449 lines
61 KiB
XML
449 lines
61 KiB
XML
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<mods:title id="E0DB464599866C5128A8A8B939AB752E">Description of a new flower fly species of the Copestylum vagum group (Diptera Syrphidae) from pristine Amazonian rainforests of Colombia and Suriname</mods:title>
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<mods:namePart id="EA2263C7140A1DB5EB5BEE893BF5C045">Montoya, Augusto León</mods:namePart>
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<mods:affiliation id="4ED74B8EC2C7D8116CDCFF3CC335F26F">Universidad de Antioquia, Instituto de Biología, Grupo de Entomología - GEAU, Calle 67 53 - 108, AA 1226 Medellín, Colombia. aleon. montoya @ udea. edu. co; https: // orcid. org / 0000 - 0003 - 3307 - 034 X</mods:affiliation>
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<mods:namePart id="2413EB0BD9F624B4E8572EF8A7E9807A">Parada-Marín, Henry Mauricio</mods:namePart>
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<mods:affiliation id="A890C4D9007BF78BE4736EADA93984F6">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEU-, Laboratório de Entomologia, Av, 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia. & h. parada @ udla. edu. co; https: // orcid. org / 0000 - 0002 - 5387 - 4495</mods:affiliation>
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<mods:nameIdentifier id="E1C34DC7637989DFC10572F1B0610788" type="email">h.parada@udla.edu.co</mods:nameIdentifier>
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<mods:namePart id="4C386D0B857BDFD39B93838ED4CDC679">Ramos-Pastrana, Yardany</mods:namePart>
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<mods:affiliation id="7B1308A5EA02B227C1394FBC20EB1101">Universidad de Antioquia, Instituto de Biología, Grupo de Entomología - GEAU, Calle 67 53 - 108, AA 1226 Medellín, Colombia. aleon. montoya @ udea. edu. co; https: // orcid. org / 0000 - 0003 - 3307 - 034 X & Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEU-, Laboratório de Entomologia, Av, 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia.</mods:affiliation>
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<mods:nameIdentifier id="F843547A0DF1EDFDFECBD79C8F4B0B60" type="email">aleon.montoya@udea.edu.co</mods:nameIdentifier>
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<mods:date id="769990B2941179501DF6EB566AC32351">2022</mods:date>
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<treatment id="03824077FFE0FFCEFF75FF654BF555A4" ID-DOI="http://doi.org/10.5281/zenodo.5851016" ID-GBIF-Taxon="192128721" ID-Zenodo-Dep="5851016" LSID="urn:lsid:plazi:treatment:03824077FFE0FFCEFF75FF654BF555A4" httpUri="http://treatment.plazi.org/id/03824077FFE0FFCEFF75FF654BF555A4" lastPageId="8" lastPageNumber="409" pageId="3" pageNumber="404">
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<taxonomicName id="4C2B8AE2FFE0FFC5FF75FF654AAC548D" ID-CoL="9HZSD" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[151,390,222,249]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
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<emphasis id="B95F2D73FFE0FFC5FF75FF654AAC548D" bold="true" box="[151,390,222,249]" italics="true" pageId="3" pageNumber="404">Copestylum enriquei</emphasis>
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<taxonomicNameLabel id="A26C9008FFE0FFC5FE6FFF654ACD548D" box="[397,487,222,248]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C331A2EAFFE0FFC5FF75FEB94A215569" box="[151,267,258,284]" pageId="3" pageNumber="404" type="description">
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<paragraph id="8B94F161FFE0FFC5FF75FEB94A215569" blockId="3.[151,487,222,284]" box="[151,267,258,284]" pageId="3" pageNumber="404">
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(
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<figureCitation id="1310EDE4FFE0FFC5FF7DFEB94A295569" box="[159,259,258,284]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1354,1378]" captionStartId-1="4.[151,250,1853,1877]" captionStartId-2="5.[151,250,1328,1352]" captionStartId-3="8.[151,250,1019,1043]" captionTargetBox-0="[189,1396,373,1329]" captionTargetBox-1="[175,1411,993,1828]" captionTargetBox-2="[169,1428,273,1305]" captionTargetBox-3="[153,1436,490,995]" captionTargetId-0="figure-40@3.[189,1398,373,1329]" captionTargetId-1="figure-379@4.[175,1411,993,1828]" captionTargetId-2="figure-61@5.[158,1428,273,1305]" captionTargetId-3="figure-153@8.[152,1436,489,996]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="8" captionText-0="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." captionText-1="FIGURE 2. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890). A. Abdomen, lateral view; B. Abdomen, ventral view; C. Whole genitalia including epandrium, cercus, and surstylus, lateral view, detail of the dorsal extension of the epandrium (white arrow); D. Epandrium, dorsal view, detail of the dorsal extension (white arrow); E. Hypandrium, ventral view. Abbreviations used in male genitalia structures are as follows: Ah = Aedeagal hood; Cer = Cercus; Epd = Epandrium; Hyp = Hypandrium; Sa= Surstylar apodeme; Sl= Superior lobe; Sur = Surstyle." captionText-2="FIGURE 3. Copestylum enriquei sp. nov., female, paratype (RMNH-collection). A. Head, frontal view; B. Head, lateral view, C. Habitus, lateral view; D. Thorax and abdomen, dorsal view." captionText-3="FIGURE 4. A. Habitat of Copestylum enriquei sp. nov. in a pristine forest at Florencia, Caquetá in Colombia. The specimens were collected using a canopy Malaise trap (Rafael & Gorayeb 1982); B. Geographical distribution map showing the collection localities of Copestylum enriquei sp. nov. in the Amazon rainforest of Colombia and Suriname (Red triangle)." figureDoi-0="http://doi.org/10.5281/zenodo.5848271" figureDoi-1="http://doi.org/10.5281/zenodo.5848273" figureDoi-2="http://doi.org/10.5281/zenodo.5848275" figureDoi-3="http://doi.org/10.5281/zenodo.5848277" httpUri-0="https://zenodo.org/record/5848271/files/figure.png" httpUri-1="https://zenodo.org/record/5848273/files/figure.png" httpUri-2="https://zenodo.org/record/5848275/files/figure.png" httpUri-3="https://zenodo.org/record/5848277/files/figure.png" pageId="3" pageNumber="404">Figs 1–4</figureCitation>
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)
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</paragraph>
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<taxonomicName id="4C2B8AE2FFE0FFC5FF75FEFC4A25552A" ID-CoL="62N9P" authorityName="Macquart" authorityYear="1846" box="[151,271,327,351]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B95F2D73FFE0FFC5FF75FEFC4A25552A" box="[151,271,327,351]" italics="true" pageId="3" pageNumber="404">Copestylum</emphasis>
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</taxonomicName>
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SUR-04 in
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<bibRefCitation id="EFBA8C90FFE0FFC5FE6BFEFC490B552A" author="Reemer, M." box="[393,545,327,351]" pageId="3" pageNumber="404" pagination="97 - 142" refId="ref12659" refString="Reemer, M. (2016) Syrphidae (Diptera) of Surinam: Eristalinae and synthesis. Tijdschrift voor Entomologie, 159, 97 - 142. https: // doi. org / 10.1163 / 22119434 - 15902002" type="journal article" year="2016">Reemer (2016)</bibRefCitation>
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<emphasis id="B95F2D73FFE0FFC5FF75FA624A695186" bold="true" box="[151,323,1496,1523]" pageId="3" pageNumber="404">Type material.</emphasis>
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<materialsCitation id="3B43FB3CFFE0FFC5FEAAFA634A2152F1" ID-GBIF-Occurrence="3436730323" collectingDate="2016-12-21" collectingDateMax="2017-01-04" collectingDateMin="2016-12-21" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana" country="Colombia" elevation="716" latitude="1.746287" location="Original" longLatPrecision="1" longitude="-75.62779" municipality="Adult" pageId="3" pageNumber="404" specimenCount="2" stateProvince="Caqueta" typeStatus="holotype">
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<typeStatus id="54904FC3FFE0FFC5FEAAFA634AC35187" box="[328,489,1496,1522]" pageId="3" pageNumber="404" type="holotype">HOLOTYPE</typeStatus>
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.
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<collectingMunicipality id="6BF06B1BFFE0FFC5FE10FA6249185186" box="[498,562,1497,1523]" pageId="3" pageNumber="404">Adult</collectingMunicipality>
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male, pinned, deposited at the LEUA collection.
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<location id="8EF4A7BAFFE0FFC5FBAFFA624F805186" LSID="urn:lsid:plazi:treatment:03824077FFE0FFCEFF75FF654BF555A4:8EF4A7BAFFE0FFC5FBAFFA624F805186" box="[1101,1194,1497,1523]" country="Colombia" latitude="1.746287" longLatPrecision="1" longitude="-75.62779" municipality="Adult" name="Original" pageId="3" pageNumber="404" stateProvince="Caqueta">Original</location>
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label: “
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<collectingCountry id="F33CB1F1FFE0FFC5FB1FFA624EB25186" box="[1277,1432,1497,1523]" name="Colombia" pageId="3" pageNumber="404">COLOMBIA</collectingCountry>
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,
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<collectingRegion id="49EF3F83FFE0FFC5FF75FA464BDC5262" box="[151,246,1533,1559]" country="Colombia" name="Caqueta" pageId="3" pageNumber="404">Caquetá</collectingRegion>
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, Florencia, Vda.[Vereda] Paraíso, Fca.[Finca] Paraíso” / “
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,
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<geoCoordinate id="EE1F97A6FFE0FFC5FBC3FA474F8E5262" box="[1057,1188,1532,1559]" degrees="75.627790" direction="west" orientation="longitude" pageId="3" pageNumber="404" precision="1" value="-75.62779">-75.627790</geoCoordinate>
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,
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<quantity id="4CD35C84FFE0FFC5FB56FA474FD35262" box="[1204,1273,1532,1559]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="3" pageNumber="404" unit="m" value="716.0">
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</quantity>
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[eters], Forest, trampa Malaise dosel” / “
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<date id="FF95D7A1FFE0FFC5FE27F99A4962524E" box="[453,584,1569,1595]" pageId="3" pageNumber="404" value="2016-12-21">
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<collectingDate id="EFD12E49FFE0FFC5FE27F99A4962524E" box="[453,584,1569,1595]" pageId="3" pageNumber="404" value="2016-12-21">21.xii.2016</collectingDate>
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</date>
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-
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<date id="FF95D7A1FFE0FFC5FDB2F99A4996524E" box="[592,700,1569,1595]" pageId="3" pageNumber="404" value="2017-01-04">
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<collectingDate id="EFD12E49FFE0FFC5FDB2F99A4996524E" box="[592,700,1569,1595]" pageId="3" pageNumber="404" value="2017-01-04">04.i.2017</collectingDate>
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</date>
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,
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<collectorName id="26DE94B7FFE0FFC5FD29F99A4881524E" box="[715,939,1569,1595]" pageId="3" pageNumber="404">Y. Ramos-Pastrana</collectorName>
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”. “
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<typeStatus id="54904FC3FFE0FFC5FC2FF99A4F4E524E" box="[973,1124,1569,1595]" pageId="3" pageNumber="404" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="4C2B8AE2FFE0FFC5FB60F99A4E45524F" ID-CoL="9HZSD" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[1154,1391,1569,1595]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
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<emphasis id="B95F2D73FFE0FFC5FB60F99A4E45524F" box="[1154,1391,1569,1595]" italics="true" pageId="3" pageNumber="404">Copestylum enriquei</emphasis>
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</taxonomicName>
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<emphasis id="B95F2D73FFE0FFC5FA99F99A4BE2522A" bold="true" pageId="3" pageNumber="404">
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<taxonomicNameLabel id="A26C9008FFE0FFC5FA99F99A4BE2522A" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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/
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<bibRefCitation id="EFBA8C90FFE0FFC5FF3FF9FE482E522A" author="Parada-Marin, H. M. & Montoya, A. L. & Ramos-Pastrana, Y." box="[221,772,1605,1631]" pageId="3" pageNumber="404" pagination="162 - 165" refId="ref12544" refString="Parada-Marin, H. M., Montoya, A. L. & Ramos-Pastrana, Y. (2021) New record of Cepa apeca (Diptera, Syrphidae, Eristalinae, Merodontini) in the Andean-Amazonian region of Colombia and expansion of its geographic range. Acta Amazonica, 51, 162 - 165. https: // doi. org / 10.1590 / 1809 - 4392202003841" type="journal article" year="2021">Montoya, Parada-Marín & Ramos-Pastrana 2021</bibRefCitation>
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” [red, handwritten except first line] (
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–00000035890, dissected)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B43FB3CFFE0FFC5FEFBF9D34801529A" ID-GBIF-Occurrence="3436730331" collectingDate="1963-05-28" collectingDateMax="2001-09-14" collectingDateMin="1963-05-28" collectorName="P. H. van Doesburg Jr. " country="Suriname" elevation="35" location="Mapane area" pageId="3" pageNumber="404" specimenCount="2" specimenCount-female="2" stateProvince="Para" typeStatus="paratype">
|
||
<emphasis id="B95F2D73FFE0FFC5FEFBF9D34AEE52F7" bold="true" box="[281,452,1640,1666]" pageId="3" pageNumber="404">
|
||
<typeStatus id="54904FC3FFE0FFC5FEFBF9D34A9552F7" box="[281,447,1640,1666]" pageId="3" pageNumber="404" type="paratype">PARATYPES</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="F33CB1F1FFE0FFC5FE2DF9D2494E52F6" box="[463,612,1641,1667]" name="Suriname" pageId="3" pageNumber="404">SURINAME</collectingCountry>
|
||
,
|
||
