381,1407]" name="Germany" pageId="19" pageNumber="20">Germany</collectingCountry>
|
||
;
|
||
<emphasis id="B96F4876FF82FF92FE5EFA9FBDC7C87E" box="[495,640,1382,1407]" italics="true" pageId="19" pageNumber="20">serpentinum</emphasis>
|
||
Zone,
|
||
<collectingRegion id="49DF5A86FF82FF92FD6CFA9FBCD7C87E" box="[733,912,1381,1408]" country="Germany" name="Niedersachsen" pageId="19" pageNumber="20">Lower Saxony</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30CD4F4FF82FF92FC10FA9CBB55C87E" box="[929,1042,1381,1407]" name="Germany" pageId="19" pageNumber="20">Germany</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C1BEFE7FF82FF92FB92FA9FBA41C87E" authority="Subzone" authorityName="Subzone" box="[1059,1286,1381,1407]" class="Cephalopoda" family="Hildoceratidae" genus="Cleviceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="exaratum">
|
||
<emphasis id="B96F4876FF82FF92FB92FA9FBBD5C87E" box="[1059,1170,1382,1407]" italics="true" pageId="19" pageNumber="20">exaratum</emphasis>
|
||
Subzone
|
||
</taxonomicName>
|
||
, Western Pomerania,
|
||
<collectingCountry id="F30CD4F4FF82FF92FEF8FA73BEFDC8A5" box="[329,442,1418,1444]" name="Germany" pageId="19" pageNumber="20">Germany</collectingCountry>
|
||
; Toarcian, Whitby,
|
||
<collectingCountry id="F30CD4F4FF82FF92FD2FFA73BD8BC8A5" box="[670,716,1418,1444]" name="United Kingdom" pageId="19" pageNumber="20">UK</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |