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<mods:title id="D453CE8B8CE75491C3995605EEF65E90">A review of the families and genera of the hyperiidean amphipod superfamily Phronimoidea Bowman &amp; Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea)</mods:title>
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<treatment id="03931615EC78FFF8FEDFFD2CFAB8FE57" ID-DOI="http://doi.org/10.5281/zenodo.5029968" ID-GBIF-Taxon="184952200" ID-Zenodo-Dep="5029968" LSID="urn:lsid:plazi:treatment:03931615EC78FFF8FEDFFD2CFAB8FE57" httpUri="http://treatment.plazi.org/id/03931615EC78FFF8FEDFFD2CFAB8FE57" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="C320F488EC78FFF9FEDFFD2CFD46FD40" box="[264,724,724,750]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B85A703EC78FFF9FEDFFD2CFD46FD40" blockId="8.[264,724,724,750]" box="[264,724,724,750]" pageId="8" pageNumber="9">
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<emphasis id="B94E7B11EC78FFF9FEDFFD2CFD46FD40" bold="true" box="[264,724,724,750]" pageId="8" pageNumber="9">
Family
<taxonomicName id="4C3ADC80EC78FFF9FEB0FD2CFD46FD40" ID-CoL="7NKG2" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[359,724,724,750]" class="Malacostraca" family="Phronimidae" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">
PHRONIMIDAE
<bibRefCitation id="EFABDAF2EC78FFF9FD9EFD2CFD46FD40" author="Dana, J. D." box="[585,724,724,750]" pageId="8" pageNumber="9" pagination="297 - 316" refId="ref25481" refString="Dana, J. D. (1852) On the classification of the Crustacea Choristopoda or Tetradecapoda. American Journal of Sciences and Arts, series 2, 14 (41), 297 - 316." type="journal article" year="1852">Dana, 1852</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="8B85A703EC78FFF9FEDFFCE7FEEDFC96" blockId="8.[264,1323,799,1545]" box="[264,383,799,824]" pageId="8" pageNumber="9">
<emphasis id="B94E7B11EC78FFF9FEDFFCE7FEEDFC96" box="[264,383,799,824]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B85A703EC78FFF9FEEFFCBFFB92FA4F" blockId="8.[264,1323,799,1545]" pageId="8" pageNumber="9">
Body length
<quantity id="4CC20AE6EC78FFF9FE02FCBFFDCBFCCF" box="[469,601,839,865]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="mm" value="25.0" valueMax="40.0" valueMin="10.0">1040 mm</quantity>
, cuticle transparent. Head subconical, widest dorsally, narrowing and prolonged ventrally, about twice as deep as long. Eyes occupying most of head surface, divided into larger dorsal part and smaller ventrolateral part. Pereonites all separate (
<taxonomicName id="4C3ADC80EC78FFF9FEADFC47FE7CFC76" authorityName="Latreille" authorityYear="1802" box="[378,494,959,984]" class="Malacostraca" family="Phronimidae" genus="Phronima" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FEADFC47FE7CFC76" box="[378,494,959,984]" italics="true" pageId="8" pageNumber="9">Phronima</emphasis>
</taxonomicName>
), or pereonites 1 &amp; 2 partially fused (
<taxonomicName id="4C3ADC80EC78FFF9FC7CFC47FBAEFC76" authorityName="Claus" authorityYear="1871" box="[939,1084,959,984]" class="Malacostraca" family="Phronimidae" genus="Phronimella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FC7CFC47FBAEFC76" box="[939,1084,959,984]" italics="true" pageId="8" pageNumber="9">Phronimella</emphasis>
</taxonomicName>