<collectingRegion id="49EF3F83FFE0FFC5FD96F9D2498352F6" box="[628,681,1641,1667]" country="Suriname" name="Para" pageId="3" pageNumber="404">Para</collectingRegion>
|
||
,
|
||
<location id="8EF4A7BAFFE0FFC5FD55F9D2486452F6" LSID="urn:lsid:plazi:treatment:03824077FFE0FFCEFF75FF654BF555A4:8EF4A7BAFFE0FFC5FD55F9D2486452F6" box="[695,846,1641,1667]" country="Suriname" name="Mapane area" pageId="3" pageNumber="404" stateProvince="Para">Mapane area</location>
|
||
, 5.466 666, -54.683333,
|
||
<quantity id="4CD35C84FFE0FFC5FB89F9D34F8052F6" box="[1131,1194,1640,1667]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="404" unit="m" value="35.0">
|
||
<elevation id="00061652FFE0FFC5FB89F9D34F8052F6" box="[1131,1194,1640,1667]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="404" unit="m" value="35.0">35 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF95D7A1FFE0FFC5FB58F9D24E0052F6" box="[1210,1322,1641,1667]" pageId="3" pageNumber="404" value="1963-05-28">
|
||
<collectingDate id="EFD12E49FFE0FFC5FB58F9D24E0052F6" box="[1210,1322,1641,1667]" pageId="3" pageNumber="404" value="1963-05-28">28.v.1963</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DE94B7FFE0FFC5FAD8F9D24A0D52D2" pageId="3" pageNumber="404">P.H. van Doesburg Jr.</collectorName>
|
||
, Leg. (RMNH-collection,
|
||
<specimenCount id="9D2D3AE8FFE0FFC5FDB2F936499D52DD" box="[592,695,1677,1704]" pageId="3" pageNumber="404" type="female">1 female</specimenCount>
|
||
);
|
||
<collectingRegion id="49EF3F83FFE0FFC5FD29F936487052D2" box="[715,858,1677,1703]" country="Suriname" name="Brokopondo" pageId="3" pageNumber="404">Brokopondo</collectingRegion>
|
||
, Brownsberg, 4.933 333, -55.166667,
|
||
<quantity id="4CD35C84FFE0FFC5FAE0F9364E6E52D2" box="[1282,1348,1677,1703]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="3" pageNumber="404" unit="m" value="189.0">
|
||
<elevation id="00061652FFE0FFC5FAE0F9364E6E52D2" box="[1282,1348,1677,1703]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="3" pageNumber="404" unit="m" value="189.0">189m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="26DE94B7FFE0FFC5FAB2F9364EB652D2" box="[1360,1436,1677,1703]" pageId="3" pageNumber="404">Nature</collectorName>
|
||
reserve, mainly with primary forest,
|
||
<date id="FF95D7A1FFE0FFC5FDA5F90A483252BE" box="[583,792,1713,1739]" pageId="3" pageNumber="404" value="2001-08-31" valueMax="2001-09-14" valueMin="2001-08-31">
|
||
<collectingDate id="EFD12E49FFE0FFC5FDA5F90A483252BE" box="[583,792,1713,1739]" pageId="3" pageNumber="404" value="2001-08-31" valueMax="2001-09-14" valueMin="2001-08-31">31.viii–14.ix.2001</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26DE94B7FFE0FFC5FCB9F90A48DB52BE" box="[859,1009,1713,1739]" pageId="3" pageNumber="404">A. Gangadin</collectorName>
|
||
,
|
||
<collectorName id="26DE94B7FFE0FFC5FBE3F90A4F1E52BE" box="[1025,1076,1713,1739]" pageId="3" pageNumber="404">Leg.</collectorName>
|
||
(RMNH-collection,
|
||
<specimenCount id="9D2D3AE8FFE0FFC5FAC4F90A4EBB52B9" box="[1318,1425,1713,1740]" pageId="3" pageNumber="404" type="female">1 female</specimenCount>
|
||
).
|
||
<collectorName id="26DE94B7FFE0FFC5FF75F96E4A2E529A" box="[151,260,1749,1775]" pageId="3" pageNumber="404">Identified</collectorName>
|
||
as
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FECBF96E4A87529A" ID-CoL="62N9P" authorityName="Macquart" authorityYear="1846" box="[297,429,1749,1775]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F2D73FFE0FFC5FECBF96E4A87529A" box="[297,429,1749,1775]" italics="true" pageId="3" pageNumber="404">Copestylum</emphasis>
|
||
</taxonomicName>
|
||
SUR-04 by
|
||
<collectorName id="26DE94B7FFE0FFC5FDD8F96E49A3529A" box="[570,649,1749,1775]" pageId="3" pageNumber="404">Menno</collectorName>
|
||
<bibRefCitation id="EFBA8C90FFE0FFC5FD73F96E4801529A" author="Reemer, M." box="[657,811,1749,1775]" pageId="3" pageNumber="404" pagination="97 - 142" refId="ref12659" refString="Reemer, M. (2016) Syrphidae (Diptera) of Surinam: Eristalinae and synthesis. Tijdschrift voor Entomologie, 159, 97 - 142. https: // doi. org / 10.1163 / 22119434 - 15902002" type="journal article" year="2016">Reemer 2016</bibRefCitation>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B94F161FFE0FFC5FF25F94349FE5366" blockId="3.[151,1437,1496,2027]" box="[199,724,1784,1811]" pageId="3" pageNumber="404">
|
||
<emphasis id="B95F2D73FFE0FFC5FF25F9434A335366" bold="true" box="[199,281,1784,1811]" italics="true" pageId="3" pageNumber="404">Length</emphasis>
|
||
(n= 1). Body,
|
||
<quantity id="4CD35C84FFE0FFC5FE5FF942493C5366" box="[445,534,1785,1811]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="404" unit="mm" value="6.8">6.8 mm</quantity>
|
||
; Wings,
|
||
<quantity id="4CD35C84FFE0FFC5FD95F94349FA5366" box="[631,720,1784,1811]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="404" unit="mm" value="5.5">5.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331A2EAFFE0FFC2FF25F8A74AA354A0" lastPageId="4" lastPageNumber="405" pageId="3" pageNumber="404" type="diagnosis">
|
||
<paragraph id="8B94F161FFE0FFC2FF25F8A74AA354A0" blockId="3.[151,1437,1496,2027]" lastBlockId="4.[151,1437,151,969]" lastPageId="4" lastPageNumber="405" pageId="3" pageNumber="404">
|
||
<emphasis id="B95F2D73FFE0FFC5FF25F8A74AF85343" bold="true" box="[199,466,1820,1846]" pageId="3" pageNumber="404">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FE3EF8A649E25343" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[476,712,1821,1847]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE0FFC5FE3EF8A649E25343" box="[476,712,1821,1847]" italics="true" pageId="3" pageNumber="404">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE0FFC5FD31F8A6481B5342" bold="true" box="[723,817,1821,1847]" pageId="3" pageNumber="404">
|
||
<taxonomicNameLabel id="A26C9008FFE0FFC5FD31F8A6481B5342" box="[723,817,1821,1847]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
shares the general morphological characters of other known species in the
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FE6BF8FA4ADE532F" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[393,500,1857,1883]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="vagum">
|
||
