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<mods:title id="836A3760276564D71957493121B8A83C">On the identity of Sundathelphusa philippina (von Martens, 1868) (Decapoda: Brachyura: Gecarcinucidae) from the Philippines, with descriptions of two new species</mods:title>
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<treatment id="C6560C5A6B29FF8FE7FEFB6AFDA0F8EE" ID-DOI="http://doi.org/10.5281/zenodo.5466959" ID-GBIF-Taxon="156192031" ID-Zenodo-Dep="5466959" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE" httpUri="http://treatment.plazi.org/id/C6560C5A6B29FF8FE7FEFB6AFDA0F8EE" lastPageId="6" lastPageNumber="321" pageId="1" pageNumber="316">
<subSubSection id="06E5EEC76B29FF88E7FEFB6AFD6FFB7C" box="[151,737,1223,1250]" pageId="1" pageNumber="316" type="nomenclature">
<paragraph id="4E40BD4C6B29FF88E7FEFB6AFD6FFB7C" blockId="1.[151,737,1223,1283]" box="[151,737,1223,1250]" pageId="1" pageNumber="316">
<heading id="15080A206B29FF88E7FEFB6AFD6FFB7C" bold="true" box="[151,737,1223,1250]" fontSize="11" level="1" pageId="1" pageNumber="316" reason="1">
<taxonomicName id="89FFC6CF6B29FF88E7FEFB6AFD6FFB7C" ID-CoL="53GC2" authority="(von Martens, 1868)" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[151,737,1223,1250]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B29FF88E7FEFB6AFD6FFB7C" bold="true" box="[151,737,1223,1250]" pageId="1" pageNumber="316">
<emphasis id="7C8B615E6B29FF88E7FEFB6AFE51FB7F" bold="true" box="[151,479,1223,1249]" italics="true" pageId="1" pageNumber="316">Sundathelphusa philippina</emphasis>
(von
<bibRefCitation id="2A6EC0BD6B29FF88E54CFB65FD57FB7C" author="Martens, E. von" box="[549,729,1224,1250]" pageId="1" pageNumber="316" pagination="608 - 615" refId="ref9072" refString="Martens, E. von (1868) Ueber einige neue crustaceen. Monatsberichte der Koniglichen Preussische Akademie der Akademie der Wissenschaften zu Berlin, Sitzung der Physikalisch-mathematische Klasse, 1868, 608 - 615." type="journal article" year="1868">Martens, 1868</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="06E5EEC76B29FF88E7FEFB47FDAAFA9D" box="[151,548,1258,1283]" pageId="1" pageNumber="316" type="description">
<paragraph id="4E40BD4C6B29FF88E7FEFB47FDAAFA9D" blockId="1.[151,737,1223,1283]" box="[151,548,1258,1283]" pageId="1" pageNumber="316">
(
<figureCitation id="D6C4A1C96B29FF88E7F6FB47FE85FA9D" box="[159,267,1258,1283]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1875,1897]" captionStartId-1="4.[151,250,1799,1821]" captionStartId-2="5.[151,250,1509,1531]" captionTargetBox-0="[151,1434,481,1854]" captionTargetBox-1="[198,1324,189,1779]" captionTargetBox-2="[150,1436,171,1487]" captionTargetId-0="figure@3.[151,1435,481,1854]" captionTargetId-1="figure@4.[192,1328,183,1779]" captionTargetId-2="figure@5.[150,1436,171,1488]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." captionText-1="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." captionText-2="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale." httpUri-0="https://zenodo.org/record/2637368/files/figure.png" httpUri-1="https://zenodo.org/record/2637370/files/figure.png" httpUri-2="https://zenodo.org/record/2637372/files/figure.png" pageId="1" pageNumber="316">Figs. 13</figureCitation>
;
<figureCitation id="D6C4A1C96B29FF88E67EFB47FE45FA9C" box="[279,459,1258,1283]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="1" pageNumber="316">6A, C, E, G, I, J</figureCitation>
;
<figureCitation id="D6C4A1C96B29FF88E6BEFB46FD94FA9D" box="[471,538,1259,1283]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: AF = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="1" pageNumber="316">9A, B</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="6EEF9A626B29FF88E7FEFA9FFC2AFAD6" box="[151,932,1330,1352]" pageId="1" pageNumber="316">
<taxonomicName id="89FFC6CF6B29FF88E7FEFA9FFDE4FAD6" authority="von Martens, 1868: 608" authorityName="von Martens" authorityPageNumber="608" authorityYear="1868" box="[151,618,1330,1352]" class="Malacostraca" family="Potamidae" genus="Thelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B29FF88E7FEFA9FFEE1FAD6" box="[151,367,1330,1352]" italics="true" pageId="1" pageNumber="316">Thelphusa philippina</emphasis>
von
<treatmentCitation id="CF5E9B5D6B29FF88E6C8FA9EFDE4FAD6" author="Martens, E. von" box="[417,618,1330,1352]" page="608" pageId="1" pageNumber="316" year="1868">
<bibRefCitation id="2A6EC0BD6B29FF88E6C8FA9EFDB9FAD6" author="Martens, E. von" box="[417,567,1330,1352]" pageId="1" pageNumber="316" pagination="608 - 615" refId="ref9072" refString="Martens, E. von (1868) Ueber einige neue crustaceen. Monatsberichte der Koniglichen Preussische Akademie der Akademie der Wissenschaften zu Berlin, Sitzung der Physikalisch-mathematische Klasse, 1868, 608 - 615." type="journal article" year="1868">Martens, 1868</bibRefCitation>
: 608
</treatmentCitation>
</taxonomicName>
; A. Milne-Edwards 1869: 168.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E40BD4C6B29FF88E7FEFAFFFCFDFAF9" blockId="1.[151,1435,1330,1536]" box="[151,883,1361,1383]" pageId="1" pageNumber="316">
<treatmentCitationGroup id="6EEF9A626B29FF88E7FEFAFFFCFDFAF9" box="[151,883,1361,1383]" pageId="1" pageNumber="316">
<taxonomicName id="89FFC6CF6B29FF88E7FEFAFFFD95FAF9" authority="Ortmann 1897" authorityName="Ortmann" authorityYear="1897" box="[151,539,1361,1383]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="philippinum" subGenus="Potamonautes">
<emphasis id="7C8B615E6B29FF88E7FEFAFFFF7DFAF9" box="[151,243,1362,1383]" italics="true" pageId="1" pageNumber="316">Potamon</emphasis>
(
<emphasis id="7C8B615E6B29FF88E668FAFFFE1CFAF9" box="[257,402,1362,1383]" italics="true" pageId="1" pageNumber="316">Potamonautes</emphasis>
)
<emphasis id="7C8B615E6B29FF88E6C8FAFCFD95FAF9" box="[417,539,1361,1383]" italics="true" pageId="1" pageNumber="316">philippinum</emphasis>
</taxonomicName>
<treatmentCitation id="CF5E9B5D6B29FF88E55CFAFCFD72FAF9" author="Ortmann, A. E." box="[565,764,1361,1383]" page="304" pageId="1" pageNumber="316" year="1897">
<bibRefCitation id="2A6EC0BD6B29FF88E55CFAFCFD44FAF9" author="Ortmann, A. E." box="[565,714,1361,1383]" pageId="1" pageNumber="316" pagination="258 - 372" refId="ref9729" refString="Ortmann, A. E. (1897) Carcinologische Studien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (3), 258 - 372, pl. 17." type="journal article" year="1897">Ortmann 1897</bibRefCitation>
: 304
</treatmentCitation>
, 307 (part).
</treatmentCitationGroup>
</paragraph>
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<taxonomicName id="89FFC6CF6B29FF88E7FEFADCFEFDFA18" box="[151,371,1392,1414]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="philippinum">
<emphasis id="7C8B615E6B29FF88E7FEFADCFEFDFA18" box="[151,371,1392,1414]" italics="true" pageId="1" pageNumber="316">Potamon philippinum</emphasis>
</taxonomicName>
—De Man 1898: 37 (part); 1902: 558.
</paragraph>
<paragraph id="4E40BD4C6B29FF88E7FEFA3DFD70FA3A" blockId="1.[151,1435,1330,1536]" box="[151,766,1422,1445]" pageId="1" pageNumber="316">
<treatmentCitationGroup id="6EEF9A626B29FF88E7FEFA3DFD70FA3A" box="[151,766,1422,1445]" pageId="1" pageNumber="316">
<taxonomicName id="89FFC6CF6B29FF88E7FEFA3DFE53FA3A" box="[151,477,1422,1445]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="philippinus" subGenus="Potamon">
<emphasis id="7C8B615E6B29FF88E7FEFA3DFF7DFA3B" box="[151,243,1424,1445]" italics="true" pageId="1" pageNumber="316">Potamon</emphasis>
(
<emphasis id="7C8B615E6B29FF88E668FA3DFED2FA3B" box="[257,348,1424,1445]" italics="true" pageId="1" pageNumber="316">Potamon</emphasis>
)
<emphasis id="7C8B615E6B29FF88E602FA23FE53FA3A" box="[363,477,1422,1444]" italics="true" pageId="1" pageNumber="316">philippinus</emphasis>
</taxonomicName>
<treatmentCitation id="CF5E9B5D6B29FF88E69FFA23FD36FA3A" author="Rathbun, M. J." box="[502,696,1422,1444]" page="304" pageId="1" pageNumber="316" year="1904">
<bibRefCitation id="2A6EC0BD6B29FF88E69FFA23FD06FA3A" author="Rathbun, M. J." box="[502,648,1422,1444]" pageId="1" pageNumber="316" pagination="225 - 312" refId="ref9767" refString="Rathbun, M. J. (1904) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, 4, 225 - 312, pls. 9 - 18." type="journal article" year="1904">Rathbun 1904</bibRefCitation>
: 304
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="4E40BD4C6B29FF88E7FEFA00FB7EFA7C" blockId="1.[151,1435,1330,1536]" pageId="1" pageNumber="316">
<treatmentCitationGroup id="6EEF9A626B29FF88E7FEFA00FB7EFA7C" pageId="1" pageNumber="316">
<taxonomicName id="89FFC6CF6B29FF88E7FEFA00FE26FA5D" ID-CoL="53GC2" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[151,424,1453,1475]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B29FF88E7FEFA00FE26FA5D" box="[151,424,1453,1475]" italics="true" pageId="1" pageNumber="316">Sundathelphusa philippina</emphasis>
</taxonomicName>
-
<treatmentCitation id="CF5E9B5D6B29FF88E6D7FA03FDC0FA5D" author="Bott, R." box="[446,590,1453,1475]" page="78" pageId="1" pageNumber="316" year="1970">
<bibRefCitation id="2A6EC0BD6B29FF88E6D7FA03FDA6FA5D" author="Bott, R." box="[446,552,1453,1475]" pageId="1" pageNumber="316" pagination="1 - 338" refId="ref8072" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Abhandlungen der Senckenbergischen Naturforschenden Gesselschaft, 526, 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott 1970</bibRefCitation>
: 78
</treatmentCitation>
, pl. 28 fig. 57, pl. 32 figs. 13;
<treatmentCitation id="CF5E9B5D6B29FF88E4FAFA00FBDDFA5D" author="Cabrera, B. D." box="[915,1107,1453,1475]" page="602" pageId="1" pageNumber="316" year="1975">
<bibRefCitation id="2A6EC0BD6B29FF88E4FAFA00FB91FA5D" author="Cabrera, B. D." box="[915,1055,1453,1475]" pageId="1" pageNumber="316" pagination="602" refId="ref8149" refString="Cabrera, B. D. (1975) Sundathelphusa philippina (Martens 1868) the new name of the crustacean intermediate host of mammalian lung fluke (Paragonimus) in the Philippines. Southeast Asian Journal of Tropical Medicine and Public Health, 6 (4), 602." type="journal article" year="1975">Cabrera 1975</bibRefCitation>
: 602
</treatmentCitation>
;
<treatmentCitation id="CF5E9B5D6B29FF88E337FA00FA9BFA5D" author="Takeda, M." box="[1118,1301,1453,1475]" page="101" pageId="1" pageNumber="316" year="1987">
<bibRefCitation id="2A6EC0BD6B29FF88E337FA00FB6BFA5D" author="Takeda, M." box="[1118,1253,1453,1475]" pageId="1" pageNumber="316" pagination="89 - 106" refId="ref9849" refString="Takeda, M. (1987) Freshwater crabs from Peninsu 1 ar Malaysia, with notes on some species from the Philippines. In: Kawashima, K. (Ed.), Paragonimus in Asia: Biology, Genetic Variation and Speciation. Paragonimus Research Report, Kyushu University, 1, pp. 89 - 106." type="journal article" year="1987">Takeda 1987</bibRefCitation>
: 101
</treatmentCitation>
(part), pl. 12 (top);
<treatmentCitation id="CF5E9B5D6B29FF88E66CFA60FE50FA7C" author="Ng, P. K. L. &amp; Sket, B." box="[261,478,1484,1506]" page="697" pageId="1" pageNumber="316" year="1996">
<bibRefCitation id="2A6EC0BD6B29FF88E66CFA60FE20FA7C" author="Ng, P. K. L. &amp; Sket, B." box="[261,430,1484,1506]" pageId="1" pageNumber="316" pagination="695 - 706" refId="ref9587" refString="Ng, P. K. L. &amp; Sket, B. (1996) The freshwater crab fauna (Crustacea: Decapoda: Brachyura) of the Philippines. IV. On the collection of Parathelphusidae from Bohol. Proceeding of the Biological Society of Washington, 109, 695 - 706." type="journal article" year="1996">Ng &amp; Sket 1996</bibRefCitation>
: 697
</treatmentCitation>
(in part);
<treatmentCitation id="CF5E9B5D6B29FF88E52DFA60FD75FA7C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[580,763,1484,1506]" page="73" pageId="1" pageNumber="316" year="2008">
<bibRefCitation id="2A6EC0BD6B29FF88E52DFA60FD57FA7C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[580,729,1484,1506]" pageId="1" pageNumber="316" pagination="1 - 286" refId="ref9531" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="7C8B615E6B29FF88E500FA60FD12FA7C" box="[617,668,1484,1506]" italics="true" pageId="1" pageNumber="316">et al.</emphasis>
2008
</bibRefCitation>
: 73
</treatmentCitation>
(list);
<treatmentCitation id="CF5E9B5D6B29FF88E457FA61FBEBFA7C" author="Mendoza, J. C. E. &amp; Naruse, T." box="[830,1125,1484,1506]" page="63" pageId="1" pageNumber="316" year="2010">
<bibRefCitation id="2A6EC0BD6B29FF88E457FA61FBCCFA7C" author="Mendoza, J. C. E. &amp; Naruse, T." box="[830,1090,1484,1506]" pageId="1" pageNumber="316" pagination="61 - 70" refId="ref9108" refString="Mendoza, J. C. E. &amp; Naruse, T. (2010) A new species of riverine crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from northeastern Luzon, Philippines. Philippine Journal of Science, 139 (1), 61 - 70." type="journal article" year="2010">Mendoza &amp; Naruse 2010</bibRefCitation>
: 63
</treatmentCitation>
(in part; list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="4E40BD4C6B29FF88E7FEFA47FBB0F99E" blockId="1.[151,1435,1330,1536]" box="[151,1086,1514,1536]" pageId="1" pageNumber="316">
<treatmentCitationGroup id="6EEF9A626B29FF88E7FEFA47FBB0F99E" box="[151,1086,1514,1536]" pageId="1" pageNumber="316">
<taxonomicName id="89FFC6CF6B29FF88E7FEFA47FCFBF99E" ID-CoL="53GC6" authority="Stasolla, Abbarchi &amp; Innocenti, 2015: 449" authorityName="Stasolla, Abbarchi &amp; Innocenti" authorityPageNumber="449" authorityYear="2015" box="[151,885,1514,1536]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="316" phylum="Arthropoda" rank="species" species="spelaeophila">
<emphasis id="7C8B615E6B29FF88E7FEFA47FE4FF99E" box="[151,449,1514,1536]" italics="true" pageId="1" pageNumber="316">Sundathelphusa spelaeophila</emphasis>
<treatmentCitation id="CF5E9B5D6B29FF88E6AFFA47FCFBF99E" ID-DOI="http://doi.org/10.5281/zenodo.5466957" ID-Zenodo-Dep="5466957" LSID="urn:lsid:zoobank.org:act:DAB6C5CD-5C5A-4291-8389-CA7FB3E88626" author="Stasolla, G. &amp; Abbarchi, A. &amp; Innocenti, G." box="[454,885,1514,1536]" httpUri="http://treatment.plazi.org/id/0398FD3B9706FFDBFEEBFEA90192F8C7" page="449" pageId="1" pageNumber="316" year="2015">
<bibRefCitation id="2A6EC0BD6B29FF88E6AFFA47FCCDF99E" author="Stasolla, G. &amp; Abbarchi, A. &amp; Innocenti, G." box="[454,835,1514,1536]" pageId="1" pageNumber="316" pagination="448 - 453" refId="ref9800" refString="Stasolla, G., Abbarchi, A. &amp; Innocenti, G. (2015) Sundathelphusa spelaeophila, a new species of cavernicolous crab from Samar, Philippines (Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 63, 448 - 453." type="journal article" year="2015">Stasolla, Abbarchi &amp; Innocenti, 2015</bibRefCitation>
: 449
</treatmentCitation>
</taxonomicName>
, figs. 1AC, 2AE.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="06E5EEC76B29FF8BE7FEF980FB32FE83" lastPageId="2" lastPageNumber="317" pageId="1" pageNumber="316" type="materials_examined">
<paragraph id="4E40BD4C6B29FF8BE7FEF980FB32FE83" blockId="1.[151,1437,1581,2038]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="317" pageId="1" pageNumber="316">
<emphasis id="7C8B615E6B29FF88E7FEF980FEF1F9D8" bold="true" box="[151,383,1581,1606]" pageId="1" pageNumber="316">Material examined</emphasis>
.