). Coxae fused with pereonites. Antennae 1 reduced to two articles in females; in males composed of threearticulate peduncle, large callynophore with dense, two­field brush of aesthestascs medially followed by multi­articulate flagellum. Antennae 2 reduced to small tubercle in females; in males reduced to two small articles in
<taxonomicName id="4C3ADC80EC78FFF9FCADFBA7FB85FBD6" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[890,1047,1119,1144]" class="Malacostraca" family="Phronimidae" genus="Phronima" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sedentaria">
<emphasis id="B94E7B11EC78FFF9FCADFBA7FB85FBD6" box="[890,1047,1119,1144]" italics="true" pageId="8" pageNumber="9">P. sedentaria</emphasis>
</taxonomicName>
, multi­articulate in all other species. Mandibles without palp in both sexes; molar well developed. Maxillae 1 with palp and well­developed outer lobe; inner lobe absent. Maxillae 2 bilobed, welldeveloped. Maxilliped with slender outer lobes; inner lobe very reduced in
<taxonomicName id="4C3ADC80EC78FFF9FB45FB2FFAB6FB5E" authorityName="Claus" authorityYear="1871" box="[1170,1316,1239,1264]" class="Malacostraca" family="Phronimidae" genus="Phronimella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FB45FB2FFAB6FB5E" box="[1170,1316,1239,1264]" italics="true" pageId="8" pageNumber="9">Phronimella</emphasis>
</taxonomicName>
, about half­length (or longer) than outer lobes in
<taxonomicName id="4C3ADC80EC78FFF9FC8EFB07FC5FFAB6" authorityName="Latreille" authorityYear="1802" box="[857,973,1279,1304]" class="Malacostraca" family="Phronimidae" genus="Phronima" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FC8EFB07FC5FFAB6" box="[857,973,1279,1304]" italics="true" pageId="8" pageNumber="9">Phronima</emphasis>
</taxonomicName>
. Gnathopods 1 &amp; 2 weakly cheliform, or simple, with complex dactylus. Pereopods 3, 4, 6 &amp; 7 simple; P3 &amp; 4 usually the longest. Pereopod 5 ending in distinct, broad subchela (
<taxonomicName id="4C3ADC80EC78FFF9FC23FAB7FBFAFAC6" authorityName="Latreille" authorityYear="1802" box="[1012,1128,1359,1384]" class="Malacostraca" family="Phronimidae" genus="Phronima" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FC23FAB7FBFAFAC6" box="[1012,1128,1359,1384]" italics="true" pageId="8" pageNumber="9">Phronima</emphasis>
</taxonomicName>
), or less perfect, slender subchela (
<taxonomicName id="4C3ADC80EC78FFF9FE0BFA8FFDFFFA3E" authorityName="Claus" authorityYear="1871" box="[476,621,1399,1424]" class="Malacostraca" family="Phronimidae" genus="Phronimella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FE0BFA8FFDFFFA3E" box="[476,621,1399,1424]" italics="true" pageId="8" pageNumber="9">Phronimella</emphasis>
</taxonomicName>
). Uropods slender with articulated endopods and exopods. Uropod 2 smaller than others, rudimentary in female
<taxonomicName id="4C3ADC80EC78FFF9FC4CFA67FBBFFA16" authorityName="Claus" authorityYear="1871" box="[923,1069,1439,1464]" class="Malacostraca" family="Phronimidae" genus="Phronimella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FC4CFA67FBBFFA16" box="[923,1069,1439,1464]" italics="true" pageId="8" pageNumber="9">Phronimella</emphasis>
</taxonomicName>
. Telson very small, rounded. Gills on pereonites 46. Oostegites on pereonites 25.