<emphasis id="B95F2D73FFE0FFC5FE6BF8FA4ADE532F" box="[393,500,1857,1883]" italics="true" pageId="3" pageNumber="404">C. vagum</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FDA8F8FA4819532F" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[586,819,1857,1883]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE0FFC5FDA8F8FA4819532F" box="[586,819,1857,1883]" italics="true" pageId="3" pageNumber="404">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE0FFC5FCD8F8FA48BE532E" bold="true" box="[826,916,1857,1883]" pageId="3" pageNumber="404">
|
||
<taxonomicNameLabel id="A26C9008FFE0FFC5FCD8F8FA48BE532E" box="[826,916,1857,1883]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a green yellowish fly, similar in appearance to
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FF5AF8DE4A0D530B" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[184,295,1893,1919]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="vagum">
|
||
<emphasis id="B95F2D73FFE0FFC5FF5AF8DE4A0D530B" box="[184,295,1893,1919]" italics="true" pageId="3" pageNumber="404">C. vagum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FEDAF8DE4AF6530B" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[312,476,1893,1919]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="musicanum">
|
||
<emphasis id="B95F2D73FFE0FFC5FEDAF8DE4AF6530B" box="[312,476,1893,1919]" italics="true" pageId="3" pageNumber="404">C. musicanum</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2B8AE2FFE0FFC5FDC3F8DE4980530B" authorityName="Ricarte & Rotheray" authorityYear="2015" box="[545,682,1893,1919]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="tenorium">
|
||
<emphasis id="B95F2D73FFE0FFC5FDC3F8DE4980530B" box="[545,682,1893,1919]" italics="true" pageId="3" pageNumber="404">C. tenorium</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs by the gena and face separated by a very broad brown vitta; scutum orange except for the wide medial vitta, which is dark and metallic, ending before the prescutellar region, with the apical margin M-shaped; tibiae dark-brown, except yellow on basal 1/4. Epandrium and cercus black, contrasting with the colour of hypandrium and surstylus, which are orange; epandrium with a dorsal extension, in addition to the surstylus L-shaped, with two pairs of rounded ridges in the dorsal edge, similar to small deer antlers in velvet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331A2EAFFE7FFC3FF25FF654954548C" lastPageId="5" lastPageNumber="406" pageId="4" pageNumber="405" type="description">
|
||
<paragraph id="8B94F161FFE7FFC2FF25FF654A0757BC" blockId="4.[151,1437,151,969]" pageId="4" pageNumber="405">
|
||
<emphasis id="B95F2D73FFE7FFC2FF25FF6549A0548C" bold="true" box="[199,650,222,249]" pageId="4" pageNumber="405">
|
||
Description (
|
||
<typeStatus id="54904FC3FFE7FFC2FE81FF654AE5548D" box="[355,463,222,248]" pageId="4" pageNumber="405" type="holotype">holotype</typeStatus>
|
||
). MALE. Head
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1310EDE4FFE7FFC2FD7EFF644835548C" box="[668,799,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="4" pageNumber="405">Figs 1A, C</figureCitation>
|
||
, and D): Face yellowish-orange without a clear central vitta; gena and face separated by a very broad brown vitta; gena orange, orange pruinose, and pilose; dorsal margin of eyes sloping and yellow pilose; lunule brownish-orange; antenna orange, scape and pedicel orange setulose; arista orange, orange pilose.
|
||
<emphasis id="B95F2D73FFE7FFC2FE7EFEF04ADF5510" bold="true" box="[412,501,331,357]" pageId="4" pageNumber="405">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="1310EDE4FFE7FFC2FDE7FEF049AD5510" box="[517,647,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="4" pageNumber="405">Figs 1B–D</figureCitation>
|
||
): Scutum orange except for the wide medial vitta, which is dark and metallic, ending before the prescutellar region, with the apical margin M-shaped (
|
||
<figureCitation id="1310EDE4FFE7FFC2FBC7FED44F5155FC" box="[1061,1147,367,393]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="4" pageNumber="405">Fig. 1C</figureCitation>
|
||
); scutum brown pilose on the median dark area, yellow pilose along lateral margins, with pile not much longer at the rear margin; lateral setae orange: two above wing insertion and two on postalar callus; pleuron extensively yellow; scutellum yellow with pre-apical depression, yellow pilose; marginal setae orange-yellow, with two dark brown setae.
|
||
<emphasis id="B95F2D73FFE7FFC2FB2EFE614E275581" bold="true" box="[1228,1293,474,500]" pageId="4" pageNumber="405">Wing</emphasis>
|
||
(
|
||
<figureCitation id="1310EDE4FFE7FFC2FAFEFE604EB25580" box="[1308,1432,475,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="4" pageNumber="405">Figs 1B–E</figureCitation>
|
||
,
|
||
<figureCitation id="1310EDE4FFE7FFC2FF75FE444B96566C" box="[151,188,511,537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1877]" captionTargetBox="[175,1411,993,1828]" captionTargetId="figure-379@4.[175,1411,993,1828]" captionTargetPageId="4" captionText="FIGURE 2. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890). A. Abdomen, lateral view; B. Abdomen, ventral view; C. Whole genitalia including epandrium, cercus, and surstylus, lateral view, detail of the dorsal extension of the epandrium (white arrow); D. Epandrium, dorsal view, detail of the dorsal extension (white arrow); E. Hypandrium, ventral view. Abbreviations used in male genitalia structures are as follows: Ah = Aedeagal hood; Cer = Cercus; Epd = Epandrium; Hyp = Hypandrium; Sa= Surstylar apodeme; Sl= Superior lobe; Sur = Surstyle." figureDoi="http://doi.org/10.5281/zenodo.5848273" httpUri="https://zenodo.org/record/5848273/files/figure.png" pageId="4" pageNumber="405">2B</figureCitation>
|
||
): Hyaline except for the brown-yellowish pterostigmal spot, costal cell yellowish, bare; wing microtrichose, except for some extensively bare areas basally and centrally, cell bm extensively bare, cell dm bare on anterior 1/3, cell r only slightly bare on basal 1/3 or less, cell CuP bare on anterior 1/3; alula microtrichose, calypter yellowishwhite, border brownish-orange, fringe brownish-orange; plumula yellow-orange; halter yellow-orange, capitulum white.