<materialsCitation id="FE97B7116B29FF88E6FAF983FA13F9D8" ID-GBIF-Occurrence="2235592109" box="[403,1437,1581,1606]" collectionCode="ZMB" collectorName="F. Jagor." location="Samar Island" pageId="1" pageNumber="316" specimenCode="ZMB 1055" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="914403EE6B29FF88E6FAF983FD89F9D8" box="[403,519,1582,1606]" pageId="1" pageNumber="316" type="lectotype">Lectotype</typeStatus>
:
<specimenCount id="58F976C56B29FF88E57EF983FDA2F9DB" box="[535,556,1582,1605]" pageId="1" pageNumber="316" type="male"></specimenCount>
(42.8 ×
<quantity id="890710A96B29FF88E5F4F983FC85F9D8" box="[669,779,1582,1606]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="1" pageNumber="316" unit="mm" value="34.8">34.8 mm</quantity>
),
<specimenCode id="1E5915376B29FF88E44DF983FC29F9D8" box="[804,935,1581,1606]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="1" pageNumber="316" type="Museum">ZMB 1055</specimenCode>
a, Lokilokon,
<location id="4B20EB976B29FF88E327F983FB67F9D8" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E327F983FB67F9D8" box="[1102,1257,1581,1606]" name="Samar Island" pageId="1" pageNumber="316">Samar Island</location>
, coll.
<collectorName id="E30AD89A6B29FF88E250F983FA13F9D8" box="[1337,1437,1582,1606]" pageId="1" pageNumber="316">F. Jagor.</collectorName>
</materialsCitation>
<typeStatus id="914403EE6B29FF88E7FEF9FCFEB1F9F4" box="[151,319,1617,1642]" pageId="1" pageNumber="316" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="58F976C56B29FF88E620F9FFFE0DF9F4" box="[329,387,1618,1642]" pageId="1" pageNumber="316" type="male">4♂♂</specimenCount>
, 1 young
<materialsCitation id="FE97B7116B29FF88E699F9FFFC29F9F4" ID-GBIF-Occurrence="2235592102" box="[496,935,1617,1642]" collectionCode="ZMB" collectorName="F. Jagor." location="Samar Island" pageId="1" pageNumber="316" specimenCode="ZMB 1055" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E699F9FFFD8BF9F7" box="[496,517,1618,1641]" pageId="1" pageNumber="316" type="female"></specimenCount>
,
<specimenCode id="1E5915376B29FF88E57AF9FFFD1EF9F4" box="[531,656,1617,1642]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="1" pageNumber="316" type="Museum">ZMB 1055</specimenCode>
b, same data as lectotype
</materialsCitation>
;
<materialsCitation id="FE97B7116B29FF88E4DAF9FFFC99F92C" ID-GBIF-Occurrence="2235592103" collectionCode="ZMB" collectorName="Calbiga &amp; Lokilokon &amp; Loquilocun &amp; Samar &amp; Calbigaufluss &amp; Loquilocum &amp; F. Jagor." country="Philippinen, Insel" location="Samar Island" pageId="1" pageNumber="316" specimenCode="ZMB 1050" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E4DAF9FFFC52F9F7" box="[947,988,1618,1642]" pageId="1" pageNumber="316" type="male">1 ♂</specimenCount>
(51.4 ×
<quantity id="890710A96B29FF88E352F9FFFB2CF9F4" box="[1083,1186,1618,1642]" metricMagnitude="-2" metricUnit="m" metricValue="4.279999999999999" pageId="1" pageNumber="316" unit="mm" value="42.8">42.8 mm</quantity>
),
<specimenCode id="1E5915376B29FF88E3DCF9FFFABDF9F4" box="[1205,1331,1617,1642]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="1" pageNumber="316" type="Museum">ZMB 1050</specimenCode>
,
<collectorName id="E30AD89A6B29FF88E257F9FCFA16F9F4" box="[1342,1432,1617,1642]" pageId="1" pageNumber="316">Calbiga</collectorName>
, stream near
<collectorName id="E30AD89A6B29FF88E64FF9D8FE15F910" box="[294,411,1653,1678]" pageId="1" pageNumber="316">Lokilokon</collectorName>
(formerly
<collectorName id="E30AD89A6B29FF88E572F9D8FD10F910" box="[539,670,1653,1678]" pageId="1" pageNumber="316">Loquilocun</collectorName>
)
<location id="4B20EB976B29FF88E5C6F9DBFCCBF910" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E5C6F9DBFCCBF910" box="[687,837,1653,1678]" country="Philippinen, Insel" name="Samar Island" pageId="1" pageNumber="316">Samar Island</location>
(original label:
<collectingCountry id="36E8FDDC6B29FF88E36AF9D8FB45F910" box="[1027,1227,1653,1678]" isEnumeration="true" pageId="1" pageNumber="316">Philippinen, Insel</collectingCountry>
<collectorName id="E30AD89A6B29FF88E3BFF9DBFA91F910" box="[1238,1311,1654,1678]" pageId="1" pageNumber="316">Samar</collectorName>
,
<collectorName id="E30AD89A6B29FF88E247F9D8FF42F92C" pageId="1" pageNumber="316">Calbigaufluss</collectorName>
bei
<collectorName id="E30AD89A6B29FF88E795F934FE04F92C" box="[252,394,1689,1714]" pageId="1" pageNumber="316">Loquilocum</collectorName>
), coll.
<collectorName id="E30AD89A6B29FF88E6BDF937FDBCF92C" box="[468,562,1690,1714]" pageId="1" pageNumber="316">F. Jagor.</collectorName>
Additional material
</materialsCitation>
:
<materialsCitation id="FE97B7116B29FF88E448F937FDA7F948" ID-GBIF-Occurrence="2235592110" collectingDate="1941-12-04" collectorName="G. E. Edano" country="Philippinen, Insel" county="Samar Island" location="Sarawang" municipality="Matuginao" pageId="1" pageNumber="316" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E448F937FCEFF92F" box="[801,865,1690,1714]" pageId="1" pageNumber="316" type="female">2 ♀♀</specimenCount>
, NMCR 239,
<location id="4B20EB976B29FF88E496F937FBFEF92C" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E496F937FBFEF92C" box="[1023,1136,1690,1714]" country="Philippinen, Insel" county="Samar Island" municipality="Matuginao" name="Sarawang" pageId="1" pageNumber="316">Sarawang</location>
,
<collectingMunicipality id="AE2427366B29FF88E312F934FB79F92C" box="[1147,1271,1689,1714]" pageId="1" pageNumber="316">Matuginao</collectingMunicipality>
,
<collectingCounty id="A721C5C06B29FF88E26BF937FA16F92C" box="[1282,1432,1689,1714]" pageId="1" pageNumber="316">Samar Island</collectingCounty>
, coll.
<collectorName id="E30AD89A6B29FF88E7A6F913FEDCF948" box="[207,338,1725,1750]" pageId="1" pageNumber="316">G.E. Edano</collectorName>
,
<date id="3A419B8C6B29FF88E634F913FDA7F948" box="[349,553,1725,1750]" pageId="1" pageNumber="316" value="1941-12-04">
<collectingDate id="2A0562646B29FF88E634F913FDA7F948" box="[349,553,1725,1750]" pageId="1" pageNumber="316" value="1941-12-04">4 December 1941</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="FE97B7116B29FF88E55CF913FBF4F964" ID-GBIF-Occurrence="2235592129" collectingDate="2006-08-01" collectionCode="ZRC" collectorName="D. E. Husana" country="Philippinen, Insel" county="Samar Island" latitude="11.779767" location="Lobo Cave" longLatPrecision="1" longitude="124.92886" municipality="Western Samar province" pageId="1" pageNumber="316" specimenCode="ZRC 2017.1062" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E55CF913FDEEF94B" box="[565,608,1726,1750]" pageId="1" pageNumber="316" type="male">1 ♂</specimenCount>
(44.9 ×
<quantity id="890710A96B29FF88E5A8F910FCA4F948" box="[705,810,1725,1750]" metricMagnitude="-2" metricUnit="m" metricValue="3.59" pageId="1" pageNumber="316" unit="mm" value="35.9">35.9 mm</quantity>
),
<specimenCode id="1E5915376B29FF88E454F913FC7BF948" box="[829,1013,1725,1750]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="316">ZRC 2017.1062</specimenCode>
,
<location id="4B20EB976B29FF88E368F910FB0EF948" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E368F910FB0EF948" box="[1025,1152,1725,1750]" country="Philippinen, Insel" county="Samar Island" latitude="11.779767" longLatPrecision="1" longitude="124.92886" municipality="Western Samar province" name="Lobo Cave" pageId="1" pageNumber="316">Lobo Cave</location>
,
<location id="4B20EB976B29FF88E3E5F910FB68F948" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E3E5F910FB68F948" box="[1164,1254,1725,1750]" country="Philippinen, Insel" county="Samar Island" latitude="11.779767" longLatPrecision="1" longitude="124.92886" municipality="Western Samar province" name="Jiabong" pageId="1" pageNumber="316">Jiabong</location>
,
<collectingMunicipality id="AE2427366B29FF88E39BF913FF72F964" pageId="1" pageNumber="316">Western Samar province</collectingMunicipality>
,
<collectingCounty id="A721C5C06B29FF88E660F94FFE2FF964" box="[265,417,1761,1786]" pageId="1" pageNumber="316">Samar Island</collectingCounty>
,
<geoCoordinate id="2BCBDB8B6B29FF88E6C6F94CFDB2F964" box="[431,572,1761,1786]" direction="north" orientation="latitude" pageId="1" pageNumber="316" precision="1" value="11.779767">11°46.786N</geoCoordinate>
<geoCoordinate id="2BCBDB8B6B29FF88E52FF94CFD6DF964" box="[582,739,1761,1786]" direction="east" orientation="longitude" pageId="1" pageNumber="316" precision="1" value="124.92886">124°55.732E</geoCoordinate>
, coll.