</paragraph>
<paragraph id="8B85A703EC78FFF9FEDFFA17FD74F9A7" blockId="8.[264,1323,799,1545]" box="[264,742,1519,1545]" pageId="8" pageNumber="9">
Two genera:
<taxonomicName id="4C3ADC80EC78FFF9FE76FA17FD87F9A6" authorityName="Latreille" authorityYear="1802" box="[417,533,1519,1544]" class="Malacostraca" family="Phronimidae" genus="Phronima" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FE76FA17FD87F9A6" box="[417,533,1519,1544]" italics="true" pageId="8" pageNumber="9">Phronima</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3ADC80EC78FFF9FD99FA17FD4DF9A6" authorityName="Claus" authorityYear="1871" box="[590,735,1519,1544]" class="Malacostraca" family="Phronimidae" genus="Phronimella" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC78FFF9FD99FA17FD4DF9A6" box="[590,735,1519,1544]" italics="true" pageId="8" pageNumber="9">Phronimella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B85A703EC78FFF9FEDFF9C7FEFCF9F6" blockId="8.[264,1324,1599,1745]" box="[264,366,1599,1624]" pageId="8" pageNumber="9">
<emphasis id="B94E7B11EC78FFF9FEDFF9C7FEFCF9F6" box="[264,366,1599,1624]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
</paragraph>
<paragraph id="8B85A703EC78FFF9FEEFF99FFBC9F97F" blockId="8.[264,1324,1599,1745]" pageId="8" pageNumber="9">
<bibRefCitation id="EFABDAF2EC78FFF9FEEFF99FFE51F92F" author="Shih, C. - T." box="[312,451,1639,1665]" pageId="8" pageNumber="9" pagination="1 - 100" refId="ref29613" refString="Shih, C. - T. (1969) The systematics and biology of the family Phronimidae (Crustacea: Amphipoda). Dana Reports, No. 74, 1 - 100." type="book chapter" year="1969">Shih (1969)</bibRefCitation>
has revised this family. Additions and modifications have been made by
<bibRefCitation id="EFABDAF2EC78FFF9FEDFF977FE38F907" author="Laval, P." box="[264,426,1679,1705]" pageId="8" pageNumber="9" pagination="347 - 362" refId="ref27668" refString="Laval, P. (1968 b) Observations sur la biologie de Phronima curvipes Voss. (Amphipode Hyperide). Cahiers de Biologie Marine, 9, 347 - 362." type="journal article" year="1968">Laval (1968b</bibRefCitation>
,
<bibRefCitation id="EFABDAF2EC78FFF9FE6EF977FE67F907" author="Laval, P." box="[441,501,1679,1705]" pageId="8" pageNumber="9" pagination="47 - 57" refId="ref27701" refString="Laval, P. (1970) Sur les Phronimidae de l'ocean Indien et de l'ocean Pacifique avec la validation de Phronima bucephala Giles, 1887 comme espece distincte de P. colletti Bov., 1887 (Crustaces Amphipodes). Cahiers de I'Office de la Recherche Scientifique et Technique Outre-Mer serie Oceanographie, 8, 47 - 57." type="journal article" year="1970">1970</bibRefCitation>
) and
<bibRefCitation id="EFABDAF2EC78FFF9FDECF977FD5CF907" author="Shih, C. - T." box="[571,718,1679,1705]" pageId="8" pageNumber="9" pagination="25 - 45" refId="ref29648" refString="Shih, C. - T. (1971 a) Phronimidae (Amphipoda, Hyperiidea) of the South Pacific Ocean. Crustaceana, 20 (1), 25 - 45." type="journal article" year="1971">Shih (1971a</bibRefCitation>
,
<bibRefCitation id="EFABDAF2EC78FFF9FD0AF977FCBBF907" author="Shih, C. - T." box="[733,809,1679,1705]" pageId="8" pageNumber="9" pagination="298 - 300" refId="ref29682" refString="Shih, C. - T. (1971 b) Note on Phronima affinis Vosseler, 1901 (Amphipoda, Phronimidae). Crustaceana, 20 (3), 298 - 300." type="journal article" year="1971">1971b</bibRefCitation>
,
<bibRefCitation id="EFABDAF2EC78FFF9FCE0F977FCE7F907" author="Shih, C. - T." box="[823,885,1679,1705]" pageId="8" pageNumber="9" pagination="322 - 335" refId="ref29717" refString="Shih, C. - T. (1991) Description of two new species of Phronima Latreille, 1802 (Amphipoda: Hyperiidea) with a key to all species of the genus. Journal of Crustacean Biology, 11 (2), 322 - 335." type="journal article" year="1991">1991</bibRefCitation>
), and additional taxonomic information for the Australian fauna is provided by
<bibRefCitation id="EFABDAF2EC78FFF9FCDDF94FFC3FF97F" author="Zeidler, W." box="[778,941,1719,1745]" pageId="8" pageNumber="9" pagination="1 - 93" refId="ref31612" refString="Zeidler, W. (1978) Hyperiidea (Crustacea: Amphipoda) from Queensland waters. Australian Journal of Zoology, Supplementary Series, No. 59, 1 - 93." type="book chapter" year="1978">Zeidler (1978</bibRefCitation>
,
<bibRefCitation id="EFABDAF2EC78FFF9FC6EF94FFB91F97F" author="Zeidler, W." box="[953,1027,1719,1745]" pageId="8" pageNumber="9" pagination="85 - 133" refId="ref31645" refString="Zeidler, W. (1992 a) Hyperiid amphipods (Crustacea: Amphipoda: Hyperiidea) collected recently from eastern Australian waters. Records of the Australian Museum, 44 (1), 85 - 133." type="journal article" year="1992">1992a</bibRefCitation>
, 1998).