|
||
<emphasis id="B95F2D73FFE7FFC2FF00FD344A3D56DD" box="[226,279,655,680]" italics="true" pageId="4" pageNumber="405">Legs</emphasis>
|
||
(
|
||
<figureCitation id="1310EDE4FFE7FFC2FEC7FD344ACF56DC" box="[293,485,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="4" pageNumber="405">Figs 1A–B, D–E</figureCitation>
|
||
): coxa and trochanter yellow, metafemur extensively orange; tibia dark-brown, except yellow on basal 1/4, black pilose on these areas; tarsomeres 1 and 2 orange, tarsomeres 3–5 black, black pilose.
|
||
<emphasis id="B95F2D73FFE7FFC2FF75FD6D4A265685" bold="true" box="[151,268,726,752]" pageId="4" pageNumber="405">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1310EDE4FFE7FFC2FEFEFD6C4A965684" box="[284,444,727,753]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="4" pageNumber="405">Figs 1B, D–E</figureCitation>
|
||
,
|
||
<figureCitation id="1310EDE4FFE7FFC2FE2BFD6C493B5684" box="[457,529,727,753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1877]" captionTargetBox="[175,1411,993,1828]" captionTargetId="figure-379@4.[175,1411,993,1828]" captionTargetPageId="4" captionText="FIGURE 2. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890). A. Abdomen, lateral view; B. Abdomen, ventral view; C. Whole genitalia including epandrium, cercus, and surstylus, lateral view, detail of the dorsal extension of the epandrium (white arrow); D. Epandrium, dorsal view, detail of the dorsal extension (white arrow); E. Hypandrium, ventral view. Abbreviations used in male genitalia structures are as follows: Ah = Aedeagal hood; Cer = Cercus; Epd = Epandrium; Hyp = Hypandrium; Sa= Surstylar apodeme; Sl= Superior lobe; Sur = Surstyle." figureDoi="http://doi.org/10.5281/zenodo.5848273" httpUri="https://zenodo.org/record/5848273/files/figure.png" pageId="4" pageNumber="405">2A–B</figureCitation>
|
||
): Mostly yellow-greenish, orange-yellow pilose, first tergum 1 yellow-greenish; terga 2–4 greenish, brownish infuscated apically; terga 3–4 black pilose on apical margin; S1–4 yellowish-green and extensively yellow pilose; male genitalia elongated, extending over the fourth sternum, with epandrium and cercus black, epandrium in lateral view with a dorsal extension, cercus in lateral view, globular and round-tipped; surstylus in lateral view, L-shaped, with two pairs of rounded ridges in the dorsal edge, similar to small deer antlers in velvet; superior lobe (Apex of hypandrium) orange, hook-shaped, with a kinked apex strongly curved downward (
|
||
<figureCitation id="1310EDE4FFE7FFC2FF7DFC144A0B57BC" box="[159,289,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1877]" captionTargetBox="[175,1411,993,1828]" captionTargetId="figure-379@4.[175,1411,993,1828]" captionTargetPageId="4" captionText="FIGURE 2. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890). A. Abdomen, lateral view; B. Abdomen, ventral view; C. Whole genitalia including epandrium, cercus, and surstylus, lateral view, detail of the dorsal extension of the epandrium (white arrow); D. Epandrium, dorsal view, detail of the dorsal extension (white arrow); E. Hypandrium, ventral view. Abbreviations used in male genitalia structures are as follows: Ah = Aedeagal hood; Cer = Cercus; Epd = Epandrium; Hyp = Hypandrium; Sa= Surstylar apodeme; Sl= Superior lobe; Sur = Surstyle." figureDoi="http://doi.org/10.5281/zenodo.5848273" httpUri="https://zenodo.org/record/5848273/files/figure.png" pageId="4" pageNumber="405">Figs 2D–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B94F161FFE6FFC3FF25FF2D4954548C" blockId="5.[151,1437,150,249]" pageId="5" pageNumber="406">
|
||
<emphasis id="B95F2D73FFE6FFC3FF25FF2D4A1554C5" bold="true" box="[199,319,150,176]" pageId="5" pageNumber="406">FEMALE</emphasis>
|
||
(
|
||
<figureCitation id="1310EDE4FFE6FFC3FEB2FF2C4AF854C4" box="[336,466,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1328,1352]" captionTargetBox="[169,1428,273,1305]" captionTargetId="figure-61@5.[158,1428,273,1305]" captionTargetPageId="5" captionText="FIGURE 3. Copestylum enriquei sp. nov., female, paratype (RMNH-collection). A. Head, frontal view; B. Head, lateral view, C. Habitus, lateral view; D. Thorax and abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5848275" httpUri="https://zenodo.org/record/5848275/files/figure.png" pageId="5" pageNumber="406">Figs 3A–D</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFBA8C90FFE6FFC3FE02FF2C499E54C4" author="Reemer, M." box="[480,692,151,177]" pageId="5" pageNumber="406" pagination="97 - 142" refId="ref12659" refString="Reemer, M. (2016) Syrphidae (Diptera) of Surinam: Eristalinae and synthesis. Tijdschrift voor Entomologie, 159, 97 - 142. https: // doi. org / 10.1163 / 22119434 - 15902002" type="journal article" year="2016">Reemer 2016: 100</bibRefCitation>
|
||
, Fig. 15, dorsal habitus). Similar to male except for usual sexual dimorphism, and differing in vertex with a complete brownish transverse macula, extensively dark brown to blackhaired in this area; sternites 4 and 5 yellow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331A2EAFFE6FFC3FF25FA214F1253FC" pageId="5" pageNumber="406" type="etymology">
|
||
<paragraph id="8B94F161FFE6FFC3FF25FA2148915230" blockId="5.[151,1437,1434,2037]" pageId="5" pageNumber="406">
|
||
<emphasis id="B95F2D73FFE6FFC3FF25FA214A6451C1" bold="true" box="[199,334,1434,1460]" pageId="5" pageNumber="406">Etymology.</emphasis>
|
||
The epithet ‘enriquei’ is a patronym given in memory of Carlos Enrique Giraldo, the grandfather of the first author, a notable, worthy, and noble person of sweet spirit who was like an oak, a very happy man who taught me (ALM) to enjoy and marvel at the small things of the natural world. For him, this beautiful species that flies high as his unforgettable smile and kind way of being, always ready to provide a space and welcome in his home in Anserma,
|
||
<collectingRegion id="49EF3F83FFE6FFC3FE89F9914A905231" box="[363,442,1578,1604]" country="Colombia" name="Caldas" pageId="5" pageNumber="406">Caldas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33CB1F1FFE6FFC3FE27F99149195231" box="[453,563,1578,1604]" name="Colombia" pageId="5" pageNumber="406">Colombia</collectingCountry>
|
||
that was always open to everyone.