<collectorName id="E30AD89A6B29FF88E442F94FFC31F967" box="[811,959,1762,1785]" pageId="1" pageNumber="316">D.E. Husana</collectorName>
,
<date id="3A419B8C6B29FF88E4A5F94FFBF4F964" box="[972,1146,1761,1786]" pageId="1" pageNumber="316" value="2006-08-01">
<collectingDate id="2A0562646B29FF88E4A5F94FFBF4F964" box="[972,1146,1761,1786]" pageId="1" pageNumber="316" value="2006-08-01">1 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="FE97B7116B29FF88E3EEF94FFC02F8DC" ID-GBIF-Occurrence="2235592131" collectingDate="2006-08-07" collectionCode="ZRC" collectorName="D. E. Husana" country="Philippinen, Insel" county="Samar Island" location="Panghuyaman Cave" municipality="Western Samar province" pageId="1" pageNumber="316" specimenCode="ZRC 2017.1058" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E3EEF94FFB3BF967" box="[1159,1205,1762,1786]" pageId="1" pageNumber="316" type="male">1 ♂</specimenCount>
,
<specimenCount id="58F976C56B29FF88E3ADF94FFA8BF967" box="[1220,1285,1762,1786]" pageId="1" pageNumber="316" type="female">2 ♀♀</specimenCount>
(35.7 ×
<quantity id="890710A96B29FF88E202F94FFF4BF883" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="1" pageNumber="316" unit="mm" value="28.7">28.7 mm</quantity>
, 39.9 ×
<quantity id="890710A96B29FF88E645F8ABFE18F883" box="[300,406,1798,1822]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="1" pageNumber="316" unit="mm" value="31.8">31.8 mm</quantity>
, 47.6 ×
<quantity id="890710A96B29FF88E694F8ABFDE7F880" box="[509,617,1798,1822]" metricMagnitude="-2" metricUnit="m" metricValue="3.79" pageId="1" pageNumber="316" unit="mm" value="37.9">37.9 mm</quantity>
),
<specimenCode id="1E5915376B29FF88E5E9F8ABFCB2F880" box="[640,828,1797,1822]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="316">ZRC 2017.1058</specimenCode>
,
<location id="4B20EB976B29FF88E422F8A8FBBFF880" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E422F8A8FBBFF880" box="[843,1073,1797,1822]" country="Philippinen, Insel" county="Samar Island" municipality="Western Samar province" name="Panghuyaman Cave" pageId="1" pageNumber="316">Panghuyaman Cave</location>
,
<location id="4B20EB976B29FF88E328F8ABFB51F880" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E328F8ABFB51F880" box="[1089,1247,1797,1822]" country="Philippinen, Insel" county="Samar Island" municipality="Western Samar province" name="Daram Island" pageId="1" pageNumber="316">Daram Island</location>
,
<collectingMunicipality id="AE2427366B29FF88E387F8ABFF77F8DC" pageId="1" pageNumber="316">Western Samar province</collectingMunicipality>
(west of
<collectingCounty id="A721C5C06B29FF88E60AF887FE75F8DC" box="[355,507,1833,1858]" pageId="1" pageNumber="316">Samar Island</collectingCounty>
), coll.
<collectorName id="E30AD89A6B29FF88E52FF887FD56F8DF" box="[582,728,1834,1857]" pageId="1" pageNumber="316">D.E. Husana</collectorName>
,
<date id="3A419B8C6B29FF88E58AF887FC02F8DC" box="[739,908,1833,1858]" pageId="1" pageNumber="316" value="2006-08-07">
<collectingDate id="2A0562646B29FF88E58AF887FC02F8DC" box="[739,908,1833,1858]" pageId="1" pageNumber="316" value="2006-08-07">7 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="FE97B7116B29FF88E4F1F884FEB6F814" ID-GBIF-Occurrence="2235592136" collectionCode="ZRC" collectorName="Brgy San Rafael &amp; Hinabangan &amp; A. C. Diesmos" country="Philippinen, Insel" location="Samar Island" pageId="1" pageNumber="316" specimenCode="ZRC 2016.0107" specimenCount="5" specimenCount-male="5" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E4F1F884FC59F8DF" box="[920,983,1833,1858]" pageId="1" pageNumber="316" type="male">5 ♂♂</specimenCount>
(30.8 ×
<quantity id="890710A96B29FF88E35EF887FB10F8DF" box="[1079,1182,1834,1858]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="1" pageNumber="316" unit="mm" value="24.8">24.8 mm</quantity>
, 20.3 ×
<quantity id="890710A96B29FF88E390F884FAD1F8DF" box="[1273,1375,1833,1858]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="1" pageNumber="316" unit="mm" value="16.7">16.7 mm</quantity>
, 18.0 × 15.0 mm, 16.9 ×
<quantity id="890710A96B29FF88E606F8E3FE5BF8FB" box="[367,469,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="1" pageNumber="316" unit="mm" value="13.9">13.9 mm</quantity>
, 14.1 ×
<quantity id="890710A96B29FF88E546F8E3FD16F8F8" box="[559,664,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="1" pageNumber="316" unit="mm" value="12.3">12.3 mm</quantity>
),
<specimenCode id="1E5915376B29FF88E5C2F8E3FCECF8F8" box="[683,866,1869,1894]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="316">ZRC 2016.0107</specimenCode>
,
<collectorName id="E30AD89A6B29FF88E407F8E3FBA7F8F8" box="[878,1065,1869,1894]" pageId="1" pageNumber="316">Brgy San Rafael</collectorName>
,
<collectorName id="E30AD89A6B29FF88E35DF8E0FB30F8F8" box="[1076,1214,1869,1894]" pageId="1" pageNumber="316">Hinabangan</collectorName>
,
<location id="4B20EB976B29FF88E3A3F8E3FAEEF8F8" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E3A3F8E3FAEEF8F8" box="[1226,1376,1869,1894]" country="Philippinen, Insel" name="Samar Island" pageId="1" pageNumber="316">Samar Island</location>
, coll.
<collectorName id="E30AD89A6B29FF88E7FEF8DFFEB6F814" box="[151,312,1905,1930]" pageId="1" pageNumber="316">A.C. Diesmos</collectorName>
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;
<materialsCitation id="FE97B7116B29FF88E62DF8DFFC3BF814" ID-GBIF-Occurrence="2235592119" box="[324,949,1905,1930]" collectionCode="MNHN" country="Philippinen, Insel" location="Samar Island" pageId="1" pageNumber="316" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E62DF8DFFE1CF814" box="[324,402,1905,1930]" pageId="1" pageNumber="316" type="male">1 male</specimenCount>
,
<collectionCode id="28EE25896B29FF88E6F4F8DFFE7DF817" box="[413,499,1906,1929]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="316">MNHN</collectionCode>
B28509, in cave,
<location id="4B20EB976B29FF88E5D6F8DFFCDBF814" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E5D6F8DFFCDBF814" box="[703,853,1905,1930]" country="Philippinen, Insel" name="Samar Island" pageId="1" pageNumber="316">Samar Island</location>
, no date
</materialsCitation>
;
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<specimenCount id="58F976C56B29FF88E4A8F8DFFC65F817" box="[961,1003,1906,1930]" pageId="1" pageNumber="316" type="female">1 ♀</specimenCount>
,
<specimenCode id="1E5915376B29FF88E490F8DFFB3EF814" box="[1017,1200,1905,1930]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="316">ZRC 2016.0109</specimenCode>
,
<location id="4B20EB976B29FF88E3D5F8DCFA9DF814" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E3D5F8DCFA9DF814" box="[1212,1299,1905,1930]" country="Philippinen, Insel" municipality="Leyte" name="Baybay" pageId="1" pageNumber="316">Baybay</location>
,
<collectingMunicipality id="AE2427366B29FF88E277F8DFFAEFF814" box="[1310,1377,1906,1930]" pageId="1" pageNumber="316">Leyte</collectingMunicipality>
, coll.
<collectorName id="E30AD89A6B29FF88E7FEF83BFEB9F830" box="[151,311,1941,1966]" pageId="1" pageNumber="316">A.C. Diesmos</collectorName>
,
<date id="3A419B8C6B29FF88E62BF83BFE5AF830" box="[322,468,1942,1966]" pageId="1" pageNumber="316" value="2002-08">
<collectingDate id="2A0562646B29FF88E62BF83BFE5AF830" box="[322,468,1942,1966]" pageId="1" pageNumber="316" value="2002-08">August 2002</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="58F976C56B29FF88E6B6F83BFD87F833" box="[479,521,1942,1966]" pageId="1" pageNumber="316" type="male">1 ♂</specimenCount>
(33.8 ×
<quantity id="890710A96B29FF88E50FF83BFD40F830" box="[614,718,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="1" pageNumber="316" unit="mm" value="31.3">31.3 mm</quantity>
),
<materialsCitation id="FE97B7116B29FF88E589F83BFB3EF830" ID-GBIF-Occurrence="2235592124" box="[736,1200,1941,1966]" collectionCode="ZRC" country="Philippinen, Insel" pageId="1" pageNumber="316" specimenCode="ZRC 2017.1259, 1" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E589F83BFC87F833" box="[736,777,1942,1966]" pageId="1" pageNumber="316" type="female">1 ♀</specimenCount>
(38.2 ×
<quantity id="890710A96B29FF88E40EF83BFC40F830" box="[871,974,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="316" unit="mm" value="31.1">31.1 mm</quantity>
),
<specimenCode id="1E5915376B29FF88E488F83BFB19F830" box="[993,1175,1941,1966]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="316">ZRC 2017.1259</specimenCode>
,
<specimenCount id="58F976C56B29FF88E3CBF83BFB3EF830" box="[1186,1200,1942,1966]" pageId="1" pageNumber="316" type="female">
<specimenCode id="1E5915376B29FF88E3CBF83BFB3EF830" box="[1186,1200,1942,1966]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="316">1</specimenCode>
</specimenCount>
</materialsCitation>
<materialsCitation id="FE97B7116B29FF88E3DFF83BFBB0F84C" ID-GBIF-Occurrence="2235592113" collectingDate="1985-09-19" collectorName="S. Shokita" country="Philippinen, Insel" county="Leyte Island" location="Bari" municipality="Jaro" pageId="1" pageNumber="316" specimenCount="1" typeStatus="paralectotype">
♀ (39.1 ×
<quantity id="890710A96B29FF88E240F83BFA1EF830" box="[1321,1424,1942,1966]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="1" pageNumber="316" unit="mm" value="31.3">31.3 mm</quantity>
), NSMT-Cr25886,
<location id="4B20EB976B29FF88E632F814FE00F84C" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E632F814FE00F84C" box="[347,398,1977,2002]" country="Philippinen, Insel" county="Leyte Island" municipality="Jaro" name="Bari" pageId="1" pageNumber="316">Bari</location>
,
<collectingMunicipality id="AE2427366B29FF88E6F1F817FE44F84F" box="[408,458,1978,2001]" pageId="1" pageNumber="316">Jaro</collectingMunicipality>
,
<collectingCounty id="A721C5C06B29FF88E6BDF817FDECF84C" box="[468,610,1977,2002]" pageId="1" pageNumber="316">Leyte Island</collectingCounty>
, coll.
<collectorName id="E30AD89A6B29FF88E5CDF817FC99F84C" box="[676,791,1977,2002]" pageId="1" pageNumber="316">S. Shokita</collectorName>
<emphasis id="7C8B615E6B29FF88E474F817FCDAF84C" box="[797,852,1977,2002]" italics="true" pageId="1" pageNumber="316">et al.</emphasis>
,
<date id="3A419B8C6B29FF88E40BF817FBB0F84C" box="[866,1086,1977,2002]" pageId="1" pageNumber="316" value="1985-09-19">
<collectingDate id="2A0562646B29FF88E40BF817FBB0F84C" box="[866,1086,1977,2002]" pageId="1" pageNumber="316" value="1985-09-19">19 September 1985</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="FE97B7116B29FF88E320F817FB10F868" ID-GBIF-Occurrence="2235592107" collectingDate="1950-03" collectorName="G. Edano" country="Philippinen, Insel" county="Leyte Island" latitude="10.95" location="Kasarugingan" longLatPrecision="1296" longitude="124.683334" municipality="Ormoc" pageId="1" pageNumber="316" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E320F817FBFCF84F" box="[1097,1138,1978,2002]" pageId="1" pageNumber="316" type="male">1 ♂</specimenCount>
(19.5 ×
<quantity id="890710A96B29FF88E3B9F814FAB9F84C" box="[1232,1335,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="316" unit="mm" value="15.8">15.8 mm</quantity>
), NMCR 13,
<location id="4B20EB976B29FF88E7ABF870FEEEF868" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B29FF88E7ABF870FEEEF868" box="[194,352,2013,2038]" country="Philippinen, Insel" county="Leyte Island" latitude="10.95" longLatPrecision="1296" longitude="124.683334" municipality="Ormoc" name="Kasarugingan" pageId="1" pageNumber="316">Kasarugingan</location>
,
<collectingMunicipality id="AE2427366B29FF88E605F873FE33F868" box="[364,445,2014,2038]" pageId="1" pageNumber="316">Ormoc</collectingMunicipality>
,
<collectingCounty id="A721C5C06B29FF88E6A0F873FDD6F868" box="[457,600,2013,2038]" pageId="1" pageNumber="316">Leyte Island</collectingCounty>
, ca.
<geoCoordinate id="2BCBDB8B6B29FF88E5E5F870FD68F868" box="[652,742,2013,2038]" direction="north" orientation="latitude" pageId="1" pageNumber="316" precision="925" value="10.95">10°57'N</geoCoordinate>
<geoCoordinate id="2BCBDB8B6B29FF88E587F873FCD8F868" box="[750,854,2014,2038]" direction="east" orientation="longitude" pageId="1" pageNumber="316" precision="925" value="124.683334">124°41'E</geoCoordinate>
, coll.
