treatments-xml/data/A7/1E/87/A71E87EE8B72FFCCDDF8C58EFB9CFCAE.xml

<document id="2EE9D2AC9DDEC18D63B574BD7E44FB57" ID-CLB-Dataset="29802" ID-DOI="10.11646/zootaxa.4442.1.3" ID-GBIF-Dataset="51315b1a-dcc4-41e2-aff7-f90478fed976" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301390" ID-ZooBank="973323F0-BA57-4ACE-9DB6-0C0237B2E487" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530293601868" checkinUser="plazi" docAuthor="Attasopa, Korrawat, Bänziger, Hans, Disayathanoowat, Terd &amp; Packer, Laurence" docDate="2018" docId="A71E87EE8B72FFCCDDF8C58EFB9CFCAE" docLanguage="en" docName="zootaxa.4442.1.3.pdf" docOrigin="Zootaxa 4442 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidotrigona satun Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="74" masterDocId="5B27FF968B71FFC7DD6FC678FF8BFFBA" masterDocTitle="A new species of Lepidotrigona (Hymenoptera: Apidae) from Thailand with the description of males of L. flavibasis and L. doipaensis and comments on asymmetrical genitalia in bees" masterLastPageNumber="82" masterPageNumber="63" pageNumber="66" updateTime="1698551722399" updateUser="ExternalLinkService">
<mods:mods id="5391FC471BF23276EF5D9823D31A6C94" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="61145EA5686FD393F68BECD2967F596E">
<mods:title id="2A6E57936543CB1A8E838DB8E163C8B5">A new species of Lepidotrigona (Hymenoptera: Apidae) from Thailand with the description of males of L. flavibasis and L. doipaensis and comments on asymmetrical genitalia in bees</mods:title>
</mods:titleInfo>
<mods:name id="4A366792FAD984B2E02F8B306397548E" type="personal">
<mods:role id="FFEEE5350B494F2A44C48427372C4090">
<mods:roleTerm id="14F1110870EEB3CAF9A13A5E6F2C52D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="162020E1529C3DE7214104E6D5869FA3">Attasopa, Korrawat</mods:namePart>
</mods:name>
<mods:name id="EAB313AA64E9CDB7F642A2042D4BE024" type="personal">
<mods:role id="B3F26128654D3D8EECD71010178E1BE8">
<mods:roleTerm id="EC99E33B390E14F5FE1EC2FC6A866650">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F236A8999F30415B8030C2A563CC3064">Bänziger, Hans</mods:namePart>
</mods:name>
<mods:name id="7DD2BB1251E7503EBCF3FC079FD13845" type="personal">
<mods:role id="149A45C7C941730EF0F955FA29DE6F75">
<mods:roleTerm id="2473F1F58E09A9367AEBE828494249AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5398C3353116C8F6CB5C093DE97EC5E2">Disayathanoowat, Terd</mods:namePart>
</mods:name>
<mods:name id="85BD3654887CC4141403C7A19C992BBE" type="personal">
<mods:role id="3C52E824C96C717CCCA1C6293FD9E7B3">
<mods:roleTerm id="42D893EB72A75954E33064973CA32857">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B6FCB49C0C67D34D40C91D93836DFCD">Packer, Laurence</mods:namePart>
</mods:name>
<mods:typeOfResource id="BB0563C792DCDA67F634A457C6709456">text</mods:typeOfResource>
<mods:relatedItem id="A7DC38613B197077FEE71AC25CF7FB4F" type="host">
<mods:titleInfo id="F83D87B2B474EA6D1F21363F5557B23E">
<mods:title id="2292E9AD1B98A57924CAF4CD45EF89B4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4B4B91F8167EB3F2B89A114E5AD8BE1A">
<mods:date id="E03D56520B8AA7E6290A5496A1042ABB">2018</mods:date>
<mods:detail id="5C58BAA2A60264A831C2B155674BFBA3" type="pubDate">
<mods:number id="D1ED169A2B3BF308D66C5AD4F22CA311">2018-06-29</mods:number>
</mods:detail>
<mods:detail id="B4E8C30390DA5832B6E044CDC064DFB0" type="volume">
<mods:number id="61A51F1E19FABB6539FD40DD87BA70C0">4442</mods:number>
</mods:detail>
<mods:detail id="084F28B77E1CBAE68BB73C51E28CADE6" type="issue">
<mods:number id="F7EE53CF4231ACF2B225F7556006985F">1</mods:number>
</mods:detail>
<mods:extent id="BF9748FBF91EAD9AEC93ECBE4A0F23AC" unit="page">
<mods:start id="3F5933E4968FC6B826EF07FCFD5B640E">63</mods:start>
<mods:end id="8DDF5573DC0333758B0A34F1211C0962">82</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8AEF9FAF2D46BD0B2085CBA4654C0D2A">journal article</mods:classification>
<mods:identifier id="12243B4AC6EA72C0FB6E5830AF99C2DF" type="CLB-Dataset">29802</mods:identifier>
<mods:identifier id="049FF956665B8B3DB0D84547C84B89D1" type="DOI">10.11646/zootaxa.4442.1.3</mods:identifier>
<mods:identifier id="CB89DDC0F0A511C1A7F77E35A9CC3833" type="GBIF-Dataset">51315b1a-dcc4-41e2-aff7-f90478fed976</mods:identifier>
<mods:identifier id="889A216A8A0E1DD9658A33448E9EF721" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E50D4513DE0CB8941EA089DABADCAFAA" type="Zenodo-Dep">1301390</mods:identifier>
<mods:identifier id="52A510650C513E5FB6FE27C0C7A95941" type="ZooBank">973323F0-BA57-4ACE-9DB6-0C0237B2E487</mods:identifier>
</mods:mods>
<treatment id="A71E87EE8B72FFCCDDF8C58EFB9CFCAE" ID-DOI="http://doi.org/10.5281/zenodo.5977703" ID-GBIF-Taxon="146043611" ID-Zenodo-Dep="5977703" LSID="urn:lsid:plazi:treatment:A71E87EE8B72FFCCDDF8C58EFB9CFCAE" httpUri="http://treatment.plazi.org/id/A71E87EE8B72FFCCDDF8C58EFB9CFCAE" lastPageId="11" lastPageNumber="74" pageId="3" pageNumber="66">
<subSubSection id="67AD65738B72FFC4DDF8C58EFE70FB88" pageId="3" pageNumber="66" type="nomenclature">
<paragraph id="2F0836F88B72FFC4DDF8C58EFD70FBAA" blockId="3.[151,763,1014,1074]" box="[151,763,1014,1040]" pageId="3" pageNumber="66">
<heading id="744081948B72FFC4DDF8C58EFD70FBAA" bold="true" box="[151,763,1014,1040]" fontSize="11" level="1" pageId="3" pageNumber="66" reason="1">
<emphasis id="1DC3EAEA8B72FFC4DDF8C58EFD70FBAA" bold="true" box="[151,763,1014,1040]" pageId="3" pageNumber="66">
<taxonomicName id="E8B74D7B8B72FFC4DDF8C58EFD3AFBAA" ID-CoL="3TDY3" authority="Attasopa and Banziger" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[151,689,1014,1040]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B72FFC4DDF8C58EFE07FBAA" bold="true" box="[151,396,1014,1040]" italics="true" pageId="3" pageNumber="66">Lepidotrigona satun</emphasis>
Attasopa and Bänziger
</taxonomicName>
<taxonomicNameLabel id="06F057918B72FFC4DFD6C58EFD70FBAA" box="[697,763,1014,1040]" pageId="3" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2F0836F88B72FFC4DDF8C261FE70FB88" blockId="3.[151,763,1014,1074]" box="[151,507,1049,1074]" pageId="3" pageNumber="66">
<figureCitation id="B78C2A7D8B72FFC4DDF8C261FF6DFB88" box="[151,230,1049,1074]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Figs. 2</figureCitation>
: A1–4, 3: A1–A4 &amp; 4–6
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B72FFC3DDF8C219FD12FEA6" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66" type="diagnosis">
<paragraph id="2F0836F88B72FFC4DDF8C219FAA4FA58" blockId="3.[151,1437,1121,2010]" pageId="3" pageNumber="66">
<emphasis id="1DC3EAEA8B72FFC4DDF8C219FE9AFBC0" bold="true" box="[151,273,1121,1146]" pageId="3" pageNumber="66">Diagnosis.</emphasis>
<taxonomicName id="E8B74D7B8B72FFC4DC75C219FD8AFBC3" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[282,513,1121,1146]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DC75C219FD8AFBC3" box="[282,513,1121,1146]" italics="true" pageId="3" pageNumber="66">Lepidotrigona satun</emphasis>
</taxonomicName>
is a member of the “ 
<taxonomicName id="E8B74D7B8B72FFC4DF96C219FCD0FBC0" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[761,859,1121,1146]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="1DC3EAEA8B72FFC4DF96C219FCD0FBC0" box="[761,859,1121,1146]" italics="true" pageId="3" pageNumber="66">ventralis</emphasis>
</taxonomicName>
” species group based primarily on size: body and forewing length each less than 
<quantity id="E84F9B1D8B72FFC4DF67C2FDFDDBFB27" box="[520,592,1157,1182]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="66" unit="mm" value="5.0">5 mm</quantity>
. It is the only species in the group known from the lower peninsula of 
<collectingCountry id="57A076688B72FFC4DDF8C2D1FF75FB78" box="[151,254,1193,1218]" name="Thailand" pageId="3" pageNumber="66">Thailand</collectingCountry>
. Males can be differentiated from those of the other two species of the “ 
<taxonomicName id="E8B74D7B8B72FFC4D929C2D1FB23FB78" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1094,1192,1193,1218]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="1DC3EAEA8B72FFC4D929C2D1FB23FB78" box="[1094,1192,1193,1218]" italics="true" pageId="3" pageNumber="66">ventralis</emphasis>
</taxonomicName>
” group confirmed as occurring in 
<collectingCountry id="57A076688B72FFC4DC42C2B5FE1AFB5C" box="[301,401,1229,1254]" name="Thailand" pageId="3" pageNumber="66">Thailand</collectingCountry>
(
<taxonomicName id="E8B74D7B8B72FFC4DCCAC2B6FDBAFB5C" box="[421,561,1229,1254]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4DCCAC2B6FDBAFB5C" box="[421,561,1229,1254]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E8B74D7B8B72FFC4DF00C2B6FC83FB5C" box="[623,776,1229,1254]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4DF00C2B6FC83FB5C" box="[623,776,1229,1254]" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