|
||
</paragraph>
|
||
<paragraph id="8B94F161FFE6FFC3FF25F9F54F1253FC" blockId="5.[151,1437,1434,2037]" pageId="5" pageNumber="406">
|
||
<emphasis id="B95F2D73FFE6FFC3FF25F9F54A97521D" bold="true" box="[199,445,1614,1640]" pageId="5" pageNumber="406">Taxonomic remarks.</emphasis>
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FE21F9F54981521D" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[451,683,1614,1640]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE6FFC3FE21F9F54981521D" box="[451,683,1614,1640]" italics="true" pageId="5" pageNumber="406">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE6FFC3FD50F9F54821521D" bold="true" box="[690,779,1614,1640]" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel id="A26C9008FFE6FFC3FD50F9F54821521D" box="[690,779,1614,1640]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
run to
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FCB8F9F44F2F521D" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[858,1029,1614,1641]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="chapadensis">
|
||
<emphasis id="B95F2D73FFE6FFC3FCB8F9F44F2F521D" box="[858,1029,1614,1641]" italics="true" pageId="5" pageNumber="406">C. chapadensis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FBC8F9F44F9F521D" authorityName="Curran" authorityYear="1930" box="[1066,1205,1614,1641]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bequaerti">
|
||
<emphasis id="B95F2D73FFE6FFC3FBC8F9F44F9F521D" box="[1066,1205,1614,1641]" italics="true" pageId="5" pageNumber="406">C. bequaerti</emphasis>
|
||
</taxonomicName>
|
||
in the key of
|
||
<bibRefCitation id="EFBA8C90FFE6FFC3FAADF9F54BF152F9" author="Curran, C. H." pageId="5" pageNumber="406" pagination="1 - 23" refId="ref11428" refString="Curran, C. H. (1930) New species of Volucellinae from America (Syrphidae, Diptera). American Museum Novitates, 413, 1 - 23." type="journal article" year="1930">Curran (1930</bibRefCitation>
|
||
; couplet 9, p. 6), based on the gena and face separated by a very broad brownish vitta. This work covered most species belonging to the
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FE4EF92C493252C5" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[428,536,1687,1713]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="vagum">
|
||
<emphasis id="B95F2D73FFE6FFC3FE4EF92C493252C5" box="[428,536,1687,1713]" italics="true" pageId="5" pageNumber="406">C. vagum</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FD8CF92D487C52C5" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[622,854,1686,1712]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE6FFC3FD8CF92D487C52C5" box="[622,854,1686,1712]" italics="true" pageId="5" pageNumber="406">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE6FFC3FCBCF92D489D52C5" bold="true" box="[862,951,1686,1712]" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel id="A26C9008FFE6FFC3FCBCF92D489D52C5" box="[862,951,1686,1712]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
would be identified as
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FB5CF92C4E7452C5" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[1214,1374,1687,1713]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="musicanum">
|
||
<emphasis id="B95F2D73FFE6FFC3FB5CF92C4E7452C5" box="[1214,1374,1687,1713]" italics="true" pageId="5" pageNumber="406">C. musicanum</emphasis>
|
||
</taxonomicName>
|
||
in an unpublished draft key by F.C. Thompson (couplet 13), as it has the tibiae much darker than the femora, terminalia large, cercus elongate, hook-like, extending over the fourth sternum. Using the key of
|
||
<bibRefCitation id="EFBA8C90FFE6FFC3FB80F9654E14528C" author="Ricarte, A. & Marcos-Garcia, M. A. & Hancock, E. G. & Rotheray, G. E." box="[1122,1342,1758,1785]" pageId="5" pageNumber="406" pagination="0142441" refId="ref12857" refString="Ricarte, A., Marcos-Garcia, M. A., Hancock, E. G. & Rotheray, G. E. (2015) Neotropical Copestylum Macquart (Diptera: Syrphidae) breeding in fruits and flowers, including 7 new species. PLOS ONE, 10 (11), e 0142441. https: // doi. org / 10.1371 / journal. pone. 0142441" type="journal article" year="2015">
|
||
Ricarte
|
||
<emphasis id="B95F2D73FFE6FFC3FB58F9644FC1528D" box="[1210,1259,1758,1784]" italics="true" pageId="5" pageNumber="406">et al</emphasis>
|
||
. (2015
|
||
</bibRefCitation>
|
||
; couplet 3, p. 52),
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FEE0F8B84AAA5369" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[258,384,1795,1821]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE6FFC3FEE0F8B84AAA5369" box="[258,384,1795,1821]" italics="true" pageId="5" pageNumber="406">C. enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE6FFC3FE65F8B94ACB5369" bold="true" box="[391,481,1794,1820]" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel id="A26C9008FFE6FFC3FE65F8B94ACB5369" box="[391,481,1794,1820]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
keys out close to
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FD48F8B8486C5369" authorityName="Ricarte & Rotheray" authorityYear="2015" box="[682,838,1795,1821]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="araceorum">
|
||
<emphasis id="B95F2D73FFE6FFC3FD48F8B8486C5369" box="[682,838,1795,1821]" italics="true" pageId="5" pageNumber="406">C. araceorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FC9EF8B84F2F5369" authorityName="Ricarte & Hancock" authorityYear="2015" box="[892,1029,1794,1821]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="willistoni">
|
||
<emphasis id="B95F2D73FFE6FFC3FC9EF8B84F2F5369" box="[892,1029,1794,1821]" italics="true" pageId="5" pageNumber="406">C. willistoni</emphasis>
|
||
</taxonomicName>
|
||
, based on the terga 3−4 with yellow marks or completely black, however,
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FDD9F89C49925335" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[571,696,1831,1857]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE6FFC3FDD9F89C49925335" box="[571,696,1831,1857]" italics="true" pageId="5" pageNumber="406">C. enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE6FFC3FD5DF89D48325335" bold="true" box="[703,792,1830,1856]" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel id="A26C9008FFE6FFC3FD5DF89D48325335" box="[703,792,1830,1856]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
lacks a median vitta on the face.