<collectorName id="E30AD89A6B29FF88E4F2F873FB8BF868" box="[923,1029,2013,2038]" pageId="1" pageNumber="316">G. Edano</collectorName>
,
<date id="3A419B8C6B29FF88E378F870FB10F868" box="[1041,1182,2013,2038]" pageId="1" pageNumber="316" value="1950-03">
<collectingDate id="2A0562646B29FF88E378F870FB10F868" box="[1041,1182,2013,2038]" pageId="1" pageNumber="316" value="1950-03">March 1950</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="FE97B7116B29FF88E3C3F873FA0DF868" ID-GBIF-Occurrence="2235592108" box="[1194,1411,2014,2038]" collectionCode="NSMT" country="Philippinen, Insel" pageId="1" pageNumber="316" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paralectotype">
<specimenCount id="58F976C56B29FF88E3C3F873FB63F86B" box="[1194,1261,2014,2038]" pageId="1" pageNumber="316" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="58F976C56B29FF88E393F873FAAAF86B" box="[1274,1316,2014,2038]" pageId="1" pageNumber="316" type="female">1 ♀</specimenCount>
,
<collectionCode id="28EE25896B29FF88E25DF873FA0DF868" box="[1332,1411,2014,2038]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="1" pageNumber="316">NSMT</collectionCode>
</materialsCitation>
,
<specimenCount id="58F976C56B29FF8BE2E7F873FF4FFF31" lastPageId="2" lastPageNumber="317" pageId="1" pageNumber="316" type="male">3 ♂♂</specimenCount>
(33.8 ×
<quantity id="890710A96B2AFF8BE64FFF35FE00FF31" box="[294,398,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" pageId="2" pageNumber="317" unit="mm" value="27.1">27.1 mm</quantity>
, 51.3 ×
<quantity id="890710A96B2AFF8BE687FF35FDD9FF31" box="[494,599,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.04" pageId="2" pageNumber="317" unit="mm" value="40.4">40.4 mm</quantity>
, 53.4 ×
<quantity id="890710A96B2AFF8BE5DEFF3AFCAFFF2E" box="[695,801,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="2" pageNumber="317" unit="mm" value="42.5">42.5 mm</quantity>
),
<materialsCitation id="FE97B7116B2AFF8BE45FFF35FB56FF4B" ID-GBIF-Occurrence="2235592133" collectingDate="1985-08-19" collectionCode="ZRC" collectorName="Camarines Sur &amp; Luzon Island &amp; N. Gapas" country="Philippinen, Insel" latitude="13.433333" location="Binang-kaan River" longLatPrecision="1291" longitude="123.51667" municipality="Buhi" pageId="2" pageNumber="317" specimenCode="ZRC 2009.0409" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B2AFF8BE45FFF35FCECFF31" box="[822,866,152,176]" pageId="2" pageNumber="317" type="female">1 ♀</specimenCount>
(51.7 ×
<quantity id="890710A96B2AFF8BE4A1FF35FBBCFF2E" box="[968,1074,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.22" pageId="2" pageNumber="317" unit="mm" value="42.2">42.2 mm</quantity>
),
<specimenCode id="1E5915376B2AFF8BE32EFF35FA8EFF2E" box="[1095,1280,152,176]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="317">ZRC 2009.0409</specimenCode>
,
<location id="4B20EB976B2AFF8BE264FF3AFF59FF4B" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B2AFF8BE264FF3AFF59FF4B" country="Philippinen, Insel" latitude="13.433333" longLatPrecision="1291" longitude="123.51667" municipality="Buhi" name="Binang-kaan River" pageId="2" pageNumber="317">Binang-kaan River</location>
,
<collectingMunicipality id="AE2427366B2AFF8BE78BFF11FE95FF4B" box="[226,283,188,213]" pageId="2" pageNumber="317">Buhi</collectingMunicipality>
, ca.
<geoCoordinate id="2BCBDB8B6B2AFF8BE624FF11FE29FF4B" box="[333,423,188,213]" direction="north" orientation="latitude" pageId="2" pageNumber="317" precision="925" value="13.433333">13°26'N</geoCoordinate>
<geoCoordinate id="2BCBDB8B6B2AFF8BE6C7FF11FD9BFF4A" box="[430,533,188,212]" direction="east" orientation="longitude" pageId="2" pageNumber="317" precision="925" value="123.51667">123°31'E</geoCoordinate>
,
<collectorName id="E30AD89A6B2AFF8BE548FF11FD46FF4A" box="[545,712,188,213]" pageId="2" pageNumber="317">Camarines Sur</collectorName>
,
<collectorName id="E30AD89A6B2AFF8BE5BAFF10FCE7FF4B" box="[723,873,188,213]" hasSafeAffix="true" pageId="2" pageNumber="317">Luzon Island</collectorName>
, coll.
<collectorName id="E30AD89A6B2AFF8BE4C5FF10FB99FF4A" box="[940,1047,188,212]" pageId="2" pageNumber="317">N. Gapas</collectorName>
,
<date id="3A419B8C6B2AFF8BE34BFF11FB56FF4B" box="[1058,1240,188,213]" pageId="2" pageNumber="317" value="1985-08-19">
<collectingDate id="2A0562646B2AFF8BE34BFF11FB56FF4B" box="[1058,1240,188,213]" pageId="2" pageNumber="317" value="1985-08-19">19 August 1985</collectingDate>
</date>
</materialsCitation>
; 1 young
<specimenCount id="58F976C56B2AFF8BE22FFF10FAD5FF4A" box="[1350,1371,189,212]" pageId="2" pageNumber="317" type="male"></specimenCount>
(15.3 × 13.0 mm), 1 young
<materialsCitation id="FE97B7116B2AFF8BE6C0FF4DFCC0FE83" ID-GBIF-Occurrence="2235592122" collectingDate="1985-08-19" collectionCode="ZRC" collectorName="Camarines Sur &amp; Luzon Island &amp; N. Gapas" country="Philippinen, Insel" latitude="13.433333" location="Borabod" longLatPrecision="1291" longitude="123.51667" municipality="Buhi" pageId="2" pageNumber="317" specimenCode="ZRC 2017.1059" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="58F976C56B2AFF8BE6C0FF4DFE30FF69" box="[425,446,224,247]" pageId="2" pageNumber="317" type="female"></specimenCount>
(20.4 ×
<quantity id="890710A96B2AFF8BE546FF72FD13FF66" box="[559,669,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="2" pageNumber="317" unit="mm" value="16.9">16.9 mm</quantity>
),
<specimenCode id="1E5915376B2AFF8BE5DFFF4DFCFAFF66" box="[694,884,223,248]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="317">ZRC 2017.1059</specimenCode>
,
<location id="4B20EB976B2AFF8BE4ECFF72FC68FF66" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B2AFF8BE4ECFF72FC68FF66" box="[901,998,223,248]" country="Philippinen, Insel" latitude="13.433333" longLatPrecision="1291" longitude="123.51667" municipality="Buhi" name="Borabod" pageId="2" pageNumber="317">Borabod</location>
stream,
<collectingMunicipality id="AE2427366B2AFF8BE33BFF72FB05FF66" box="[1106,1163,223,248]" pageId="2" pageNumber="317">Buhi</collectingMunicipality>
, ca.
<geoCoordinate id="2BCBDB8B6B2AFF8BE3A0FF72FAADFF66" box="[1225,1315,223,248]" direction="north" orientation="latitude" pageId="2" pageNumber="317" precision="925" value="13.433333">13°26'N</geoCoordinate>
<geoCoordinate id="2BCBDB8B6B2AFF8BE259FF4DFA19FF66" box="[1328,1431,224,248]" direction="east" orientation="longitude" pageId="2" pageNumber="317" precision="925" value="123.51667">123°31'E</geoCoordinate>
,
<collectorName id="E30AD89A6B2AFF8BE7FEFEA9FEB0FE82" box="[151,318,260,285]" pageId="2" pageNumber="317">Camarines Sur</collectorName>
,
<collectorName id="E30AD89A6B2AFF8BE620FEA8FE51FE83" box="[329,479,260,285]" hasSafeAffix="true" pageId="2" pageNumber="317">Luzon Island</collectorName>
, coll.
<collectorName id="E30AD89A6B2AFF8BE54BFEA8FD00FE82" box="[546,654,260,284]" pageId="2" pageNumber="317">N. Gapas</collectorName>
,
<date id="3A419B8C6B2AFF8BE5F1FEA9FCC0FE83" box="[664,846,260,285]" pageId="2" pageNumber="317" value="1985-08-19">
<collectingDate id="2A0562646B2AFF8BE5F1FEA9FCC0FE83" box="[664,846,260,285]" pageId="2" pageNumber="317" value="1985-08-19">19 August 1985</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="FE97B7116B2AFF8BE430FEA9FB36FE83" ID-GBIF-Occurrence="2235592134" box="[857,1208,260,285]" country="Philippines" location="All" pageId="2" pageNumber="317" specimenCount="1" typeStatus="paralectotype">
<location id="4B20EB976B2AFF8BE430FEA9FCF3FE83" LSID="urn:lsid:plazi:treatment:C6560C5A6B29FF8FE7FEFB6AFDA0F8EE:4B20EB976B2AFF8BE430FEA9FCF3FE83" box="[857,893,260,285]" country="Philippines" name="All" pageId="2" pageNumber="317">All</location>
locations in the
<collectingCountry id="36E8FDDC6B2AFF8BE35FFEA9FB36FE83" box="[1078,1208,260,285]" name="Philippines" pageId="2" pageNumber="317">Philippines</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="06E5EEC76B2AFF8BE7AEFE8AFB14F9EA" pageId="2" pageNumber="317" type="description">
<paragraph id="4E40BD4C6B2AFF8BE7AEFE8AFAECFC1E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
<emphasis id="7C8B615E6B2AFF8BE7AEFE8AFEDFFEDE" bold="true" box="[199,337,295,320]" pageId="2" pageNumber="317">Description</emphasis>
. Carapace trapezoidal, widest at anterior quarter, dorsal surface convex longitudinally, dorsoventrally depressed, regions distinct (
<figureCitation id="D6C4A1C96B2AFF8BE5FDFEE1FCA8FEFA" box="[660,806,332,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1A, B</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE453FEE1FCD2FEFA" box="[826,860,332,356]" captionStart="FIGURE 3" captionStartId="5.[151,250,1509,1531]" captionTargetBox="[150,1436,171,1487]" captionTargetId="figure@5.[150,1436,171,1488]" captionTargetPageId="5" captionText="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale." httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="2" pageNumber="317">3E</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE406FEE1FC34FEFA" box="[879,954,332,357]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="2" pageNumber="317">6A, E</figureCitation>
). Frontal region sloping anteroventrally; anterolateral regions inflated dorsolaterally, smooth to gently cristate; cervical grooves deep; H-shaped gastric groove deep; epigastric cristae distinct, edges rough, separated by distinct median furrow; postorbital cristae, sharp but low; epigastric and postorbital cristae not confluent; epibranchial teeth and postorbital cristae not confluent, separated by gaps (
<figureCitation id="D6C4A1C96B2AFF8BE618FE71FE62FE6A" box="[369,492,476,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Fig. 1A, B</figureCitation>
). Frontal margin broadly protruded, two lobes clearly separated with broad median concavity; external orbital tooth produced anteriorly, outer margin longer than inner margin; epibranchial tooth distinct, triangular, well separated from external orbital tooth, tapering anteriorly, surface flat, margin not curving dorsally; anterolateral margin convex, crest low, not clearly visible even when viewed laterally, not clearly demarcated from posterolateral margin; posterolateral margin straight, converging gradually towards posterior margin of carapace (
<figureCitation id="D6C4A1C96B2AFF8BE617FD22FD8AFD39" box="[382,516,655,680]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1A, B</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE57BFD22FDF2FD36" box="[530,636,655,680]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="2" pageNumber="317">6A, C, G</figureCitation>
). Frontal median triangle complete; dorsal and lateral margins distinct, smooth; dorsal margin more produced anteriorly than lateral margins (
<figureCitation id="D6C4A1C96B2AFF8BE4C3FD19FC71FD52" box="[938,1023,692,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="2" pageNumber="317">Fig. 2F</figureCitation>
); orbit well demarcated; supraorbital margin smooth; infraorbital margin protruded anteriorly granulated; outer edge reaching, fused with anterolateral margin; suborbital and subbranchial regions covered with scattered oblique long, short striae; pterygostomial region smooth with oblique ridges on upper outer part (
<figureCitation id="D6C4A1C96B2AFF8BE449FCB2FC08FCA6" box="[800,902,799,824]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1C</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE4FCFCB2FC36FCA6" box="[917,952,799,824]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="2" pageNumber="317">6E</figureCitation>
). Posterior margin of epistome with three lobes, median lobe large, subtriangular, more produced; lateral lobes wider and protruded, placed more posteriorly than median lobe (
<figureCitation id="D6C4A1C96B2AFF8BE60EFCCAFE47FC1E" box="[359,457,871,896]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1C</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE6BFFCC5FE72FC1E" box="[470,508,872,896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="2" pageNumber="317">2G</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE56EFCCAFDA3FC1E" box="[519,557,871,896]" captionStart="FIGURE 6" captionStartId="10.[151,250,1855,1877]" captionTargetBox="[150,1436,265,1833]" captionTargetId="figure@10.[150,1436,265,1833]" captionTargetPageId="10" captionText="FIGURE 6. A, C, E, G, I, J, Sundathelphusa philippina (von Martens, 1868), ♂ (44.9 × 35.9 mm), ZRC 2017.1062, Western Samar; B, D, F, H, K, L, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254.102, Cebu. A, B, tilted frontal view of dorsal surface of carapace showing striations on anterolateral regions; C, D, lateral view of cephalothorax showing anterolateral margin and curvature of external orbital tooth; E, F, frontal view of cephalothorax showing convexity of gastric and branchial regions; G, H, posterior margin of epistome showing median lobe; I, K, ventral view of left G1; J, L, dorsal view of left G1. Scales = 2.0 mm." httpUri="https://zenodo.org/record/2637378/files/figure.png" pageId="2" pageNumber="317">6G</figureCitation>
). Eyes well developed, occupying entire orbit (
<figureCitation id="D6C4A1C96B2AFF8BE354FCCAFB0BFC1E" box="[1085,1157,871,896]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1</figureCitation>
AC; 6A, C, E, G).