</paragraph>
</subSubSection>
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<paragraph id="8B85A703EC79FFF8FEDFFEE3FCF9FE9B" blockId="9.[264,875,283,309]" box="[264,875,283,309]" pageId="9" pageNumber="10">
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<emphasis id="B94E7B11EC79FFF8FEDFFEE3FCF9FE9B" bold="true" box="[264,875,283,309]" pageId="9" pageNumber="10">
Key to the genera of the family
<taxonomicName id="4C3ADC80EC79FFF8FD46FEE3FCF9FE9B" ID-CoL="7NKG2" authorityName="Dana" authorityYear="1852" box="[657,875,283,309]" class="Malacostraca" family="Phronimidae" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">PHRONIMIDAE</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320F488EC79FFF8FEDFFE9FFAB8FE57" pageId="9" pageNumber="10" type="key">
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1. Pereonites 1 &amp; 2 separate. Pereopod 5 with distinct broad subchela. Uropod 2 welldeveloped ..........................................................................
<taxonomicName id="4C3ADC80EC79FFF8FBD3FE77FABBFE07" ID-CoL="7PRLX" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[1028,1321,399,425]" class="Malacostraca" family="Phronimidae" genus="Phronima" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC79FFF8FBD3FE77FBEAFE06" box="[1028,1144,399,424]" italics="true" pageId="9" pageNumber="10">Phronima</emphasis>
<bibRefCitation id="EFABDAF2EC79FFF8FBA9FE77FABBFE07" author="Latreille, P. A." box="[1150,1321,399,425]" pageId="9" pageNumber="10" refId="ref27430" refString="Latreille, P. A. (1802) Histoire naturelle, generale et particuliere des Crustaces et des Insectes. 4 Vols. Paris." type="book" year="1802">Latreille, 1802</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="30CB0436EC79FFF8FEDFFE4FFAB8FE57" pageId="9" pageNumber="10">
­ Pereonites 1 &amp; 2 partially fused. Pereopod 5 with slender subchela. Uropod 2 rudimentary or absent .................................................................
<taxonomicName id="4C3ADC80EC79FFF8FBD1FE27FAB8FE57" ID-CoL="7PRLY" authority="Claus, 1871" authorityName="Claus" authorityYear="1871" box="[1030,1322,479,505]" class="Malacostraca" family="Phronimidae" genus="Phronimella" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B94E7B11EC79FFF8FBD1FE27FB05FE56" box="[1030,1175,479,504]" italics="true" pageId="9" pageNumber="10">Phronimella</emphasis>
<bibRefCitation id="EFABDAF2EC79FFF8FB49FE27FAB8FE57" author="Claus, C." box="[1182,1322,479,505]" pageId="9" pageNumber="10" pagination="149 - 157" refId="ref25189" refString="Claus, C. (1871) Untersuchungen uber den Bau und der Verwandtschaft der Hyperiden. Nachrichten von der Konigliche Gesellschaft der Wissenschaften und der Georg-Augustus-Universitat zu Gottingen, Yahre 1871, 149 - 157." type="journal article" year="1871">Claus, 1871</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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