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FB6EF89D4E5E5335" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[1164,1396,1830,1856]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFE6FFC3FB6EF89D4E5E5335" box="[1164,1396,1830,1856]" italics="true" pageId="5" pageNumber="406">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFE6FFC3FA99F89D4BE25311" bold="true" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel id="A26C9008FFE6FFC3FA99F89D4BE25311" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also morphologically similar to
|
||
<taxonomicName id="4C2B8AE2FFE6FFC3FDA9F8F049FA5311" authorityName="Ricarte & Rotheray" authorityYear="2015" box="[587,720,1867,1893]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="tenorium">
|
||
<emphasis id="B95F2D73FFE6FFC3FDA9F8F049FA5311" box="[587,720,1867,1893]" italics="true" pageId="5" pageNumber="406">C. tenorium</emphasis>
|
||
</taxonomicName>
|
||
from which it differs by the diagnostic characters listed in
|
||
<tableCitation id="C6A9C4DAFFE6FFC3FABCF8F14B8F53FD" captionStart="TABLE 1" captionStartId="6.[151,244,464,491]" captionTargetPageId="6" captionText="TABLE 1. Morphological differences between males of Copestylum enriquei sp. nov. and similar species of the Copestylum vagum group." httpUri="http://table.plazi.org/id/DF54A1E9FFE5FFC0FF75FE6B4D81559E" pageId="5" pageNumber="406" tableUuid="DF54A1E9FFE5FFC0FF75FE6B4D81559E">Table 1</tableCitation>
|
||
(See also “Taxonomic notes” under each species in
|
||
<bibRefCitation id="EFBA8C90FFE6FFC3FD0CF8D5489753FC" author="Ricarte, A. & Marcos-Garcia, M. A. & Hancock, E. G. & Rotheray, G. E." box="[750,957,1902,1929]" pageId="5" pageNumber="406" pagination="0142441" refId="ref12857" refString="Ricarte, A., Marcos-Garcia, M. A., Hancock, E. G. & Rotheray, G. E. (2015) Neotropical Copestylum Macquart (Diptera: Syrphidae) breeding in fruits and flowers, including 7 new species. PLOS ONE, 10 (11), e 0142441. https: // doi. org / 10.1371 / journal. pone. 0142441" type="journal article" year="2015">
|
||
Ricarte
|
||
<emphasis id="B95F2D73FFE6FFC3FCA4F8D4485D53FD" box="[838,887,1902,1928]" italics="true" pageId="5" pageNumber="406">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
and keys).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331A2EAFFE6FFCEFF25F8294A6A5534" lastPageId="8" lastPageNumber="409" pageId="5" pageNumber="406" type="distribution">
|
||
<paragraph id="8B94F161FFE6FFCEFF25F8294A6A5534" blockId="5.[151,1437,1434,2037]" lastBlockId="8.[151,1437,151,465]" lastPageId="8" lastPageNumber="409" pageId="5" pageNumber="406">
|
||
<emphasis id="B95F2D73FFE6FFC3FF25F8294A4B53D9" bold="true" box="[199,353,1938,1964]" pageId="5" pageNumber="406">Distribution.</emphasis>
|
||
The male
|
||
<typeStatus id="54904FC3FFE6FFC3FE35F829491353D9" box="[471,569,1938,1964]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
|
||
was collected between
|
||
<date id="FF95D7A1FFE6FFC3FCA0F8294F3D53D9" box="[834,1047,1938,1964]" pageId="5" pageNumber="406" value="2016-12-21">21 December 2016</date>
|
||
and
|
||
<date id="FF95D7A1FFE6FFC3FBAFF8294E3E53D9" box="[1101,1300,1938,1964]" pageId="5" pageNumber="406" value="2017-02-04">
|
||
04 February
|
||
<quantity id="4CD35C84FFE6FFC3FB3EF8294E3E53D9" box="[1244,1300,1938,1964]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="5" pageNumber="406" unit="in" value="2017.0">2017</quantity>
|
||
</date>
|
||
in a canopy Malaise trap (Rafael & Gorayeb 1982) in a pristine rainforest patch of a conflict territory of limited access and political unrest in
|
||
<collectingRegion id="49EF3F83FFE6FFC3FEE3F8614A4A5381" box="[257,352,2010,2036]" country="Colombia" name="Caqueta" pageId="5" pageNumber="406">Caquetá</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33CB1F1FFE6FFC3FE8EF8614AF05381" box="[364,474,2010,2036]" name="Colombia" pageId="5" pageNumber="406">Colombia</collectingCountry>
|
||
(
|
||
<figureCitation id="1310EDE4FFE6FFC3FE08F861496C5381" box="[490,582,2010,2036]" captionStart="FIGURE 4" captionStartId="8.[151,250,1019,1043]" captionTargetBox="[153,1436,490,995]" captionTargetId="figure-153@8.[152,1436,489,996]" captionTargetPageId="8" captionText="FIGURE 4. A. Habitat of Copestylum enriquei sp. nov. in a pristine forest at Florencia, Caquetá in Colombia. The specimens were collected using a canopy Malaise trap (Rafael & Gorayeb 1982); B. Geographical distribution map showing the collection localities of Copestylum enriquei sp. nov. in the Amazon rainforest of Colombia and Suriname (Red triangle)." figureDoi="http://doi.org/10.5281/zenodo.5848277" httpUri="https://zenodo.org/record/5848277/files/figure.png" pageId="5" pageNumber="406">Fig. 4A</figureCitation>
|
||
). The type locality is located at the eastern slope of the Colombian Oriental Cordillera at an altitude of 716 meters above sea level (m a.s.l). Two additional female specimens (
|
||
<typeStatus id="54904FC3FFEBFFCEFAC4FF2C4EBF54C4" box="[1318,1429,151,177]" pageId="8" pageNumber="409" type="paratype">paratypes</typeStatus>
|
||
) were collected in two sampling events on
|
||
<date id="FF95D7A1FFEBFFCEFD8FFF00482854A0" box="[621,770,187,213]" pageId="8" pageNumber="409" value="1963-05-28">
|
||
<collectingDate id="EFD12E49FFEBFFCEFD8FFF00482854A0" box="[621,770,187,213]" pageId="8" pageNumber="409" value="1963-05-28">28 May 1963</collectingDate>
|
||
</date>
|
||
and between
|
||
<date id="FF95D7A1FFEBFFCEFC7FFF004E3554A0" box="[925,1311,187,213]" pageId="8" pageNumber="409" value="2001-08-31" valueMax="2001-09-14" valueMin="2001-08-31">
|
||
<collectingDate id="EFD12E49FFEBFFCEFC7FFF004E3554A0" box="[925,1311,187,213]" pageId="8" pageNumber="409" value="2001-08-31" valueMax="2001-09-14" valueMin="2001-08-31">
|
||
31 August and 14 September
|
||