</paragraph>
<paragraph id="4E40BD4C6B2AFF8BE7AEFC21FBBBFC72" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
Ischium of third maxilliped rectangular, with distinct submedian sulcus close to mesial margin; merus quadrate, anteroexternal angle convex, anterior margin slightly concave; tip of exopod reaching midpoint of outer margin of merus, with long flagellum reaching beyond mesial margin (
<figureCitation id="D6C4A1C96B2AFF8BE4D8FC79FBA6FC72" box="[945,1064,980,1005]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Fig. 1C, E</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2AFF8BE7AEFC5AFCD4FBE2" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
Adult male chelipeds stout, subequal; dorsal margin of merus serrated, dorsal margin with distinct subdistal tooth; carpus with strong distal inner angle, flattened dorsoventrally, laterally fringed with proximal teeth; palm equal in length with finger; ventral margin granulated; fingers robust, cutting edges with teeth of various sizes, largest medially, smaller on distal and proximal parts (
<figureCitation id="D6C4A1C96B2AFF8BE591FBC9FCC2FBE2" box="[760,844,1124,1149]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2AFF8BE7AEFB2AFD2AFACA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
Ambulatory legs not elongate (
<figureCitation id="D6C4A1C96B2AFF8BE55FFB2AFD10FB3E" box="[566,670,1159,1184]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1A</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE5C6FB25FC8BFB3E" box="[687,773,1160,1184]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: AF = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="2" pageNumber="317">9A, B</figureCitation>
), second leg longest; anterior margin of merus serrated, without subdistal tooth or spine, posterior margins smooth; carpus short, with longitudinal submedian ridge on dorsal and ventral surfaces of all legs except on fourth leg that lacks ventral ridge, with barely visible dorsal ridge, widened distally, outer margins indistinctly serrated; propodus with rows of spines on inner and outer margins, shorter on outer margin; dactylus with rows of spines on all margins, spines of both outer and inner margins of dactylus almost equal in length (
<figureCitation id="D6C4A1C96B2AFF8BE56AFA91FDE8FACA" box="[515,614,1340,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1A</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE51AFA91FD19FACA" box="[627,663,1340,1364]" captionStart="FIGURE 9" captionStartId="14.[151,250,1020,1042]" captionTargetBox="[161,1428,171,999]" captionTargetId="figure@14.[161,1428,171,999]" captionTargetPageId="14" captionText="FIGURE 9. Right third and fourth ambulatory legs. A, B, Sundathelphusa philippina (von Martens, 1868), ♂ (51.3 × 40.4 mm), ZRC 2009.0409, Luzon; C, D, S. cebu sp. nov., holotype ♂ (38.1 × 31.2 mm), ZRC 2017.1254, Cebu; E, F, S. quirino sp. nov., holotype ♂ (49.3 × 37.5 mm), NMCR 50701, Quirino. A, C, E, third ambulatory leg; B, D, F, fourth ambulatory leg. Scales: AF = 5.0 mm." httpUri="https://zenodo.org/record/2637384/files/figure.png" pageId="2" pageNumber="317">9B</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2AFF8BE7AEFAF2FB22F996" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
Male sternopleonal cavity reaching to level of proximal quarter of coxae of chelipeds (
<figureCitation id="D6C4A1C96B2AFF8BE3C2FAF2FA88FAE6" box="[1195,1286,1375,1400]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Fig. 1D</figureCitation>
). Adult male pleon narrow, T-shaped; somite 1 very short, proximal and distal margins sinuous; somite 2 transversely subrectangular; somites 35 narrow gradually; lateral margins of somite 3 convex, lateral margins of somites 4 and 5 slightly concave; somite 6 rectangular, longer than broad, lateral margins slightly concave; telson subtriangular, longer than broad, lateral margin concave medially, rounded distally (
<figureCitation id="D6C4A1C96B2AFF8BE4CAFA42FB89F996" box="[931,1031,1519,1544]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Figs. 1D</figureCitation>
;
<figureCitation id="D6C4A1C96B2AFF8BE37AFA5DFB11F996" box="[1043,1183,1520,1544]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="2" pageNumber="317">2E, U, V, W</figureCitation>
).
</paragraph>
<paragraph id="4E40BD4C6B2AFF8BE7AEF9B9FB14F9EA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
G1 relatively slender; subterminal segment, almost straight, tapering; terminal segment obliquely bent outwards at midpoint, outer margin concave, tapering, cylindrical, slightly setose (
<figureCitation id="D6C4A1C96B2AFF8BE338F99AFB2FF9CE" box="[1105,1185,1591,1616]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="2" pageNumber="317">Figs. 2</figureCitation>
AD, HS;
<date id="3A419B8C6B2AFF8BE245F99AFACBF9CE" box="[1324,1349,1591,1616]" pageId="2" pageNumber="317">6I</date>
, J). G2 shorter than G1, terminal segment long, about half length of subterminal segment (
<figureCitation id="D6C4A1C96B2AFF8BE35FF9F1FB00F9EA" box="[1078,1166,1628,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1821]" captionTargetBox="[198,1324,189,1779]" captionTargetId="figure@4.[192,1328,183,1779]" captionTargetPageId="4" captionText="FIGURE 2. Sundathelphusa philippina (von Martens, 1868). AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm." httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="2" pageNumber="317">Fig. 2T</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="06E5EEC76B2AFF8FE7AEF9D2FDA0F8EE" lastPageId="6" lastPageNumber="321" pageId="2" pageNumber="317" type="discussion">
<paragraph id="4E40BD4C6B2AFF8BE7AEF9D2FEFCF8D3" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="317">
<emphasis id="7C8B615E6B2AFF8BE7AEF9D2FEBAF906" bold="true" box="[199,308,1663,1688]" pageId="2" pageNumber="317">Remarks</emphasis>
. Von
<bibRefCitation id="2A6EC0BD6B2AFF8BE615F92CFDA2F906" author="Martens, E. von" box="[380,556,1663,1688]" pageId="2" pageNumber="317" pagination="608 - 615" refId="ref9072" refString="Martens, E. von (1868) Ueber einige neue crustaceen. Monatsberichte der Koniglichen Preussische Akademie der Akademie der Wissenschaften zu Berlin, Sitzung der Physikalisch-mathematische Klasse, 1868, 608 - 615." type="journal article" year="1868">Martens (1868)</bibRefCitation>
described
<taxonomicName id="89FFC6CF6B2AFF8BE5C5F92DFCB0F906" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[684,830,1663,1688]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="317" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2AFF8BE5C5F92DFCB0F906" box="[684,830,1663,1688]" italics="true" pageId="2" pageNumber="317">S. philippina</emphasis>
</taxonomicName>
from an unspecified number of specimens from the islands of
<collectingRegion id="8C3B73AE6B2AFF8BE662F909FEDCF922" box="[267,338,1700,1724]" country="Philippines" name="Samar" pageId="2" pageNumber="317">Samar</collectingRegion>
and Luzon. All the specimens he examined are, therefore,
<typeStatus id="914403EE6B2AFF8BE482F908FBDFF923" box="[1003,1105,1701,1725]" pageId="2" pageNumber="317" type="syntype">syntypes</typeStatus>
.
<bibRefCitation id="2A6EC0BD6B2AFF8BE332F908FA87F922" author="Bott, R." box="[1115,1289,1700,1724]" pageId="2" pageNumber="317" pagination="1 - 338" refId="ref8072" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Abhandlungen der Senckenbergischen Naturforschenden Gesselschaft, 526, 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott (1970: 79)</bibRefCitation>
designated a male measuring 43 ×
<quantity id="890710A96B2AFF8BE6EEF965FE5BF97E" box="[391,469,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="317" unit="mm" value="33.0">33 mm</quantity>
from the type series (ZMB 1055) as the
<typeStatus id="914403EE6B2AFF8BE4F1F96AFB8CF97E" box="[920,1026,1735,1760]" pageId="2" pageNumber="317" type="lectotype">lectotype</typeStatus>
. The bottle catalogued as ZMB 1055 contains a 42.8 ×
<quantity id="890710A96B2AFF8BE632F941FE31F89A" box="[347,447,1772,1796]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="2" pageNumber="317" unit="mm" value="34.8">34.8 mm</quantity>
male specimen from “Loquilocun” (in
<collectingRegion id="8C3B73AE6B2AFF8BE41CF941FC30F89A" box="[885,958,1772,1796]" country="Philippines" name="Samar" pageId="2" pageNumber="317">Samar</collectingRegion>
) (
<figureCitation id="D6C4A1C96B2AFF8BE4BDF941FBA3F89A" box="[980,1069,1772,1797]" captionStart="FIGURE 1" captionStartId="3.[151,250,1875,1897]" captionTargetBox="[151,1434,481,1854]" captionTargetId="figure@3.[151,1435,481,1854]" captionTargetPageId="3" captionText="FIGURE 1. Sundathelphusa philippina (von Martens, 1868). A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela." httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="2" pageNumber="317">Fig. 1A</figureCitation>
, CF) with the label “
<typeStatus id="914403EE6B2AFF8BE249F941FA00F89B" box="[1312,1422,1772,1797]" pageId="2" pageNumber="317" type="lectotype">lectotype</typeStatus>
”, and it agrees well with the figures and measurements given by
<bibRefCitation id="2A6EC0BD6B2AFF8BE4EEF8BCFBB3F8B6" author="Bott, R." box="[903,1085,1808,1832]" pageId="2" pageNumber="317" pagination="1 - 338" refId="ref8072" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Abhandlungen der Senckenbergischen Naturforschenden Gesselschaft, 526, 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott (1970: 79)</bibRefCitation>
for the
<typeStatus id="914403EE6B2AFF8BE3C8F8A2FA85F8B6" box="[1185,1291,1807,1832]" pageId="2" pageNumber="317" type="lectotype">lectotype</typeStatus>
, and is here recognized as such.
</paragraph>
<paragraph id="4E40BD4C6B2AFF8AE7AEF8FAFB81FE83" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="318" pageId="2" pageNumber="317">
The
<typeStatus id="914403EE6B2AFF8BE793F8F4FEA4F8EF" box="[250,298,1881,1905]" pageId="2" pageNumber="317">type</typeStatus>
locality “Loquilocun” is a small village (also known as barangay) in Paranas town (formerly known as “Wright” town) upstream of Ulot river in
<collectingRegion id="8C3B73AE6B2AFF8BE589F8D1FCA9F80A" box="[736,807,1916,1940]" country="Philippines" name="Samar" pageId="2" pageNumber="317">Samar</collectingRegion>
and has the following coordinates:
<geoCoordinate id="2BCBDB8B6B2AFF8BE391F8D1FA16F80A" box="[1272,1432,1916,1940]" direction="north" orientation="latitude" pageId="2" pageNumber="317" precision="1" value="11.808333">11°4830.0”N</geoCoordinate>
,
<geoCoordinate id="2BCBDB8B6B2AFF8BE7FEF80DFECDF827" box="[151,323,1952,1977]" direction="east" orientation="longitude" pageId="2" pageNumber="317" precision="1" value="125.10667">125°0624.0”E</geoCoordinate>
. The spelling of the village has already changed to “Lokilokon” in recent years; hence the old name cannot be found in modern maps and digital references. As for the data on the old labels, we note that it is a common practice in the
<collectingCountry id="36E8FDDC6B2BFF8AE6CEFF3AFDABFF2E" box="[423,549,151,176]" name="Philippines" pageId="3" pageNumber="318">Philippines</collectingCountry>
to name a stream or a river after the name of a town or a village. We are of the opinion that the name “Calbiga” on the old Jagor labels is imprecise because Lokilokon is actually located at Paranas town, which is about
<quantity id="890710A96B2BFF8AE685FF4DFDBBFF66" box="[492,565,223,248]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="318" unit="km" value="30.0">30 km</quantity>
away. We believe that the collector F. Jagor used this name only because he had passed Calbiga town on his way to Lokilokon village during his expedition
</paragraph>
<paragraph id="4E40BD4C6B2BFF8AE7AEFE8AFDC0FE33" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="318">
The specimen from Camarines Sur, a province located in the Bicol peninsula in the southeastern part of Luzon Island, just north of Samar Island, could be a different species from
<taxonomicName id="89FFC6CF6B2BFF8AE4E4FEE0FB92FEFB" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[909,1052,332,357]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2BFF8AE4E4FEE0FB92FEFB" box="[909,1052,332,357]" italics="true" pageId="3" pageNumber="318">S. philippina</emphasis>
</taxonomicName>
given its geographic location. But until comparative specimens become available, we regard the
<taxonomicName id="89FFC6CF6B2BFF8AE406FEC2FBACFE16" authorityName="Bott" authorityYear="1969" box="[879,1058,367,392]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="7C8B615E6B2BFF8AE406FEC2FBACFE16" box="[879,1058,367,392]" italics="true" pageId="3" pageNumber="318">Sundathelphusa</emphasis>
</taxonomicName>
populations in the two adjacent islands as one taxon for the time being.
</paragraph>
<caption id="1A80EDC46B2BFF8AE7FEF8FEFB01F838" httpUri="https://zenodo.org/record/2637368/files/figure.png" pageId="3" pageNumber="318" startId="3.[151,250,1875,1897]" targetBox="[151,1434,481,1854]" targetPageId="3">
<paragraph id="4E40BD4C6B2BFF8AE7FEF8FEFB01F838" blockId="3.[151,1436,1875,1958]" pageId="3" pageNumber="318">
<emphasis id="7C8B615E6B2BFF8AE7FEF8FEFE9AF8F6" bold="true" box="[151,276,1875,1897]" pageId="3" pageNumber="318">FIGURE 1.</emphasis>
<taxonomicName id="89FFC6CF6B2BFF8AE672F8FEFC89F8F7" authority="(von Martens, 1868)" authorityName="von Martens" authorityYear="1868" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[283,775,1875,1897]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="318" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2BFF8AE672F8FEFDA2F8F7" box="[283,556,1875,1897]" italics="true" pageId="3" pageNumber="318">Sundathelphusa philippina</emphasis>
(von Martens, 1868)
</taxonomicName>
. A, CF, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; B, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar. A, overall habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, ventral view of cephalothorax; E, left third maxilliped; F, outer view of left chela.
</paragraph>
</caption>
<caption id="1A80EDC46B2CFF8DE7FEF8AAFC9CF86A" httpUri="https://zenodo.org/record/2637370/files/figure.png" pageId="4" pageNumber="319" startId="4.[151,250,1799,1821]" targetBox="[198,1324,189,1779]" targetPageId="4">
<paragraph id="4E40BD4C6B2CFF8DE7FEF8AAFC9CF86A" blockId="4.[151,1436,1799,2036]" pageId="4" pageNumber="319">
<emphasis id="7C8B615E6B2CFF8DE7FEF8AAFE9DF882" bold="true" box="[151,275,1799,1821]" pageId="4" pageNumber="319">FIGURE 2.</emphasis>
<taxonomicName id="89FFC6CF6B2CFF8DE670F8AAFC8DF883" authority="(von Martens, 1868)" authorityName="von Martens" authorityYear="1868" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[281,771,1799,1821]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="319" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2CFF8DE670F8AAFDA7F883" box="[281,553,1799,1821]" italics="true" pageId="4" pageNumber="319">Sundathelphusa philippina</emphasis>
(von Martens, 1868)
</taxonomicName>
. AG, paralectotype ♂ (51.4 × 42.8 mm), ZMB 1050, Samar; H K, lectotype ♂ (42.8 × 34.8 mm), ZMB 1055, Samar; LO, U, ♂ (33.8 × 27.2 mm), ZRC 2017.1059, Leyte; PT, V, ♂ (51.3 × 40.2 mm), ZRC 2009.409, Luzon; W, ♂ (33.7 × 26.8 mm), ZRC 2009.409, Luzon. A, H, dorsal view of right G1; B, I, ventral view of right G1; C, J, dorsal view of terminal segment of right G1; D, K, ventral view of terminal segment of right G1; L, P, ventral view of left G1; M, Q, dorsal view of left G1; N, R, ventral view of terminal segment of left G1; O, S, dorsal view of terminal segment of left G1; E, male pleonal somites 36 and telson; F, frontal median triangle; G, posterior margin of epistome; T, right G2; UW, male pleonal somites 46 and telson. Scales: A, B, H, I, L, M, P, Q = 1.0 mm; U, W = 2.0 mm; E = 5.0 mm; F, G, V = 3.0 mm; C, D, J, K, N, O, R, S = 0.5 mm.
</paragraph>
</caption>
<caption id="1A80EDC46B2DFF8CE7FEFA48FC05F9A6" httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="5" pageNumber="320" startId="5.[151,250,1509,1531]" targetBox="[150,1436,171,1487]" targetPageId="5">
<paragraph id="4E40BD4C6B2DFF8CE7FEFA48FC05F9A6" blockId="5.[151,1436,1509,1592]" pageId="5" pageNumber="320">
<emphasis id="7C8B615E6B2DFF8CE7FEFA48FE9DFA64" bold="true" box="[151,275,1509,1531]" pageId="5" pageNumber="320">FIGURE 3.</emphasis>
<taxonomicName id="89FFC6CF6B2DFF8CE673FA48FC8DFA65" authority="(von Martens, 1868)" authorityName="von Martens" authorityYear="1868" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[282,771,1509,1531]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2DFF8CE673FA48FDA4FA65" box="[282,554,1509,1531]" italics="true" pageId="5" pageNumber="320">Sundathelphusa philippina</emphasis>
(von Martens, 1868)
</taxonomicName>
, variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale.
</paragraph>
</caption>
<paragraph id="4E40BD4C6B2DFF8CE7AEF9F2FDDBF841" blockId="5.[151,1436,1631,2016]" pageId="5" pageNumber="320">
The taxonomic history of
<taxonomicName id="89FFC6CF6B2DFF8CE686F9CDFDF1F9E6" baseAuthorityName="von Martens" baseAuthorityYear="1868" box="[495,639,1631,1656]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2DFF8CE686F9CDFDF1F9E6" box="[495,639,1631,1656]" italics="true" pageId="5" pageNumber="320">S. philippina</emphasis>
</taxonomicName>
is confusing as various authors have mixed up material together.
<bibRefCitation id="2A6EC0BD6B2DFF8CE205F9CDFE82F902" author="Bott, R." pageId="5" pageNumber="320" pagination="1 - 338" refId="ref8072" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Abhandlungen der Senckenbergischen Naturforschenden Gesselschaft, 526, 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott (1970: 79)</bibRefCitation>
stated that only specimens from
<collectingRegion id="8C3B73AE6B2DFF8CE513F929FD4DF902" box="[634,707,1668,1692]" country="Philippines" name="Samar" pageId="5" pageNumber="320">Samar</collectingRegion>
, Mariveles and
<collectingRegion id="8C3B73AE6B2DFF8CE41AF92EFC39F902" box="[883,951,1667,1692]" country="Philippines" name="Bohol" pageId="5" pageNumber="320">Bohol</collectingRegion>
can be confidently referred to
<taxonomicName id="89FFC6CF6B2DFF8CE264F929FA12F902" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1293,1436,1667,1692]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2DFF8CE264F929FA12F902" box="[1293,1436,1667,1692]" italics="true" pageId="5" pageNumber="320">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2DFF8CE7FEF905FEA9F95E" box="[151,295,1703,1728]" pageId="5" pageNumber="320" sensu="stricto">sensu stricto</taxonomicNameLabel>
. Material that had been identified as this species by
<bibRefCitation id="2A6EC0BD6B2DFF8CE41FF905FBBBF95E" author="Burger, O." box="[886,1077,1704,1728]" pageId="5" pageNumber="320" pagination="1 - 7" refId="ref8109" refString="Burger, O. (1894) Beitrage zur Kenntniss der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (1), 1 - 7, pl. 1." type="journal article" year="1894">Bürger (1894: 4)</bibRefCitation>
, De Man (1898: 437),
<bibRefCitation id="2A6EC0BD6B2DFF8CE251F905FE92F97A" author="Ortmann, A. E." pageId="5" pageNumber="320" pagination="258 - 372" refId="ref9729" refString="Ortmann, A. E. (1897) Carcinologische Studien. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (3), 258 - 372, pl. 17." type="journal article" year="1897">Ortmann (1897: 304)</bibRefCitation>
,
<bibRefCitation id="2A6EC0BD6B2DFF8CE641F966FD9FF97A" author="Rathbun, M. J." box="[296,529,1739,1764]" pageId="5" pageNumber="320" pagination="225 - 312" refId="ref9767" refString="Rathbun, M. J. (1904) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, 4, 225 - 312, pls. 9 - 18." type="journal article" year="1904">Rathbun (1904: 304)</bibRefCitation>
and
<bibRefCitation id="2A6EC0BD6B2DFF8CE52EF966FC80F97A" author="Balss, H." box="[583,782,1739,1764]" pageId="5" pageNumber="320" pagination="177 - 184" refId="ref7960" refString="Balss, H. (1934) Mitteilungen von der Wallacea-Expedition Woltereck. Mitteilung XI: Die Dekapoden. Zoologischer Anzeiger, 106 (7 - 8), 177 - 184." type="journal article" year="1934">Balss (1934: 177)</bibRefCitation>
from Luzon, Negros and other locations were mixed, and/ or have been confused with
<taxonomicName id="89FFC6CF6B2DFF8CE6B5F942FBC6F896" authority="(H. Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[476,1096,1775,1800]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="grapsoides">
<emphasis id="7C8B615E6B2DFF8CE6B5F942FC9DF896" box="[476,787,1775,1800]" italics="true" pageId="5" pageNumber="320">Sundathelphusa grapsoides</emphasis>
(H.
<bibRefCitation id="2A6EC0BD6B2DFF8CE420F942FBCEF896" author="Milne Edwards, H." box="[841,1088,1775,1800]" pageId="5" pageNumber="320" pagination="163 - 228" refId="ref9316" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodides. Annales des Sciences Naturelles, Series 3, 20, 163 - 228." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
instead (see also
<bibRefCitation id="2A6EC0BD6B2DFF8CE27FF942FA16F896" author="Balss, H." box="[1302,1432,1775,1800]" pageId="5" pageNumber="320" pagination="143 - 197" refId="ref7993" refString="Balss, H. (1937) Potamoniden (Dekapoda Brachyura) der Philippinen und des Malayischen Archipels. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 34, 143 - 197. https: // doi. org / 10.1002 / iroh. 19370340111" type="journal article" year="1937">Balss 1937</bibRefCitation>
: 153).
<bibRefCitation id="2A6EC0BD6B2DFF8CE7B2F8B9FED1F8B2" author="Bott, R." box="[219,351,1812,1836]" pageId="5" pageNumber="320" pagination="1 - 338" refId="ref8072" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Abhandlungen der Senckenbergischen Naturforschenden Gesselschaft, 526, 1 - 338, pls. 1 - 58." type="journal article" year="1970">Bott (1970)</bibRefCitation>
also synonymized
<taxonomicName id="89FFC6CF6B2DFF8CE554F8B8FBF8F8B2" authority="Rathbun, 1904" authorityName="Rathbun" authorityYear="1904" box="[573,1142,1811,1836]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="montanoanus" subGenus="Potamon">
<emphasis id="7C8B615E6B2DFF8CE554F8B8FD2CF8B2" box="[573,674,1813,1836]" italics="true" pageId="5" pageNumber="320">Potamon</emphasis>
(
<emphasis id="7C8B615E6B2DFF8CE5DDF8B8FC97F8B2" box="[692,793,1813,1836]" italics="true" pageId="5" pageNumber="320">Potamon</emphasis>
)
<emphasis id="7C8B615E6B2DFF8CE445F8B8FC4DF8B2" box="[812,963,1813,1836]" italics="true" pageId="5" pageNumber="320">montanoanus</emphasis>
<bibRefCitation id="2A6EC0BD6B2DFF8CE4A5F8BEFBF8F8B2" author="Rathbun, M. J." box="[972,1142,1811,1836]" pageId="5" pageNumber="320" pagination="225 - 312" refId="ref9767" refString="Rathbun, M. J. (1904) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, 4, 225 - 312, pls. 9 - 18." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="89FFC6CF6B2DFF8CE3DFF8B8FE00F8CE" authority="Rathbun, 1904" authorityName="Rathbun" authorityYear="1904" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="mistio" subGenus="Potamon">
<emphasis id="7C8B615E6B2DFF8CE3DFF8B8FA95F8B2" box="[1206,1307,1813,1836]" italics="true" pageId="5" pageNumber="320">Potamon</emphasis>
(
<emphasis id="7C8B615E6B2DFF8CE244F8B8FA1CF8B2" box="[1325,1426,1813,1836]" italics="true" pageId="5" pageNumber="320">Potamon</emphasis>
)
<emphasis id="7C8B615E6B2DFF8CE7FEF894FF55F8CE" box="[151,219,1849,1872]" italics="true" pageId="5" pageNumber="320">mistio</emphasis>
<bibRefCitation id="2A6EC0BD6B2DFF8CE78CF89AFE00F8CE" author="Rathbun, M. J." box="[229,398,1847,1872]" pageId="5" pageNumber="320" pagination="225 - 312" refId="ref9767" refString="Rathbun, M. J. (1904) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, 4, 225 - 312, pls. 9 - 18." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
</taxonomicName>
, under
<taxonomicName id="89FFC6CF6B2DFF8CE68FF895FDF9F8CE" authorityName="Husana &amp; Ng" authorityYear="2019" box="[486,631,1847,1872]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2DFF8CE68FF895FDF9F8CE" box="[486,631,1847,1872]" italics="true" pageId="5" pageNumber="320">S. philippina</emphasis>
</taxonomicName>
, although both species are from mountains in the island of Mindanao which is far south from his localities.
<bibRefCitation id="2A6EC0BD6B2DFF8CE53BF8F1FC80F8EA" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[594,782,1883,1908]" pageId="5" pageNumber="320" pagination="1 - 286" refId="ref9531" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="7C8B615E6B2DFF8CE517F8F0FD37F8EA" box="[638,697,1883,1908]" italics="true" pageId="5" pageNumber="320">et al.</emphasis>
(2008)
</bibRefCitation>
treated both species as distinct from
<taxonomicName id="89FFC6CF6B2DFF8CE3ABF8F1FADDF8EA" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1218,1363,1883,1908]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2DFF8CE3ABF8F1FADDF8EA" box="[1218,1363,1883,1908]" italics="true" pageId="5" pageNumber="320">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2DFF8CE237F8F1FF53F806" pageId="5" pageNumber="320" sensu="stricto">sensu stricto</taxonomicNameLabel>
although no reasons were provided (see also
<bibRefCitation id="2A6EC0BD6B2DFF8CE59CF8D2FB93F806" author="Mendoza, J. C. E. &amp; Naruse, T." box="[757,1053,1919,1944]" pageId="5" pageNumber="320" pagination="61 - 70" refId="ref9108" refString="Mendoza, J. C. E. &amp; Naruse, T. (2010) A new species of riverine crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Gecarcinucidae) from northeastern Luzon, Philippines. Philippine Journal of Science, 139 (1), 61 - 70." type="journal article" year="2010">Mendoza &amp; Naruse 2010</bibRefCitation>
).
<bibRefCitation id="2A6EC0BD6B2DFF8CE35AF8D2FAB6F806" author="Mendoza, J. C. E. &amp; Sy, E. Y." box="[1075,1336,1919,1944]" pageId="5" pageNumber="320" pagination="1039 - 1053" refId="ref9164" refString="Mendoza, J. C. E. &amp; Sy, E. Y. (2017) Sundathelphusa miguelito, a new species of freshwater crab from the southern Philippines (Brachyura, Gecarcinucidae). In: Magalhaes, C., Becker, C., Davie, P. J. F., Klimpel, S., Martinez-Arbizu, P. &amp; Sonnewald, M. (Eds.), Special issue dedicated to the memory of Michael Turkay (1948 - 2015). Crustaceana, 90 (7 - 10), pp. 1039 - 1053. https: // doi. org / 10.1163 / 15685403 - 00003638" type="journal article" year="2017">Mendoza &amp; Sy (2017)</bibRefCitation>
recently described a new
<taxonomicName id="89FFC6CF6B2DFF8CE63EF80EFD84F822" authorityName="Bott" authorityYear="1969" box="[343,522,1955,1980]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="7C8B615E6B2DFF8CE63EF80EFD84F822" box="[343,522,1955,1980]" italics="true" pageId="5" pageNumber="320">Sundathelphusa</emphasis>
</taxonomicName>
species from Mindanao and redescribed
<taxonomicName id="89FFC6CF6B2DFF8CE488F809FBCAF822" authorityName="Rathbun" authorityYear="1904" box="[993,1092,1956,1980]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="mistio">
<emphasis id="7C8B615E6B2DFF8CE488F809FBCAF822" box="[993,1092,1956,1980]" italics="true" pageId="5" pageNumber="320">S. mistio</emphasis>
</taxonomicName>
and
<taxonomicName id="89FFC6CF6B2DFF8CE314F809FABCF822" authorityName="Rathbun" authorityYear="1904" box="[1149,1330,1956,1980]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="montanoanus">
<emphasis id="7C8B615E6B2DFF8CE314F809FABCF822" box="[1149,1330,1956,1980]" italics="true" pageId="5" pageNumber="320">S. montanoanus</emphasis>
</taxonomicName>
from the
<typeStatus id="914403EE6B2DFF8CE7FEF865FF5CF87E" box="[151,210,1992,2016]" pageId="5" pageNumber="320">types</typeStatus>
and showed they are distinct taxa.
</paragraph>
<paragraph id="4E40BD4C6B2EFF8FE7AEFF35FE54FD86" blockId="6.[151,1437,151,1904]" pageId="6" pageNumber="321">
<bibRefCitation id="2A6EC0BD6B2EFF8FE7AEFF35FE13FF2E" author="Burger, O." box="[199,413,152,176]" pageId="6" pageNumber="321" pagination="1 - 7" refId="ref8109" refString="Burger, O. (1894) Beitrage zur Kenntniss der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (1), 1 - 7, pl. 1." type="journal article" year="1894">Bürgers (1894: 4)</bibRefCitation>
record of “
<emphasis id="7C8B615E6B2EFF8FE54EFF3AFC87FF2E" box="[551,777,151,176]" italics="true" pageId="6" pageNumber="321">
Telphusa
<taxonomicName id="89FFC6CF6B2EFF8FE5FFFF3AFC87FF2E" authorityName="Husana &amp; Ng" authorityYear="2019" box="[662,777,151,176]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">philippina</taxonomicName>
</emphasis>
” is a subsequent record from von Martens and does not constitute an original description as indicated in
<bibRefCitation id="2A6EC0BD6B2EFF8FE5BBFF10FC05FF4A" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[722,907,188,213]" pageId="6" pageNumber="321" pagination="1 - 286" refId="ref9531" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="7C8B615E6B2EFF8FE596FF10FCB5FF4B" box="[767,827,188,213]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
(2008
</bibRefCitation>
: 73), and so poses no homonymy issues. We examined the extant
<bibRefCitation id="2A6EC0BD6B2EFF8FE6F9FF4DFDBAFF66" author="Burger, O." box="[400,564,224,248]" pageId="6" pageNumber="321" pagination="1 - 7" refId="ref8109" refString="Burger, O. (1894) Beitrage zur Kenntniss der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (1), 1 - 7, pl. 1." type="journal article" year="1894">Bürger (1894)</bibRefCitation>
material in the ZMB; his material from Agno River in Luzon are all
<taxonomicName id="89FFC6CF6B2EFF8FE2EEFF4DFE9CFE83" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="grapsoides">
<emphasis id="7C8B615E6B2EFF8FE2EEFF4DFE9CFE83" italics="true" pageId="6" pageNumber="321">S. grapsoides</emphasis>
</taxonomicName>
. The material he listed from
<collectingRegion id="8C3B73AE6B2EFF8FE518FEA9FD3BFE83" box="[625,693,260,285]" country="Philippines" name="Bohol" pageId="6" pageNumber="321">Bohol</collectingRegion>
was referred to a new species,
<taxonomicName id="89FFC6CF6B2EFF8FE359FEA8FB08FE83" authorityName="Ng &amp; Sket" authorityYear="1996" box="[1072,1158,260,285]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="boex">
<emphasis id="7C8B615E6B2EFF8FE359FEA8FB08FE83" box="[1072,1158,260,285]" italics="true" pageId="6" pageNumber="321">S. boex</emphasis>
</taxonomicName>
, by
<bibRefCitation id="2A6EC0BD6B2EFF8FE3D6FEA8FA16FE83" author="Ng, P. K. L. &amp; Sket, B." box="[1215,1432,260,285]" pageId="6" pageNumber="321" pagination="695 - 706" refId="ref9587" refString="Ng, P. K. L. &amp; Sket, B. (1996) The freshwater crab fauna (Crustacea: Decapoda: Brachyura) of the Philippines. IV. On the collection of Parathelphusidae from Bohol. Proceeding of the Biological Society of Washington, 109, 695 - 706." type="journal article" year="1996">Ng &amp; Sket (1996)</bibRefCitation>
. Although
<bibRefCitation id="2A6EC0BD6B2EFF8FE663FE85FE55FEDE" author="Ng, P. K. L. &amp; Sket, B." box="[266,475,295,320]" pageId="6" pageNumber="321" pagination="695 - 706" refId="ref9587" refString="Ng, P. K. L. &amp; Sket, B. (1996) The freshwater crab fauna (Crustacea: Decapoda: Brachyura) of the Philippines. IV. On the collection of Parathelphusidae from Bohol. Proceeding of the Biological Society of Washington, 109, 695 - 706." type="journal article" year="1996">Ng &amp; Sket (1996)</bibRefCitation>
treated the material from
<collectingRegion id="8C3B73AE6B2EFF8FE465FE8AFCC9FEDE" box="[780,839,295,320]" country="Philippines" name="Cebu" pageId="6" pageNumber="321">Cebu</collectingRegion>
as “
<taxonomicName id="89FFC6CF6B2EFF8FE417FE85FB9EFEDE" authorityName="Husana &amp; Ng" authorityYear="2019" box="[894,1040,295,320]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE417FE85FB9EFEDE" box="[894,1040,295,320]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
”, we now show that they actually belong to a separate species, here described as
<taxonomicName id="89FFC6CF6B2EFF8FE5D7FEE0FC9CFEFB" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[702,786,332,357]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="cebu" status="sp. nov.">
<emphasis id="7C8B615E6B2EFF8FE5D7FEE0FC9CFEFB" box="[702,786,332,357]" italics="true" pageId="6" pageNumber="321">S. cebu</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B2EFF8FE475FEE1FCF7FEFB" bold="true" box="[796,889,332,357]" pageId="6" pageNumber="321">
<taxonomicNameLabel id="67B8DC256B2EFF8FE475FEE1FCF7FEFB" box="[796,889,332,357]" pageId="6" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see discussion later). Bürgers specimen from Mariveles (in Luzon) is actually an undescribed small species, which will be described at a later stage. The specimen figured by him (
<bibRefCitation id="2A6EC0BD6B2EFF8FE6A2FE39FD7DFE32" author="Burger, O." box="[459,755,404,429]" pageId="6" pageNumber="321" pagination="1 - 7" refId="ref8109" refString="Burger, O. (1894) Beitrage zur Kenntniss der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (1), 1 - 7, pl. 1." type="journal article" year="1894">Bürger, 1894: pl. 1 fig. 3</bibRefCitation>
) is clearly not
<taxonomicName id="89FFC6CF6B2EFF8FE4C0FE38FBB5FE33" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[937,1083,404,429]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE4C0FE38FBB5FE33" box="[937,1083,404,429]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2EFF8FE32FFE38FB5AFE33" box="[1094,1236,404,429]" pageId="6" pageNumber="321" sensu="stricto">sensu stricto</taxonomicNameLabel>
as understood at present and resembles the new species from Mariveles. But because he did not indicate the origins of the specimen figured, we are unsure of its identity. As far as we know,
<bibRefCitation id="2A6EC0BD6B2EFF8FE444FE70FC43FE6A" author="Burger, O." box="[813,973,476,501]" pageId="6" pageNumber="321" pagination="1 - 7" refId="ref8109" refString="Burger, O. (1894) Beitrage zur Kenntniss der Gattung Telphusa. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (1), 1 - 7, pl. 1." type="journal article" year="1894">Bürger (1894)</bibRefCitation>
did not have specimens of
<taxonomicName id="89FFC6CF6B2EFF8FE262FE70FA12FE6B" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1291,1436,476,501]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE262FE70FA12FE6B" box="[1291,1436,476,501]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2EFF8FE7FEFDADFEACFD86" box="[151,290,511,536]" pageId="6" pageNumber="321" sensu="stricto">sensu stricto</taxonomicNameLabel>
as defined here.
</paragraph>
<paragraph id="4E40BD4C6B2EFF8FE7AEFD89FD6FFD36" blockId="6.[151,1437,151,1904]" pageId="6" pageNumber="321">
The species named as “
<taxonomicName id="89FFC6CF6B2EFF8FE6BEFD88FC67FDA3" authority="Martens, 1869" authorityName="Martens" authorityYear="1869" box="[471,1001,548,573]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina" subGenus="Telphusa">
<emphasis id="7C8B615E6B2EFF8FE6BEFD88FDB2FDA2" box="[471,572,549,572]" italics="true" pageId="6" pageNumber="321">Potamon</emphasis>
(
<emphasis id="7C8B615E6B2EFF8FE527FD89FD3CFDA3" box="[590,690,548,573]" italics="true" pageId="6" pageNumber="321">Telphusa</emphasis>
)
<emphasis id="7C8B615E6B2EFF8FE5ACFD89FCB6FDA3" box="[709,824,548,573]" italics="true" pageId="6" pageNumber="321">philippina</emphasis>
Martens, 1869
</taxonomicName>
”, in
<bibRefCitation id="2A6EC0BD6B2EFF8FE348FD89FA80FDA3" author="Estampador, E. P." box="[1057,1294,548,573]" pageId="6" pageNumber="321" pagination="1 - 125" refId="ref8451" refString="Estampador, E. P. (1959) Revised check list of Philippine crustacean decapods. Natural and Applied Science Bulletin, 17, 1 - 125." type="journal article" year="1959">Estampadors (1959)</bibRefCitation>
checklist of the decapod crustaceans of the
<collectingCountry id="36E8FDDC6B2EFF8FE561FDEAFD08FDFE" box="[520,646,583,608]" name="Philippines" pageId="6" pageNumber="321">Philippines</collectingCountry>
is definitely not
<taxonomicName id="89FFC6CF6B2EFF8FE43AFDE5FC68FDFE" authorityName="Martens" authorityYear="1869" box="[851,998,583,608]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE43AFDE5FC68FDFE" box="[851,998,583,608]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
because his localities were all in the northern region of Luzon Island including Camiguin, an islet north of Luzon, not to be confused with the more famous Camiguin island located north of Mindanao.
</paragraph>
<paragraph id="4E40BD4C6B2EFF8FE7AEFD18FE78FC8B" blockId="6.[151,1437,151,1904]" pageId="6" pageNumber="321">
In addition, we have specimens from
<collectingRegion id="8C3B73AE6B2EFF8FE517FD19FCCAFD53" box="[638,836,692,717]" country="Philippines" name="Quirino" pageId="6" pageNumber="321">Quirino province</collectingRegion>
, in northern Luzon, which superficially resemble
<taxonomicName id="89FFC6CF6B2EFF8FE2EEFD18FE84FD6E" authorityName="Martens" authorityYear="1869" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE2EEFD18FE84FD6E" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
but are distinct in their carapace and ambulatory morphology as well as G1 characters. They are here described as
<taxonomicName id="89FFC6CF6B2EFF8FE64EFD50FE18FC8A" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[295,406,765,788]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="quirino" status="sp. nov.">
<emphasis id="7C8B615E6B2EFF8FE64EFD50FE18FC8A" box="[295,406,765,788]" italics="true" pageId="6" pageNumber="321">S. quirino</emphasis>
</taxonomicName>
<emphasis id="7C8B615E6B2EFF8FE6F4FD51FE78FC8B" bold="true" box="[413,502,764,789]" pageId="6" pageNumber="321">
<taxonomicNameLabel id="67B8DC256B2EFF8FE6F4FD51FE78FC8B" box="[413,502,764,789]" pageId="6" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="4E40BD4C6B2EFF8FE7AEFCB2FD9FFB3E" blockId="6.[151,1437,151,1904]" pageId="6" pageNumber="321">
The good series of specimens of
<taxonomicName id="89FFC6CF6B2EFF8FE538FC8DFD6BFCA6" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[593,741,799,824]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE538FC8DFD6BFCA6" box="[593,741,799,824]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2EFF8FE599FC8DFC0EFCA6" box="[752,896,799,824]" pageId="6" pageNumber="321" sensu="stricto">sensu stricto</taxonomicNameLabel>
from Samar Island allow us to determine the variation in this species. Significant allometric changes are often observed with increasing size, in particular with the form of the carapace. In
<taxonomicName id="89FFC6CF6B2EFF8FE6BCFCC5FDEBFC1E" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[469,613,871,896]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE6BCFCC5FDEBFC1E" box="[469,613,871,896]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2EFF8FE505FCC5FD75FC1E" box="[620,763,871,896]" pageId="6" pageNumber="321" sensu="stricto">sensu stricto</taxonomicNameLabel>
, smaller specimens (e.g., 14.1 ×
<quantity id="890710A96B2EFF8FE305FCC5FB5CFCE1" box="[1132,1234,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="6" pageNumber="321" unit="mm" value="12.3">12.3 mm</quantity>
, ZRC 2016.0107) have a subquadrate carapace with the regions of the dorsal surface (notably the branchial region) relatively less convex and the surface more rugose (
<figureCitation id="D6C4A1C96B2EFF8FE508FC02FD4CFC56" box="[609,706,943,968]" captionStart="FIGURE 3" captionStartId="5.[151,250,1509,1531]" captionTargetBox="[150,1436,171,1487]" captionTargetId="figure@5.[150,1436,171,1488]" captionTargetPageId="5" captionText="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale." httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="6" pageNumber="321">Fig. 3A</figureCitation>
). The G1s of these smaller males, although appearing to be functional, are also usually relatively straighter, with the distal half less distinctly curved (
<figureCitation id="D6C4A1C96B2EFF8FE3FBFC79FB65FC72" box="[1170,1259,980,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1509,1531]" captionTargetBox="[150,1436,171,1487]" captionTargetId="figure@5.[150,1436,171,1488]" captionTargetPageId="5" captionText="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale." httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="6" pageNumber="321">Fig. 3B</figureCitation>
). As specimens get larger (e.g., 30.8 ×
<quantity id="890710A96B2EFF8FE6FFFC5AFD84FB91" box="[406,522,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="2.475" pageId="6" pageNumber="321" unit="mm" value="24.75">24.75 mm</quantity>
, ZRC 2016.0107), the anterolateral and branchial regions gradually become more inflated with the surface smoother, and the carapace becomes more transversely ovate with the posterolateral margins more prominently convergent (
<figureCitation id="D6C4A1C96B2EFF8FE537FB92FD35FBC6" box="[606,699,1087,1112]" captionStart="FIGURE 3" captionStartId="5.[151,250,1509,1531]" captionTargetBox="[150,1436,171,1487]" captionTargetId="figure@5.[150,1436,171,1488]" captionTargetPageId="5" captionText="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale." httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="6" pageNumber="321">Fig. 3A</figureCitation>
). In these larger males, the G1 is also relatively stouter and the distal half becomes prominently curved (
<figureCitation id="D6C4A1C96B2EFF8FE519FBC9FD45FBE2" box="[624,715,1124,1149]" captionStart="FIGURE 3" captionStartId="5.[151,250,1509,1531]" captionTargetBox="[150,1436,171,1487]" captionTargetId="figure@5.[150,1436,171,1488]" captionTargetPageId="5" captionText="FIGURE 3. Sundathelphusa philippina (von Martens, 1868), variation in male carapace and G1 form. A, A, 14.1 × 12.3 mm; B, 16.9 × 13.9 mm; C, 18.0 × 15.0 mm; D, D, 20.3 × 16.7 mm; E, E, 30.8 × 24.8 mm [all ZRC 2016.0107, Samar]. AE, overall habitus; A, D, E, left G1. AE same scale; A, D, E same scale." httpUri="https://zenodo.org/record/2637372/files/figure.png" pageId="6" pageNumber="321">Fig. 3B</figureCitation>
). The changes in carapace morphology are also evident in the female specimens examined here.
</paragraph>
<paragraph id="4E40BD4C6B2EFF8FE7AEFB01FE4DF9B2" blockId="6.[151,1437,151,1904]" pageId="6" pageNumber="321">
The presently observed size-related changes in carapace shape, surface morphology and G1 structure indicate that
<taxonomicName id="89FFC6CF6B2EFF8FE7A5FB62FC50FB76" authority="Stasolla, Abbarchi &amp; Innocenti, 2015" authorityName="Stasolla, Abbarchi &amp; Innocenti" authorityYear="2015" box="[204,990,1231,1256]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="spelaeophila">
<emphasis id="7C8B615E6B2EFF8FE7A5FB62FD94FB76" box="[204,538,1231,1256]" italics="true" pageId="6" pageNumber="321">Sundathelphusa spelaeophila</emphasis>
<bibRefCitation id="2A6EC0BD6B2EFF8FE54FFB62FC50FB76" author="Stasolla, G. &amp; Abbarchi, A. &amp; Innocenti, G." box="[550,990,1231,1256]" pageId="6" pageNumber="321" pagination="448 - 453" refId="ref9800" refString="Stasolla, G., Abbarchi, A. &amp; Innocenti, G. (2015) Sundathelphusa spelaeophila, a new species of cavernicolous crab from Samar, Philippines (Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 63, 448 - 453." type="journal article" year="2015">Stasolla, Abbarchi &amp; Innocenti, 2015</bibRefCitation>
</taxonomicName>
, is a junior subjective synonym of
<taxonomicName id="89FFC6CF6B2EFF8FE2EEFB7DFE84FA93" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE2EEFB7DFE84FA93" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2EFF8FE678FB58FE10FA93" box="[273,414,1268,1293]" pageId="6" pageNumber="321" sensu="stricto">sensu stricto</taxonomicNameLabel>
. The former species was described from four relatively small males (
<typeStatus id="914403EE6B2EFF8FE3C9FB59FA88FA93" box="[1184,1286,1268,1293]" pageId="6" pageNumber="321" type="holotype">holotype</typeStatus>
, 15.9 by
<quantity id="890710A96B2EFF8FE202FB59FF4BFAAE" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="6" pageNumber="321" unit="mm" value="14.1">14.1 mm</quantity>
, MZUF 3920;
<typeStatus id="914403EE6B2EFF8FE603FAB4FE56FAAF" box="[362,472,1305,1329]" pageId="6" pageNumber="321" type="paratype">paratypes</typeStatus>
, 11.7 ×
<quantity id="890710A96B2EFF8FE559FABAFD1AFAAE" box="[560,660,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="6" pageNumber="321" unit="mm" value="10.6">10.6 mm</quantity>
MZUF 4273, 15.4 ×
<quantity id="890710A96B2EFF8FE4E8FAB5FC6BFAAE" box="[897,997,1304,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="6" pageNumber="321" unit="mm" value="12.9">12.9 mm</quantity>
NMCR 40102, 16.8 ×
<quantity id="890710A96B2EFF8FE38CFAB5FAC7FAAE" box="[1253,1353,1304,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="6" pageNumber="321" unit="mm" value="14.7">14.7 mm</quantity>
MZUF 3927) from Can Gortio Cave and “SNAZ 1” Cave, Barangay Matalud, San Jorge municipality,
<collectingRegion id="8C3B73AE6B2EFF8FE38BFA91FA16FACB" box="[1250,1432,1340,1365]" country="Philippines" name="Samar" pageId="6" pageNumber="321">Samar province</collectingRegion>
,
<collectingCountry id="36E8FDDC6B2EFF8FE7FEFAF2FE97FAE6" box="[151,281,1375,1400]" name="Philippines" pageId="6" pageNumber="321">Philippines</collectingCountry>
. The type locality of
<taxonomicName id="89FFC6CF6B2EFF8FE56AFACDFD23FAE6" authorityName="Stasolla, Abbarchi &amp; Innocenti" authorityYear="2015" box="[515,685,1375,1400]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="spelaeophila">
<emphasis id="7C8B615E6B2EFF8FE56AFACDFD23FAE6" box="[515,685,1375,1400]" italics="true" pageId="6" pageNumber="321">S. spelaeophila</emphasis>
</taxonomicName>
is actually only about 15 kilometers north of Lobo Cave where one of the specimens of
<taxonomicName id="89FFC6CF6B2EFF8FE6EAFA28FD9AFA03" authorityName="Husana &amp; Ng" authorityYear="2019" box="[387,532,1412,1437]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE6EAFA28FD9AFA03" box="[387,532,1412,1437]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
(ZRC 2017.1062) was collected. We also obtained specimens of
<taxonomicName id="89FFC6CF6B2EFF8FE263FA28FA12FA03" authority="Husana &amp; Ng, 2019" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1290,1436,1412,1437]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" sensu="stricto" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE263FA28FA12FA03" box="[1290,1436,1412,1437]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67B8DC256B2EFF8FE7FEFA04FEADFA5E" box="[151,291,1447,1472]" pageId="6" pageNumber="321" sensu="stricto">sensu stricto</taxonomicNameLabel>
from the shallow part of the caves (Lobo and Daram Caves), similar to the habitat of
<taxonomicName id="89FFC6CF6B2EFF8FE399FA05FA12FA5E" authorityName="Stasolla, Abbarchi &amp; Innocenti" authorityYear="2015" box="[1264,1436,1447,1472]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="spelaeophila">
<emphasis id="7C8B615E6B2EFF8FE399FA05FA12FA5E" box="[1264,1436,1447,1472]" italics="true" pageId="6" pageNumber="321">S. spelaeophila</emphasis>
</taxonomicName>
reported by
<bibRefCitation id="2A6EC0BD6B2EFF8FE674FA61FD89FA7B" author="Stasolla, G. &amp; Abbarchi, A. &amp; Innocenti, G." box="[285,519,1484,1509]" pageId="6" pageNumber="321" pagination="448 - 453" refId="ref9800" refString="Stasolla, G., Abbarchi, A. &amp; Innocenti, G. (2015) Sundathelphusa spelaeophila, a new species of cavernicolous crab from Samar, Philippines (Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 63, 448 - 453." type="journal article" year="2015">
Stasolla
<emphasis id="7C8B615E6B2EFF8FE612FA60FE3DFA7B" box="[379,435,1484,1509]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
(2015)
</bibRefCitation>
. The map for the type locality of the
<taxonomicName id="89FFC6CF6B2EFF8FE4C8FA60FBC5FA7B" authorityName="Stasolla, Abbarchi &amp; Innocenti" authorityYear="2015" box="[929,1099,1484,1509]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="spelaeophila">
<emphasis id="7C8B615E6B2EFF8FE4C8FA60FBC5FA7B" box="[929,1099,1484,1509]" italics="true" pageId="6" pageNumber="321">S. spelaeophila</emphasis>
</taxonomicName>
has an inaccuracy; the correct location of Lobo cave is actually that indicated as Langun cave on their map (
<bibRefCitation id="2A6EC0BD6B2EFF8FE343FA42FA81F996" author="Stasolla, G. &amp; Abbarchi, A. &amp; Innocenti, G." box="[1066,1295,1519,1544]" pageId="6" pageNumber="321" pagination="448 - 453" refId="ref9800" refString="Stasolla, G., Abbarchi, A. &amp; Innocenti, G. (2015) Sundathelphusa spelaeophila, a new species of cavernicolous crab from Samar, Philippines (Decapoda: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 63, 448 - 453." type="journal article" year="2015">
Stasolla
<emphasis id="7C8B615E6B2EFF8FE3E4FA5CFB47F996" box="[1165,1225,1519,1544]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
2015
</bibRefCitation>
: fig. 3) (see
<bibRefCitation id="2A6EC0BD6B2EFF8FE7FEF9B8FEE0F9B2" author="Husana, D. E. M. &amp; Naruse, T. &amp; Kase, T." box="[151,366,1556,1581]" pageId="6" pageNumber="321" pagination="419 - 427" refId="ref8622" refString="Husana, D. E. M., Naruse, T. &amp; Kase, T. (2009) Two new cavernicolous species of the genus Sundathelphusa from Western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology, 29 (3), 419 - 427. https: // doi. org / 10.1651 / 08 - 3081.1" type="journal article" year="2009">
Husana
<emphasis id="7C8B615E6B2EFF8FE79BF9B8FEA4F9B3" box="[242,298,1556,1581]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
2009
</bibRefCitation>
: fig. 1).
</paragraph>
<paragraph id="4E40BD4C6B2EFF8FE7AEF99AFDA0F8EE" blockId="6.[151,1437,151,1904]" pageId="6" pageNumber="321">
The presence of
<taxonomicName id="89FFC6CF6B2EFF8FE6FAF995FDA6F9CE" authorityName="Husana &amp; Ng" authorityYear="2019" box="[403,552,1591,1616]" class="Malacostraca" family="Potamidae" genus="Potamon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE6FAF995FDA6F9CE" box="[403,552,1591,1616]" italics="true" pageId="6" pageNumber="321">S. philippina</emphasis>
</taxonomicName>
in caves is not significant as the species is clearly not a stygobiont, not possessing any of the attributes associated with such fauna (see
<bibRefCitation id="2A6EC0BD6B2EFF8FE4C2F9F1FBDBF9EB" author="Holthuis, L. B." box="[939,1109,1628,1653]" pageId="6" pageNumber="321" pagination="589 - 615" refId="ref8561" refString="Holthuis, L. B. (1986) Decapoda. In: Botosaneanu, L. (Ed.), Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting Subterranean Waters (Including the Marine Interstitial). E. J. Brill, Leiden, pp. 589 - 615." type="book chapter" year="1986">Holthuis 1986</bibRefCitation>
;
<bibRefCitation id="2A6EC0BD6B2EFF8FE301F9F1FB71F9EA" author="Guinot, D." box="[1128,1279,1628,1653]" pageId="6" pageNumber="321" pagination="3 - 40" refId="ref8480" refString="Guinot, D. (1988) Les crabes cavernicoles du monde. Memoires de Biospeologie, 15, 3 - 40." type="journal article" year="1988">Guinot 1988</bibRefCitation>
,
<bibRefCitation id="2A6EC0BD6B2EFF8FE278F9F1FAC3F9EA" author="Guinot, D." box="[1297,1357,1628,1652]" pageId="6" pageNumber="321" pagination="165 - 179" refId="ref8503" refString="Guinot, D. (1994) Decapoda Brachyura. In: Juberthie, C. &amp; Decou, V. (Eds.), Encyclopaedia Biospeologica. Tome 1. Societe de Biospeologie, CNRS, Moulis, Ariege, and Academie Roumaine, Bucarest, pp. 165 - 179, pl. 3." type="journal article" year="1994">1994</bibRefCitation>
). True stygobiont species of
<taxonomicName id="89FFC6CF6B2EFF8FE6E5F9D2FDB1F906" authorityName="Bott" authorityYear="1969" box="[396,575,1663,1688]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="7C8B615E6B2EFF8FE6E5F9D2FDB1F906" box="[396,575,1663,1688]" italics="true" pageId="6" pageNumber="321">Sundathelphusa</emphasis>
</taxonomicName>
with loss of carapace pigmentation, reduced eyes and elongated ambulatory legs are known from the
<collectingCountry id="36E8FDDC6B2EFF8FE6D1F909FDB8F923" box="[440,566,1700,1725]" name="Philippines" pageId="6" pageNumber="321">Philippines</collectingCountry>
(see
<bibRefCitation id="2A6EC0BD6B2EFF8FE51CF908FCC1F922" author="Husana, D. E. M. &amp; Naruse, T. &amp; Kase, T." box="[629,847,1700,1725]" pageId="6" pageNumber="321" pagination="419 - 427" refId="ref8622" refString="Husana, D. E. M., Naruse, T. &amp; Kase, T. (2009) Two new cavernicolous species of the genus Sundathelphusa from Western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology, 29 (3), 419 - 427. https: // doi. org / 10.1651 / 08 - 3081.1" type="journal article" year="2009">
Husana
<emphasis id="7C8B615E6B2EFF8FE5B8F908FC82F923" box="[721,780,1700,1725]" italics="true" pageId="6" pageNumber="321">et al.</emphasis>
2009
</bibRefCitation>
;
<bibRefCitation id="2A6EC0BD6B2EFF8FE434F909FBB2F922" author="Takeda, M. &amp; Ng, P. K. L." box="[861,1084,1700,1725]" pageId="6" pageNumber="321" pagination="1123 - 1127" refId="ref9910" refString="Takeda, M. &amp; Ng, P. K. L. (2001) The freshwater crab fauna (Crustacea, Brachyura) of the Philippines. VI. A new cavernicolous crab from Mindanao. Zoological Science, 18 (8), 1123 - 1127. https: // doi. org / 10.2108 / zsj. 18.1123" type="journal article" year="2001">Takeda &amp; Ng 2001</bibRefCitation>
).
<taxonomicName id="89FFC6CF6B2EFF8FE338F909FA0EF923" authorityName="Husana &amp; Ng" authorityYear="2019" box="[1105,1408,1700,1725]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="7C8B615E6B2EFF8FE338F909FA0EF923" box="[1105,1408,1700,1725]" italics="true" pageId="6" pageNumber="321">Sundathelphusa philippina</emphasis>
</taxonomicName>
is clearly an epigeal species that roams around surface streams at night and just occasionally wanders into cave habitats for shelter during the day. From our extensive field collections and investigations, we observed that the habitat range of this species is from the shallow part of the caves in karstic areas to small surface streams with stagnant to slow-moving freshwater. They mainly stay under the submerged rocks, logs or leaves in the water and only come out in the open to forage.
</paragraph>
</subSubSection>
</treatment>
</document>