) based upon external morphology as follows: margin of mesoscutum of 
<taxonomicName id="E8B74D7B8B72FFC4DC26C28AFE2FFAB3" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[329,420,1266,1289]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DC26C28AFE2FFAB3" box="[329,420,1266,1289]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
with plumose, scale-like, yellow hairs (
<figureCitation id="B78C2A7D8B72FFC4DE0AC289FC27FAB0" box="[869,940,1265,1290]" captionStart="FIGURE 4" captionStartId="7.[151,250,1999,2021]" captionTargetBox="[185,1409,193,1978]" captionTargetId="figure@7.[185,1409,193,1978]" captionTargetPageId="7" captionText="FIGURE 4. Lepidotrigona satun n. sp., dorsal habitus. (A) male holotype and (B) worker paratype." httpUri="https://zenodo.org/record/1301398/files/figure.png" pageId="3" pageNumber="66">Fig. 4</figureCitation>
: A) (no such hairs in the other two species). S4 of 
<taxonomicName id="E8B74D7B8B72FFC4DDB1C36EFEB1FA97" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[222,314,1302,1325]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DDB1C36EFEB1FA97" box="[222,314,1302,1325]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
is angularly emarginate apicomedially (
<figureCitation id="B78C2A7D8B72FFC4DE67C36DFCDBFA94" box="[776,848,1301,1326]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: A1) (convex medially and bisinuate laterally in 
<taxonomicName id="E8B74D7B8B72FFC4D8EAC36EFE89FAE8" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4D8EAC36EFE89FAE8" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
, slightly concave in 
<taxonomicName id="E8B74D7B8B72FFC4DC86C342FDF5FAE8" box="[489,638,1337,1362]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4DC86C342FDF5FAE8" box="[489,638,1337,1362]" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
). The apicosubmedial lobes of S 
<quantity id="E84F9B1D8B72FFC4DE83C341FB9CFAE8" box="[1004,1047,1337,1362]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="66" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="E8B74D7B8B72FFC4D971C342FBF3FAEB" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1054,1144,1338,1361]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4D971C342FBF3FAEB" box="[1054,1144,1338,1361]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
are apically rounded each with 4–7 thick, long setae (
<figureCitation id="B78C2A7D8B72FFC4DCABC325FD81FACC" box="[452,522,1373,1398]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: A2) (the lobes are pointed in the other two species and bear only 1–2 setae which are very short in 
<taxonomicName id="E8B74D7B8B72FFC4DC0BC3FAFE7BFA20" box="[356,496,1409,1434]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4DC0BC3FAFE7BFA20" box="[356,496,1409,1434]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B72FFC4DF6CC3F9FDC7FA20" box="[515,588,1409,1434]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: B2) or with one very long and, if present, a second much shorter in 
<taxonomicName id="E8B74D7B8B72FFC4D8EAC3FAFE84FA04" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4D8EAC3FAFE84FA04" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B72FFC4DC71C3DDFEEFFA04" box="[286,356,1445,1470]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: C2)). S5 gradulus does not touch the antecosta in 
<taxonomicName id="E8B74D7B8B72FFC4DECDC3DEFC77FA07" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[930,1020,1446,1469]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DECDC3DEFC77FA07" box="[930,1020,1446,1469]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B72FFC4D963C3DDFBD9FA04" box="[1036,1106,1445,1470]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: A2) whereas it does touch it in the other two species, briefly in 
<taxonomicName id="E8B74D7B8B72FFC4DF73C3B2FD2FFA58" box="[540,676,1481,1506]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4DF73C3B2FD2FFA58" box="[540,676,1481,1506]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B72FFC4DFDBC3B1FD72FA58" box="[692,761,1481,1506]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: B2) and extensively in 
<taxonomicName id="E8B74D7B8B72FFC4D966C3B2FB15FA58" box="[1033,1182,1481,1506]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4D966C3B2FB15FA58" box="[1033,1182,1481,1506]" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B72FFC4D9C2C3B1FB79FA58" box="[1197,1266,1481,1506]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
: C2).
</paragraph>
<paragraph id="2F0836F88B72FFC4DDA8C395FB80F900" blockId="3.[151,1437,1121,2010]" pageId="3" pageNumber="66">
Males can also be differentiated from those of these other two species because of the very short malar space of 
<taxonomicName id="E8B74D7B8B72FFC4DDF8C06AFF7AF993" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[151,241,1554,1577]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DDF8C06AFF7AF993" box="[151,241,1554,1577]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
, at most 0.16 times as long as the length of the first flagellomere whereas in the other two species it is at least 0.27 times as long as the first flagellomere. The ratio of the forewing diagonal to the length of the 2nd submarginal cell also varies among the three: it is largest (ḵ2.3) in 
<taxonomicName id="E8B74D7B8B72FFC4DEC4C022FB83F9CB" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[939,1032,1626,1649]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DEC4C022FB83F9CB" box="[939,1032,1626,1649]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
, but at most 
<quantity id="E84F9B1D8B72FFC4D9C8C022FB7CF9C8" box="[1191,1271,1625,1650]" metricMagnitude="-2" metricUnit="m" metricValue="5.3594" pageId="3" pageNumber="66" unit="in" value="2.11">2.11 in</quantity>
the other two species (shortest - 1.94 to 2.05 - in 
<taxonomicName id="E8B74D7B8B72FFC4DF41C006FD4EF92C" box="[558,709,1661,1686]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4DF41C006FD4EF92C" box="[558,709,1661,1686]" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
and intermediate - 2.05 to 2.11 - in 
<taxonomicName id="E8B74D7B8B72FFC4D901C006FB71F92C" box="[1134,1274,1661,1686]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4D901C006FB71F92C" box="[1134,1274,1661,1686]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
) (see 
<tableCitation id="623503438B72FFC4D853C005FA01F92C" box="[1340,1418,1661,1686]" captionStart="TABLE 1" captionStartId="16.[151,239,1012,1035]" captionText="TABLE 1. Measurement of malar space length (MSL), F1 length (F1L), metatibia width (MTW), forewing diagonal (FWD), and 2nd submarginal cell (SM2L) ratios that separate L. satun n. sp., L. flavibasis, and L. doipaensis, expressed as range (Average±SD). Holotype ratios in bold." pageId="3" pageNumber="66">Table 1</tableCitation>
). The genitalia are also very different in the three species (see below and 
<figureCitation id="B78C2A7D8B72FFC4DED6C0D9FC76F900" box="[953,1021,1697,1722]" captionStart="FIGURE 3" captionStartId="6.[151,250,1918,1940]" captionTargetBox="[229,1364,221,1877]" captionTargetId="figure@6.[201,1389,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Male genital capsules, labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.-GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to dorsal, ventral, lateral, and caudal aspect respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the right penis valve." httpUri="https://zenodo.org/record/1301396/files/figure.png" pageId="3" pageNumber="66">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="2F0836F88B72FFC3DDA8C0BDFD12FEA6" blockId="3.[151,1437,1121,2010]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66">
Workers of 
<taxonomicName id="E8B74D7B8B72FFC4DC24C0BEFE2FF967" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[331,420,1734,1757]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DC24C0BEFE2FF967" box="[331,420,1734,1757]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
can be differentiated from all others in the 
<taxonomicName id="E8B74D7B8B72FFC4DEE2C0BEFB86F964" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[909,1037,1733,1758]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="1DC3EAEA8B72FFC4DEE2C0BEFB86F964" box="[909,1037,1733,1758]" italics="true" pageId="3" pageNumber="66">L. ventralis</emphasis>
</taxonomicName>
species group, except 
<taxonomicName id="E8B74D7B8B72FFC4D863C0BEFA1EF964" box="[1292,1429,1733,1758]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4D863C0BEFA1EF964" box="[1292,1429,1733,1758]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
, due to the coloration of the pubescence. 
<taxonomicName id="E8B74D7B8B72FFC4DF33C091FCB4F8BB" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[604,831,1769,1794]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DF33C091FCB4F8BB" box="[604,831,1769,1794]" italics="true" pageId="3" pageNumber="66">Lepidotrigona satun</emphasis>
</taxonomicName>
has dark hairs on the tibiae and pale yellowish brown to pale brown hairs on the vertex and mostly pale hairs (except posteriorly) on the mesoscutellum; no other member of the 
<taxonomicName id="E8B74D7B8B72FFC4DC2FC149FE29F8F0" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[320,418,1841,1866]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="1DC3EAEA8B72FFC4DC2FC149FE29F8F0" box="[320,418,1841,1866]" italics="true" pageId="3" pageNumber="66">ventralis</emphasis>
</taxonomicName>
species group (except 
<taxonomicName id="E8B74D7B8B72FFC4DFCEC14AFCA2F8F0" box="[673,809,1841,1866]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4DFCEC14AFCA2F8F0" box="[673,809,1841,1866]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
) has this combination: 
<taxonomicName id="E8B74D7B8B72FFC4D940C14AFB20F8F0" baseAuthorityName="Strand" baseAuthorityYear="1913" box="[1071,1195,1841,1866]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="hoozana">
<emphasis id="1DC3EAEA8B72FFC4D940C14AFB20F8F0" box="[1071,1195,1841,1866]" italics="true" pageId="3" pageNumber="66">L. hoozana</emphasis>
</taxonomicName>
and 
<taxonomicName id="E8B74D7B8B72FFC4D98EC14AFAEBF8F0" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1249,1376,1841,1866]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="ventralis">
<emphasis id="1DC3EAEA8B72FFC4D98EC14AFAEBF8F0" box="[1249,1376,1841,1866]" italics="true" pageId="3" pageNumber="66">L. ventralis</emphasis>
</taxonomicName>
have some or entirely (respectively) whitish hairs on the metatibia, whereas the other four species have brown to black hairs; 
<taxonomicName id="E8B74D7B8B72FFC4DDB2C102FEDDF828" baseAuthorityName="Cockerell" baseAuthorityYear="1929" box="[221,342,1914,1938]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="arcifera">
<emphasis id="1DC3EAEA8B72FFC4DDB2C102FEDDF828" box="[221,342,1914,1938]" italics="true" pageId="3" pageNumber="66">L. arcifera</emphasis>
</taxonomicName>
has pale hairs on the pro- and mesotibiae whereas 
<taxonomicName id="E8B74D7B8B72FFC4DEF4C102FC7DF82B" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[923,1014,1914,1937]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4DEF4C102FC7DF82B" box="[923,1014,1914,1937]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
, 
<taxonomicName id="E8B74D7B8B72FFC4D96AC102FB04F828" box="[1029,1167,1913,1938]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4D96AC102FB04F828" box="[1029,1167,1913,1938]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E8B74D7B8B72FFC4D9A7C102FAD4F828" box="[1224,1375,1913,1938]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4D9A7C102FAD4F828" box="[1224,1375,1913,1938]" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
have dark hairs; 
<taxonomicName id="E8B74D7B8B72FFC4DC7BC1E6FE22F80C" box="[276,425,1949,1974]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B72FFC4DC7BC1E6FE22F80C" box="[276,425,1949,1974]" italics="true" pageId="3" pageNumber="66">L. doipaensis</emphasis>
</taxonomicName>
has black hairs on the vertex and mesoscutellum, whereas 
<taxonomicName id="E8B74D7B8B72FFC4D957C1E6FB4BF80C" box="[1080,1216,1949,1974]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B72FFC4D957C1E6FB4BF80C" box="[1080,1216,1949,1974]" italics="true" pageId="3" pageNumber="66">L. flavibasis</emphasis>
</taxonomicName>
and 
<taxonomicName id="E8B74D7B8B72FFC4D99AC1E6FAC4F80F" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1269,1359,1950,1973]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B72FFC4D99AC1E6FAC4F80F" box="[1269,1359,1950,1973]" italics="true" pageId="3" pageNumber="66">L. satun</emphasis>
</taxonomicName>
do not have these hairs black but yellowish grey or variously pale to dark brown, respectively; the thickest of these hairs are about 
<quantity id="E84F9B1D8B75FFC3DC64C6E0FE0AFF15" box="[267,385,152,176]" metricMagnitude="-6" metricUnit="m" metricValue="7" pageId="4" pageNumber="67" unit="mm" value="0.007">0.007 mm</quantity>
, 
<quantity id="E84F9B1D8B75FFC3DCE0C6E0FD8FFF15" box="[399,516,152,176]" metricMagnitude="-6" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="67" unit="mm" value="0.003">0.003 mm</quantity>
and 
<quantity id="E84F9B1D8B75FFC3DF2FC6EFFD3FFF15" box="[576,692,151,176]" metricMagnitude="-6" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="67" unit="mm" value="0.005">0.005 mm</quantity>
wide, respectively. Workers of 
<taxonomicName id="E8B74D7B8B75FFC3D94CC6E1FB0BFF0A" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1059,1152,153,176]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B75FFC3D94CC6E1FB0BFF0A" box="[1059,1152,153,176]" italics="true" pageId="4" pageNumber="67">L. satun</emphasis>
</taxonomicName>
and 
<taxonomicName id="E8B74D7B8B75FFC3D9D3C6E1FAC3FF0A" box="[1212,1352,151,176]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B75FFC3D9D3C6E1FAC3FF0A" box="[1212,1352,151,176]" italics="true" pageId="4" pageNumber="67">L. flavibasis</emphasis>
</taxonomicName>
can be readily differentiated by the ratio of lengths of the metatibia and 2nd submarginal cell, at least 
<quantity id="E84F9B1D8B75FFC3D9A7C6C4FA9DFF6F" box="[1224,1302,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" pageId="4" pageNumber="67" unit="in" value="1.21">1.21 in</quantity>
<taxonomicName id="E8B74D7B8B75FFC3D871C6C5FAF3FF6E" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1310,1400,189,212]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B75FFC3D871C6C5FAF3FF6E" box="[1310,1400,189,212]" italics="true" pageId="4" pageNumber="67">L. satun</emphasis>
</taxonomicName>
, at most 
<quantity id="E84F9B1D8B75FFC3DDB4C698FEBBFF42" box="[219,304,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.4892" pageId="4" pageNumber="67" unit="in" value="0.98">0.98 in</quantity>
<taxonomicName id="E8B74D7B8B75FFC3DC51C699FE44FF42" box="[318,463,223,248]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B75FFC3DC51C699FE44FF42" box="[318,463,223,248]" italics="true" pageId="4" pageNumber="67">L. flavibasis</emphasis>
</taxonomicName>
and intermediate 
<quantity id="E84F9B1D8B75FFC3DFC2C6A7FCCAFF42" box="[685,833,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.7178" metricValueMax="2.7686" metricValueMin="2.667" pageId="4" pageNumber="67" unit="in" value="1.07" valueMax="1.09" valueMin="1.05">1.05–1.09 in</quantity>
<taxonomicName id="E8B74D7B8B75FFC3DE21C699FC60FF42" box="[846,1003,223,248]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B75FFC3DE21C699FC60FF42" box="[846,1003,223,248]" italics="true" pageId="4" pageNumber="67">L. doipaensis</emphasis>
</taxonomicName>
. There are additional morphometric differences among the three species (
<tableCitation id="623503438B75FFC3DF5BC77CFD03FEA6" box="[564,648,260,285]" captionStart="TABLE 1" captionStartId="16.[151,239,1012,1035]" captionText="TABLE 1. Measurement of malar space length (MSL), F1 length (F1L), metatibia width (MTW), forewing diagonal (FWD), and 2nd submarginal cell (SM2L) ratios that separate L. satun n. sp., L. flavibasis, and L. doipaensis, expressed as range (Average±SD). Holotype ratios in bold." pageId="4" pageNumber="67">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="7BC866708B75FFC3DDF8C03EFD20F902" httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="4" pageNumber="67" startId="4.[151,250,1606,1628]" targetBox="[160,1427,354,1563]" targetPageId="4">
<paragraph id="2F0836F88B75FFC3DDF8C03EFD20F902" blockId="4.[151,1436,1606,1720]" pageId="4" pageNumber="67">
<emphasis id="1DC3EAEA8B75FFC3DDF8C03EFE9DF9E1" bold="true" box="[151,278,1606,1628]" pageId="4" pageNumber="67">FIGURE 2.</emphasis>
Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to 
<taxonomicName id="E8B74D7B8B75FFC3D921C03FFB28F9E6" authority="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer, 2018" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1102,1187,1607,1628]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B75FFC3D921C03FFB28F9E6" box="[1102,1187,1607,1628]" italics="true" pageId="4" pageNumber="67">L. satun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F057918B75FFC3D9C2C03EFB62F9E1" box="[1197,1257,1606,1627]" pageId="4" pageNumber="67" rank="species">
<emphasis id="1DC3EAEA8B75FFC3D9C2C03EFB62F9E1" bold="true" box="[1197,1257,1606,1627]" pageId="4" pageNumber="67">n. sp.</emphasis>
</taxonomicNameLabel>
(Paratype, H.B.- GP3311), 
<taxonomicName id="E8B74D7B8B75FFC3DD90C01EFEF7F9C0" box="[255,380,1636,1659]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B75FFC3DD90C01EFEF7F9C0" box="[255,380,1636,1659]" italics="true" pageId="4" pageNumber="67">L. flavibasis</emphasis>
</taxonomicName>
(H.B.-GP3322), and 
<taxonomicName id="E8B74D7B8B75FFC3DF34C01EFD68F9C0" box="[603,739,1636,1659]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B75FFC3DF34C01EFD68F9C0" box="[603,739,1636,1659]" italics="true" pageId="4" pageNumber="67">L. doipaensis</emphasis>
</taxonomicName>
(H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses.
</paragraph>
</caption>
<subSubSection id="67AD65738B75FFCCDDA8C0A7FA9DFEFA" lastPageId="11" lastPageNumber="74" pageId="4" pageNumber="67" type="description">
<paragraph id="2F0836F88B75FFC3DDA8C0A7FD9AF942" blockId="4.[151,1437,1758,2036]" box="[199,529,1758,1784]" pageId="4" pageNumber="67">
<emphasis id="1DC3EAEA8B75FFC3DDA8C0A7FD9AF942" bold="true" box="[199,529,1758,1784]" pageId="4" pageNumber="67">
Description. 
<emphasis id="1DC3EAEA8B75FFC3DC30C0A6FD82F94D" bold="true" box="[351,521,1758,1783]" italics="true" pageId="4" pageNumber="67">
<typeStatus id="F00C885A8B75FFC3DC30C0A6FE4CF94D" box="[351,455,1758,1783]" pageId="4" pageNumber="67" type="holotype">Holotype</typeStatus>
Male
</emphasis>
.
</emphasis>
</paragraph>
<paragraph id="2F0836F88B75FFC2DDA8C17CFC2DFF6E" blockId="4.[151,1437,1758,2036]" lastBlockId="5.[151,1437,151,1904]" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
<emphasis id="1DC3EAEA8B75FFC3DDA8C17CFEBDF8A1" box="[199,310,1796,1819]" italics="true" pageId="4" pageNumber="67">Structure:</emphasis>
Body length 
<quantity id="E84F9B1D8B75FFC3DCA1C17BFDBFF8A1" box="[462,564,1795,1820]" metricMagnitude="-3" metricUnit="m" metricValue="4.67" pageId="4" pageNumber="67" unit="mm" value="4.67">4.67 mm</quantity>
. Head wider than long; head width 
<quantity id="E84F9B1D8B75FFC3DEADC17CFBA2F8A1" box="[962,1065,1796,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" pageId="4" pageNumber="67" unit="mm" value="1.77">1.77 mm</quantity>
; head length 
<quantity id="E84F9B1D8B75FFC3D9D3C17CFAAFF8A1" box="[1212,1316,1796,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="4" pageNumber="67" unit="mm" value="1.31">1.31 mm</quantity>
; eye width (in side view) 
<quantity id="E84F9B1D8B75FFC3DC54C15FFE29F885" box="[315,418,1831,1856]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="67" unit="mm" value="0.51">0.51 mm</quantity>
; gena width 
<quantity id="E84F9B1D8B75FFC3DF5CC15FFD10F885" box="[563,667,1831,1855]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="4" pageNumber="67" unit="mm" value="0.18">0.18 mm</quantity>
; interocellar distance 
<quantity id="E84F9B1D8B75FFC3DEFDC15FFC71F885" box="[914,1018,1831,1855]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="67" unit="mm" value="0.31">0.31 mm</quantity>
; ocellocular distance 
<quantity id="E84F9B1D8B75FFC3D981C15FFADDF885" box="[1262,1366,1831,1855]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="67" unit="mm" value="0.17">0.17 mm</quantity>
; inner orbits strongly converging below, upper interorbital distance 
<quantity id="E84F9B1D8B75FFC3DE27C134FC25F8D9" box="[840,942,1868,1892]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="4" pageNumber="67" unit="mm" value="0.98">0.98 mm</quantity>
, lower interorbital distance 
<quantity id="E84F9B1D8B75FFC3D985C133FAD9F8D9" box="[1258,1362,1867,1892]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="67" unit="mm" value="0.68">0.68 mm</quantity>
; scape length excluding basal bulb 
<quantity id="E84F9B1D8B75FFC3DCBEC117FDBCF83D" box="[465,567,1903,1927]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="67" unit="mm" value="0.48">0.48 mm</quantity>
, with basal bulb 
<quantity id="E84F9B1D8B75FFC3DF9CC117FCD2F83D" box="[755,857,1903,1928]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="4" pageNumber="67" unit="mm" value="0.57">0.57 mm</quantity>
; 1 
<taxonomicName id="E8B74D7B8B75FFC3DE00C117FB92F832" box="[879,1049,1903,1928]" pageId="4" pageNumber="67" rank="subSpecies" subSpecies="flagellomere">st flagellomere</taxonomicName>
length 
<quantity id="E84F9B1D8B75FFC3D903C117FB58F83D" box="[1132,1235,1903,1927]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="67" unit="mm" value="0.09">0.09 mm</quantity>
; 2nd flagellomere length 
<quantity id="E84F9B1D8B75FFC3DD8BC1ECFEC1F811" box="[228,330,1940,1964]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="67" unit="mm" value="0.17">0.17 mm</quantity>
, width 
<quantity id="E84F9B1D8B75FFC3DCF2C1ECFD8FF811" box="[413,516,1940,1964]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="67" unit="mm" value="0.13">0.13 mm</quantity>
; 3rd subequal to 2nd; 4th slightly longer than broad; malar space length 
<quantity id="E84F9B1D8B75FFC3D85EC1ECFA13F811" box="[1329,1432,1940,1964]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="67" unit="mm" value="0.01">0.01 mm</quantity>
; mandible with subapical tooth. Forewing length (measured from humeral angle to apex) 
<quantity id="E84F9B1D8B75FFC3D917C1CFFB55F875" box="[1144,1246,1975,1999]" metricMagnitude="-3" metricUnit="m" metricValue="4.03" pageId="4" pageNumber="67" unit="mm" value="4.03">4.03 mm</quantity>
, width 
<quantity id="E84F9B1D8B75FFC3D85EC1CFFA13F875" box="[1329,1432,1975,2000]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="4" pageNumber="67" unit="mm" value="1.52">1.52 mm</quantity>
; forewing diagonal 
<quantity id="E84F9B1D8B75FFC3DC04C1A4FE48F849" box="[363,451,2012,2036]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="67" unit="mm" value="1.1">1.1 mm</quantity>
, length of 1 
<taxonomicName id="E8B74D7B8B75FFC3DF2AC1A4FD66F84E" box="[581,749,2011,2036]" pageId="4" pageNumber="67" rank="subSpecies" subSpecies="submarginal">st submarginal</taxonomicName>
cell 
<quantity id="E84F9B1D8B75FFC3DE4BC1A3FC01F849" box="[804,906,2011,2036]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="67" unit="mm" value="0.35">0.35 mm</quantity>
, 2nd submarginal cell length 
<quantity id="E84F9B1D8B75FFC3D9BAC1A4FAB0F849" box="[1237,1339,2012,2036]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="4" pageNumber="67" unit="mm" value="0.47">0.47 mm</quantity>
, vein Rs reaching stigma, 1 
<taxonomicName id="E8B74D7B8B74FFC2DC07C6E0FE66FF15" box="[360,493,152,176]" pageId="5" pageNumber="68" rank="subSpecies" subSpecies="recurrent">st recurrent</taxonomicName>
vein at approximately mid-length of 2nd submarginal cell. Metatibia length 
<quantity id="E84F9B1D8B74FFC2D804C6E0FF4EFF6E" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="5" pageNumber="68" unit="mm" value="1.34">1.34 mm</quantity>
, width 
<quantity id="E84F9B1D8B74FFC2DC78C6C4FEF5FF6E" box="[279,382,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="68" unit="mm" value="0.52">0.52 mm</quantity>
; metabasitarsus length 
<quantity id="E84F9B1D8B74FFC2DFEDC6C4FD63FF6E" box="[642,744,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="68" unit="mm" value="0.51">0.51 mm</quantity>
, width 
<quantity id="E84F9B1D8B74FFC2DE54C6C4FC29FF6E" box="[827,930,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="68" unit="mm" value="0.34">0.34 mm</quantity>
.
</paragraph>
<paragraph id="2F0836F88B74FFC2DDA8C6A7FB30FF42" blockId="5.[151,1437,151,1904]" box="[199,1211,223,248]" pageId="5" pageNumber="68">Metasomal sterna and terminalia are described from paratypes in a separate section below.</paragraph>
<paragraph id="2F0836F88B74FFC2DDA8C77CFA98FD86" blockId="5.[151,1437,151,1904]" pageId="5" pageNumber="68">
<emphasis id="1DC3EAEA8B74FFC2DDA8C77CFE7BFEA6" box="[199,496,260,285]" italics="true" pageId="5" pageNumber="68">Coloration of integument:</emphasis>
Head black except as follows: dorsal surface of scape, most of pedicel, proximal half of F1, and small marks towards base and apex of mandible dark brown; supraclypeal area and clypeus brown; labrum, most of mandible, ventral surface of scape, apical mark on pedicel and the rest of F1 yellowish brown. Mesosoma brown except pronotal lobe, mesoscutum, metepisternum and sides of metanotum black and metanotum medially yellowish brown; fore and mid legs pale brown except coxae, trochanters proximally and femora dorsally brown; hind leg brown except tarsus yellowish brown; wings hyaline to pale brown, wing veins and stigma pale brown except veins C, R, M, Rs, and R1 brownish black. Metasomal terga amber brown except T1 with semicircular brown band darker towards sides on disc, basal depression translucent, posterior marginal zone translucent amber, T4–T5 brown; S1 and S2 anteriorly whitish translucent, S3 with lateral brown spot.
</paragraph>
<paragraph id="2F0836F88B74FFC2DDA8C43FFEFAFAB7" blockId="5.[151,1437,151,1904]" pageId="5" pageNumber="68">
<emphasis id="1DC3EAEA8B74FFC2DDA8C43FFEDFFDDA" box="[199,340,583,608]" italics="true" pageId="5" pageNumber="68">Pubescence:</emphasis>
Most of face from lower margin of vertex to apex of clypeus with short plumose dense appressed yellowish white hairs; labrum, ventral scape, most portion of genal area, and lower edge of mandible with pale yellowish brown hairs longest at apex of labrum and lower mandible; ventral surface of scape with very short pale brown hairs; vertex with erect long pale yellowish brown hairs, longest hairs of vertex approximately one half as long as scape. Mesoscutum with long and very short pale yellowish brown hairs, the longest hairs approximately one quarter as long as scape; margins of mesoscutum with distinct band of short plumose scale-like yellow hairs interrupted anteromedially and posteriorly; mesoscutellum with pale yellowish brown hairs, progressively longer from anterior to posterior; mesepisternum, metepisternum and metanotum with pale brown hairs, somewhat darker anteroventrally on mesepisternum; posteroventral area of mesepisternum with long white hairs, pronotal lobe and mesepisternum anterodorsally with yellow plumose scale-like hairs; metapostnotum without hairs, lateral surface of propodeum with minute dense white hairs; coxa and trochanter of all legs with long pale brownish white hairs, ventral surface of all femora with pale brown hairs, remaining femora with pale brown hairs intermixed with dark brown hairs, protibia with pale brown hairs intermixed with few short dark brown hairs, the dorsal and anterior mesotibia with dark brown hairs, rest of mesotibia with pale yellowish brown hairs, metatibia dorsally with hairs variable, mostly brown, sometimes dark brown or greyish yellow, or intermixed; protarsus with yellowish brown hairs, meso- and metatarsi with yellowish brown hairs intermixed with few dark brown hairs, posterior surface of metatibia with short yellowish brown keirotrichia. Posterolateral margin of T1 and posterior and lateral parts of T2–T4 with minute sparse pale brownish white hairs; T3–T6 laterally with fine pale brown hairs progressively longer from T3–T6; T7 with pale brown hairs most dense on posterior margin, the longest hairs as long as the longest hairs of T6.
</paragraph>
<paragraph id="2F0836F88B74FFC2DDA8C360FE80FA7A" blockId="5.[151,1437,151,1904]" pageId="5" pageNumber="68">
<emphasis id="1DC3EAEA8B74FFC2DDA8C360FE11FA8A" box="[199,410,1303,1328]" italics="true" pageId="5" pageNumber="68">Surface sculpture:</emphasis>
Head and most of mesosoma with fine dense punctures; metapostnotum with coarse crowded sharp-edged punctures, diameters 2–2.5 times those of mesoscutum; all legs with fine sparse punctures; metasomal terga shining, T1 with few coarse punctures on posterolateral brown band, T2–T7 with few fine shallow punctures restricted to the posterior margin; metasomal sterna somewhat shining with sparse fine shallow punctures.
</paragraph>
<paragraph id="2F0836F88B74FFC2DDA8C3B4FA04FA5F" blockId="5.[151,1437,151,1904]" box="[199,1423,1483,1509]" pageId="5" pageNumber="68">
<emphasis id="1DC3EAEA8B74FFC2DDA8C3B4FD98FA5E" bold="true" box="[199,531,1483,1509]" pageId="5" pageNumber="68">
Description. 
<emphasis id="1DC3EAEA8B74FFC2DC30C3B4FD98FA5E" bold="true" box="[351,531,1483,1508]" italics="true" pageId="5" pageNumber="68">
<typeStatus id="F00C885A8B74FFC2DC30C3B4FE4DFA59" box="[351,454,1484,1507]" pageId="5" pageNumber="68" type="paratype">Paratype</typeStatus>
Males
</emphasis>
</emphasis>
, measured as range (Average±SD, n=10). As in the 
<typeStatus id="F00C885A8B74FFC2D934C3B4FB36FA5F" box="[1115,1213,1484,1509]" pageId="5" pageNumber="68" type="holotype">holotype</typeStatus>
except as follows:
</paragraph>
<paragraph id="2F0836F88B74FFC2DDA8C388FC05F8CA" blockId="5.[151,1437,151,1904]" pageId="5" pageNumber="68">
<emphasis id="1DC3EAEA8B74FFC2DDA8C388FEBDF9BD" box="[199,310,1520,1543]" italics="true" pageId="5" pageNumber="68">Structure:</emphasis>
Body length 4.33–4.85 (4.58±0.15) mm. Head width 1.72–1.77 (1.75±0.02) mm; head length 1.24– 1.34 (1.32±0.03) mm; inner orbits converging below, upper interorbital distance 0.94–1.01 (0.97±0.03) mm, lower interorbital distance 0.67–0.73 (0.69±0.02) mm; eye width 0.48–0.54 (0.52±0.02) mm; gena width 0.14–0.18 (0.17±0.02) mm; interocellar distance 0.28–0.31 (0.29±0.01) mm; ocellocular distance 0.1 8–0.22 (0.2±0.01) mm; scape length excluding basal bulb 0.47–0.52 (0.49±0.01) mm, with basal bulb 0.56–0.59 (0.58±0.01) mm); 1 
<taxonomicName id="E8B74D7B8B74FFC2D8E9C0F8FEAFF907" pageId="5" pageNumber="68" rank="subSpecies" subSpecies="flagellomere">st flagellomere</taxonomicName>
length 0.09–0.11 (0.1±0.01) mm; 2nd flagellomere length 0.15–0.17 (0.16±0.01) mm, width 0.12– 0.14 (0.13±0.004); 3rd subequal to 2nd; 4th slightly longer than broad; malar space length 0.01–0.02 (0.01±0.002) mm. Forewing length 3.94–4.09 (4.01±0.05) mm, width 1.47–1.52 (1.49±0.02); forewing diagonal 1.09–1.18 (1.15±0.03) mm, length of 1 
<taxonomicName id="E8B74D7B8B74FFC2DCB9C168FDF4F892" box="[470,639,1807,1832]" pageId="5" pageNumber="68" rank="subSpecies" subSpecies="submarginal">st submarginal</taxonomicName>
cell 0.3–0.34 (0.32±0.01) mm, 2nd submarginal cell length 0.46–0.52 (0.49±0.01) mm. Metatibia length 1.34–1.43 (1.39±0.03) mm, width 0.47–0.53 (0.5±0.02) mm. Metabasitarsus length 0.47–0.55 (0.5±0.03) mm, width 0.29–0.33 (0.31±0.01) mm.
</paragraph>
<caption id="7BC866708B77FFC1DDF8C106FE73F84A" httpUri="https://zenodo.org/record/1301396/files/figure.png" pageId="6" pageNumber="69" startId="6.[151,250,1918,1940]" targetBox="[229,1364,221,1877]" targetPageId="6">
<paragraph id="2F0836F88B77FFC1DDF8C106FE73F84A" blockId="6.[151,1436,1918,2032]" pageId="6" pageNumber="69">
<emphasis id="1DC3EAEA8B77FFC1DDF8C106FE99F829" bold="true" box="[151,274,1918,1940]" pageId="6" pageNumber="69">FIGURE 3.</emphasis>
Male genital capsules, labelled as: (A), (B), and (C) refer to 
<taxonomicName id="E8B74D7B8B77FFC1DE13C107FC46F82E" authority="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer, 2018" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[892,973,1919,1940]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B77FFC1DE13C107FC46F82E" box="[892,973,1919,1940]" italics="true" pageId="6" pageNumber="69">L. satun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F057918B77FFC1DEBBC107FB86F82E" box="[980,1037,1919,1940]" pageId="6" pageNumber="69" rank="species">
<emphasis id="1DC3EAEA8B77FFC1DEBBC107FB86F82E" bold="true" box="[980,1037,1919,1940]" pageId="6" pageNumber="69">n. sp.</emphasis>
</taxonomicNameLabel>
(Paratype, H.B.-GP3311), 
<taxonomicName id="E8B74D7B8B77FFC1D84FC107FA17F82E" box="[1312,1436,1918,1940]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B77FFC1D84FC107FA17F82E" box="[1312,1436,1918,1940]" italics="true" pageId="6" pageNumber="69">L. flavibasis</emphasis>
</taxonomicName>
(H.B.-GP3322), and 
<taxonomicName id="E8B74D7B8B77FFC1DC1EC1E6FE70F809" box="[369,507,1949,1971]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B77FFC1DC1EC1E6FE70F809" box="[369,507,1949,1971]" italics="true" pageId="6" pageNumber="69">L. doipaensis</emphasis>
</taxonomicName>
(H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to dorsal, ventral, lateral, and caudal aspect respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the right penis valve.
</paragraph>
</caption>
<caption id="7BC866708B76FFC0DDF8C1B7FB07F85F" box="[151,1164,1999,2021]" httpUri="https://zenodo.org/record/1301398/files/figure.png" pageId="7" pageNumber="70" startId="7.[151,250,1999,2021]" targetBox="[185,1409,193,1978]" targetPageId="7">
<paragraph id="2F0836F88B76FFC0DDF8C1B7FB07F85F" blockId="7.[151,1164,1999,2021]" box="[151,1164,1999,2021]" pageId="7" pageNumber="70">
<emphasis id="1DC3EAEA8B76FFC0DDF8C1B7FE99F85E" bold="true" box="[151,274,1999,2021]" pageId="7" pageNumber="70">FIGURE 4.</emphasis>
<taxonomicName id="E8B74D7B8B76FFC0DC76C1B7FE6CF85F" authority="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer, 2018" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[281,487,1999,2021]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="70" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B76FFC0DC76C1B7FE6CF85F" box="[281,487,1999,2021]" italics="true" pageId="7" pageNumber="70">Lepidotrigona satun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F057918B76FFC0DC82C1B7FDADF85E" box="[493,550,1999,2020]" pageId="7" pageNumber="70" rank="species">
<emphasis id="1DC3EAEA8B76FFC0DC82C1B7FDADF85E" bold="true" box="[493,550,1999,2020]" pageId="7" pageNumber="70">n. sp.</emphasis>
</taxonomicNameLabel>
, dorsal habitus. (A) male holotype and (B) worker paratype.
</paragraph>
</caption>
<caption id="7BC866708B79FFCFDDF8C0E7FB06F90F" box="[151,1165,1695,1718]" httpUri="https://zenodo.org/record/1301400/files/figure.png" pageId="8" pageNumber="71" startId="8.[151,250,1695,1717]" targetBox="[151,1436,180,1686]" targetPageId="8">
<paragraph id="2F0836F88B79FFCFDDF8C0E7FB06F90F" blockId="8.[151,1165,1695,1718]" box="[151,1165,1695,1718]" pageId="8" pageNumber="71">
<emphasis id="1DC3EAEA8B79FFCFDDF8C0E7FE99F90F" bold="true" box="[151,274,1695,1717]" pageId="8" pageNumber="71">FIGURE 5.</emphasis>
<taxonomicName id="E8B74D7B8B79FFCFDC76C0E7FE6CF90C" authority="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer, 2018" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[281,487,1695,1718]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B79FFCFDC76C0E7FE6CF90C" box="[281,487,1695,1718]" italics="true" pageId="8" pageNumber="71">Lepidotrigona satun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F057918B79FFCFDC82C0D8FDADF90F" box="[493,550,1696,1717]" pageId="8" pageNumber="71" rank="species">
<emphasis id="1DC3EAEA8B79FFCFDC82C0D8FDADF90F" bold="true" box="[493,550,1696,1717]" pageId="8" pageNumber="71">n. sp.</emphasis>
</taxonomicNameLabel>
, lateral habitus. (A) male holotype and (B) worker paratype.
</paragraph>
</caption>
<paragraph id="2F0836F88B79FFCFDDA8C0A4FAD4F83C" blockId="8.[151,1437,1756,2033]" pageId="8" pageNumber="71">
<emphasis id="1DC3EAEA8B79FFCFDDA8C0A4FE60F94F" box="[199,491,1756,1781]" italics="true" pageId="8" pageNumber="71">Coloration of integument:</emphasis>
Ventral scape usually yellowish brown rarely brown on distal half, yellowish brown area on pedicel and 1 
<taxonomicName id="E8B74D7B8B79FFCFDCECC179FDA5F8A3" box="[387,558,1792,1817]" pageId="8" pageNumber="71" rank="subSpecies" subSpecies="flagellomere">st flagellomere</taxonomicName>
varies in size, position, and sometimes brown; pale marking on disc of tegula varies in size, seldom absent; mesepisternum sometimes dark brown; propodeum sometimes dark brown or more or less black; all coxae sometimes brown, femora and tibiae vary in darkness of brown; semicircular brown band of T1 varies slightly in width, sometimes darker than other terga; T2–T7 sometimes darker than in the 
<typeStatus id="F00C885A8B79FFCFD99AC115FAD0F83C" box="[1269,1371,1901,1926]" pageId="8" pageNumber="71" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="2F0836F88B79FFCFDDA8C1E8FDF5F84B" blockId="8.[151,1437,1756,2033]" pageId="8" pageNumber="71">
<emphasis id="1DC3EAEA8B79FFCFDDA8C1E8FEDFF813" box="[199,340,1936,1961]" italics="true" pageId="8" pageNumber="71">Pubescence:</emphasis>
Long hairs on vertex, mesoscutum and mesoscutellum sometimes slightly paler and longer than the 
<typeStatus id="F00C885A8B79FFCFDDAFC1CDFEAEF874" box="[192,293,1973,1998]" pageId="8" pageNumber="71" type="holotype">holotype</typeStatus>
; yellow plumose scale-like hairs on mesepisternum vary in density; minute hairs on visible portion of metasomal terga vary in length and density.
</paragraph>
<caption id="7BC866708B78FFCEDDF8C1B7FB3FF85F" box="[151,1204,1999,2021]" httpUri="https://zenodo.org/record/1301402/files/figure.png" pageId="9" pageNumber="72" startId="9.[151,250,1999,2021]" targetBox="[181,1408,197,1978]" targetPageId="9">
<paragraph id="2F0836F88B78FFCEDDF8C1B7FB3FF85F" blockId="9.[151,1204,1999,2021]" box="[151,1204,1999,2021]" pageId="9" pageNumber="72">
<emphasis id="1DC3EAEA8B78FFCEDDF8C1B7FE99F85E" bold="true" box="[151,274,1999,2021]" pageId="9" pageNumber="72">FIGURE 6.</emphasis>
<taxonomicName id="E8B74D7B8B78FFCEDC76C1B7FE6CF85F" authority="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer, 2018" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[281,487,1999,2021]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B78FFCEDC76C1B7FE6CF85F" box="[281,487,1999,2021]" italics="true" pageId="9" pageNumber="72">Lepidotrigona satun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F057918B78FFCEDC82C1B7FDADF85E" box="[493,550,1999,2020]" pageId="9" pageNumber="72" rank="species">
<emphasis id="1DC3EAEA8B78FFCEDC82C1B7FDADF85E" bold="true" box="[493,550,1999,2020]" pageId="9" pageNumber="72">n. sp.</emphasis>
</taxonomicNameLabel>
, head, frontal view. (A) male holotype and (B) worker paratype.
</paragraph>
</caption>
<paragraph id="2F0836F88B7BFFCDDDA8C6E0FC43FF0A" blockId="10.[151,1437,151,2013]" box="[199,968,151,176]" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C6E0FE18FF0A" box="[199,403,151,176]" italics="true" pageId="10" pageNumber="73">Surface sculpture:</emphasis>
Fine shallow punctures on T2–T7 vary in density.
</paragraph>
<paragraph id="2F0836F88B7BFFCDDDA8C6C4FB89FF6E" blockId="10.[151,1437,151,2013]" box="[199,1026,187,213]" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C6C4FDA6FF6E" bold="true" box="[199,557,187,213]" pageId="10" pageNumber="73">
Description. 
<emphasis id="1DC3EAEA8B7BFFCDDC30C6C4FDA6FF6E" bold="true" box="[351,557,187,212]" italics="true" pageId="10" pageNumber="73">
<typeStatus id="F00C885A8B7BFFCDDC30C6C4FE4DFF69" box="[351,454,188,211]" pageId="10" pageNumber="73" type="paratype">Paratype</typeStatus>
Workers
</emphasis>
</emphasis>
, measured as range (Average±SD, n=10).
</paragraph>
<paragraph id="2F0836F88B7BFFCDDDA8C698FDDEFEA7" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="73">
As in the 
<typeStatus id="F00C885A8B7BFFCDDC5DC6A7FE1FFF42" box="[306,404,223,248]" pageId="10" pageNumber="73" type="holotype">holotype</typeStatus>
except for usual sexual dimorphism in flagellomeres, metasomal segment number, presence of corbicula, terminalia, and as follows:
</paragraph>
<paragraph id="2F0836F88B7BFFCDDDA8C750FB7FFD4A" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C750FEBDFE85" box="[199,310,296,319]" italics="true" pageId="10" pageNumber="73">Structure:</emphasis>
Body length 3.94–4.39 (4.16±0.15) mm. Head wider than long; head width 1.76–1.85 (1.81±0.03) mm; head length 1.38–1.45 (1.41±0.02) mm, inner orbits slightly diverging for upper half, converging below, upper interorbital distance 1.08–1.13 (1.11±0.02) mm, lower interorbital distance 0.97–1.03 (0.99±0.02) mm; eye width 0.41–0.48 (0.45±0.02) mm; gena width 0.2–0.27 (0.25±0.02) mm; interocellar distance 0.2 7–0.32 (0.29±0.02) mm; ocellocular 0.26–0.3 (0.28±0.01) mm; scape length excluding basal bulb 0.62–0.67 (0.65±0.01) mm, with basal bulb 0.72–0.75 (0.74±0.01) mm; 1 
<taxonomicName id="E8B74D7B8B7BFFCDDF76C7A4FD4FFE4F" box="[537,708,476,501]" pageId="10" pageNumber="73" rank="subSpecies" subSpecies="flagellomere">st flagellomere</taxonomicName>
length 0.11–0.13 (0.12±0.01) mm; 2nd flagellomere length 0.1– 0.13 (0.11±0.01) mm, width 0.11–0.13 (0.13±0.01); 3rd approximate subequal to 2nd; 4th as long as broad; malar space length 0.08–0.11 (0.09±0.01) mm; mandible with two subapical teeth. Forewing length 3.97–4.09 (4.04±0.04) mm, width 1.51–1.54 (1.52±0.01) mm; forewing diagonal 1.15–1.19 (1.17±0.01) mm, length of 1 
<taxonomicName id="E8B74D7B8B7BFFCDD8E9C430FEAAFD3F" pageId="10" pageNumber="73" rank="subSpecies" subSpecies="submarginal">st submarginal</taxonomicName>
cell 0.3–0.34 (0.32±0.01) mm, 2nd submarginal cell length 0.46–0.52 (0.49±0.01) mm, 1 
<taxonomicName id="E8B74D7B8B7BFFCDD877C414FA17FD3E" box="[1304,1436,620,644]" pageId="10" pageNumber="73" rank="subSpecies" subSpecies="recurrent">st recurrent</taxonomicName>
vein at approximately before middle-length of 2nd submarginal cell. Metatibia length 1.45–1.51(1.49±0.02) mm, width 0.6–0.63 (0.62±0.01) mm; metabasitarsus length 0.57–0.66 (0.61±0.03), width 0.38–0.41 (0.4±0.01). S1 emarginate posteromedially, posterior margin of S2–S5 with shallow broad emargination, S6 entire.
</paragraph>
<paragraph id="2F0836F88B7BFFCDDDA8C484FAFFFC72" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C484FE79FCAE" box="[199,498,764,789]" italics="true" pageId="10" pageNumber="73">Coloration of integument:</emphasis>
Labrum, scape basal bulb, distal and basal marks on scape and 1 
<taxonomicName id="E8B74D7B8B7BFFCDD981C484FA17FCAF" box="[1262,1436,764,789]" pageId="10" pageNumber="73" rank="subSpecies" subSpecies="flagellomere">st flagellomere</taxonomicName>
medially light brown; rest of antenna dark brown (scape and pedicel sometimes black). Mesosoma black except mesoscutellum and tegula dark brown with paler brown disc; all legs black to dark brown, probasitarsus dark to yellowish brown, apex of metatibia and metabasitarsus sometimes brown, tarsi of all legs light brown except slightly darker posteriorly. Semicircular brown band on disc of T1 usually wider laterally than in male; T2–T5 black, sometimes pregradular area of T2 brown; T6 usually black except apex pale, seldom wholly pale brown.
</paragraph>
<paragraph id="2F0836F88B7BFFCDDDA8C5ACFDF2FA5E" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C5ACFEDFFC57" box="[199,340,980,1005]" italics="true" pageId="10" pageNumber="73">Pubescence:</emphasis>
Plumose appressed hairs on face absent from upper frons to vertex, occiput and upper genal area; scape ventrally with few very short pale brown hairs; vertex with erect pale brown hairs shorter and slightly darker than those of male, mesoscutum with sparse short pale brown hairs, medial interruption absent on distinct yellow band; mesoscutellum with long pale yellowish grey hairs variously intermixed with pale to dark brown (but not black) hairs, the thickest hairs about 
<quantity id="E84F9B1D8B7BFFCDDF09C21CFD6AFBC6" box="[614,737,1124,1149]" metricMagnitude="-6" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="73" unit="mm" value="0.005">0.005 mm</quantity>
wide (about 1/10 of the gonostylar width of the male), mesepisternum with fine brown hairs, with long white hairs ventrally, patches of yellow plumose scale-like hairs on mesepisternum anterodorsally and pronotal lobe usually larger than in males; metepisternum and lateral surface of propodeum with fine short pale yellowish white hairs (sometimes yellowish brown); all femora dorsally with dark brown hairs though paler posteriorly, with white (sometimes pale brown) hairs ventrally; pro- and mesotibiae with dark brown hairs intermixed with pale brown hairs posteriorly, metatibia with dark brown hairs dorsally, tips of hairs slightly paler, ventrally with paler hairs intermixed with a few dark hairs, posteriorly with whitish brown keirotrichia; all basitarsi with dark brown intermixed with yellowish brown hairs, remaining tarsomeres with yellowish brown hairs sometimes intermixed with few brown hairs; T1–T4 with minute hairs slightly darker than in male; T3–T6 towards sides with dark brown hairs; S1–S6 with minute pale brown hairs progressively longer from anterior to posterior on each sternum.
</paragraph>
<paragraph id="2F0836F88B7BFFCDDDA8C388FE7AF9CF" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C388FE1FF9B2" box="[199,404,1519,1544]" italics="true" pageId="10" pageNumber="73">Surface sculpture:</emphasis>
Punctures on mesoscutellum, metanotum, and metepisternum slightly larger than on rest of mesosoma; all legs mostly with fine punctures intermixed with few coarse punctures, all femora with punctures similar to those of metapostnotum though slightly smaller; anterior surface of metatibia usually shining for ~apical 2/3, shining area varies in size.
</paragraph>
<paragraph id="2F0836F88B7BFFCCDDA8C007FA9DFEFA" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1437,151,789]" lastPageId="11" lastPageNumber="74" pageId="10" pageNumber="73">
<emphasis id="1DC3EAEA8B7BFFCDDDA8C007FD2DF922" bold="true" box="[199,678,1663,1688]" pageId="10" pageNumber="73">Male Metasomal Sterna and Genitalia.</emphasis>
In the following, description of genitalic features is based upon dissections of 
<typeStatus id="F00C885A8B7BFFCDDC52C0DDFE16F907" box="[317,413,1701,1725]" pageId="10" pageNumber="73" type="paratype">paratype</typeStatus>
males from all four nests (see material examined). S4–S7 weakly sclerotized except S4 posterolaterally, S5 laterally and posteriorly brown (
<figureCitation id="B78C2A7D8B7BFFCDDF87C0BFFCA4F95A" box="[744,815,1735,1760]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="10" pageNumber="73">Fig. 2</figureCitation>
: A1–4); S4 angularly emarginate apicomedially, depth of emargination ~0.33 times length of sternum, emargination separating two broadly rounded lobes, more strongly sclerotized area of S4 with fine dense hairs, longest and densest lateral to apex of lobe; gradulus of S4 weakly convex close to anterior margin of S4 (
<figureCitation id="B78C2A7D8B7BFFCDDF3BC14CFD11F8F6" box="[596,666,1844,1869]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="10" pageNumber="73">Fig. 2</figureCitation>
: A1); S5 with four apical lobes, long submedian lobes separated by deep U-shaped emargination, depth of emargination ~0.63 times medial length of S5, short lateral lobe separated from medial lobe by shallow concavity, medial lobes bearing ~4–7 thick setae apically, gradulus of S5 transverse medially, near anterior margin (
<figureCitation id="B78C2A7D8B7BFFCDDC94C1D8FDCBF802" box="[507,576,1952,1977]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="10" pageNumber="73">Fig. 2</figureCitation>
: A2); S6 apical margin weakly produced medially, weakly concave laterally (
<figureCitation id="B78C2A7D8B7BFFCDDDF0C1BCFF6DF866" box="[159,230,1988,2013]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="10" pageNumber="73">Fig. 2</figureCitation>
: A3); 
<geoCoordinate id="4A83503F8B7BFFCDDC5FC1BCFE0CF867" box="[304,391,1988,2013]" direction="south" orientation="latitude" pageId="10" pageNumber="73" precision="9" value="-7.026">S7 1.56</geoCoordinate>
times as wide as long, apical margin convex throughout (
<figureCitation id="B78C2A7D8B7BFFCDD94EC1BCFBE3F866" box="[1057,1128,1988,2013]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="10" pageNumber="73">Fig. 2</figureCitation>
: A4). Genital capsule light brown except black for apical 3/5 of penis valve; penis valve curved ventrally, somewhat hook-like, curvature strongest on basal third to half, then almost straight, the right valve tends to overlap the left one somewhat and its apex is slightly posterior to that of the left valve. Gonostylus not sclerotized, narrow, at least 12 times as long as wide (
<quantity id="E84F9B1D8B7AFFCCDDB2C77CFE08FEA6" box="[221,387,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="7.65" metricValueMax="7.8" metricValueMin="7.5" pageId="11" pageNumber="74" unit="mm" value="0.765" valueMax="0.78" valueMin="0.75">0.75–0.78 mm</quantity>
long and 
<quantity id="E84F9B1D8B7AFFCCDC98C77CFD3CFEA6" box="[503,695,260,285]" metricMagnitude="-5" metricUnit="m" metricValue="5.1" metricValueMax="5.8" metricValueMin="4.3999999999999995" pageId="11" pageNumber="74" unit="mm" value="0.051" valueMax="0.058" valueMin="0.044">0.044–0.058 mm</quantity>
wide), slightly expanded for apical 1/3, medial surface setose on apical half, hairs denser towards apex, lateral surface with few short hairs in apical half (
<figureCitation id="B78C2A7D8B7AFFCCD918C75FFB37FEFA" box="[1143,1212,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1918,1940]" captionTargetBox="[229,1364,221,1877]" captionTargetId="figure@6.[201,1389,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Male genital capsules, labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.-GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to dorsal, ventral, lateral, and caudal aspect respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the right penis valve." httpUri="https://zenodo.org/record/1301396/files/figure.png" pageId="11" pageNumber="74">Fig. 3</figureCitation>
: A1–4).
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B7AFFCCDDA8C734FEBBFE32" pageId="11" pageNumber="74" type="etymology">
<paragraph id="2F0836F88B7AFFCCDDA8C734FEBBFE32" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="74">
<emphasis id="1DC3EAEA8B7AFFCCDDA8C734FEC5FEDF" bold="true" box="[199,334,332,357]" pageId="11" pageNumber="74">Etymology.</emphasis>
The specific epithet refers to the province in 
<collectingCountry id="57A076688B7AFFCCDE28C734FC20FEDF" box="[839,939,332,357]" name="Thailand" pageId="11" pageNumber="74">Thailand</collectingCountry>
where the species was collected; it is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B7AFFCCDDA8C7ECFB9CFCAE" pageId="11" pageNumber="74" type="materials_examined">
<paragraph id="2F0836F88B7AFFCCDDA8C7ECFAD1FE6A" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="74">
<emphasis id="1DC3EAEA8B7AFFCCDDA8C7ECFE27FE17" bold="true" box="[199,428,404,429]" pageId="11" pageNumber="74">Material examined.</emphasis>
<emphasis id="1DC3EAEA8B7AFFCCDCDBC7ECFD93FE17" box="[436,536,404,429]" italics="true" pageId="11" pageNumber="74">Holotype</emphasis>
. 
<materialsCitation id="9FDF3CA58B7AFFCCDF48C7ECFADDFE6A" ID-GBIF-Occurrence="1852083324" collectingDate="2017-04-27" collectionCode="QSBG" collectorName="Nest TB &amp; H. Banziger" country="Thailand" elevation="100" location="Satun" pageId="11" pageNumber="74" specimenCount="1" stateProvince="Satun" typeStatus="holotype">
(
<collectorName id="8242532E8B7AFFCCDF41C7ECFD04FE16" box="[558,655,404,428]" pageId="11" pageNumber="74">Nest TB</collectorName>
2, male, to be deposited in 
<collectionCode id="49A6AE3D8B7AFFCCDED5C7ECFB8EFE16" box="[954,1029,404,428]" pageId="11" pageNumber="74">QSBG</collectionCode>
) labelled as follows: “Thaleban Nat. Park/ 
<collectingRegion id="ED73F81A8B7AFFCCDDB8C7C0FE9EFE6A" box="[215,277,440,464]" country="Thailand" name="Satun" pageId="11" pageNumber="74">Satun</collectingRegion>
Prov., 
<elevation id="A49AD1CB8B7AFFCCDC0CC7C0FE2CFE6A" box="[355,423,440,464]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="74" unit="m" value="100.0">
<quantity id="E84F9B1D8B7AFFCCDC0CC7C0FE2CFE6A" box="[355,423,440,464]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="74" unit="m" value="100.0">100 m</quantity>
</elevation>
/ 
<collectingDate id="4B4DE9D08B7AFFCCDCD9C7CFFDA3FE6A" box="[438,552,439,464]" pageId="11" pageNumber="74" value="2017-04-27">
<date id="5B0910388B7AFFCCDCD9C7CFFDA3FE6A" box="[438,552,439,464]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
</collectingDate>
/ ex nest TB2/ S. 
<collectingCountry id="57A076688B7AFFCCDF89C7C0FCFFFE6A" box="[742,884,440,464]" name="Thailand" pageId="11" pageNumber="74">THAILAND</collectingCountry>
/ 
<collectorName id="8242532E8B7AFFCCDEECC7C0FB83FE6A" box="[899,1032,439,464]" pageId="11" pageNumber="74">H. Bänziger</collectorName>
leg prep det”, “ 
<typeStatus id="F00C885A8B7AFFCCD9DDC7C0FAC7FE6A" box="[1202,1356,440,464]" pageId="11" pageNumber="74" type="holotype">HOLOTYPE</typeStatus>
”
</materialsCitation>
.
</paragraph>
<paragraph id="2F0836F88B7AFFCCDDA8C7A5FB3EFDDA" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="74">
<emphasis id="1DC3EAEA8B7AFFCCDDA8C7A5FEF5FDA2" italics="true" pageId="11" pageNumber="74">
<typeStatus id="F00C885A8B7AFFCCDDA8C7A5FEB2FE4E" box="[199,313,477,500]" pageId="11" pageNumber="74" type="paratype">Paratypes</typeStatus>
. All bearing labels that state “ 
<typeStatus id="F00C885A8B7AFFCCDFEFC7A4FC84FE49" box="[640,783,476,499]" pageId="11" pageNumber="74" type="paratype">PARATYPE</typeStatus>
” and the same locality as those of the 
<typeStatus id="F00C885A8B7AFFCCD9DEC7A4FA84FE4F" box="[1201,1295,476,501]" pageId="11" pageNumber="74" type="holotype">holotype</typeStatus>
except dates and nests as follows:
</emphasis>
Measured 
<typeStatus id="F00C885A8B7AFFCCDC96C478FDE9FDA2" box="[505,610,512,536]" pageId="11" pageNumber="74" type="paratype">paratypes</typeStatus>
(included in description and 
<tableCitation id="623503438B7AFFCCDEC9C787FC7DFDA2" box="[934,1014,511,536]" captionStart="TABLE 1" captionStartId="16.[151,239,1012,1035]" captionText="TABLE 1. Measurement of malar space length (MSL), F1 length (F1L), metatibia width (MTW), forewing diagonal (FWD), and 2nd submarginal cell (SM2L) ratios that separate L. satun n. sp., L. flavibasis, and L. doipaensis, expressed as range (Average±SD). Holotype ratios in bold." pageId="11" pageNumber="74">Table 1</tableCitation>
) labelled as follows: 
<date id="5B0910388B7AFFCCD989C787FAD2FDA2" box="[1254,1369,511,536]" pageId="11" pageNumber="74" value="2017-04-26">26.iv.2017</date>
(Nest TB1, 
<specimenCount id="39B1FD718B7AFFCCDDB9C45CFEB6FD87" box="[214,317,548,573]" pageId="11" pageNumber="74" type="worker">1 worker</specimenCount>
); 
<date id="5B0910388B7AFFCCDC3FC45CFE48FD87" box="[336,451,548,573]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB1, 
<specimenCount id="39B1FD718B7AFFCCDF23C45CFD10FD87" box="[588,667,548,573]" pageId="11" pageNumber="74" type="male">1 male</specimenCount>
); 
<date id="5B0910388B7AFFCCDFC1C45CFCAAFD87" box="[686,801,548,573]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB2, 
<specimenCount id="39B1FD718B7AFFCCDEC5C45CFB8AFD87" box="[938,1025,548,573]" pageId="11" pageNumber="74" type="male">3 males</specimenCount>
, 
<specimenCount id="39B1FD718B7AFFCCD962C45CFBF4FD87" box="[1037,1151,548,573]" pageId="11" pageNumber="74" type="worker">3 workers</specimenCount>
); 
<date id="5B0910388B7AFFCCD9FDC45CFA8EFD87" box="[1170,1285,548,573]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB3, 
<specimenCount id="39B1FD718B7AFFCCD8E1C45CFF51FDDA" pageId="11" pageNumber="74" type="male">3 males</specimenCount>
, 
<specimenCount id="39B1FD718B7AFFCCDD8BC430FEDDFDDA" box="[228,342,583,608]" pageId="11" pageNumber="74" type="worker">3 workers</specimenCount>
); 
<date id="5B0910388B7AFFCCDC07C43FFE50FDDA" box="[360,475,583,608]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB4, 
<specimenCount id="39B1FD718B7AFFCCDF0DC430FD31FDDA" box="[610,698,583,608]" pageId="11" pageNumber="74" type="male">3 males</specimenCount>
, 
<specimenCount id="39B1FD718B7AFFCCDFABC430FCBDFDDA" box="[708,822,583,608]" pageId="11" pageNumber="74" type="worker">2 workers</specimenCount>
); 
<date id="5B0910388B7AFFCCDE27C43FFC30FDDA" box="[840,955,583,608]" pageId="11" pageNumber="74" value="2017-04-29">29.iv.2017</date>
(Nest TB1, 
<specimenCount id="39B1FD718B7AFFCCD92DC430FB22FDDA" box="[1090,1193,583,608]" pageId="11" pageNumber="74" type="worker">1 worker</specimenCount>
).
</paragraph>
<paragraph id="2F0836F88B7AFFCCDDA8C414FB9CFCAE" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="74">
Non-measured paratypes: 
<date id="5B0910388B7AFFCCDC82C414FDE8FD3F" box="[493,611,620,645]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB1, 2 males), 
<date id="5B0910388B7AFFCCDE34C414FC5AFD3F" box="[859,977,620,645]" pageId="11" pageNumber="74" value="2017-04-26">26.iv.2017</date>
(Nest TB1, 2 workers), 
<date id="5B0910388B7AFFCCD98DC414FAD2FD3F" box="[1250,1369,620,645]" pageId="11" pageNumber="74" value="2017-04-29">29.iv.2017</date>
(Nest TB1, 2 workers), 
<date id="5B0910388B7AFFCCDC01C4F7FE6FFD12" box="[366,484,655,680]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB2, 17 males, 14 workers; 
<date id="5B0910388B7AFFCCDEFAC4F7FB80FD12" box="[917,1035,655,680]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB3, 7 males, 2 workers); 
<date id="5B0910388B7AFFCCDDF8C4CCFE86FD77" box="[151,269,692,717]" pageId="11" pageNumber="74" value="2017-04-27">27.iv.2017</date>
(Nest TB4, 11 males and 2 workers). Genitalia examined: GP 3304, 3305, 3306, 3308, 3309, 3310, 3311, same nest as holotype; GP 3307, 3312, 3313, 3314, 3315, 3316, 3317, paratypes of nest TB4; GP 3318, 3319, 3320, 3321, 3322, paratypes of nest TB3; GP 3323, paratype of nest TB1.
</paragraph>
</subSubSection>
</treatment>
</document>