<quantity id="4CD35C84FFEBFFCEFB05FF004E3554A0" box="[1255,1311,187,213]" metricMagnitude="1" metricUnit="m" metricValue="5.08254" pageId="8" pageNumber="409" unit="in" value="2001.0">2001</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
in a nature reserve and primary forest in
|
||
<collectingMunicipality id="6BF06B1BFFEBFFCEFE3DFF64495A548C" box="[479,624,223,249]" pageId="8" pageNumber="409">Northeastern</collectingMunicipality>
|
||
<collectingCountry id="F33CB1F1FFEBFFCEFD94FF6449CB548C" box="[630,737,223,249]" name="Suriname" pageId="8" pageNumber="409">Suriname</collectingCountry>
|
||
(
|
||
<collectingRegion id="49EF3F83FFEBFFCEFD0DFF6448CC548F" box="[751,998,223,250]" isEnumeration="true" pageId="8" pageNumber="409">Para and Brokopondo</collectingRegion>
|
||
) between 35 and
|
||
<elevation id="00061652FFEBFFCEFB4AFF644E03548C" box="[1192,1321,223,249]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="8" pageNumber="409" unit="m" value="189.0">
|
||
<quantity id="4CD35C84FFEBFFCEFB4AFF644FC5548C" box="[1192,1263,223,249]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="8" pageNumber="409" unit="m" value="189.0">189 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(
|
||
<figureCitation id="1310EDE4FFEBFFCEFADAFF644EA5548C" box="[1336,1423,223,249]" captionStart="FIGURE 4" captionStartId="8.[151,250,1019,1043]" captionTargetBox="[153,1436,490,995]" captionTargetId="figure-153@8.[152,1436,489,996]" captionTargetPageId="8" captionText="FIGURE 4. A. Habitat of Copestylum enriquei sp. nov. in a pristine forest at Florencia, Caquetá in Colombia. The specimens were collected using a canopy Malaise trap (Rafael & Gorayeb 1982); B. Geographical distribution map showing the collection localities of Copestylum enriquei sp. nov. in the Amazon rainforest of Colombia and Suriname (Red triangle)." figureDoi="http://doi.org/10.5281/zenodo.5848277" httpUri="https://zenodo.org/record/5848277/files/figure.png" pageId="8" pageNumber="409">Fig. 4B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C2B8AE2FFEBFFCEFF75FEB84AAA5569" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[151,384,259,285]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFEBFFCEFF75FEB84AAA5569" box="[151,384,259,285]" italics="true" pageId="8" pageNumber="409">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFEBFFCEFE6BFEB84ACE5568" bold="true" box="[393,484,259,285]" pageId="8" pageNumber="409">
|
||
<taxonomicNameLabel id="A26C9008FFEBFFCEFE6BFEB84ACE5568" box="[393,484,259,285]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is probably endemic to the
|
||
<location id="8EF4A7BAFFEBFFCEFCC2FEB8488B5568" LSID="urn:lsid:plazi:treatment:03824077FFE0FFCEFF75FF654BF555A4:8EF4A7BAFFEBFFCEFCC2FEB8488B5568" box="[800,929,259,285]" country="Suriname" municipality="Northeastern" name="Amazonian" pageId="8" pageNumber="409" stateProvince="Para and Brokopondo">Amazonian</location>
|
||
rainforest of
|
||
<collectingCountry id="F33CB1F1FFEBFFCEFBD9FEB84F835568" box="[1083,1193,259,285]" name="Colombia" pageId="8" pageNumber="409">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33CB1F1FFEBFFCEFB01FEB84E645568" box="[1251,1358,259,285]" name="Suriname" pageId="8" pageNumber="409">Suriname</collectingCountry>
|
||
on the
|
||
<collectingCountry id="F33CB1F1FFEBFFCEFF75FE9C4BC45534" box="[151,238,295,321]" name="Guyana" pageId="8" pageNumber="409">Guyana</collectingCountry>
|
||
shield.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331A2EAFFEBFFCEFF25FEF14BF555A4" pageId="8" pageNumber="409" type="biology_ecology">
|
||
<paragraph id="8B94F161FFEBFFCEFF25FEF14BF555A4" blockId="8.[151,1437,151,465]" pageId="8" pageNumber="409">
|
||
<emphasis id="B95F2D73FFEBFFCEFF25FEF14A025510" bold="true" box="[199,296,330,357]" pageId="8" pageNumber="409">Biology.</emphasis>
|
||
<taxonomicName id="4C2B8AE2FFEBFFCEFED2FEF049335511" authority="Montoya & Parada-Marín & Ramos-Pastrana, 2022" authorityName="Montoya & Parada-Marín & Ramos-Pastrana" authorityYear="2022" box="[304,537,331,357]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="B95F2D73FFEBFFCEFED2FEF049335511" box="[304,537,331,357]" italics="true" pageId="8" pageNumber="409">Copestylum enriquei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F2D73FFEBFFCEFDC3FEF049515510" bold="true" box="[545,635,331,357]" pageId="8" pageNumber="409">
|
||
<taxonomicNameLabel id="A26C9008FFEBFFCEFDC3FEF049515510" box="[545,635,331,357]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is only known from pristine patches in the lowlands of the Amazonian rainforest of
|
||
<collectingCountry id="F33CB1F1FFEBFFCEFECFFED44AB155FC" box="[301,411,367,393]" name="Colombia" pageId="8" pageNumber="409">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33CB1F1FFEBFFCEFE35FED4496855FC" box="[471,578,367,393]" name="Suriname" pageId="8" pageNumber="409">Suriname</collectingCountry>
|
||
(
|
||
<figureCitation id="1310EDE4FFEBFFCEFDB6FED4498555FC" box="[596,687,367,393]" captionStart="FIGURE 4" captionStartId="8.[151,250,1019,1043]" captionTargetBox="[153,1436,490,995]" captionTargetId="figure-153@8.[152,1436,489,996]" captionTargetPageId="8" captionText="FIGURE 4. A. Habitat of Copestylum enriquei sp. nov. in a pristine forest at Florencia, Caquetá in Colombia. The specimens were collected using a canopy Malaise trap (Rafael & Gorayeb 1982); B. Geographical distribution map showing the collection localities of Copestylum enriquei sp. nov. in the Amazon rainforest of Colombia and Suriname (Red triangle)." figureDoi="http://doi.org/10.5281/zenodo.5848277" httpUri="https://zenodo.org/record/5848277/files/figure.png" pageId="8" pageNumber="409">Fig. 4B</figureCitation>
|
||
). The specimens were collected using a canopy Malaise trap and sweeping net, which could suggest that the species is associated with both canopy and ground levels in conserved forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |